Zootaxa: New Fossil Rhopalids (Heteroptera: Coreoidea) from the Middle

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Zootaxa: New Fossil Rhopalids (Heteroptera: Coreoidea) from the Middle

Zootaxa 1384: 41–58 (2006) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ ZOOTAXA 1384 Copyright © 2006 Magnolia Press ISSN 1175-5334 (online edition) New fossil rhopalids (Heteroptera: Coreoidea) from the Middle Jurassic of Inner Mongolia, China YUNZHI YAO1, WANZHI CAI1, 2, 3, DONG REN 2, 3 & CHUNGKUN SHIH 2 1Department of Entomology, China Agricultural University, Yuanmingyuan West Road, Beijing 100094, China. 2Key Lab of Insect Evolution & Environmental Changes, Capital Normal University, Beijing 100037, China. 3Corresponding author Abstract Three new genera and three species of fossil rhopalid, Originicorizus pyriformis gen. & sp. nov., Quatlocellus liae gen. & sp. nov., and Grandicaputus bipunctatus gen. & sp. nov., are described and illustrated. They were discovered from the Middle Jurassic Jiulongshan Formation of Eastern Inner Mongolia, China. The present discovery demonstrates that the early diversification of rhopalids was well underway by the Middle Jurassic. Key words: Heteroptera, Coreoidea, Rhopalidae, fossil, Middle Jurassic, Jiulongshan Formation, Daohugou, China Introduction The extant Rhopalidae contains two subfamilies, the Serinethinae with 3 genera, and the Rhopalinae with 17 genera (Göllner-Scheiding 1983), and most of them occur in the Pale- arctic Region (Schaefer & Chopra 1982, Schaefer 1992). In China, 36 species of extant rhopalids have been described and most species are distributed in the Inner Mongolia and Xinjiang Autonomous Regions (Liu et al. 1994, Hsiao et al. 1977, Nonnaizab et al. 1988). Only two fossil genera and species of the family Rhopalidae have been recorded from China prior to this study: Miracorizus punctatus Yao, Cai & Ren 2006 and Longiclavula calvata Yao, Cai & Ren 2006 from the same locality and stratum as the new species described here. According to the distribution pattern of extant rhopalids and fossil data, the most parsimonious hypothesis is that the basal species of this family initially radiated in Northeast China. Accepted by C. Schaefer: 11 Oct. 2006; published: 18 Dec. 2006 41.

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