a

eqt ]D pe^JeseJdere suaulceds ..p[,{\,, s.u,{\org qBnoqUV (]rsr^ uElluJlsnv s.u,{\org Jo s[E]ep rol '0961 '996I 686I acucllE^puu S86l ,{eFaqqutrJ uruels a8prqrngees) 90g I ol I0g l uro{ prlB4snvol 'lsEel 'tErlt 1rsr,lsrq 3ur:np pelcelloo aq qJrq.\\IErJol€ur ,lplr.r,, pesnosle eq tE sesucettlos ur olqpqoJd ',roleq s(Ueaslr pessncsrpeq IIr,r sr'Je,\e,roH suorlduJsapsrq e:uderdol sueurrcedspele,\Dlnc '(s861 pesn u,\\org tuql asoddnsot elq€uosEersl ll ,{eFaqquyqapl 'pue18ugu:eqlnos ur ereq,\\esle 'u?pruC put:) ,tra) cruulog 1u,(oaeqt le uortE^rtJncur stupld JoI suorldr:csappaprloJd ..srsue,re) '(I6Z snuoH,, sV pes€qeJe,r ,{eqt|?uetuul trq,r uo u,{\ou{ lou sr,r tr esneceq,tlurDur :g/6I utlsptr{ '86t pur 9Zn-57,V :SL6I utl'Suylapll) sauEu aseqtJo €ruos3ur,{grd{1 ur lsed eqt ur perelunocueueeq 'DJoulZJDru 'DpnlJ 'D.rotlq e^Eq sertlncuJrc DtD)lns V pux D aqJlnd V'uolLrouDpw V V V V 'DJDuiJo ',{leutEu',reu V'DiDlD'V'sttolnJlJ, y sp pequcsepe.rs,{\ urll?ltsnv ulog s?rJeds?uru ..srsua,\\e)snuoH,, s,uolrv Io Z uoqrpaur u,rorg ueqog {q pelueseldDrJrJyJo saroedsEg eq} JO

uo!lJnporlul

'Jod€d 'JE'' snor,\erdE ur peryrd,{1oauuaaq pvq'A)uoluaoJq DuDluaotq V=) Lijt2ltlJ V 'serJeds .V euo NSCI te uorlcelloJqBNcI trruBIllrIN eqt ur luuld pele,trllncu uo p"rJrdhoeu s ototuto olrq,{\ suortcelloc .,plr^\,, s,u,roJg ruo4 u€soqJ ueeq spq (ltuttio.lq V=) suoln)DD y roJ ed,{loau 'Dtotlq 'V 'otop 'V B :oiorlns V puE qpqrlnd V'uolt:roLtrypw V'(Duoluttw V=) DwalSrow V :lurJelE[l .,plr,r,, s,u,roJgJo srsEq3q] uo pegrd,{lolceleJeq oJr serupuserceds u,rolg Sur,rolloj ?qJ '(NgC) urlqnC le lunr]Dq]eHIe uortEN eql ^q peturnrsr uotcallof, slr0 :60gI pul]909I uoe^\]eq,{\a) lE sluuydpele,lqlncruo4ser3eds,/JrJys,u,\\orgJosueurrcadsepeuroq^'^\o)tureuep:u8e'qe1qc1,rtr 'Ja^a,\\oq'sr urErllrad,{q uorlcelloJ c eleql lurrelurued,{1 su pepluBaJaq upJ qcrq,r luelx€ sueulceds ',{lpuoJos petu^Dlnrou a:u aroqtturp sJpeddr tr pupl8ug ur uorlurrllnc ur sluelduo;; lo/put 'g0gI puu prleJtsny l08l ue?^\leq ruo.qsSuueqluB .,p1,lr,, s.u,rorg a r'p3seq ale,r,{eqt qcrq^\ uo I€rJelurueql Surp:ufai seDululrasunaJad\ eJeql 'flslrC suostel umul o,\\1ro; sualqord peluese:dsuq sela?ds eseql 'ts€d .+ouorluorJrd{t eql uI .(srsue,{\e)snuoH,,Io uoqrps puoces s,uotrv'Jd\ ur,?rrJyJo ssrJeds ,reueuru peqsrlqnd u,{\org ueqou (S661)gIl-f0I:(I)0TDrsl,{nN,,srssa,$e)sntroH,,s.uotrvIoZ 'U ,urlsEtr uorlrpeu l s?urBurrrDJy,reu srqJo uonErrlrd{taqt u,nolglrJqou ue,roJ S d puE g tr

lJBrlsqv

zsl9 lrllcllsnv urelso^l orro:) tol xog od 'unurqlaH lucue8ruxw pur-I pueuortD^rssuoJ Jo tuoulrrdec urrpltsnv uJels.^\

uB,roJ s dpue urlsulr'{ug

.(srsuaa{.aysnFoH,, s.uo}rY .u,rorg Jo z uolllpe ur sarrrBum)Drv /Jreuslq Jo uorl€)!JJd,{1eq1 lraqog

(S661)8 10t (l) ut nr!i{nN NuytsiaVol. 10.No. I (1995)

Natural History Museum, London (BM) and elsewhere(lide Powell & Morley 1976),there seemsto be no surviving cultivated material that can definitely be said to have been used by Brown for the "Hortus Kewensis". There is, however, a hithedo little-known collection of plants madeby William McNab from the plants grown at Kew Gardens. The collection, which is housed at the National Herbarium, Glasnevin, Dublin (DBN), is valuablebecause it containswhat appearsto be the only specimensofthe cultivated plantsBrown described,and assuch they help to identify the taxa, aswell as being potentially availableas neotypes.

The purposeof this paper is to determinewhat Acacla specimensRobert Brown usedto compile "Hortus his descriptionsof new speciesin Kewensis", and to selecttypes from among them This task has been facilitated by an examinationof Robert Blown's unpublishedmanuscdpt descliptions "wild" "wild" at BM of both andcultivated plants, and his collectionof specimensat BM, andMcNab's cultivated matedal at DBN.

