Checklist1 of Vascular Flora Cuesta Ridge East from Highway 101 to Upper Lopez Canyon San Luis Obispo County, (09 June 2019)

David J. Keil Robert F. Hoover Herbarium Biological Sciences Department California Polytechnic State University San Luis Obispo, California

Scientific Name Common Name Family Rare n Acer macrophyllum big- maple SAPINDACEAE v n ❀ Achillea millefolium yarrow o n ❀ Acmispon americanus var. americanus Spanish-clover FABACEAE o n ❀ Acmispon glaber var. glaber common deerweed FABACEAE v n Acmispon junceus var. biolettii Biolett's rush deerweed FABACEAE v n Acmispon parviflorus miniature deervetch FABACEAE v n ❀ Acmispon strigosus strigose deer-vetch FABACEAE v

1 Please notify the author of additions or corrections to this list ([email protected]). ❀ — See Wildflowers of San Luis Obispo, California, second edition (2018) for photograph. Most are illustrated in the first edition as well; old names for some species in square brackets. n — California native i — exotic species, introduced to California, naturalized or waif. v — documented by one or more specimens (Consortium of California Herbaria record; specimen in OBI; or collection that has not yet been accessioned) o — observed during field surveys; no voucher specimen known R—California Rare Rank Scientific Name Common Name Family Rare n ❀ Acourtia microcephala sacapelote ASTERACEAE o n ❀ Adenostoma fasciculatum var. chamise ROSACEAE v fasciculatum n Adiantum aleuticum five finger fern PTERIDACEAE o n Adiantum jordanii California maidenhair fern PTERIDACEAE v n ❀ Agoseris grandiflora var. grandiflora large-flowered mountain- ASTERACEAE v dandelion n Agrostis exarata western bentgrass, spike POACEAE v bentgrass n Agrostis hooveri Hoover's bentgrass POACEAE 1B.2 v n Allophyllum glutinosum sticky false-gilia POLEMONIACEAE v i Amaryllis belladonna naked ladies AMARYLLIDACEAE o n Amorpha californica var. californica false indigo-bush FABACEAE v n Amsinckia intermedia common fiddleneck BORAGINACEAE v n madioides woodland tarweed ASTERACEAE v i Anthriscus caucalis bur-chervil APIACEAE v n ❀ Antirrhinum kelloggii climbing snapdragon PLANTAGINACEAE v n ❀ Antirrhinum thompsonii bush snapdragon PLANTAGINACEAE v n Aphanes occidentalis western parsley-piert ROSACEAE v n Aphyllon tuberosum broomrape OROBANCHACEAE v n Aralia californica California spikenard, elk-clover ARALIACEAE v n ❀ menziesii madrone v n glandulosa subsp. Cushing manzanita ERICACEAE v cushingiana n bigberry manzanita ERICACEAE v n ❀ Arctostaphylos luciana Santa Lucia manzanita ERICACEAE 1B.2 v n ❀ Artemisia californica California sagebrush ASTERACEAE v n ❀ Artemisia douglasiana mugwort ASTERACEAE o n ❀ Asclepias eriocarpa woolly-pod milkweed, kotolo APOCYNACEAE o n ❀ Asclepias fascicularis narrow-leaved milkweed APOCYNACEAE o Scientific Name Common Name Family Rare i Avena barbata slender wild oats POACEAE o i Avena fatua common wild oats POACEAE o n ❀ Baccharis pilularis coyote bush ASTERACEAE o n ❀ Barbarea orthoceras American wintercress BRASSICACEAE o i