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"'cientific f>apers -,!!;ll,.---------------------------- Natural History Museum The University of Kansas 16 April 2003 Number 27:1-41 An Annotated Taxonomic Catalogue of the Hoplandriini of the World (Insecta: Coleoptera: Staphylinidae: Aleocharinae)1 By RODNEY s. HANLEY2 Division of Entomology, Natural History Museum and Biodiversity Research Center, The University of Kansas Lawrence, Kansas 66045-2109, U.S.A. CONTENTS ABSTRACT ................................................................................................................ 2 INTRODUCTION ..................................................................................................... 2 ACKNOWLEDGMENTS .............................................................................................. 3 MATERIALS AND METHODS .............................................................................. 3 ANNOTATED TAXONOMIC CATALOG ............................................................ 4 HorLANDRIINI .•...•...............•.........•..•..........•.............•....••......••......•••••..•.........•.•... 4 HOPLANDRIINA ...................................................................................................... 4 HoPLANDRIA .......................................................................................................... 4 SUBGENUS ARRHENANDRIA ..................................................................................... 4 SUBGENUS GENOS EMA ............................................................................................ 5 SuBGENus HoPLANDRIA .......................................... ; .............................................. 5 SUBGENUS LOPHOMUCTER ....................................................................................... 7 SUBGENUS PLATONI CA ............................................................................................ 9 lNCERTAE SEDIS . ........... ..... ... ... .. ... ... .. .... .. .. ..... ...... •. ...... .......... .... ... .. ... .... 11 LEPTANDRIA ....................................................................................... ···•••·•••......... 18 HELICONANDRIA ............................................,: ...................................................... 19 PLATANDRIINA ..................................................................................................... 20 1 Contribution Number 3301 from the Division of Entomology, Natural History Museum and Biodiversity Research Center, and Department of Ecology and Evolutionary Biology, The University of Kansas. 2 Present Address: Earth Systems Science Institute, Box 9007, University of North Dakota, Grand Forks, North Dakota 58202-9007,USA. E-mail: [email protected] © Natural History Museum, The University of Kansas ISSN No. 1094-0782 2 SCIENTIFIC PAPERS, NATURAL HISTORY MUSEUM, THE UNIVERSITY OF KANSAS DITROPANDRIA ••.•.....•.•••.••....•..•••.•...•..•.•....••.....•....••....•.•••..•....••.•....•....•............•..... 20 SUBGENUS DJTROPANDRIA ....................................................................................... 20 SUBGENUS PEDANDRIA ............................................................................................ 20 LIGULATA ................................................................................................................ 21 MICROLIA .•••••.•••..••.••.•..•..•.•..•....•......· .••.• ~ ...••....•..•••.•...•..............•.........••............•...• 21 0MOPLANDRIA ....................................................................................................... 22 PAROPLANDRIA ............................................................................, .......................... 23 PLATANDRIA .••.••..••.•....•.••.•....••.•..•.....•.•....•.••.......••.•.•...••.....•.•.•........•.........•......••... 23 PSEUDOPLANDRIINA ............................................................................................... 26 PsEUDOPLANDRIA ................................................................................................... 26 HorLANDRIINI INCERTAE SEDIS .............................................................................. 34 ACANTOXYURA ....................................................................................; .•..•.....•....•.. 34 ALEOCHANDRIA ••••••··••·••••··•··•·•·••···•····•··•··•··••····•••••·•···•····•••·······•··•····•··•·•·•···••···•···35 ALLOPLANDRIA ....................................................................................................... 35 BESSOGLOSSA ........................................................................... ···•·······.................... 35 BRACHIDAMORPHA .................................................................................................. 