Robert Brown and the "Hortus Kewensis"

"Hortus 1. Brown's involvement in Aiton's Kewensis" ed. 2

.,Hortus Aiton's Kewensis,,was publishedin two editiors, the first in 1789in tbree volumes and "Hortus the secondbetween 1810-1813 in five volumes. Britten (1912)remarked that the title, Kewensis,,,was misleadingbecause the work treatedplants cultiYatednot only at the Royal Botanic Gardensat Kew, but also at Chelsea,Upton and Islington gardens. Indeed, as noted by Mabberley (1985), this work was really a guide to the plantsin cultivation in southernEngland at that time. This "an is not at vadance with the prefaceof edition 2, where the purposeis describedas attempt .... to trace back, as far as possible,how long eachplant hasbeen cultivated in the British gardensand to fix ...... rhe epoch of ils introduc(ion.'

"Hortus The secondedition of Aiton's Kewensis" was edited initially by Dryander but after his task was assumedby RobertBrown. The natuleofBrown's involvementwas that ofsoientific deaththe "I editor, rather than the pre printing sort ofediting. In a letter to de Candolle in 1817,Brown wlote must take this oppodunity ofstating that I am neitherdeserving ofpraise or blame for any pafi ofthat work unlesswhat I put my name to ... the Manuscript was preparedfor publication befbre it came into my hands, and all I had time to do was to attempt to saveit tiom very grosselTors & to add a certain porlion oforiginal information in eachofthe 3last volumes [i e commencingwith page 175 of vol.31."(Mabberley 1985).

"Hortus 2. Authorship ofAcacia names published in Kewensis"

"Hortus Aiton, in a postscripton pages531-532 of volune 5 of Kewensis"ed 2, statedthat not all of Brown's contributions to this work are indicatedin the volumes;however, those that ale bear "Brown the notation mss". In the caseof Acoci0, this notation appea$ at the end of the desoriptive text of each of the nine new species,as well as at the end of three of the four new combination tllat were efTected,namely, A. decipiens(Koenig ) R Br.' A. nigricans (Labill) R Br' and A' sophorae (Labill.) R. Br. That Brown was indeedthe author ofthe nine new Acacia speciesis supportedalso by the factthat in J.J.Bennett's ( 1867:471-510) publication of Brown'smiscellaneous contributions he includedthe descriptionsof thesespecies. Bennett was in a peculiarlygood positionto know DreciselvwhatBrown had contributedbecause he wasBrown's long-timefriend andpelsonal assistant a

'seuo peqsqqndsrq qlr.n suoqducsep ldrrcsnurur s,u,{\olg qclyur ot sn .ro;f;essaceu ueeq sEq lr uell g ro uA\org reqlre ,{q pepr,tordtou ara,r saureusarceds paqsllqnd eqt areq,\\se.uelsur uI auuu sarcodspaqsrlqnd eql qlr,t suo4dr.rcseplduJsnu€ur eql petElouuu 'sescc suq (uetlrrg {lqnunserd) auoeruos ,4uuuru1 lunoccepeqsrlqnd eql ur asoqtuorJ }uoJaJJrpele ro luesqureqlre fllsoru eru sleqlrdeserceds eql pue ,.rorr?JryJo trpd sEpolte.u eru exet eqJ paqucsep 11]ueluureql Jo ecrnoseqt ol etuoleJor17 sapnlcur puE (suerurcedspal€Arllnc JoJ uortduasapIo stBp eqt ,{lquurnse:dpuc suaurroeds .,pp,r,, :o; uortcollocJoelep eqt) pelcp fllunsn sl uorldu.sopq.ug

tu?ld p?l€^rllnc E.ro!.plr,r,! e ol sJeleJuoqdr:csep u,troul ,{1uoaqt:aqtarl,\\ tll sr?leco,lrnba sr 1rrioJ,ttouopw V JoI eltqt :suortduoseplucld pelu^qlnc Kluooylaqtlndypue o|olllJ V'oJoLfiqyrol:uonduJsep luuld..plr,t,, u,{1uo sl-ro|nctco y to1'.s1ue1d 'DiDwD 'DlDlD pelu^Dlncpur? .,pll,{\,, t{loq fo suottducsepeJE eJeq! olDJlns V pul p7ou!8tDtuV V V :o; :suoqdrrcsep;oslas qloq ur poluas3tdate::,l- onoty Jo serceds,'rleueuru )^.u.{ orgJo llBlou tuql ,noqsytrgtusldlrcsnuulx're^a,roH.,plr,r,,eqluroJ.+peesurorSu,ror8sluvydpets^qlncuopesuq'€xel euus aql eq ol pe^artaqeq teq,rJo suotdursep;o 1esreqtouu pe:rda:d puulSugot guruJntoruodn puu pe1celloceq sercedseql Jo ,{ueruJo suonldursepporuderd u,roJg BllersnV ur s1e,rc:1srq Suunq

zreoey;o suorldl.rcsapldlJrsnuuur s.u,{roJB

'.,,{qqnJqS,, sE,r pue .(asnoHueerD,, e ur u,rol8 sE,^q.pa tBr{tSurtouap sloqur,{s puv 'poued Suue,nog aql 'sercadseql ro; etuuuuoururoc eql pepnlcuran3o1o1o:d aqt ur uoquurJoJur Jeqto 96/l uI peanpo4ur,{lsnour,(uouu su,n 1r suDln)eo y .roJpua reII€iN uqof ,{q,.808I tnoqu,, sc,('lt uolLrouDlala y JoJtnq pooC Joled ^q t08I sE,r s[0 sesEclsoru ur :elqrsuodseruosred eqt Io ?urcu eql qlr,^\Suop ue,lr8 osle se,r uoDuArlJncolur uorlcnpo.uursll Jo etcp oqt sercadsqJEo.loJ