Brachypodium hybridum false brome POACEAE o i ❀ Brassica nigra black mustard BRASSICACEAE v n Bromus carinatus var. carinatus California brome POACEAE v i Bromus diandrus ripgut brome POACEAE v n Bromus grandis tall brome POACEAE v i Bromus hordeaceus soft chess brome POACEAE o i Bromus madritensis foxtail chess POACEAE v n Bromus pseudolaevipes woodland brome POACEAE v i Bromus rubens red brome POACEAE v n Calandrinia breweri Brewer's calandrinia MONTIACEAE 4.2 v n ❀ Calandrinia menziesii [Calandrinia ciliata] red maids MONTIACEAE v n ❀ Calochortus albus fairy lantern LILIACEAE v n ❀ Calystegia macrostegia subsp. cyclostegia coast morning glory CONVOLVULACEAE v n Calystegia purpurata subsp. purpurata Pacific morning glory CONVOLVULACEAE v n Camissoniopsis hirtella hairy suncup ONAGRACEAE v n ❀ Cardamine californica milk maids, rain bells, BRASSICACEAE v California toothwort i ❀ Carduus pycnocephalus Italian thistle ASTERACEAE v n Carex globosa round-fruited sedge CYPERACEAE v n ❀ Castilleja affinis subsp. affinis coast Indian paintbrush OROBANCHACEAE v n Ceanothus oliganthus var. sorediatus jim brush RHAMNACEAE v n ❀ Ceanothus papillosus wartleaf ceanothus RHAMNACEAE v i ❀ Centaurea melitensis tocalote, Maltese star thistle ASTERACEAE v i ❀ Centaurea solstitialis yellow star thistle ASTERACEAE o i Cerastium glomeratum annual mouse-ear chickweed CARYOPHYLLACEAE v Scientific Name Common Name Family Rare i ❀ Chondrilla juncea skeleton weed ASTERACEAE o n Chorizanthe staticoides Turkish-rugging spineflowe POLYGONACEAE v n ❀ Cirsium occidentale var. lucianum Cuesta Ridge thistle ASTERACEAE 1B.2 v i ❀ Cirsium vulgare bull thistle ASTERACEAE o n ❀ Clarkia purpurea subsp. quadrivulnera fourspot clarkia ONAGRACEAE v n ❀ Clarkia unguiculata elegant clarkia ONAGRACEAE v n Claytonia parviflora subsp. parviflora narrow-leaved miner's lettuce MONTIACEAE v n Claytonia perfoliata subsp. mexicana common miner's lettuce MONTIACEAE v n ❀ Claytonia perfoliata subsp. perfoliata common miner's lettuce MONTIACEAE v n ❀ Clematis lasiantha virgin's bower, pipestems RANUNCULACEAE v n ❀ Collinsia heterophylla common Chinese houses PLANTAGINACEAE v i ❀ Conium maculatum poison hemlock APIACEAE o n ❀ Corethrogyne filaginifolia California-aster ASTERACEAE o n ❀ Cornus sericea subsp. occidentalis creek dogwood CORNACEAE v n ❀ Crassula connata common pygmyweed CRASSULACEAE o n ❀ Crocanthemum scoparium California rush-rose CISTACEAE v n ❀ Croton setiger turkey-mullein EUPHORBIACEAE o n ❀ Cryptantha clevelandii var. florosa coastal cryptantha BORAGINACEAE v n Cryptantha corollata coast range cryptantha BORAGINACEAE v n Cryptantha intermedia var. intermedia Nievitas cryptantha BORAGINACEAE v n Cryptantha muricata var. jonesii Jones's cryptantha BORAGINACEAE v n Cuscuta californica var. californica common chaparral dodder CONVOLVULACEAE v n ❀ Cuscuta subinclusa canyon dodder CONVOLVULACEAE v n Cyperus eragrostis common umbrella sedge CYPERACEAE v i Dactylis glomerata orchard grass POACEAE v n Delphinium nudicaule canyon