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7,&;%#.H&'%?#*%( Existen varios grupos bajo sospecha de ser pan-archosauria: Los marinos Thallatosauria, Ichthyopterygia, Sauropterygia; y los arborícolas Megalancosaurus, Longisquama.

Ej. Megalancosaurus comparte con archosauria la perdida del foramen entepicondylar del húmero y la perdida del tarsal distal V.

Longisquama: comparte con archosauria dos corridas de escamas dorsales

Sin embargo en otros aspectos son plesiomórficos, y tanto Longisquama como Megalancosaurus comparten synapomorfías con Coelurosauravus, un diápsido basal, en un grupo Avicephala, fuera de Sauria.

Tanto Thalattosauria como Sauropterygia comparten una articulacion mandibular posterior al occipital y un rostro alargado (mordida amplia), otros rasgos sin publicar (John Merck, tesis PhD Yale)

Askeptosaurus Los pan-archosaurios más universalmente reconocidos como tales son los Choristodera, que presentan huesos nasales más largos que los frontales, y más de 6 vertebras cervicales Archosauromorpha: ACMR de protorosauria, archosauria y todos sus descendientes

Prolacerta El cuello arcosauromorfo: -Centros inclinados (vert 2-6) -Prezygapofises largas (verts 3-7) -costillas cervicales en forma de “arado” (3-6)

Más semejantes a archosauria en que se pierde el hueso septomaxilar (y vomeronasal asociado tb??)

Posición elevada de la cabeza inadecuada para sabo-olisquear?

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Q"/"'2#%(%/2"*2%4( PROTEROSUCHIDAE Apertura pineal: primitivo!

Archosaurus Proterosuchidae Archosauriformes (thecodontia, apo-archosauria: fenestra anteorbital)

-Dientes con bordes aserrados -Diente caniniforme en el dentario -fenestra mandibular (desde Erythrosuchidae: ausente en proterosuchidae) -Orbita mas alta que ancha, con el globo ocular en la parte superior

-Mandíbulas macrocarnívoras: presas de mayor tamaño, fundamentalmente otros vertebrados. Matar y engullir algo de mayor tamaño a)! La mandibula esta desplazada ventralmente para aumentar la “bocanada” b)! Musculos mandibulares modificados, más profundos al interior del cráneo, quizás provoca la génesis de una osificación lateroesfenoide anterior al neurocráneo c)! Estos grandes musculos se insertan en una enorme depresion mandibular con una fenestra lateral d)! Músculo pterygoideo se extiende a la parte superior del paladar, debajo del ojo

Craneo macrocarnivoro; Hocico largo que es alto, angosto y triangular en seccion transversal . El resto del craneo en seccion transversal es trapezoidal (de la orbita hacia atras)

“meatcleaver face” desarrollada por varios macrocarnivoros, depredadores de tetrápodos (T%.*%("4(O/%4($"4(2#*+'*.&(5+'(2#"5,#%/&(H%."/('?(%,%#*.*[/(4&'((E%G#'%(+"&'.5$4(.&5&("4(>?'-18,@%#)!$"(q@y(5($"( G%#A&@(G&'(.#+/"&'($"(%4A?/&'("#I2H#&'?.H*$%"(%4.%/C%3%/(5+'($"(s(5"2#&($"(4%#A&@(T%."('?(%,%#*.*[/(4%(E"/"'2#%( 5%/$*3?4%#(h,*.%($"(4&'(7#.H&'%?#*%@(P4(#%/A&($"(5&F*5*"/2&($"(4%'("R2#"5*$%$"'("'(5%I
('?A*#*"/$&(?/%(,&'*34"( .%,%.*$%$(E%.?42%>F%(,%#%(?/%(,&'2?#%(5+'("#A?*$%@(P4(EN5?#('"(2&#/%('*A5&*$%4("/(.&/2&#/&(I(%,%#"."(?/('*>&($"( */'"#.*[/(5?'.?4%#1("4(.?%#2&(2#&.H%/2"#1(H%.*%(4%(.%#%(,&'2"#*&#($"4(2"#.*&(,#&R*5%4($"4(EN5?#!

L?%#2&(2#&.H%/2"#(

Femur de archosaurio derivado Q"/"'2#%(5%/$*3?4%#( (dinosaurio, Eoraptor)