'(suDlnJIJD yroJ).,uosratEd rur?rllrlAleuoloJ seleld qlnos,&eN,, pur (uo{,rouDp.a V roJ) '(Dtptlns ..puBlsls.ueuerc uuL,,'(otDwto y JoI)puulloH,reNJo tsuoc qtnos,! Vpueo aqtlnd V 'otoul8tD1u 'DtDllD 'Drotl[lq 'o1o1o V V V y to1)..puplloH ,reN Jo lsEoctse,r-qtnos,, :e:e,nsarcads ,{\eueuru eq} roJ ue^13selrs 3ur1ca11oc rnoj eq1 sercedseqt;o tdacuo. srq ol patnqgtuoceauq flelns tsnu luuetcurJo spuqt1;oq'DrcLflq y;o sdeq;adpuu a1ru1rs y,1o uoqdrrcsepaqt ur ,{1uope,t1o,r.ur a:a,n suerurcedspet"^Dlnf, puE..ptr,{\,,qtoq lEql ull?uec,{lquuosue: oq uPc e,{\alrqlA sen8olotold aql Surruderdur suortcelloJ ..pll^\,, s1qtrro4 rlup esn prp oq tuelxe teq,r ot ssesspol Surldueuu ur seuroJftlncuJrp €qI ,{lluenberJur,{luo s}uaullpoJt,,srsue,re) snuoH,, eqt ur suoqcelloJ .,pll^\,, stq uro:; ,{pca:rpuoqpuJoJur pesn eluq o} sJeedd€u,llolg',troleq suorssn3srpsarcads eql ur luepr,te 'Ja^e^\oH euocaq llr,{\ sB uoDetrerdreturrulrurrs € peldope(0661) dsr.rJ:,,plr,r,, eql ur sercedseseql patJellocu,l\org lEql uueurot oraqpateJdretur sr srql ,,bsg'u'notg yaqoy,, uoqulou aq1,(q pe,trollog 'slJDlliJtJD y JoJlde3xa'pu€ ue,tr8se,tr serceds qcue;o ecuuua,lordeql u,roJg Jo l€rll lJeJ ur su^\ uorlnqrjtuoc eql lvql seleJrpur'pelouFpueJlE sE'qcrqn\.,ssu u,roJg,, uorlEloueql , q pe,r,r.o11oysv.n 'rlrJ1r.r 'y 'y uoDducsepqceE l€uatuurSuue,\\ou uo ^lolospospq uaaq e^€q ol tuees puuuolikouolaw 'puE ;o uoqdecxe eql q1,,'\ Jeuq ,{ra,rare,,n oep)V Jo s€rceds,. eu euru aql Jo suorlducsepeq1

..slsua^\axsngoH,, ur rzrrnyJo suogdgrsaps,u,uo.rg'g

saruBurrrrry s,u^\orglaqou ur^\o:) sdpue urtsgw xg NultsirVol. 10.No l(lqqs)

The specimens

1. Robert Brown's collection of "wild" specimensat BM

As alreadynoted RobertBrown wasin Australiafrom l80l to 1805,during which time he collected assiduouslyin rnanyparts ofthe continent. Although many ofhis bestspecimens fiom the southcoast of Austnlia, and all the living plants, were destroyedwith the sinking of the Porpoise, he reached Englandwithout havinglost any one speciesaitogether (Stearn 1960). Survivingspecimens were studied by Brown at the BM and after his deathwere bequeathedto his successor,J.J. Bennett,who stipulatedin his will thatthe first setshoulcl go to BM andduplicates to K andE. Edwards(1976) relatedthat JamesBdtten subsequentlycompiled a manuscriptlist of the species,ananged by the numbers which he assignedto each species;it is thesenumbers that are often erroneouslycited as Bennett or Brown numbgrs.

2. Is there a collection of cultivated plants used by Brown?

"Hortus Severalauthors have statedthat a collection of cultivatedplants was at BM on which the Kewensis" was based:James Britten ( 1905)in discussingthe importantcollections at herb.BM noted that "The most importantcollection ofcultivated plantsis, however,that from the Royal Gardens,Kew, which containsthe typesofthe numerousspecies described by Banks' librariansSolander and Dryander (helpedin the secondedition by Brown) in Aiton's HortusKewer'sis, the MS. originaldesariptions of theseand of a large number of other plants in the Sloaneand Banksianherbaria are preservedin the DepaftmentofBotany." Lasegue(18,15) in his classiccompilation ofinformation aboutcollections "An then in existcncestated (translation) herbariumcomposed of plantscultivated at the Royal Garden at Kew and descdbedh the Hoftus Kewenslsis found in a specialAiton collection which is part of the Banks herbarium". Bdtten (1912),however, pointed outthatLasegue was not altogetheraccurate, "for 'carry Aiton did not possessa herbadum,it being his custom,as hasbeen already stated (p.3), to his specimensand doubts to Banks' library, where they were examinedand resolvedby Solander." "The Furthermore,Stearn ( 1981)notes that plantsraised at Kew from their seedneeded identification, which could only be done by a botanistsuch as Solanderwith both Banks' herbariumand rich library at hand. Thusspecimens grown at Kew likewisecame into the Banksian herbarium." He alsostates: "The typesofthe HortusKewensis are not at Kew, asis ofien expected,but in the DepartmentofBotany at the Natural History Museum, having come with the Banksian herbarium."

Notwithstandingthe precedingconments, we havebeen unable to locatecultivated material at BM that originated tiom Kew Gardensand formed the basisofRobefi Brown's treatmentof Acdci./.

3. William McNab's collection of cultivated specimensat DBN

An importantand until recently,little-known collection ofplants in theW.R. McNab collectionat DBN hasrelevance to the typificationof Brown'sAcdcia names (Nelson 1990). W.R. McNab was professorof botany to the Royal College of Sciencein Dublin and Scientific Superintendentof the NationalBotanic Gardens, Glasnevin. He wasthe grandson of Wiiliam McNab who wasa gardener at the Royal BotanicGardens, Kew, fiom l80l until l8l0 (Nelson1980). The Acacla specimens at DBN werecollected by William McNab (Nelson1990) between 1806 and 1809from plantsgrowing at Kew Gardensand included the following species,A. dlatd, A. armd:a (= A. paradoxa),A. bifktra, A. ciliata (= A. browniana vr. browniana),A. mqrginqta(= A. ntrttfolie), A. pulchella and, A. .sulcqta. These plants were probably grown from seedsupplied by Peter Good around 1803 and "Hortus ". were the plantsupon which RobertBrown baseddescriptions published in Kewensis a