larkspur, red larkspur RANUNCULACEAE v n ❀ Dendromecon rigida bush poppy, tree poppy PAPAVERACEAE v n ❀ Diplacus aurantiacus var. aurantiacus common bush monkeyflower PHRYMACEAE v [Mimulus aurantiacus] Scientific Name Common Name Family Rare n Diplacus aurantiacus var. longiflorus long-flowered bush PHRYMACEAE v monkeyflower n Diplacus rattanii [Mimulus rattanii] Rattan's monkeyflower PHRYMACEAE v n Drymocallis glandulosa var. glandulosa sticky cinquefoil ROSACEAE v n Dryopteris arguta California wood fern DRYOPTERIDACEAE v n Elymus condensatus giant wildrye POACEAE o n Elymus glaucus subsp. glaucus blue wildrye POACEAE v n ❀ Emmenanthe penduliflora var. penduliflora whispering bells HYDROPHYLLACEAE v n ❀ Epilobium brachycarpum tall willow-herb ONAGRACEAE o n ❀ Epilobium canum subsp. canum California-fuchsia ONAGRACEAE o n ❀ Epilobium ciliatum subsp. ciliatum common willow-herb ONAGRACEAE v i Erigeron canadensis common horseweed ASTERACEAE o n ❀ Erigeron philadelphicus Philadelphia fleabane daisy ASTERACEAE v n Eriodictyon traskiae subsp. smithii Smith's yerba santa NAMACEAE v n ❀ Eriogonum elongatum var. elongatum long-stem buckwheat POLYGONACEAE v n Eriogonum fasciculatum var. foliolosum leafy California buckwheat POLYGONACEAE o n Eriogonum nudum var. pubiflorum inflated naked buckwheat POLYGONACEAE o n ❀ Eriophyllum confertiflorum var. common golden-yarrow ASTERACEAE v confertiflorum i ❀ Erodium cicutarium redstem filaree GERANIACEAE o i Erodium moschatum greenstem filaree GERANIACEAE o n Erodium texanum Texas filaree GERANIACEAE o n Erysimum capitatum var. capitatum western wallflower BRASSICACEAE v n ❀ Erythranthe cardinalis [Mimulus cardinalis] scarlet monkeyflower PHRYMACEAE v n Erythranthe grandis large monkeyflower PHRYMACEAE v n ❀ Erythranthe guttata [Mimulus guttatus] common monkeyflower PHRYMACEAE v n ❀ Eschscholzia caespitosa Foothill poppy PAPAVERACEAE v n ❀ Eschscholzia californica California poppy PAPAVERACEAE v Scientific Name Common Name Family Rare n ❀ Eucrypta chrysanthemifolia var. sweet eucrypta HYDROPHYLLACEAE v chrysanthemifolia n Festuca elmeri Elmer's fescue POACEAE v n Festuca microstachys three-weeks fescue, small POACEAE v fescue i Festuca myuros rattail fescue POACEAE v n Festuca octoflora sixweeks fescue POACEAE v i Festuca perennis ryegrass POACEAE v n ❀ Frangula californica subsp. californica California coffeeberry RHAMNACEAE v n Galium andrewsii prickly bedstraw RUBIACEAE v n ❀ Galium aparine cleavers, common bedstraw, RUBIACEAE v goosegrass n Galium californicum subsp. flaccidum California bedstraw RUBIACEAE v n Galium porrigens var. porrigens climbing bedstraw RUBIACEAE v n Garrya veatchii southern silk-tassel bush GARRYACEAE v i Gastridium phleoides nitgrass POACEAE v i Geranium rotundifolium round-leaved geranium GERANIACEAE v n ❀ Gilia achilleifolia subsp. achilleifolia California gilia POLEMONIACEAE v n Gilia clivorum purple-spotted gilia POLEMONIACEAE v n Gilia tenuiflora subsp. amplifaucalis trumpet-throated gilia POLEMONIACEAE 4.3 v n ❀ Grindelia