Erythrosuchidae: origen de la pelvis triradiada

LH%b%#"'?.H?'(

M#&2"#&.H%5,'%( M#&2"#&.H%5,'*$%"((,#"'"/2%/(?/%(/&2%34"($*E"#"/.*%("/("4(2%5%b&($"(4&'(3#%C&'(#"',".2&(%(4%'(,*"#/%'@(

P/(4%(,%#2"(5+'(2%#$=%($"4(2#*+'*.&(2"5,#%/&(H%."('?(%,%#*.*[/(( 7'4#+8/+%#A!7'4#+8/+%#("'(."#.%/&(%(!%&'(+$"%.$1(A#?,&(B?"(*/.4?I"(%(2&$&'(4&'( $"'."/$*"/2"'("4(%/."'2#&(.&5\/(5+'(#".*"/2"(.&5,%#>$&(,&#(4&'(.&.&$#*4&'(I(4%'(%F"'@( 7'4#+8/+%#!E?"($"'.?3*"#2&(.&5&(?/%(%A#"A%.*[/($"(s^("',".=5"/"'('?3%$?42&'1("/(?/( ,#&3%34"(.&/2"R2&('&.*%4@(G%($*'5*/?.*[/($"(2%5%b&(,?"$"(#"4%.*&/%#'"(.&/(4%(5%I&#( %A*4*$%$1(5%I&#(?'&($"(4%(.&/$?.2%($"(H?=$%( Euparkeria y Proterochampsidae son más similares a Archosauria (crown) en que tienen dos filas parasagitales de osteodermos

Un sistema elaborado de musculos bajo estos osteodermos permite un “rebote” al galopar

Sin embargo aún las arterias carotidas pasan por debajo (en vez de encima) del proceso del basipterygoides, lo que les diferencia de archosauria

7#.H&'%?#*%(-.#&x/(A#&?,0(

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o bien , Pan-Aves, y Pan Crocodylia. En esta clase estudiaremos la rama Pan-crocodylia! Synapomorfías Arcosaurianas!

-Corazon: a) Ventriculos divididos b) 3 valvulas semilunares c) Valvula lateral muscular del orificio auricoloventricular derecho

-“Buche”: estomago anterior muscularizado, en el q se acumulan piedras tragadas

-Construyen nidos de vegetacion

-Cuidado del nido y de la cria

-“Canto” para atraer pareja y defender territorio

-Vocalizaciones fetales/juveniles: sincronizan eclosion, gatillan apertura del nido, provocan cuidado parental, cohesion de “bandada’ entre juveniles

-Respiran al correr: a) postura erguida, parasagital c) torso y filas de osteodermos reducen ondulaciones laterales d) pulmones grandes y compartimentalizados e) respiración con flujo unidireccional del aire

G&'(M%#%'?.H*%1(E%'(%.?+>.%'(5?I('"5"`%/2"'(%(4&'(.&.&$#*4&'(,"#&( B?"('*/("53%#A&(/&('&/('?'(%/."'2#&'@(8/%(.%#%.2"#='>.%(E+.*45"/2"( $*'>/>F%("'(4%(,&'*.*[/(#"2#%=$%($"(4%'(%,"#2?#%'(/%'%4"'1(B?"('"( "/.?"/2#%/(`?'2&(%4(E#"/2"($"(4&'(&`&'("/(?/(,#&5&/2&#*&("4"F%$&((

'"()*#+,$%&- AETOSAURIA

G&'(A*A%/2"'.&'(9%?*'?.H*%-M#"'2&'?.H*$%"0(.%#/=F&#&'1($"(,&'2?#%(2&2%45"/2"("#A?*$%(@( P'2&'(44"A%#&/(%(5"$*#(sq(5"2#&'(I(,&'"=%/(.#+/"&'(2%/(A#%/$"'(.&5&(BC+#((,-#'+'-! +/D(( RAUISUCHIA

G*/j(2&(P#I2H#&'?.H*%e((

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G&'(,&,&'%?#*$%"1(.&5&(",-.,-'21'-)!A"/"#%45"/2"('"( .&/'*$"#%/(.&5&(?/%(E%5*4*%($"(9%?*'?.H*%1(,"#&(H%I( ,#&,?"'2%'(%42"#/%>F%'(-z??40!

E,.,-#'+'-)!"'(?/(/&2%34"! 9%?*'?.H*%(H"#3=F&#&($"( LH*/%1('*/($*"/2"'1(,#&F*'2&( $"(F"4%($&#'%4( ORNITHOSUCHIDAE G&'(V#/*2H&'?.H*$%"('"(.%#%.2"#*C%/(,&#((4%(A#%/((#"$?..*[/($"('?'( "R2#"5*$%$"'(%/2"#*&#"'(I(?/(2&3*44&("/("4(.?%4("4(.%4.+/"&('"(,#&I".2%(H%.*%("4( %'2#+A%4&1(%4(#"FN'(B?"("/(4&'($"5+'(.#?#&2%#'*@(P'2%((.&/$*.*[/(('"(H%( /&53#%$&(.&5&(Y.&.&$#*4&(#"F"#>$&Z(