'i^tuuld polr^rtlncJo suorldr:osep1dr:csnuuru srq uo en8o1olo:d ...srsue,lra) ?ql lo txot ?Auducsepeql pesuq^llvnuou u,rolg snUoH,,ur paqrJrsepserceds nlca:y ieqto.+osuorssnJsrp:no{qpeluorpurstst:,esne:laqFulsudlnstouslslqJ.pelrerlsqest,ruorldr:osap .(s)ueurrceds peqsrlqndeqt q.rq^\ tuor1srssq eqt eq ol rraddulou seoptl lnq Buue,\\ol+sequJsep pur petup .srJulncrrE lou sr llluelBut,,pll^\,, Jo tunoJcEsrqJ .,,{ue}og? ,{eup,{g:elur €so(Uttrl,, peltlouuBsr lr pueJ trglusltoltlJtJD y;o uor;dr-rrs:p tduJs n u n ul u,rolg ueqog euo,{1uo sr ateqa

'NgCI tB pat€col ua3qlou e^uq suepr€C,re) tp petc^rllnc sercedssrqt 1o stueld urorgsueulceds '..srsuo,re) .(en8ololoJd snuoH,, ur u,^\org,{q pequJsepDlrD)V lo sarceds:eqto tsoul elrlull eql ur snorqBlSslelqcuutq) sacrdelelqcue:q eqt ol peuguo3eJE q.rq,r slruq etnuruIJo urnlueunpur es.reds,{ro^ u Sur,requr ,{1uoenSoloto.rd oql u-rorJtaJJrp sueuuoads esaql .,.tggl .tco ,uos{cBt tlod'BrloJeetrua Esorurw,,se u,ro:g fq pelleqBl sr leeqs slqJ tln4 Buno,{f:e,r ur euo pue ro,^\ou 'sarceds ur requnu u eqt Jo s8urreqlu8,.plr^\,! s.u,rorg Jo sueurrcodsBuuueq lE teeqsu sr eteql 'Je,\e,lroH trlu I Ig le uosratudur€rTlri&,{q petcelloJ 1er:a1eur,{uee}pJol ot elqp uaeq tou a^uq ed\

'961I ur 'pur]l?uA [,{tsnoul,{uouB olurl pecnporluruoeq p€q sercadseqt luql pruspue..sal8lA qtnos ^\aN,, tuo4 petcellotrpDq uosrattdruurllrlAleuoloClEqllEuolptuolpeleJoroslEeH.uoqducsepaqlot,.ssuru,!\oJg,,uoqptou eqt papuedde pue uaurceds Suue,lrogr pequcsspu,rorg sln)ln)Do alrrry .;o en8ololord aq1u1

'(hlg) !esuarlErlsny 00gt I ou rro]tugl SuuseqpuE ,,9,20g 1 rell, pe0rl loeqs uo ueurrceds pueq - '€0gl .uos{oul ...€rloJeBJr.re lq8r: lEJtueJ u otg U tco'd|S N uod esoruttrl,, :(ueuljceds ,upe .uotrv .I.,^& .rg.d Suue,rou)ed,(loeN (€lgt) 09t:9 puz ,\\e) sntroH ul srrBlnJIJGBITETV

'(9/61 spr?,\\pg)neuueg t I ot pequcsB,{lsnoeuoxo ueeq ue]Io 'r{Jrrl,\\'ua1rJfl e^eq'e^oqe pelou sE setupl ,{q pexrllt esoqte.tu,roleq pelro s]oqunu taeqsoql

ygq tu sueurrceds q"Ncl4l eqt 'ed,{loau .}srxe .Z ruor; ,{11unsn E patJal?se^Eq e,r ot u,rou{ sr luu?tpur ..pp,{\,,ou JI

'serceds oql Jo lsoul jo ldacuoc crurouoxelstq o1,{1pue1eur polnqrJluoc puelBru ..plL\1,,aql leql ucaruot uo{pl sr suoDcallocuErtllJlsnv sq pelrceq lpqt t?"J eqt }nq I€r.ratelllpetu^qln:) ,lred uo ,{1r:uurudpeseq are,tr suorldr:lsep ..srsua,re! snuogj, s.u,{\org }sour Jql toC eutpu aql.+o edtr;o1celeq1su 1r pelceles e^Bq a,r'enSolotord oql qlr^\ sealSesrqlJt puE.uoxul eqlJo eur€upeqsrlqncl ,ro . lducsnuerueql q1r,r u,l,io.rg,{q petu}ouupsl tuqt (Wg 1e)stsrxe uerulceds ,,plr,u,, elenbepu uu j1 1

:pelou eq plnoqs reded luese:d eql o1rulncq:ed 'uorssnJstp ,le^a,roH .(deld ,uE,roJ suoqereprsuocSur,rolloJ eq] a^oqu aql Jo ,^\or,\ur ur d) urls€tr^{) raded Sururocquo;e ur peurllno eq l1r,nsedf1jo uortceleslno Butule,\o8seldr:tuud 1u:eue8 eq1

sausu s.u^roJg Jo uorlucglddl

'ranuollJDl DwaslblrrrA roJ (066I) dsuJ ,{q euop sum su 'ad,{loeu 'V e supelceye s sr (DlDutto Io t9qt)asel euo ur pue enlu^ crJolsrqe,{aq op sueurceds eql 'Je,\e,roH Ng( pueluur ed,{1put8r:o peraprsuoceq louurc suerurcedseseql uoseoJsrql loC ..,srsue,{\e) sn1roH,,ur DtJDxVJo suoqdrrcseppeqsrlqnd eq1_ro suorlducsep ldrrcsnuuur eqt teqtre aruderd o1 suettlrc?ds ,Jo^e,roH NgC oql pesnflpnlJB u,$olgtuql eJuepr,\eou sreleql.,&oloq pessncslp eq IIr,r sE

.u seuculrrrry sur'olg uaqod uu^\o] s u pu} urlsvl{ g 1t2 NuyrsirVol 10.No. t (1995J

In the absenceofboth Patemonspecimens and ofmaterial fiom cultivatedplants it is necessaryto selecta replacementtype for the nameA . .tcicularis. Accordingly, a neotypehas been selected from "wild" among Brown's gatheringsfrom the samegeneral area cited in his manuscriptdescription of "wild" material. The slight difference between the n€otype spccimen and the protologue can be explainedby assuDringthat Brown overlookedthe sparse,rninute hairs at the apicesofthe branchlets.

