Atlas of Yorkshire Coleoptera part 7 - to

Introduction

This is Part 7 of the Atlas and covers the Families Eucinetidae, , , , and Byrrhidae.

Each species in the database is considered and in each case a distribution map representing records on the database (at March 2017) is presented. The number of records on the database for each species is given in the account in the form (a,b,c,d,e) where 'a' to 'e' are the number of records from VC61 to VC65 respectively. These figures include undated records (see comment on undated records in the paragraph below on mapping). As a recorder, I shall continue to use the vice-county recording system, as the county is thereby divided up into manageable, roughly equal, areas for recording purposes. For an explanation of the vice-county recording system, under a system devised in Watson (1883) and subsequently documented by Dandy (1969), Britain was divided into convenient recording areas ("vice-counties"). Thus Yorkshire was divided into vice-counties numbered 61 to 65 inclusive, and notwithstanding fairly recent county boundary reorganisations and changes, the vice-county system remains a constant and convenient one for recording purposes; in the text, reference to “Yorkshire” implies VC61 to VC65 ignoring modern boundary changes.

For some species there are many records, and for others only one or two. In cases where there are five species or less full details of the known records are given. Many common species have quite a high proportion of recent records. This is because the older county recorders tended not to record species they regarded as "common" quite as assiduously as we do today.

“Recent” records are those considered to be later than 1 January 2000 – this is also the break point used in the production of the distribution maps. (In previously posted parts of this Atlas, the break point for recent and otherwise was 1 January 1990).

Habitat A brief statement on habitat is included where known and relevant.

Taxon status The national status of each taxon is given, this being the status as defined in JNCC’s Recorder 6 data program. Status is included if Notable B or above. Please bear in mind that many of these statuses are now in need of revision, in the light of recent knowledge of distribution and frequency. The following status criteria are derived from Shirt (1987) and Hyman and Parsons (1992 and 1994):

RED DATA BOOK CATEGORY 1. RDB1 – ENDANGERED - Species which are known or believed to occur only as a single population within one 10km square of the OS National Grid.

RED DATA BOOK CATEGORY 2. RDB2 – VULNERABLE - Species which are declining throughout their range.

RED DATA BOOK CATEGORY 3. RDB3 – RARE - Species which are estimated to exist in only 15 or fewer 10km squares; this criterion may be relaxed where populations are likely to exist in more than 15 squares but occupy small areas of especially vulnerable habitat.

RED DATA BOOK CATEGORY I. RDB I – INDETERMINATE - Taxa considered to be endangered, Vulnerable or Rare, but where there is not enough information to say which of the three categories (RDB1 to 3) is appropriate.

RED DATA BOOK CATEGORY K. RDBK – INSUFFICIENTLY KNOWN - Taxa recently discovered or recognised in Great Britain, which may prove to be more widespread in the future (although some recent discoveries may be placed in other categories if the group to which they belong is thought to be not under-recorded).

PROVISIONAL RED DATA BOOK. Prdb - The prefix ‘p’ before any red Data Book category implies that the grading is provisional.

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Notable A – species which do not fall within the RDB categories but are thought to occur in 30 or fewer 10km squares of the National Grid, or for less well-recorded groups, within seven or fewer vice-counties.

Notable B – species which do not fall within the RDB categories but thought to occur in between 31 and 100 10km squares of the National Grid.

Nationally Notable - species which are estimated to occur within the range of 16 to 100 10km squares. (subdivision into Notable A and Notable B is not always possible because there may be insufficient information available). Now generally superseded by Nationally Scarce, and therefore no longer much used in modern literature.

The following status criteria are included in the species accounts and derive from those used in the Recorder 6 database. Please note that many of these statuses require reconsideration in the light of currently known records and distributions.

RedList_pre94-EN - Taxa in danger of extinction and whose survival is unlikely if the causal factors continue operating.. Superseded by new IUCN categories in 1994, so no longer in general use.

RedList_pre94-EX - Taxa which are no longer known to exist in the wild after repeated searches of their localities and other known likely places. Superseded by new IUCN categories in 1994, so no longer in general use.

RedList_pre94-NR - Taxa with small populations that are not at present Endangered or Vulnerable, but are at risk. (In GB, this was interpreted as species which exist in fifteen or fewer 10km squares). Superseded by new IUCN categories in 1994, so no longer in general use.

