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Home , Acourtia thurberi, Amoreuxia palmatifida, Argemone pleiacantha, Asclepias asperula, Asclepias linaria, Bouvardia ternifolia

Figure 1. View from the summit of the Sierra Juriquipa of steep slopes with woodland and a few . Photo by Ana L. Reina-G. Preliminary Flora of the Sierra Juriquipa, , by Elizabeth Makings1, Thomas R. Van Devender2, Ana Lilia Reina-Guerrero2, and Stephen F. Hale3

Abstract with oak woodland or -oak . The lowland “seas” are Sonoran and Chihuahuan desertscrub, desert , The Sierra Juriquipa mountain range is a small but important foothills thornscrub, or tropical deciduous forest. part of the in northeastern Sonora, and an area previously unexplored botanically until the Madrean Study Area and Methods Discovery Expedition (MDE) in the summer of 2017. In this This preliminary flora is based on observations from a preliminary flora, we document 282 taxa in 72 families, and scouting trip on July 14–16, and intense collecting during the 198 genera. Eleven (3.9%) are non-native. Madrean Discovery Expedition (MDE) Sierra Juriquipa on Introduction August 12–16, 2017 (Figure 1). This Sky Island is a little over an hour’s drive from Agua Prieta to the mining town of The Madrean Archipelago is located between the Sierra Madre Nacozari de García, then about 19 kilometers (12 miles) Occidental (SMO) and the Mogollon Rim in central . southeast on winding dirt roads through the small mining In this area there are 55 Sky Island isolated mountain ranges village of Santo Domingo. The range is directly south of one or complexes of several ranges connected by oak woodland of the largest copper mines in Mexico — La Mina de la corridors (Van Devender et al. 2013). Sky Islands are crowned Caridad. Our camp at Rancho Zulema in the northwestern section of Juriquipa occupied a narrow mesa with just enough 1 Arizona State University Herbarium, 734 West Alameda Drive, room for the vehicles, gear, and tents of the 35 participants. To 2 Tempe, AZ 85282. GreaterGood.org, 6262 N. Swan Rd., Suite 150, document the floristic diversity we inventoried Ranchos Tucson, AZ 85718. 3EcoPlan Associates Inc., 3610 N. Prince Village Orégano Viejo and San Felipe, and the slopes and ridgetops Place, Suite 140, Tucson, AZ 85719. continued next page www.aznativeplantsociety.org The Press Arizona Native Plant Society 33 Sierra Juriquipa continued

near Rancho Zulema, all on Ejido Santo Domingo, Municipality of Nacozari de García. The Sierra Juriquipa is approximately 6,900 hectares of rugged mountain terrain and V-shaped valleys. The highest peaks reach a little over 2,000 m. (6,561 ft.) elevation with north slopes forested by mostly and Engelmann pine (Pinus chihuahuana, and P. engelmannii,), and an occasional Arizona madrone (Arbutus arizonica) (Figure 2). Typical landscapes explored near Rancho Zulema were steep slopes with alligator juniper (Juniperus deppeana) and . Figure 2. Pine-oak forest on the summit. Photo by Ana L. Reina-G. The oak diversity was noteworthy, with seven species (, Q. chihuahuensis, Q. emoryi, Q. hypoleucoides, Q. oblongifolia, Q. toumeyi, and Q. viminea) distributed across Flora. We observed or vouchered a total of 282 taxa in 72 the range, the dominants changing frequently according to families and 198 genera. Our study includes all biases that go habitat preference. An assortment of grasses, , and along with a short, single-season window of collecting events. succulents occupy the understory including desert spoon/sotol Only 11 species (3.9%) are non-native. The most important (Dasylirion wheeleri) from the desert , and oak families are (35 taxa), Poaceae (31 taxa), and woodland species such as firecracker bush ( (25 taxa), and align with other Sonoran floras. The ternifolia) and velvetpod mimosa/gatuño (Mimosa dysocarpa). regional story is told by the second-tier families that bring out Lower elevations (~1000 m., 3,280 ft.) are foothills thornscrub the personality of the local flora. For the Sierra Juriquipa, the landscapes with mostly boat-thorn acacia/güinolo (Acacia (15 taxa), (15 taxa), cochliacantha), tepeguaje (Lysiloma watsonii), and velvet Convolvulaceae (12 taxa), (11 taxa), Cyperaceae mesquite (Prosopis velutina). (10 taxa), Cactaceae (7 taxa), (7 taxa), and continued next page