Acacla acicularisis the basename ofA. brownil (Poiret) Steud.and the complex nomenclatureof this speciesand its very close rclative, A. ulicifulia (Salisb.)Court, is discussedby Court (1957 & 1972;155-156) and by Pedley(1980:239-214).

Acacia alata R. Br. in W.T. Aiton, HortusKew.2nd edn,5;464(1813). Lectotype(here selected): "Mimosa platycaulis." ObservatoryHill, PrincessRoyal Harbour [Albany, W.A.]. In ccllibus saxosis solo saxosoprope littorae Poftus Regis Georgii III in ora australeNova Hollandia,Dec. 1801,R. Brown "Iter - upper right hand flowering specimenon sheettitled Australiense,1802-5" and bearing[Britten no.l 4314(BM); isolectotypes:K,E. Paralectotypes:Sterile and fruiting specimens mountedon sheet with lectotype(BM, K, E).

Note: In the protologueBrown describedflowering material and appendedthe notation "Brown mss" "South-west to the description; he also referred to material he had collectedfrom the coast of New Holland", and said that the specieshad beenintroduced finto England] by Peter Good in 1803.

At BM the only Robert Brown specimensseen are those cited above which representBrown's "wild" gatheringsofthe species.The specinenscomprise flowering, fruiting and sterileelemcnts and the sheetis labelled by Brown as Minora platycaulis. The DBN material of this speciesconsists of flowering and fruiting specimens,both I abelledAcqciq aLataby McNab. They were collectedin 1808 liom plants grown at Kew but there is no evidencethat Brown actually saw this material. Both these "wi1d" and cultivated specimensrepresent the sametaxon.

There are two Robert Brown manuscript descriptionsof A. qlata at BM, both under the name Mimosa pla4caulis. One describesboth flowers and fruits ofthe "wild" gatheringsreferred to above, while the other describesonly a flowering specinen in cultivation at Kew in 1806.

From comparing the two manuscriptdescriptions rvith the protologueit seemsprobable that the "Hortus Kewensis"account was abstracted from Brown's 1806description ofthe plantin cultivation at Kew. Althoughthe DBN specimensare from plantsgrown at Kew (collectedin 1808),there is no evidencethat Brown actually usedthese to preparehis publisheddescription. Therefore, because there is no cultivated material which can be regardedas a type o'i A. alata \\e have selectedthe lectotype fromamong Brown's wild' g:ttherings.

Acacia alata will be teated as comprising four varietiesin the fb hcoming "Flora of Australia" account.

Acacia armata R. Br. in W.T. Aiton, Hort. Kew. ed. 2, 5: 463 (l813). Neotype(here selected): "Acacia annata," Kew, 1809 [William McNab] (DBN, flowering specimen).

"Brown Note: In the protologueBrown describedflowering material and appendedthe notation mss" to the description;he alsoreferred to materialhe hadcollected from the "South coastof New Holland", and saidthat the specieshad been introduced [into England] by PeterGood in 1803. r

l1suas'D.tojq y truese:derpuu ,{u g ,{1cn1uorl Fur:eq1u8 ,,p1r,r,, s, u,ro.r g eq ot perunsscale ,eurlaql eqt ot'sueLrlo?ds .d JoUel reqloaqJ (puuqs.u,roJgur lou) ,..rg urol+rqV,,,{ld!urs polleqel sr puu urlsetrNaoulqot y st eurl eq1;o lq8u €ql ot uorurcodsaq1 eurl pallrlued c ,{q pelu:udessuortcalloc 3ur:o,nogo,n1 slroddns srql puu Wg )Eserrads srqt 1o ecuc:tr;ruBrs odfl;o leeqseuo ,{luo sr eleql

't08l uI pooo retad,{q [puplSugo]url psJnpolurueeq peq serceds 3ql )DqtprES puc ,LlpucttoH ^\eN JOtSr?OC lse,r-qlnos,, 3q1(uo! pelcelJoJpuq eq lurJaleuro] pelleJeJoslD oq :uorlducsepeql ol ((ssulu,ro.rg,, uorlplou eqt pepueddupuu lur.relcurBur.te,ro6 peqr:csep u,to:g en8ololo;d eq1u1 :e1otr1

uarl'sliuu'eq puB ,.s-208 r ,esuerr'rrsno,"u,, o"ou,""u,1"X"i}S.liii:Jii,,r, -tfi$liH;:J '2081 uDI '[gE6l e8prqrngapf!, y ,,{eg l L\.eruuredsg3ro A ,{).n-l] 1 ,{eg ,..uuo1erscsourrtritr,, :(pelool?sereq) ,Z.pe..ro) ecl{tolra-I(tt8l) f9t :S t.roH,uotrv J,1\ ul rg.d BrourqelJBrV 'u?soqr ueaq suq (rurq ^q patJ€[o. uaoqe^Bq 01 pa(InssB puu) qpNcj,a\truer11116 {q pelrlouuu uarurceds ',{lSurproccy .NgC Surre,nogeqt tu suaurrcedspe}u^BInc eql Io euo ,{qeureu stql ,{grd,{loau 'eluEu ol lsoqlr reprsuoce,nn lducsnuElu:opeqsrlqnd eqt ol sD.urq,{q pelDlouuu sr suorurcedseseqt .)oauou esnpJeqpuu 1419tr sBurreqle8.,p1r,r,, s,u,ro.rg Burp:nfer serluregacune:t; oteqt esnBOe{