HabDir-A2* - and plant species of Community interest (i.e. endangered, vulnerable, rare or endemic in the European Community) whose conservation requires the designation of special areas of conservation. Note that the contents of this annex have been updated in April 2003 following the Treaty of Accession.

Phenology

Details are given for the range of months in the year for which there are records, with or without a peak month of occurrence, where the number and detail of records are sufficient to give a meaningful statement.

Maps Distribution maps have been produced using Dr Alan Morton’s DMAP software (version 7.0b). It may be worthwhile to note that as DMAP is asked to map pre- and post-2000 records in this application, any undated records will not be mapped. While efforts have been made to establish dates of records, some are difficult to ascertain, and I do not want to make guesses which may be wrong.

Nomenclature The YNU database is presently held in a copy of Recorder 6, which uses a new Coleoptera checklist based on Duff (2008) and this new list incorporates a very large number of taxon name changes since the checklist of Pope (1977); there is now a second edition of the latest Coleoptera checklist (Duff, 2012) which has not yet been incorporated into Recorder 6. Occasionally I receive records using “noddy names” (see Cooter, (1999) and Key (1999) for an excellent treatment of this subject), e.g. “Snail Hunter”, “Crucifix ” and “Heath-glory ” (Cychrus caraboides, Panagaeus crux-major and nitens respectively). Such inventions were designed for use in political documents, and seem to have originated somewhere within Natural England (previously English Nature) or JNCC. For the purposes of serious science these terms are, in my opinion, either vague, ridiculous, confusing or unhelpful. I do not accept records using such names only, unless accompanied by specimens or other convincing evidence.

For the purposes of this atlas the Duff 2012 checklist will be used as the basis of the nomenclature and arrangement of species.

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Quality of the records A regrettable aspect of many records within the YNU database is the lack of detail accompanying the record, especially of accurate grid references, lack of pertinent comments as to the micro-habitat of the capture, its association with plants/, and sometimes, in the case of some modern records, a lack of care in determination, and the recording of the sex of the specimen. Old records, although sound, tend not to give much or indeed any of this detail. Old records on the database so rarely give habitat details but sometimes interpretation of the site name can give clues to the habitat type. Grid references could not be applied before 1930 anyway, and grid references do not figure in the paper archive of the YNU Coleoptera database until the late 1970s. Apparently it was not considered necessary to give that level of accuracy. Entering records into Recorder 6 requires a grid reference, and with older records I have added a grid reference where I believe this has been possible. If the site name is a vague one I have used a 10km reference, and where a 1km reference is deemed accurate enough I have included that. There can be little excuse with modern records not to include some accurate comment with the record, and at least a six-figure grid reference.

Taxonomic knowledge and expertise have increased greatly since the days of Walsh, Bayford and Stainforth. Much good literature now exists for the taxonomist in the form of dichotomous keys, and when identifying specimens, especially with critical species pairs or groups, too much reliance on the outcome of keys can be placed on a determination, without apparently considering the habitat in which the specimen was taken, or the host plant/animal. Atlases of the national distributions of various orders of are now becoming available and should always be consulted. These often give a guide to the likelihood or otherwise of a determination, especially in scarce taxa. Museum collections containing correctly determined and labelled specimens are invaluable in accurate recording; museum collections are there for our use and should be consulted. If a species is ‘new’ to a vice-county or is very significant in some other respect, a second opinion should always be sought or an acknowledged specialist in the group consulted. A voucher should always be retained by the collector if possible. The recorder can only deal with the information he is given, and often has to beware of identifications made on photographic evidence only, and identifications of ‘difficult’ species by non-specialists. It takes many years to become competent in many areas of entomology, and the recorder often has to rely on the expertise of co-workers, especially when records covering large Orders are being documented.

I have adopted a policy in my county recording whereby any received record of a species afforded Notable B or above, especially if a member of a ‘difficult’ beetle group, should be confirmed by a specialist in that group before the record is entered into the database as a correct record.

Ultimately, the responsibility for verification of records on the YNU database rests with me. With modern records received from reputable coleopterists there is little problem. Some older records requiring vouchers for confirmation have been retained in the database but flagged as requiring confirmation.