Figure 3. A. The charismatic lemmonii. Photo by Elizabeth Makings. B. Mandevilla stans. A single Arizona record in the Santa Rita Mountains. Photo by Ana L. Reina-G. 34 The Plant Press Arizona Native Plant Society Fall 2018 Figure 4. with affinities that reach Arizona. A. Hybanthus attenuatus. B. The Mexican star, Milla biflora. Photos by Elizabeth Makings.

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Pteridaceae (6 taxa) are especially important. The genera with Floristic affinities. Botanists have the tendency to take the most species were Cyperus (9), Ipomoea (7), Euphorbia interest in things that are unfamiliar, out of place, or showy (7), Quercus (7), Asclepias (6), and Solanum (6). Asclepias and irresistible. In addition, botanists love learning different milkweeds were scattered but showy and hard to miss, species and are absorbed with the biogeographical component especially the very large and charismatic Asclepias lemmonii of floristics, pointing out interesting distributions: disjuncts, (Figure 3A). However, in the Sierra Juriquipa, the oaks are the new records, endemics, range extensions, etc. The Sierra stars of the show in a classic Madrean oak woodland or Juriquipa certainly provided examples of these categories. The encinal. One or two species tend to dominate locally, but it is plants are the collective narrative of the region and there are not unusual to encounter four or more species on a single several recurring distributional themes for taxa of the Sierra slope that are, for the most part, easy to distinguish by leaf Juriquipa flora. A few are northern species at their southern shape, color, and texture, as well as habit. continued next page

Figure 5. Plants restricted to the Sky Island Region. A. Coyphantha recurvata. Photo by Stephen F. Hale. B. Mandevilla brachysiphon. Photo by Elizabeth Makings.

www.aznativeplantsociety.org The Plant Press Arizona Native Plant Society 35 Figure 6. SMO species that do not occur in Arizona. A. Penstemon companulatus. Photo by Liz Makings. B. Begonia gracilis. Photo by Stephen F. Hale.

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limits (e.g., Lathyrus lanszwertii var. arizonicus and Opuntia prevents erosion as well as enriching the microfauna diversity, chlorotica). Many more species have southern distributions but it can easily disappear with plowing and other surface that extend into Arizona from the SMO and the Sonoran Sky disturbances. Islands, such as eludens, orthocarpa, angustidens, Coryphantha recurvata (Figure 5A), Fraxinus gooddingii, Hybanthus attenuatus (Figure 4A), Mandevilla brachysiphon (Figure 5B), and Hypoxis mexicana, Mandevilla stans (Figure 3B), Milla biflora mostly occur in the Madrean Archipelago. The Sky Island (Figure 4B), Quercus viminea, Roldana hartwegii, and phytogeographic pattern is not fully appreciated and often Tripsacum lanceolatum. Other species in this category with called “Madrean,” even if the species does not occur in the more tropical affinities are widespread in thornscrub, e.g., Capsicum annuum, bicornutus, and Havardia continued next page mexicana. Bouteloua diversispicula (formerly Cathestecum brevifolium) is ubiquitous in foothills thornscrub (FTS), and Plains of Sonora desertscrub. It is only known in Arizona from the vicinity of Ragged Top Mountain west of Tucson (Wiens 2000). This dwarf, tufted, stoloniferous perennial grass plays an important but underappreciated role in arid habitats where it forms turf that

Figure 7. Mexican species not in Arizona. A. Cyclanthera minima. Photo by Stephen F. Hale. B. Tigridia pavonia, a captivating species — only briefly open. Photo by Elizabeth Makings. 36 The Plant Press Arizona Native Plant Society Fall 2018 Figure 8. Mexican species not in Arizona. A. Manihot rubricaulis. Photo by Susan D. Carnahan. B.Solanum houstonii. Photo by Stephen L. Minter.