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.y,sanads (DlDwD V u^s) oropotnd elquuu,t ,{1q8tqeues eql luaserdarsueullceds petB^plnJ pur,,plr^\,, oqt qtog l€uetBlusrql ^\DS,{Llyatru u,roJg trqt ecueprAe ou sr orsql lnq,{\e)I le urrror8s1ueld uo:3: 6081 uI petrallot ere,\\,{eqJ .quNtl,{,{q DlDwtDDtro)V pelleqtl qtgq 'ueurrJadsSuqrn{ E pu€ Bvrre,nogu 1o slsrsuocserceds slllt Io lprJaleruN{O ?rll .puu1s1 oo:e8ur;q:o e,to3,{;oruayJ ruo:g}ou lnq ,,IIA eFEJoqJuv,!uro{ DluisslsouldsDsowlWJo peas slsrl htrg le lducsnuuurs,poog:e1a4 tuql pu; a4 ,{q ue,lrBsr uorteterd.ratsr :^rql rol Uoddnsouros leq€t tioJJoJE lnoqlr,^\leeqs srql uo pelunoul uaaq seq uerurcedssrql teql alqrssodsurees puu ti uorlducsepsrql:o; qclerupoo8 e elrnbsr latqs ,..:u8uuy,,aqt uo ueutrcads tsourre,\\oleq]'le,^e,roH (,roleqaes ,.V ,sputlsl .e.r) S slepurtCpu? a,rejfp1e16 ,.JIAeBEJoqcuV,, ruoJjpaloelloo lBr.reluLu uo pesrq sE,rqJrq,i' (owtsstsoutdsosolulw sE)DlowJo .V,,pIr,tr,, go uoqducsap stq ,,{Jsnoun) .erun8el lducsnuuru ur u,\\olg ,{q polrt sr s?rltluJole,\oq8 eql Jo Jeqltou ei8uts e puu sJe,roU'spnq q1r,re:e leeqs,..ruBuu),, eqt uo uaurrJedstsourJe,{\oI eql pue teeqs.,y1 ,(ug,, eq1uo suerurcedsrnojBururtueJeqJ.selndrls,{urdsle,reg,tosluof,puusepoll,{qdpecuds{lepr^erorupuu 'stelqcup:q .Ie3[l paqqu f,lasreocBur,ruq ,{q sueurrcadsBurur€uror eq] uroq peqs,nEurjs,pem puu pnq 3uno,{ ,{Je^ur elu sueuuladsesaql leeqs ,, ru8uu;1,,eq1 uo uerurcedslsouuaddn eq1 rol qcluru poo8 B sr ,{p9,, eqt .n1owo .y teeqs,,XI uo ueurrcedspuuq lq8rr reddn eq1 1o s1|Jnqtu8 pexrurluesudar fpuoreddu steeqso,rt esoqt uo suerurcedseql ,pue1s1ooru8uel] ':eqdtcep [.V.S ,..rp8uu),, oq ot sJEeddB ol pnq qinoqllp'raqlo puE ,e^oJ.{rouel^I1 aqt l9S6l o8prqrngrpy',.V.S ,;r1,trg,, pe11"qe1 s^r euo ,slequl leeqs ..EI€t,,peJsq[rnu eJE qJrq,\\ Jo qloq .,3suerF.usnvr3]I u,roJg .u,, o1peqcutlu :edudlo sdrls uo pucq .uoxet 1lerus s.u,!\olg ur e:e saqqrcol BurloelloJaql srqtroJ pesneq lcql etupu peqsrlqnd:o eql reqlte .je^e,\\oq lducsnuEu qtr.{\u,rorg,{q p?lleqDIsr Jeq}reu :i^el]edsslqt 1o s8uueqte8 ,,plL1d,,s.u,{\o]g luaselder ,{1qe qord qcrq,r (DXopotDd.V =) DtDw.lDy Jo staeqso^\t ol1?a.roqt tr^{gtV t1r .u?^1oJ sett|rr,rr?,ry s,u,rorg]liqod s d pu]rurrsstr tr d g NuytsiaVol. 10,No. I (1995)

Iectotypico(see above).Thelabel accompanyingthese specimens is annotatedin Brown's hand. The DBN material of this speciesconsists of flowering specimens,both IabelledAcaciabifloraby McNab; they were collected in 1809from plants grown at Kew but there is no evidencethat Brown actually saw this material. This material reprcsentsA. robinde.

"Mimosa Thereis only onerelevant Robert Brown manuscdptdescription at BM. This is annotated biflora" and describesa flowering specinen in cultivation at Kew in 1807,grown from seedcollected "Port at King George'sSound" [Albany, W.A.]. Closescrutiny of this descrjptionleads us to the conclusion that the plant describedby B rown was likely A. robinae, a conclusionsupported by the fact that the DBN cultivatedspecimens are alsothis species.Moreover, the plant desctibedand illustratedas A. biflora by Wendland (1820) also seemsto be A. robinae, suggestingthat the misapplication was prevalentin gardensin Europe at the time.

The publishedoriginal descriptionol A. biflora, however,seems unlikely to have beenbased exclusively on Brown's manuscriptdescription of tbe plant cultivated at Kew. Indeed,following a careful examinationof all relevantmaterial we are of the opinion that elementsof both A. biflora and A. rctbinaeare included in the original description. As there is no cultivated material which can be "wild" regardedas type of A. bi;florawehave selectedthe lectotypefrom amongBrown's gathedngs

Acaciaciliata R. Br. in W.T. Aiton,Hort. Kew. ed.2, 5: 465 (1813).Neotype fide B.R.Maslin 1975: 425): "Acacia strigosaLink" (not in Brown's handand with no otherdetails) - lower lelt handflowering "Iter specimenon sheettitled Australiense,1802-5" and bearing [Britten no.] 4321 (BM).

"Brown Note: In the protologueBrown describedflowedng material and appendedthe notation mss" "South-west to the descdption; he also referred to material he had collected from the coast of New Holland",and saidthat the specieshad been introduced [into England] by PeterGood in 1803.