Contributors (as initials given in the text):

AG - A Godfrey MLD - M L Denton

CJ - Colin Johnson PJH - P J Hodge

EGB - E G Bayford RJM - R J Marsh

EJS - E J Smith RL - R Lawson

EWA - E W Aubrook TS - T Stainforth

GBW - GB Walsh WAE - W A Ely

HB - Harry Britten jnr WDH - W D Hincks

HP - H Pollard WJF - W J Fordham

JP - Jean Parkin

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Superfamily Fleming, 1821

Family EUCINETIDAE Lacordaire FL, 1857

EUCINETUS Germar, 1818 meridionalis (Laporte, 1836)

NO RECORDS

Subfamily CALYPTOMERINAE Crowson, 1955

CALYPTOMERUS Redtenbacher, 1849 dubius (Marsham, 1802)

Calyptomerus dubius (Marsham, 1802)

3 WATSONIAN (17,3,20,3,0) Commonly found in piles of 2 YORKSHIRE cut grass, straw bales and compost heaps 1 where the contents are warm and 0 NY NZ NZ OV SD SE SE TA beginning to putrefy. The distribution 9 tends to reflect collecting effort as the 8 species is extremely small and has to be 7 6 searched for. (ii-xi) 5 4 3 2 1

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Subfamily CLAMBINAE Fischer von Waldheim, 1821

CLAMBUS Fischer von Waldheim, 1821 armadillo (De Geer, 1774)

Clambus armadillo (De Geer, 1774)

3 WATSONIAN (8,8,5,4,2) Occasionally found in ground 2 YORKSHIRE litter, straw bales and compost heaps 1 where there is warmth. But also from 0 NY NZ NZ OV SD SE SE TA piles of ground litter in a saltmarsh. 9 Seems mainly to occur in lowland areas 8 and hardly reported from upland sites. 7 6 A very small species and has to be 5 searched for carefully. (iii-x, peak v). 4 3 2 1

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evae Endrödy-Younga, 1960

Clambus ev ae Endrödy-Younga, 1960

3 WATSONIAN (0,1,0,0,1) Apparently very scarce with 2 YORKSHIRE only two records on the YNU database: 1 Goathland (NZ8202), 13/2/1937, HB; 0 NY NZ NZ OV SD SE SE TA Foxglove Covert (SE1697), 28/5/2003, 9 RJM teste CJ. The latter record was 8 reported from woodland ground litter. 7 6 5 4 3 2 1

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gibbulus (LeConte, 1850)

Clambus gibbulus (LeConte, 1850)

3 WATSONIAN (0,0,3,0,0) Very rarely recorded with three 2 YORKSHIRE entries for South Yorkshire in the YNU 1 database: Hawks Wood (SK5281),

NY NZ NZ OV 15/6/1999, WAE; King's Wood (SK5489), 0 SD SE SE TA 9 12/6/1986, WAE; Thorne Moor (SE7116), 8 1971, PS. Because of nomenclature 7 problems only recently resolved (Duff, 6 2012:58) these records may be referrable to 5 what we used to call C. radula Endrody- 4 Younga or C.punctulum (Beck). I have seen 3 vouchers for gibbulus (as radula) from the 2 Hawks Wood and King's Wood localities, and 1 these specimens appear to be that species,

SD SE SE TA but await expert confirmation. The records 0 SJ SK SK TF 9 remain on the database as requiring confirmation. 8 post 31/12/1999 Map produced for Yorkshire Naturalists' Union by DMAP 17/2/2017 pre 31/12/1999 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4

nigrellus Reitter, 1914

Clambus nigrellus Reitter, 1914

3 WATSONIAN (0,0,0,4,1) Notable A. A very scarce 2 YORKSHIRE species noted from wet situations 1 including flood refuse and stream-side 0 NY NZ NZ OV SD SE SE TA ground litter. There are five records in 9 the YNU database: Birk Crag (SE2754), 8 10/5/2001, RJM, in ground litter by 7 6 stream; Fountains Abbey (SE2768), 5 24/8/2000, RJM, in flood refuse on 4 shingle bank of River Skell; Studley 3 Royal (SE2969), 6/7/1966, EWA; 2 Studley Royal (SE2969), 14/7/1990, 1 RJM, in flood refuse on Skell banks; 0 SD SE SE TA SJ SK SK TF Foxglove Covert (SE1697), 28/5/2003, 9 RJM. 8 post 31/12/1999 Map produced for Yorkshire Naturalists' Union by DMAP 17/2/2017 pre 31/12/1999 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4