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SMO. Other SMO species that reach Arizona include Quercus Arizona SEINet dabatases (http://swbiodiversity.org/seinet/ viminea, Roldana hartwegii, and Tripsacum lanceolatum. collections/index.php). Field observations are available in the fagaroides var. elongata is a widespread tropical species Madrean Discovery Expedition (MDE) that reaches its northwestern distributional limit in (madreandiscovery.org). thornscrub and Sonoran desertscrub in Sonora, except for a Acknowledgements single 1929 collection by Robert H. Peebles in Fresnal Canyon in the southern Baboquivari Mountains. Since then, attempts I am deeply grateful to GreaterGood.org for the financial and to relocate this population were not successful. Other SMO logistical support that enables such meaningful and plants that approach but do not occur in Arizona include important work in the field of conservation and biodiversity Begonia gracilis (Figure 6B), parviflora, Cyclanthera assessment, and I am privileged to engage with an amazing minima (Figure 7A), Penstemon campanulatus (Figure 6A), group of natural history enthusiasts, artists, NGO’s, and some and Tigridia pavonia (Figure 7B). Thornscrub species that of the best naturalists I have ever been around. My thanks go occur just south of Arizona are Manihot rubricaulis (Figure out to all the participants and staff that make these outings 8A), Milleria quinqueflora, and Solanum houstonii (Figure 8B). possible. George Ferguson made valuable and appreciated Lantana camara is a widespread tropical species reaching its edits to the manuscript. northwestern distributional limit in thornscrub and Sonoran desertscrub in Sonora and Baja . Merremia palmeri a is a showy white-flowered vine typical of thornscrub and the adjacent Plains of Sonora desertscrub. Populus monticola is a Literature Cited tropical riparian that would have Sky Islands distribution, Van Devender, T.R., S. Avila-V., M. Emerson, D. Turner, A.D. Flesch, except that it also occurs in . All in all, the and N.S. Deyo. 2013a. Biodiversity in the Madrean Archipelago of Sierra Juriquipa has a diverse collection of biogeographical Sonora, Mexico. Pp. 10–16 in G.J. Gottfried, P.F. Ffolliott, B.S. affinities. Gebow, L.G. Eskew, and L.C. Collins (compilers). Merging science While investigating previous botanical work in the Sierra and management in a rapidly changing world: Biodiversity and management of the Madrean Archipelago III and 7th Conference Juriquipa, we were quite surprised to discover that previous on Research and Resource Management in the Southwestern botanical work was non-existent — not a single herbarium Deserts. 2012 May 1–5, Tucson, AZ. Proceedings RMRS-P-67. Fort specimen had been vouchered prior to our 2017 expedition. Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky To work in an area so rich and yet so unexplored was Mountain Research Station. remarkable and gratifying. Clearly, there is still a lot to learn Wiens, J.F. 2000. Vegetation and flora of Ragged Top, Pima County, about the floras of the Sonoran Sky Island and many other Arizona. Desert Plants 16:3–32. areas in Sonora. Physical vouchers and many images are available in the Arizona State University and University of www.aznativeplantsociety.org The Plant Press Arizona Native Plant Society 37 CHECKLIST: Sierra Juriquipa page 1 of 4