ddddddddddddddddddddddddddddd( Como tortugas

Nothochampsa Notochampsa Protosuchus

Triasico: pubis aun participa del acetabulo

F+#2%$%-'21'-1(?/(<%R[/(,#&34"5+>.&J((M%#%(%4A?/&'("'(?/(6,H"/&'?.H*&1(,%#%(&2#&'1(?/( V#/*2H&'?.H*&1(I(,%#%(&2#&'1(."#.%/&(%(",-.,-'21'-! GRACILISUCHUS G&'(,#*5"#&'(.&.&$#*4&5&#E&'(3%'%4"'(-(Y6,H"/&'?.H*%Z0(.&5&(B/++/-.+%-'21'-!I!*#$.,4,-'21'-!'&/(5?I(+A*4"'1($"( "R2#"5*$%$"'("'3"42%'@(P/(B/++/-.+%-'21'-(4&'(H?"'&'(?4/%#"(I(#%$*%4"($"4(.%#,&('&/(%4%#A%$&'( SPHENOSUCHIA No hay paladar secundario

Receso del canino mandibular G&'("4"5"/2&'(.%#,%4"'(%4%#A%$&'( 2%53*N/('"(%,#".*%/("/(4&'( 6,H"/&'?.H*%! M'"?$&H"',"#&'?.H*$%"((

PROTOSUCHIA

U&2H&.H%5,'%( 2#*4&,H&'%?#?'( -,#&2&'?B?*&0(

THALATTOSUCHIA (“mesosuquia” marinos) 16 familias!

Gran reduccion del brazo en metriorhynchidae V'2"&$"#5&'(

Gran tamaño de la apertura tempora superior

G&'(<"4"&'%?#*$%"('"(#".?,"#%/($"('"$*5"/2&'( 5%#*/&'(I(,&'""/(?/(H&.*&(%4%#A%$&(.&/(,*"/2"'( ,?/C%/2"'(.4%#%5"/2"(,*'.=F&#&@(M&'""/(%5%$?#%( $N#5*.%(( G&'(K"2#*&#HI/.H*$%"("'2%3%/(%42%5"/2"("',".*%4*C%$&'("/(4%(F*$%(5%#*/%@(74(*A?%4(B?"(4&'( :.H2HI&'%?#*%1(,&'"=%/(?/%(.&4%(H"2"#&."#.%(B?"(#".&##=%("4(4[3?4&(*/E"#*&#@(M"#$*"#&/(,&#( .&5,4"2&("4("'B?"4"2&($N#5*.&@(((

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Dakosaurus (metriorhynchidae)

Hasta hace poco, solo se hacía la distinción de proto, meso y eusuquia. Sine embargo, conviene hacer la distinción de METASUCHIA para una serie de taxa muy ecomorfológicamnete diversos ubicados entre los Thalattosuchia marinos y los cocodrilos modernos. Las afinidades de estos taxa son altamente debatidas, dependiendo principalmente de la inclusividad de un grupo monofilético diverso, los Nothosuchia. Sereno y Larsson 2009 proponen una separación de Metasuchia en Nothosuchia y Neosuchia

Metasuchia: el ACMR de Notosuchus terrestris y Crocodylus niloticus, y todos sus descendientes. NOTHOSUCHIA Larsson y Sereno 2009 U&2H&'?.H*%J(,"B?"b&'(.&.&$#*4&'($"(,&'2?#%(5?I("#A?*$%1(+A*4"'1(#&'2#&(.&1(I($"/>.*[/(H"2"#&$&/2%1(,&'*34"5"/2"( H"#3=F&#%@(-"`@1(;1%<#/+,-'21'-)!$"(.H*/%0@(L&/>"/"(E%'(4%2%5"/2"($"#*F%$%'{(7#%#*,"'?.H?'('"(%,#&R*5%(%4((/&2&'?B?*&( %/."'2#%4(

G+#+%4/-'21'-!

Simosuchus Yacarenani

(sí, todavía estamos en Nothosuchia) Los Baurusuchia y Sphagesauridae son reconocidos por su cráneo profundo y diferenciación dentaria que recuerda algunos synapsidos

Baurusuchus

Sphagesaurus Armadillosuchus Sphagesauridae

Segun Sereno y Larsson, Baurusuchia y sphagesauridae son NOTOSUCHIA

Los Sebecidae son más cercanos a cocodrilos modernos G&'(<#"5%2&.H%5,'*$%"(&(6"3".*$%"('&/(A#%/$"'(.&.&$#*4&'(2"##"'2#"'1($"("R2#"5*$%$"'(4%#%A%$%'1(B?"( ,&'"=%/($*"/2#"'(,%4%/%$&'(I(%'"##%$&'(5?I('"5"`%/2"'(%(4&'($*"/2"'($"($*/&'%?#*&'(.%#/=F&#&'1(.&/$*.*[/( B?"('"(H%($"/&5*/%$&(.&5&(YC*E&$&/2"Z1(I(B?"(4?"A&("F&4?.*&/[($"(5%/"#%(.&/F"#A"/2"("/(&2#&'( L&&$#*4&'(P?.#&.&$I4*%((

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