hirsutula California gumplant ASTERACEAE v n ❀ Hazardia squarrosa var. squarrosa large-headed sawtooth ASTERACEAE v goldenbush i ❀ Helminthotheca echioides bristly oxtongue ASTERACEAE o n ❀ Hemizonia congesta subsp. luzulifolia hayfield tarweed ASTERACEAE o n ❀ Hesperocnide tenella black-haired stinging nettle URTICACEAE v n ❀ Heterotheca grandiflora telegraph weed ASTERACEAE o n Hieracium argutum southern hawkweed ASTERACEAE v i ❀ Hirschfeldia incana short-pod mustard, perennial BRASSICACEAE v mustard Scientific Name Common Name Family Rare n ❀ Holodiscus discolor ocean spray, creambush ROSACEAE v n Hosackia crassifolia var. otayensis Otay Mountain lotus FABACEAE v i ❀ Hypochaeris glabra smooth cat's-ear ASTERACEAE v n Juncus patens spreading rush JUNCACEAE v n ❀ Keckiella cordifolia red climbing-penstemon PLANTAGINACEAE v n Kopsiopsis strobilacea California ground cone OROBANCHACEAE v i ❀ Lactuca serriola prickly lettuce ASTERACEAE o n ❀ Lagophylla ramosissima common hareleaf ASTERACEAE v i Lamarckia aurea goldentop grass POACEAE v n ❀ Lathyrus vestitus var. vestitus Pacific sweet-pea FABACEAE v n ❀ Lilium pardalinum subsp. pardalinum leopard lily LILIACEAE v n ❀ Lithophragma heterophyllum hillside woodland star SAXIFRAGACEAE v n Logfia filaginoides California cotton-rose ASTERACEAE v i Logfia gallica narrow-leaved herba impia ASTERACEAE v n ❀ Lonicera hispidula California honeysuckle CAPRIFOLIACEAE v n Lupinus albifrons var. abramsii Abrams's silver bush lupine FABACEAE v n Lupinus bicolor miniature lupine FABACEAE v n Lupinus cervinus Santa Lucia lupine FABACEAE 4.3 v n Lupinus concinnus bajada lupine FABACEAE v n ❀ Lupinus hirsutissimus stinging lupine FABACEAE v n ❀ Lupinus succulentus arroyo lupine FABACEAE o n Lupinus truncatus chaparral lupine FABACEAE v i ❀ Lysimachia arvensis [Anagallis arvensis] scarlet pimpernel MYRSINACEAE o n ❀ Lysimachia latifolia [Trientalis latifolia] Pacific starflower MYRSINACEAE v n ❀ gracilis slender tarweed ASTERACEAE v n ❀ Madia sativa coast tarweed ASTERACEAE o n Malacothrix floccifera woolly-tufted malacothrix ASTERACEAE v n Malacothrix phaeocarpa Central Coast malacothrix ASTERACEAE 4.3 v n ❀ Marah fabacea manroot, wild cucumber vine CUCURBITACEAE v Scientific Name Common Name Family Rare i Medicago polymorpha California bur-clover FABACEAE o n Melica imperfecta little melic POACEAE v i ❀ Melilotus albus white sweet-clover FABACEAE o i ❀ Melilotus indicus annual sweet-clover FABACEAE o n Mentzelia micrantha chaparral blazingstar, San Luis LOASACEAE v stickleaf n Microsteris gracilis slender phlox POLEMONIACEAE v n ❀ Monardella villosa subsp. obispoensis Obispo coyote-mint LAMIACEAE o n ❀ Nemophila menziesii baby blue eyes HYDROPHYLLACEAE v n Notholithocarpus densiflorus var. densiflorus tan oak, tanbark oak FAGACEAE v n Osmorhiza berteroi mountain sweet cicely APIACEAE v n Osmorhiza brachypoda California sweet cicely APIACEAE v n ❀ Paeonia californica California peony PAEONIACEAE v n ❀ Papaver californicum fire poppy PAPAVERACEAE v n ❀ Pedicularis densiflora Indian warrior OROBANCHACEAE v n Pellaea andromedifolia coffee fern PTERIDACEAE v n Pentagramma triangularis subsp. triangularis goldback fern PTERIDACEAE v n Phacelia cicutaria var. hispida caterpillar phacelia HYDROPHYLLACEAE v n ❀ Phacelia distans fern-leaved phacelia HYDROPHYLLACEAE v n ❀ Phacelia grisea window phacelia HYDROPHYLLACEAE v n ❀ Phacelia imbricata subsp. imbricata wide-sepaled rock phacelia HYDROPHYLLACEAE v n ❀ Phacelia viscida var. viscida sticky phacelia HYDROPHYLLACEAE v i Phalaris aquatica harding grass POACEAE v n ❀ Pholistoma auritum var. auritum fiesta flower HYDROPHYLLACEAE v n Phoradendron villosum subsp. villosum oak mistletoe VISCACEAE v n ❀ Pickeringia montana var. montana chaparral pea FABACEAE v n Pinus attenuata knobcone pine PINACEAE v n Pinus coulteri Coulter pine, big-cone pine PINACEAE v n Pinus sabiniana foothill pine, gray pine PINACEAE o Scientific Name Common Name Family Rare n ❀ Plagiobothrys nothofulvus common popcorn flower, rusty BORAGINACEAE v popcorn flower n Platanus racemosa California sycamore PLATANACEAE v n ❀ Platystemon californicus cream cups PAPAVERACEAE v i Poa annua annual bluegrass POACEAE v i Poa bulbosa subsp. vivipara bulblet bluegrass POACEAE v n Poa howellii Howell's bluegrass POACEAE v n Poa secunda subsp. secunda one-sided bluegrass POACEAE v i Polygonum aviculare common knotweed POLYGONACEAE o n Polypodium californicum California polypody POLYPODIACEAE v n Polypodium calirhiza nested polypody POLYPODIACEAE v n Polystichum imbricans subsp. curtum narrowleaf swordfern DRYOPTERIDACEAE v n Populus trichocarpa black cottonwood SALICACEAE o n Prunus emarginata bitter cherry ROSACEAE v n ❀ Prunus ilicifolia var. ilicifolia holly-leaved cherry ROSACEAE v n Prunus virginiana var. demissa western choke cherry ROSACEAE v n Pseudognaphalium beneolens fragrant everlasting ASTERACEAE v n ❀ Pseudognaphalium californicum California everlasting ASTERACEAE v i Pseudognaphalium luteoalbum Jersey cudweed ASTERACEAE v n Pseudognaphalium microcephalum white everlasting, small- ASTERACEAE v headed everlasting n Pteridium aquilinum var. pubescens bracken fern DENNSTAEDTIACEAE v n Pterostegia drymarioides notchleaf POLYGONACEAE v n Quercus agrifolia var. agrifolia coast live oak FAGACEAE v n Quercus berberidifolia scrub oak FAGACEAE v n Quercus chrysolepis canyon live oak, gold-cup oak, FAGACEAE v maul oak n Quercus durata var. durata leather oak FAGACEAE v nn Quercus wislizeni var. frutescens shrubby interior live oak FAGACEAE v n ❀ Rafinesquia californica California-chicory ASTERACEAE v n Ranunculus hebecarpus slender annual buttercup RANUNCULACEAE v Scientific Name Common Name Family Rare n Rhamnus ilicifolia holly-leaved redberry RHAMNACEAE v n Rhus aromatica skunkbush ANACARDIACEAE v n Ribes amarum bitter gooseberry GROSSULARIACEAE v n ❀ Ribes malvaceum var. malvaceum common chaparral currant GROSSULARIACEAE v n ❀ Ribes sanguineum var. glutinosum pink-flowered currant GROSSULARIACEAE v n ❀ Ribes speciosum fuchsia-flowered