An asterisk (*) denotes non-native Gomphrena caespitosa Torr. Lasianthaea podocephala (A. Gray) K. status. Gomphrena nitida Rothrock Becker Gomphrena sonorae Torr. Melampodium appendiculatum B.L. Lycophytes Robins. Guilleminea densa (Humb. & Bonpl. ex SELAGINELLACEAE Willd.) Moq. Melampodium cupulatum A. Gray Melampodium longicorne A. Gray Selaginella rupincola Underwood ANACARDIACEAE Milleria quinqueflora L. Pteridophytes Rhus aromatica Aiton Porophyllum macrocephalum DC. Rhus virens Lindheimer ex A. Gray ASPLENIACEAE Psacalium decompositum (A. Gray) H.E. Toxicodendron radicans (L.) Kuntze Asplenium palmeri Maxon Robins. & Brett. APOCYNACEAE Roldana hartwegii (Benth.) H. Rob. & Asclepias asperula (Dcne.) Woods. Brettell Bommeria hispida (Mett. ex Kuhn) Symphyotrichum expansum (Poepp. ex Underwood Asclepias elata Benth. Spreng.) G.L. Nesom aurea (Poir.) Grusz & Asclepias lemmonii A. Gray lemmonii A. Gray Windham Asclepias linaria Cav. Verbesina longifolia (A. Gray) A. Gray Myriopteris lindheimeri (Hook.) J. Sm. Asclepias nummularia Torr. Zinnia peruviana (L.) L. Myriopteris wrightii (Hook.) Grusz & Asclepias ovata Steud. Zinnia zinnioides (Kunth) Olorode & A.M Windham Cynanchum ligulatum (Benth.) Woods. Torres wrightiana Hook. Mandevilla brachysiphon (Torr.) Pichon BEGONIACEAE Mandevilla stans (A. Gray) J.K. Williams Gymnosperms Begonia gracilis Vilmorin-Andrieux Matelea tristiflora (Standl.) Woodson CUPRESSACEAE Metastelma mexicanum (Brandegee) M. Juniperus deppeana Steud. Fishbein & R. Levin palmatifida Moc. & Sessé ex DC. PINACEAE ARALIACEAE BORAGINACEAE Pinus chihuahuana Engelm. Aralia humilis Cav. Lithospermum cobrense Greene Pinus engelmannii Carr. ASTERACEAE BRASSICACEAE thurberi (A. Gray) Reveal & King Hesperidanthus linearifolius (A. Gray) ACANTHACEAE Rydb. Ambrosia ambrosioides (Cav.) W.W. Elytraria imbricata (Vahl) Pers. Payne Pennellia micrantha (A. Gray) Nieuwl. Ruellia nudiflora (Engelm. & A. Gray) Ambrosia confertiflora Dc. Urban Artemisia ludoviciana Nutt. Bursera fagaroides (Kunth) Engl. var. Tetramerium nervosum Nees Baccharis salicifolia (Ruiz & Pav.) Pers. elongata McVaugh & Rzed. ADOXACEAE Baccharis sarothroides A. Gray CACTACEAE Sambucus cerulea Raf. Carphochaete bigelovii A. Gray Coryphantha recurvata (Engelm.) Britt. AMARANTHACEAE Conyza canadensis (L.) Cronquist & Rose Alternanthera caracasana Kunth Erigeron flagellaris A. Gray Cylindropuntia versicolor (Engelm. ex J. M. Coult.) Knuth Amaranthus dubius Mart. ex Thell. (new fendleri Schultz-Bip. Sonoran voucher) Echinocereus rigidissimus (Engelm.) Hieracium pringlei A. Gray Haage f. Amaranthus palmeri S. Watson Laennecia eriophylla (A. Gray) G.L. Mammillaria grahamii Engelm. Nesom