As discussedby Maslin (1975 425),the BM sheeton which the neotypsis mountedis very inailequatelylabelled; there is no indication of the origin of the (rnixed) specirnensit bearsand there is no direct indication that Robert Brown ever consultedthe material (as evidencedby the absenceof his handwriting). The rnaterialcornprises a flowering specimenofA. LuteoLqM^slrn,four flowering specimensand a fruitingspecimen ofA . brownianQH.Wendl. var. browniana, these two taxaoccur sympatrically at Albany, a locality visited by Brown in 1801/02. The lowermost flowering specimen on the sheetwas selectedas the neotypeof A. ciLiata by Maslin (loc. cil ). The DBN materiai is mountedon a sheetof flowering specimenslabelledAcacla cillala by McNab; theseate A. broh)nian.I vat. browniq a and were collected in Jll08 from plants grown at Kew but there is no evidencethat Brown actually saw this material.

There is one Robert Brown manuscriptdescription at BM which accordswell with the protologue "Mimosa" of A. ciLiata. It is simply annotated by Brown and describesa flowering specimenin cultivation at Kew in May 1806. Although this descriptionagrees quite well with the specimenselected previously as the neotype,it is impossibleto determinewitl't any degreeof certainty whether this is "wild" cultivated or material.

"Hortus From comparingthe manuscriptdescription with the protologueit seemsprobable that the Kewensis"account was abstracted from Brown's 1806description ofthe plantin cultivationat Kew. However, as there remainsso many uncertaintiesregarding the origin and authenticityof specimens at BM we are not able to improve on the original neotypitication. .eluDu :snqt u,^\ojg,{q palleqrl ole sualurcedsaseqJ srqtIo uorlucr;rd,{1 'r9€t ,htrg eql uo 3ur.Ieequ e,req,{uru qcrqm I ou lleuuag] patptouu€qtoq tu steeqso,{\t oJ€ eraq:L

'80i1I ur lellE/(\ uqol ,(q IpulrlSuao]url poJupoqurueeq prq sercadseqt teqt prespuE .,puElsI s.ueluarq .uoDducs3p uBA,,uro4 peltellocpEq eq leuetuulot perroJaloslE eH eqt ot..ssuru,rorg,! uorlutou papueddr 'sledoU ,spees,strn4 eqt pue osls ,{lqeqo.rdpue pequcsepu,no:g en8o1o1o.,de4 u1 :e1o51

'(lrn4 3uno,{ur ueuucads,WOtggt Iou uellug] Bur:uaqpuu 'esueIlErtsnv 'utlroig .>1.lOgI.uu1 ,etuurusu; ,e1dur^rl€C ,.9 208 I ratl, pe0lt teeqs Uod .,.sueJSEJeur? .a ,wg) usourtr^\tr,,(z) 0eeqslBnpr^rpur uE uo lnq Ngc oslu odftol:nl qlr,&leeqs uo pelunourueurlceds 3ur:e,to1g(l) :sed,{totcele.rudA :ad^lolrelosr:(uerurJeds Bunrnrg,llg) y997 ue}tugl Buuueq 'asueIIErtsnV L.os rell, pellq 'irdro.rgy '[tOg .qog,EruELuseJ .re,\lU] PUE.,9-208 I taaqs l ftnf .,tue,rrec,, :(pelcalesoroq) ed{lo1ce1 (9191);9y:S.Z.pr,ra).1rog.uo;ry1.111ui.Ig.U uol,{xou€lau BJoETV

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'( plllid ( tus),l/olu(ut'v =)'Jg >JotDut\rmay.e.r.uox?t eurgseqt luasa:der sueurrceds pele^IllnJ puB ..plln ,, eql qlog l?uellru srqt ^{BsFltuntJ€ udo.rg tuqt eJuepr^eou sr eJeqttnq,re){ .qENcJ,\tr lu u,no;8 s1ue1duro{ g0gl ur pelJelloc ale,$ ,(eql ,{q ,tD?.r8Jaaorrory pe11equlueurceds Suue,rog e slstsuocsol3eds srql go .sIr0 3:o leual?ruNSCI erlJ tnoquelrugop aq ot lueld pele,rrtlnceqf lo uortducs€p .11?rJel'u.r .uoH tduJsnuuursq ruo,r;3ut3pnt,elqr ssod lou sr1r ,{\e) s.u,ro:g luese:de.rf.ru ,{eqtelrq,n ispeespue slrn:; o1,{1uo sreler ,{uuq1y1u Bur,tor8 s1ue1deqt go uoqdr:rsepldrrcsnuuru srq esnpJeqs8uuorllu8,.plr^\,,s,u^\oJgeq01,{1e1r1une_ru,{eq;.ureuecunsrleeqsedf,taqtuosuerurc"ds ,re.\e,{oH .(,\\oleq 3ur:e,to1; o,tr1Sururcure: aql.;:o ur8rro eq1 ees)r,jru?Arrra .y,.plr,{\,, eq ol eulnssu o,{\ lrq,{\ uorldr.rcseps.u,rojg qlr,r Jo 1le,r ,{re,tsaeJfu ed{lolcal se pelcelesugrurceds J,a\trg eqJ '8081 uI pooC reled ,{q [puulSuAolud pecnporlurueeq pDq serceds eql luql pus pue ,..puBIIoH ^{oN Jo lsloc lso,\\-q1nos,,eql uroJJpetJellotr prq oq lErretEruot pexaJe.roslu eq :uoqducsepeqt ol ..sstuu,^^org,, uo0€tou eql pepueddepuu ler.reluur3u ue,ro11 peqrJOSop u,rorg on8olotordeql uI :eloN

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scurtu ,rirry su.rorg .tlr^\oJ .d ueqou s d pux urlsDhJ I n6 Nuytsi.rvol. 10,No. I (1995)

0 ).,Derwent,,butwith no otherdetails (specimens with matureseeds and near-mature flowers). Ina manuscriptat BM which lists the plantscollected fiom the DerwentRiver betweenFebruary and July 'Mimosa 1804,Brown citesthe following: cinerascens.In campis& ripasriviilor: frcquens";this entry may well be a referenceto the collection cited above. There are duplicatesofthis collection at both f andDB\ tex herb.W.R. McNrb,.

(2) "Mimosa cinerascens.Incampis non[?] longe a cult [?]:Port Dalrymple, Jan l804" (specimenin young fruit).