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nigriclavis Stephens, 1832

Clambus nigriclav is Stephens, 1835

3 WATSONIAN (0,0,0,4,0) A scarce ground litter 2 YORKSHIRE species, of which we have four records 1 from the YNU recording area, all noted 0 NY NZ NZ OV SD SE SE TA from flood refuse: Studley Royal Park 9 (SE2969), 6/7/1966, CJ, and 19/6/1989, 8 RJM; Willow Garth SE3173), 13/8/2002, 7 RJM, recorded from two localities 6 5 within the same site. 4 3 2 1

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pallidulus Reitter, 1911

NO RECORDS

pubescens Redtenbacher, 1849

Clambus pubescens Redtenbacher, 1849

3 WATSONIAN (12,4,18,1,1) Fairly common and 2 YORKSHIRE recorded from putrefying material 1 such as rotting potatoes and turnips, in 0 NY NZ NZ OV SD SE SE TA compost heaps and mouldy piles of 9 grass clippings. (iii-xi) 8 7 6 5 4 3 2 1

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punctulum (Beck, 1817)

Clambus punctulum (Beck, 1817)

3 WATSONIAN (3,3,20,1,0) Another widely distributed 2 YORKSHIRE and fairly common species recorded 1 from dry manure heaps, reed litter, 0 NY NZ NZ OV SD SE SE TA grass piles, and once from a dead 9 sheep. 8 7 6 5 4 3 2 1

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simsoni Blackburn, 1902

Clambus simsoni Blackburn, 1902

3 WATSONIAN (0,0,4,0,0) An Australian clambid 2 YORKSHIRE described as new to Britain in 1996 1 (Johnson, 1997). We have four records 0 NY NZ NZ OV SD SE SE TA on our database: Crookesmoor 9 (SK3387), 4/9/2009, EJS det RJM; 8 Nostell Priory (SE4117), 4/8/2003, MLD 7 det CJ; Upper Fell Grieve Wood 6 5 (SE1420), 16/8/2006, MLD; Wickersley 4 Wood (SK4891), 27/5/2005, WAE det 3 CJ. This species will probably prove to 2 be more widespread in the county. 1

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26. Family SCIRTIDAE Fleming, 1821

ELODES Latreille, 1796

[a difficult genus usually requiring dissected males for reliable identification - see Johnson (1992), and various papers by Klausnitzer, and Lohse (1979)] elongata Tournier, 1868

Elodes elongata Tournier, 1868

3 WATSONIAN (1,0,2,2,1) RedList_pre94-NR. Scarce. 2 YORKSHIRE Previously known in our fauna as E. 1 koelleri, we have six widely distributed 0 NY NZ NZ OV SD SE SE TA records. Our records indicate that the 9 species may be collected by sweeping 8 in wet places including pond margins 7 6 and river banks. 5 4 3 2 1

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minuta (Linnaeus, 1767)

Elodes minuta (Linnaeus, 1767)

3 WATSONIAN (4,12,13,12,3) Widely distributed and if 2 YORKSHIRE all our records are correct a fairly 1 common Elodes. Several other Elodes 0 NY NZ NZ OV SD SE SE TA species resemble minuta so vouchers 9 especially of males should always be 8 kept. Recorded from a variety of wet 7 and dry situations - pond margins, 6 5 woodland edges and from rowan and 4 hawthorn blossoms. No attempt has 3 been made to confirm any of our pre- 2 1980 records. Apparently with a short 1 adult period. (vi-vii). SD SE SE TA 0 SJ SK SK TF 9 8 post 31/12/1999 Map produced for Yorkshire Naturalists' Union by DMAP 17/2/2017 pre 31/12/1999 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4

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pseudominuta Klausnitzer, 1971

Elodes pseudominuta Klausnitzer, 1971

3 WATSONIAN (1,12,32,1,4) Widespread and a fairly 2 YORKSHIRE commonly recorded Elodes. All our 1 records are post-1980 and come from 0 NY NZ NZ OV SD SE SE TA reliable sources. As noted earlier, 9 vouchers should always be kept and 8 males are desirable. (v-ix, peak vi). 7 6 5 4 3 2 1