38 The Plant Press Arizona Native Plant Society Fall 2018 CHECKLIST: Sierra Juriquipa page 2 of 4

Opuntia chlorotica Engelm. & Bigelow Euphorbia cuphosperma (Engelm.) * Leucaena leucocephala (Lam.) de Wit Opuntia engelmannii Salm-Dyck Boiss. Mimosa distachya Cav. Opuntia cf. wilcoxii Britton & Rose Euphorbia heterophylla L. Mimosa dysocarpa Benth. Euphorbia hirta L. CANNABACEAE Mimosa grahamii A. Gray Euphorbia hyssopifolia L. Celtis pallida Torr. Nissolia schottii (Torr.) A. Gray Euphorbia indivisa (Engelm.) Tidestrom Celtis reticulata Torr. Parkinsonia aculeata L. Euphorbia macropus (Klotzsch & Phaseolus acutifolius A. Gray CARYPOPHYLLACEAE Garcke) Boiss. Phaseolus ritensis M.E. Jones Drymaria effusa A. Gray Manihot angustiloba (Torr.)Müll. Arg. Prosopis velutina Wooton Drymaria leptophylla (Cham. & Schlecht.) Manihot rubricaulis I.M. Johnst. Fenzl ex Rohrb. Senna hirsuta (L.) Irwin & Barneby Tragia laciniata (Torr.) Müll. Arg. Drymaria molluginea (Lag.) Didr. Zornia reticulata Sm. Tragia nepetifolia Cav. CONVOLVULACEAE FAGACEAE FABACEAE Cuscuta americana Thunb. ex Engelm. Quercus arizonica Sarg. Acacia angustissima (Mill.) Kuntze Dichondra brachypoda Wooton & Standl. Trel. Acacia cochliacantha Humb. & Bonpl. Evolvulus alsinoides (L.) L. ex Willd. Quercus emoryi Torr. Evolvulus arizonicus A. Gray Acacia farnesiana (L.) Willd. A. Camus Ipomoea capillacea (Kunth) G. Don Aeschynomene villosa Poir. Torr. Ipomoea costellata Torr. humilis Benth. var. humilis Quercus toumeyi Sarg. Ipomoea cristulata Hallier f. Calliandra humilis Benth. var. reticulata Quercus viminea Trel. Ipomoea hederacea Jacq. (A. Gray) L.D. Benson FOUQUIERIACEAE Ipomoea plummerae A. Gray Chamaecrista absus (L.) Irwin & Barneby Fouquieria splendens Engelm. Ipomoea tenuiloba Torr. Chamaecrista nictitans (L.) Moench GERANIACEAE Chamaecrista serpens (L.) Greene Ipomoea thurberi A. Gray Geranium richardsonii Fisch. & Trautv. Cologania angustifolia Kunth Merremia palmeri (S. Watson) Hallier f. Geranium wislizeni S. Watson Cologania obovata Schlecht. CRASSULACEAE JUGLANDACEAE Coursetia caribaea (Jacq.) Lavin Sedum stelliforme S. Watson Juglans major (Torr.) Heller Crotalaria pumila Blanco CUCURBITACEAE Crotalaria sagittalis L. Cucurbita digitata A. Gray Dalea versicolor Zucc. Agastache wrightii (Greenm.) Wooton & Cyclanthera minima (S. Watson) Kearns & Standl. Desmanthus bicornutus S. Watson C.E. Jones Monarda citriodora Cerb. var. Desmanthus covillei (Britt. & Rose) ERICACEAE austromontana (Epling) B.L. Turner Wiggins ex B.L. Turner Arbutus arizonica (A. Gray) Sarg. subincisa Benth. flabelliformis Kearney Arctostaphylos pungens Kunth LINACEAE Eysenhardtia polystachya (Ortega) Sarg. Linum neomexicanum Greene EUPHORBIACEAE Galactia wrightii A. Gray Acalypha neomexicana Muell. Arg. Havardia mexicana (Rose) Britton & LOASACEAE Acalypha ostryifolia Riddell ex J. M. Coult. Rose Mentzelia aspera L. Acalypha papillosa Rose Indigofera sphaerocarpa A. Gray LYTHRACEAE Lathyrus lanszwertii Kellogg var. Cnidoscolus angustidens Torr. Cuphea wrightii A. Gray arizonicus (Britton) S.L. Welsh Euphorbia bilobata Engelm. www.aznativeplantsociety.org The Plant Press Arizona Native Plant Society 39 CHECKLIST: Sierra Juriquipa page 3 of 4