There is only one Brown rnanuscriptdescription of this speciesat BM. It usesthe nameMi'?osc and describesflowering material. Although there is no direct indication as to the origin cinerlscens "wild" of the plant described("wilcl" or cultivated) it may possibly be basedon a gatheringbecause the plant is descdbedas a small or medium tlee

"Hortus It is difficult to detemine the basisof the Kewensis"descdption of A. mel4nory lon because noneof the specimensat BM or DBN is from a cultivatedplant and the soulceof the manuscript "Hortus descriptionis Lquivocal. An examinationof this descriptionshows that the Kewcnsis" account *u. not tuk"n Jirectly from it. Furthennore,it is a curious fact that there is no McNab material at DBN collected from Kew Gardens,as thefe is tbf most other Brown speciesdiscussed in this papel However,because the protologuealludes to Brown's collectionsfrom Tasmania,we considerthis material to be available fbr lectotypification. Itis noted that Pedley (1918:222) regardedBrorvn's Port Dalrymple, Jan. I 804, specirnen(in young fruit) as the holotype of A melanoryLon However' "wild" in view oi'the fact that two collectionsof material areinvolved, a lectotypeshould have been "Port selected.Because the,,Derwent" collection,unlike the one from Dalrymple", providesthe seed characterscontained in the original descdptionand is representedby a duplicatc at E, we consider it to be the better sourceof the lectotype

Acacia pulchellaR. Br. in W.T. Aiton, Hort. Kew ed.2' 5: 464 (l813) Lectotype(here selected): - ,,Mimosaarmata." King GeorgeIII SouncllAlbany, w.A.l, R. Brown lower right handllowering "Iter specimenon sbeettitled Australiense,1802-5" and bearing[Britten no.] 4322 (BM); isotype: K Paralectotypes:Sterile and fruiting specimenslnounted on sheetwith lectotype(BM)

Note: In the original descriptionofA. p ulchelktBrown describeda flowering specimen;he referred "South-west to his own manuscriptand to material he had collectedfiom the coastof New Holland" and he also statedthat the specieshacl been introduced [into England] by PeterGood in 1803.

There aretwo sheetsat BM which may havea bearjngon the typification of this name,one labelled .,Hor.t. "Mimosa (in an unknownhand) Kew. New Holl. Mr Brown" andthe other (in Brown'shand) armata.King GeorgeIII Sound". The first sheetbears a single flowering specimenand the second bearsfour specimens(cornprising both flowering and fruiting elements) The DBN material of this speciesconsists of two sheetslabelled A cacia pulchellaby McNab; in eachcase the specimenswele collected from plants grown at Kew. The flowering specimenswere collectedin 1806,the fruiting onesin 1808. There is no evidencethat Brown actually saw this matedal All specimensreferred to abo'te areA. ;tulcheLIa\at. puLchella

The onc relevantmanuscdpt description ol A. pulchellaat BM is annotatedby Brown as both ,,M. ..Minlosa congesta"ancl microphylla". It describesa flowering specimenin cultivation at Kew "Port in Mav 1806. srown liom seedsent from R. G. III" [Albany, W'A ] a

'seqalJe^ eeJqlseslrdutox Dto)lns y(€661) urfsBl{T ul,,&o3(q pessncsrpsy

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'9081 ,{BIr'{ur ,re;4 tu pale,rqlnc1uu1d e uo.rJ ueulrcedsu uo peseqlurreluru Eurre,rog ,punog s?qrJJseppuB DlDrlnsDDD7V se paruderdst Jaqlo oql ,{ur:q1y e8loeC Bur) ruoJl lurre}Bu ..plr,^^,,s.u,rorg uo pospqsr lr :slrn{ puDsra,roU qtoq sequasoppue oJDpu?losowtT,g auvu ldrrcsnueur aqt Jepunsr auo tr{g le aDJlns V Jo suolduJsrp ldrrcsnuuuLu,1\oJtI ueqo5 o,rl eJ" eleql

.l€r.relgur uox€]eruss eql lua:-JldersurultJJds palf,^rtlnJ puc ,.plr,r,,es€ql qlog sq1 ,tr.us{llen1ce u,ro]g l€ql ecuapr^eou sr ejeq} lnq,re;4 lu umorS sturld ruor.}g0gl ur pelcalloc 'sueunJeds are,lr,{eql :qeNctr^{,{qDIrJlns olJoJV palleq[l qtoq 8urre,,,rog1lurus o,n1 Suqroddns leeqs B Jo slsrsuocserrads srql go leualsur NgC eqJ DlDpunDsowl4 su u,rorg ,{q pelleq€l sr teeqseql .serceds pue sluauele 3u4rn4 pue Suus,rog qloq estrdurocsueulceds aq1 eqt;o sBuueqlef ..p1r,tr,, s.u^{oJg lueserderqorq,r eloqE pelrc esoqt ele ueessuourrceds u,{\otg lreqog,{luo eql 1a{glV

'€081ur pooD ,..pu€lloH reled ,{q [puElSuEolul] pocnpojtur uaaqpeq sercedseq] lEqt prus pup ,roN Jo tseoc tse,\\ -qtnoS,, eq] uro4 pelc€llocpuq eq lpuel€lu ot pexoJa.rosp eq :uoqducsepeql ot ,,sstuu,rolg,, uoqetou eqt pepueddupu" Irualrut 8uurruJ puu 3urro,,lrouqtoq psqrJcsapu,no:g en8o1o1o:daq] uI ;etoN

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sauJeu,rDry s,u^\orglaqou'uD^\oJ s N pur urlsrw u u NuytsiaVol. 10,No. I (1995)

Acknowledgements

We are most grateful to David Mabberley, Paul Wilson and Gillian Perry for their critical review of the manuscript. The Directorsof both the Natuml History Museum (BM) and the National Botanic Gardens,Glasnevin (DBN) arethanked for allowing us accessto their valuableRobert Brown material. We appreciatethe assistanceprovided by Roy Vickery (BM) andPeter Weston, AustralianBotanical Liaison Officer, Kew 1992-3. CharlesNelson is thankedfor his useful discussionsconcerning the McNab collections. The work was conductedwith financialsupport provided by the Australian BiolosicalResources Studv.

References

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