gooseberry GROSSULARIACEAE v n Rorippa curvisiliqua western yellowcress BRASSICACEAE v n Rosa gymnocarpa wood rose ROSACEAE v n ❀ Rosa spithamea coast ground rose ROSACEAE v n ❀ Rubus ursinus California blackberry ROSACEAE v i Rumex pulcher fiddle dock POLYGONACEAE o n Salix laevigata red willow SALICACEAE o n Salix lasiolepis arroyo willow, California SALICACEAE v pussywillow n Salvia apiana white sage LAMIACEAE v n ❀ Salvia columbariae chia LAMIACEAE v n ❀ Salvia mellifera black sage LAMIACEAE v n ❀ Sambucus nigra subsp. caerulea blue elderberry ADOXACEAE v n Sanicula crassicaulis common sanicle APIACEAE v n ❀ Scrophularia californica subsp. californica California figwort SCROPHULARIACEAE v n Scrophularia californica subsp. floribunda California figwort SCROPHULARIACEAE v n ❀ Scutellaria tuberosa chaparral skullcap LAMIACEAE v n Senecio flaccidus var. douglasii shrubby groundsel, threadleaf ASTERACEAE v groundsel n Silene multinervia S. Watson multinerved catchfly CARYOPHYLLACEAE v i ❀ Silybum marianum milk thistle ASTERACEAE o i Sisymbrium officinale common hedge mustard BRASSICACEAE v n ❀ Solanum douglasii Douglas' nightshade SOLANACEAE v i Sonchus asper subsp. asper prickly sow-thistle ASTERACEAE v Scientific Name Common Name Family Rare i ❀ Sonchus oleraceus common sow-thistle ASTERACEAE v i Sorghum halepense Johnson grass POACEAE o i Spergula arvensis corn spurrey CARYOPHYLLACEAE v i ❀ Spergularia rubra common sand-spurrey, red CARYOPHYLLACEAE v sand-spurrey n ❀ Stachys bullata coast hedge-nettle LAMIACEAE v n Stachys pycnantha short-spiked hedge-nettle LAMIACEAE v n Stephanomeria elata Santa Barbara wire-lettuce ASTERACEAE v n Stephanomeria exigua subsp. coronaria white-plume wire-lettuce ASTERACEAE v n Stipa lepida foothill needlegrass POACEAE o i Stipa miliacea var. miliacea smilo POACEAE v n Streptanthus lasiophyllus California mustard BRASSICACEAE v n ❀ Symphoricarpos mollis creeping snowberry CAPRIFOLIACEAE v n Tauschia hartwegii Hartweg's umbrellawort APIACEAE v n ❀ Thalictrum fendleri var. polycarpum tall meadow-rue RANUNCULACEAE v n Thysanocarpus curvipes subsp. curvipes common fringepod BRASSICACEAE v i Torilis arvensis common hedge-parsley APIACEAE v n ❀ Toxicodendron diversilobum poison-oak ANACARDIACEAE o n ❀ Toxicoscordion fremontii Fremont's death-camas, star- MELANTHIACEAE v zigadene n ❀ Trichostema lanatum woolly bluecurls LAMIACEAE v n Trifolium ciliolatum foothill clover FABACEAE v i ❀ Trifolium hirtum rose clover FABACEAE v n Trifolium microcephalum miniature clover FABACEAE v n ❀ Trifolium willdenovii tomcat clover FABACEAE v n ❀ Typha latifolia broad-leaved cattail TYPHACEAE o n Umbellularia californica California laurel, California bay LAURACEAE v n ❀ Uropappus lindleyi silverpuffs ASTERACEAE v n ❀ Verbena lasiostachys var. lasiostachys western vervain VERBENACEAE v n Vicia hassei slender vetch FABACEAE v Scientific Name Common Name Family Rare n ❀ Viola pedunculata Johnny jump-up, wild pansy VIOLACEAE v n Woodwardia fimbriata giant chain fern BLECHNACEAE v