MALPIGHIACEAE Schistophragma intermedium (A. Gray) Aspicarpa hirtella L.C. Rich. Pennell Buddleja parviflora Kunth MALVACEAE POLYGALACEAE SOLANACEAE Anoda cristata (L.) Schlecht. Hebecarpa obscura (Benth.) J. R. Abbott Browallia eludens R.K. Van Devender & Ayenia filiformis S. Watson POLYGONACEAE P.D. Jenkins Corchorus hirtus L. Eriogonum abertianum Torr. Capsicum annuum L. Gossypium thurberi Todaro Datura discolor Bernh. procumbens (Cav.) J.L. Gentry Sida rhombifolia L. oleracea L. Lycium berlandieri Dunal MARTYNIACEAE Portulaca suffrutescens Engelm. * Nicotiana glauca Graham Proboscidea parviflora (Wooton) Portulaca umbraticola Kunth Physalis hederifolia A. Gray Wooton & Standl. PRIMULACEAE MELIACEAE Physalis philadelphica Lam. Samolus vagans Greene Physalis pubescens L. * Melia azedarach L. RANUNCULACEAE Solanum elaeagnifolium Cav. MOLLUGINACEAE Clematis ligusticifolia Nutt. Solanum houstonii Martyn Mollugo verticillata L. Thalictrum fendleri Engelm. ex A. Gray Solanum lumholtzianum Bartlett MORACEAE RHAMNACEAE Solanum nigrescens M. Martens & Morus microphylla Buckl. Ceanothus buxifolius Willd. ex Schult.f. Galeotti NYCTAGINACEAE Condalia correllii M.C. Johnston Solanum stoloniferum Schltdl. & Bouché Allionia incarnata L. Sageretia wrightii S. Watson TALINACEAE Boerhavia coccinea P. Mill. ROSACEAE Talinum paniculatum (Jacq.) Gaertn. Boerhavia erecta L. Prunus serotina Ehrh. VERBENACEAE OLEACEAE Aloysia gratissima (Gillies & Hook.) Tronc. Fraxinus gooddingii Little Bouvardia ternifolia (Cav.) Schlecht. Lantana camara L. Crusea hispida Robinson VIOLACEAE kunthiana (Spach) Munz proliferum A. Gray Hybanthus attenuatus (Humb. & Bonpl. Oenothera tetraptera Cav. Mitracarpus hirtus (L.) DC. ex J.A. Schultes) G. K. Schulze OROBANCHACEAE Randia sonorensis Wiggins VITACEAE Castilleja tenuiflora Benth. SALICACEAE Vitis arizonica Engelm. OXALIDACEAE Populus monticola Mert. ex Loud. ZYGOPHYLLACEAE Oxalis latifolia Kunth Salix gooddingii Ball Kallstroemia grandiflora Torr. ex A. Gray SANTALACEAE Monocots Phoradendron californicum Nutt. pleiacantha Greene ASPARAGACEAE PASSIFLORACEAE Phoradendron macrophyllum (Engelm.) Cockerell Agave palmeri Engelm. Passiflora bryonioides Kunth Phoradendron serotinum (Raf.) M. C. Dasylirion wheeleri S. Watson Johnst. ssp. tomentosum (DC.) Kuijt Echeandia flavescens (J.A. & J.H. Schultes) Mecardonia procumbens (P. Mill.) Small SAPINDACEAE Cruden Milla biflora Cav. Penstemon campanulatus (Cav.) Willd. Dodonaea viscosa Jacq. continued on page 42 40 The Plant Press Arizona Native Plant Society Fall 2018 Thirty-Seven Years on a Mountain Trail continued