35 PALAEOCHARA ........................................................................................................ 35 LITERATURE CITED ............................................................................................... 36 INDEX ......................................................................................................................... 39 ABSTRACT An annotated taxonomic catalogue of the Hoplandriini of the world is presented. The proposed classification of the Hoplandriini contains three subtribes, 18 genera, seven subgenera and 214 species. Lectotypes have been designated for 90 species, and a holotype is designated for one species. Key Words: Insecta, Coleoptera, Staphylinidae, Aleocharinae, Hoplandriini, Hoplandriina, Platandriina, Pseudoplandriina, taxonomy, classification, catalog, Acantoxyura, Aleochandria, Alloplandria, Arrhenandria, Bessoglossa, Brachidamorpha, Ditropandria, Genosema, Heliconandria, Hoplandria, Leptandria, Ligulata, Lophomucter, Microlia, Omoplandria, Palaeochara, Paroplandria, Pedandria, Platandria, Platonica, Pseudoplandria. INTRODUCTION The last complete World catalogue of the Hoplandriini logenetic analysis of the Hoplandriini (Hanley, 2002a). Full was developed by Bernhauer and Scheerpeltz (1926), with bibliographic citations, including annotations, are pre- a major supplement by Scheerpeltz (1934) as part of a cata- sented for each available name. Data regarding each spe- log of the beetle family Staphylinidae. In this monumen- cies, including type specimen information and type local- tal work, the Hoplandriini was treated as a subtribe of ity, are presented. Furthermore, for each genus, complete Myrmedoniini under the name Hoplandriae and was com- annotated bibliographic citations, type species, and the posed of 11 genera and 89 species. Since then, all subse- known distribution are also given. Every effort was made quent catalogs and checklists of the Hoplandriini have been to make this a complete catalogue in that it includes every restricted to specified geographical regions, most notably published mention of each taxon. Unfortunately, certainty the Palearctic Region. In addition, many new hoplandriine in such a claim is essentially unverifiable. The likelihood genera and species have been described and numerous is present that articles that provided only lists of species changes have been made to the classification of the collected at one site or another were inadvertently omited. Hoplandriini since the original catalogue and supplement. The proposed classification of the Hoplandriini con- Until now, the classification has not been based on pat- tains 18 genera, 7 subgenera and 210 species (Table 1 ). Eight terns of relatedness, and a coherent view of evolution in genera are listed as incertae sedis but were included within the group is simply not possible. the Hoplandriini by previous authors. In most instances, The primary purpose of this catalogue is to provide a the type specimens from these taxa are housed in private bibliographic basis for the Hoplandriini and to place spe- collections and are unavailable for examination. Repeated cies into appropriate subtribes, genera and subgenera efforts to borrow material from these collections ended in based on patterns of relatedness. The taxonomic arrange- failure. In the cases of taxa described by Cameron and ment used throughout this catalogue is based on the phy- Bernhauer, type ·series generally consisted of single sped- TAXONOMIC CATALOGUE OF HOPLANDRIINI 3 Table 1. Generic classification of the Hoplandriini. comments or explanatory statements. I found numerous situations in Pace's papers where the tribal name is mis- TRIBE HoPLANDRIINI Casey placed, thereby giving the appearance of taxa being in- Subtribe Hoplandriina Casey Genus Hoplandria Kraatz cluded or excluded from particular tribes resulting in con- Subgenus Arrhenandria Genier (1 species) siderable taxonomic confusion. Every effort was made in Subgenus Genosema Notman (3 species) this study to determine accurately the tribe in which those Subgenus Hoplandria sensu stricto (3 species) Subgenus Lophomucter Notman (7 species) taxa listed as hoplandriines should be placed. Many of Subgenus Platonica Sharp (14 species) Pace's types, however, are unavailable for examination and Incertae sedis (69 species) the published descriptions are often inadequate to deter- Genus Leptandria Hanley (3 species) Genus Heliconandria Hanley (3 species) mine tribal placement. Subtribe Platandriina Hanley ACKNOWLEDGMENTS Genus Ditropandria Pace Subgenus Ditropandria sensu stricto (3 species) Subgenus Pedandria Pace (2 species) I extend my gratitude to my major professor, James S. Genus Ligulata Hanley and Ashe (1 species) Ashe (University of Kansas) for his suggestions, encour- Genus Microlia
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