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tricuspis Nyholm, 1985

Elodes tricuspis Nyholm, 1985

3 WATSONIAN (0,0,0,1,0) RedList_GB_post2001-VU. 2 YORKSHIRE There is a single record of this rare 1 species in the YNU database: Breary 0 NY NZ NZ OV SD SE SE TA Marsh (SE2541), 9/6/1999, RJM det 9 RJM teste CJ, by sweeping around an 8 old log pile near a stream. There very 7 few authenticated British records. 6 5 4 3 2 1

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ODELES Klausnitzer, 2004 marginata (Fabricius, 1798)

Odeles marginata (Fabricius, 1798)

3 WATSONIAN (2,18,43,7,11). A widespread and 2 YORKSHIRE common Elodes, recorded from wet 1 woodland edges, pool margins and wet 0 NY NZ NZ OV SD SE SE TA places generally. (iv-viii, peak vi, with a 9 curious record for xi). 8 7 6 5 4 3 2 1

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MICROCARA Thomson, C.G., 1859 testacea (Linnaeus, 1767)

Microcara testacea (Linnaeus, 1767)

3 WATSONIAN (42,10,93,32,2) Widespread and 2 YORKSHIRE common and recorded generally by 1 sweeping in reedbeds, from flowering 0 NY NZ NZ OV SD SE SE TA plants in marshes and by lagoons, 9 ponds, in fens and in wet meadows. 8 The larvae are aquatic at the roots of 7 water plants, in detritus in ditches and 6 5 rot-holes in trees as is often the case 4 with other scirtids. (v-ix, peak vi-vii) 3 2 1

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CYPHON Paykull, 1799 coarctatus Paykull, 1799

Cyphon coarctatus Paykull, 1799

3 WATSONIAN (64,61,211,75,13). Widespread and 2 YORKSHIRE very common in a wide variety of wet 1 places where the aquatic larvae 0 NY NZ NZ OV SD SE SE TA develop. Collected mainly by sweeping 9 and grubbing in marshes and fens, and 8 adults on blossoms in damp habitats. 7 (iii-x, peak vi). 6 5 4 3 2 1

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hilaris Nyholm, 1944

Cyphon hilaris Nyholm, 1844

3 WATSONIAN (5,12,64,7,2) Widely but rather thinly 2 YORKSHIRE distributed throughout. In wet places 1 including ponds, marshes, drain edges, 0 NY NZ NZ OV SD SE SE TA and by beating Salix etc in carr and in 9 peat moors. (v-x, peak vii). 8 7 6 5 4 3 2 1

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kongsbergensis Munster, 1924

NO RECORDS

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laevipennis Tournier, 1868

Cyphon laev ipennis Tournier, 1868

3 WATSONIAN (59,5,110,11,2) Our records show a 2 YORKSHIRE marked southern county distribution. 1 Early records of variabilis may refer to 0 NY NZ NZ OV SD SE SE TA this species, previously known as 9 phragmiteticola - see Kevan (1962). 8 7 6 5 4 3 2 1

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ochraceus Stephens, 1830

Cyphon ochraceus Stephens, 1830

3 WATSONIAN (21,14,37,12,1) This species exhibits a 2 YORKSHIRE somewhat eastern-half distribution in 1 the county, but fairly common and 0 NY NZ NZ OV SD SE SE TA widespread there. (iv-x, peak vi). 9 8 7 6 5 4 3 2 1

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padi (Linnaeus, 1758)

Cyphon padi (Linnaeus, 1758)

3 WATSONIAN (23,23,130,38,6) Widespread but with a 2 YORKSHIRE rather central distribution. Records hint 1 at a preference for lowland somewhat 0 NY NZ NZ OV SD SE SE TA sheltered sites. Collected by sweeping 9 in wet meadows, in richly vegetated 8 areas of moorland, carr land and in 7 reedbeds. (iv-xi, peak vi). 6 5 4 3 2 1

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palustris Thomson, C.G., 1855

Cyphon palustris Thomson, C.G., 1855

3 WATSONIAN (4,6,24,8,5) Widespread but rather 2 YORKSHIRE thinly distributed with most records 1 from the southern half of the county. 0 NY NZ NZ OV SD SE SE TA Noted from wetlands generally, 9 including marshland, well-vegetated 8 drain and pond margins, canal sides 7 6 and in reed beds. (v-ix, peak vii). 5 4 3 2 1

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pubescens (Fabricius, 1792)