descriptions is lacking. Each description here is grounded Herbarium alone. It is a mountain range that has enjoyed with some subtle explanations that bring a plant the attention of many collectors. Bertelsen has brought community to life for the reader, making it feel more botanical science in this mountain range to the next rung memorable and understandable. These distinctive on the ladder by providing context, context that can only comments are subtle but worth noting. Of Desert Scrub, come of a deep personal understanding of the data. In Bertelsen says, “Many typical desert species in the Tucson Thirty-Seven Years, we are treated to a slice of what must area are uncommon to rare, or altogether absent, in the be a much bigger pie. The data Bertelsen have amassed study area. This is probably due to a number of factors, extend well beyond what is on display in this flora. We including elevation and the absence of bajada, sandy may find ourselves a little anxious to see what comes of washes, or silty .” This last sentence describes an this seminal work in the future, and we may feel a kinship essential difference between Desert Scrub in Finger Rock with the mountain that wasn’t there before. Canyon and Desert Scrub in flatter topography. This Two Supplements: (1) Corrections and Editions and (2) nuanced descriptive writing is woven into the entire flora. Flora Nomenclature Index may be obtained upon request Thirty-Seven Years brings with it a lot of finely parsed from the author, David Bertelsen, information. [email protected]. There are almost eight thousand specimens from the Catalina Mountains in the University of Arizona a

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Nolina microcarpa S. Watson IRIDACEAE Eragrostis intermedia A.S. Hitchc. Yucca madrensis Gentry Sisyrinchium cernuum (Bickn.) Kearney Eragrostis pectinacea (Michx.) Nees ex Steud. COMMELINACEAE Tigridia pavonia (L. f.) DC. Eriochloa acuminata (J. Presl) Kunth Commelina tuberosa L. POACEAE Eriochloa lemmonii Vasey & Scribn. Commelina erecta L. Aristida adscensionis L. Heteropogon contortus (L.) P. Beauv. ex Tradescantia pinetorum Greene Aristida ternipes Cav. var. ternipes Roemer & J.A. Schultes CYPERACEAE Bouteloua curtipendula (Michx.) Torr. * Hordeum vulgare L. Bulbostylis juncoides (Vahl) Kükenth. Bouteloua diversispicula Columbus * Melinis repens (Willd.) Zizka Cyperus dipsaceus Liebamann Bouteloua hirsuta Lag. Muhlenbergia alopecuroides (Griseb.) Cyperus esculentus L. Bouteloua repens (Kunth) Scribn. & P.M. Peterson & Columbus Merr. Cyperus hermaphroditus (Jacq.) Standl. Muhlenbergia emersleyi Vasey Chloris virgata Sw. Cyperus hypopitys G. Tucker Panicum alatum var. minus (Andersson) * Cynodon dactylon (L.) Pers. F. Zuloaga & O. Morrone Cyperus manimae Kunth * Digitaria sanguinalis (L.) Scop. Panicum hirticaule J. Presl Cyperus odoratus L. Dinebra panicea ssp. brachiata (Steud.) Paspalum setaceum Michx. Cyperus pallidicolor (Kükenth.) G. Tucker P.M. Peterson & N. Snow Setaria liebmannii E. Fourn. Cyperus seslerioides Kunth Disakisperma dubium (Kunth) P.M. * Sorghum halepense (L.) Pers. Cyperus squarrosus L. Peterson & N. Snow Tripsacum lanceolatum Rupr. ex Fourn. HYPOXIDACEAE * Echinochloa colona (L.) Link * Triticum aestivum L. Hypoxis mexicana J.A. & J.H. Schultes * Eragrostis cilianensis (All.) Vignolo ex Janch. Zuloagaea bulbosa (Kunth) Bess

42 The Plant Press Arizona Native Plant Society Fall 2018