Cyphon pubescens (Fabricius, 1792)

3 WATSONIAN (1,3,2,1,0) RedList_pre94-NR. Scarce, 2 YORKSHIRE and very rarely recorded - previously 1 confused with variabilis (see Kevan, 0 NY NZ NZ OV SD SE SE TA 1962). Our records come from pond 9 margins, drain sides and Phragmites 8 beds. (v-x). 7 6 5 4 3 2 1

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punctipennis Sharp, 1873

Cyphon punctipennis Sharp, 1873

3 WATSONIAN (1,6,0,8,16) Notable-A. A species with 2 YORKSHIRE an interesting distribution in Yorkshire: 1 apart from an early VC61 record (1919, 0 NY NZ NZ OV SD SE SE TA GBW), which would require 9 confirmation, all our records (nearly all 8 post-2000) are northern. The national 7 picture indicates a mainly northern 6 5 species with a few Welsh and eastern 4 English records. (iii-xi, peak v). 3 2 1

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variabilis (Thunberg, 1787)

Cyphon v ariabilis (Thunberg, 1787)

3 WATSONIAN (31,21,154,36,12) Widely and generally 2 YORKSHIRE distributed with records noted from 1 Sphagnum pools, reed beds, carr land 0 NY NZ NZ OV SD SE SE TA drain and pond margins. One of our 9 most commonly recorded Cyphon. 8 7 (ii-xii, peaks vi and ix). 6 5 4 3 2 1

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PRIONOCYPHON Redtenbacher, 1858 serricornis (Müller, P.W.J., 1821)

Prionocyphon serricornis (Müller, P.W.J., 1821)

3 WATSONIAN (0,8,6,2,0) RedList_pre94-NR. A rarely 2 YORKSHIRE recorded beetle, apparently a specialist 1 species of water-filled rot-holes in 0 NY NZ NZ OV SD SE SE TA various deciduous trees. Collected 9 either directly from such rot-holes in 8 which the larvae develop, or adults 7 swept from nearby vegetation on their 6 5 dispersal. This habitat does not attract 4 much popular entomological attention 3 and so this cryptic species may be more 2 widespread than the records suggest. 1 (iii-xi).

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HYDROCYPHON Redtenbacher, 1858 deflexicollis (Müller, P.W.J., 1821)

Hydrocyphon deflexicollis (Müller, P.W.J., 1821)

3 WATSONIAN (1,4,0,1,6) Notable-B. Rare and mainly 2 YORKSHIRE northern in its distribution in Yorkshire, 1 this riverine species has aquatic larvae 0 NY NZ NZ OV SD SE SE TA and pupae, and the adults may be 9 collected from emergent water plants 8 7 or riverside plants and trees. (v-vii) 6 5 4 3 2 1

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SCIRTES Illiger, 1807 hemisphaericus (Linnaeus, 1758)

Scirtes hemisphaericus (Linnaeus, 1758)

3 WATSONIAN (17,12,29,17,5) Widely distributed in a 2 YORKSHIRE more or less central north-south band in 1 the county. Recorded by sweeping 0 NY NZ NZ OV SD SE SE TA pondside and marsh plants including 9 Mentha and Scirpus, occasionally in 8 ground detritus at edges of suitably 7 vegetated water bodies. (vi-ix, peak vii). 6 5 4 3 2 1

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orbicularis (Panzer, 1793)

NO RECORDS

Atlas of Yorkshire Coleoptera - part 7 - Eucinetidae to Byrrhidae page 17

Superfamily Guérin-Méneville,1843 (1834)

27. Family DASCILLIDAE 1843 (1834)

DASCILLUS Latreille, 1796 cervinus (Linnaeus, 1758)

Dascillus cerv inus (Linnaeus, 1758)

3 WATSONIAN (5,12,12,14,6) Rather uncommon but 2 YORKSHIRE very widely distrubuted. Recorded by 1 sweeping or shaking flowerheads of 0 NY NZ NZ OV SD SE SE TA various plants, including orchids, the 9 larvae feed in the roots and pupate in 8 the soil. Also recorded in the county in 7 pitfall traps in hay meadows and 6 5 grassland. (vi-viii). 4 3 2 1

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Superfamily Leach, 1815

28. Family BUPRESTIDAE Leach, 1815

Subfamily BUPRESTINAE Leach, 1815

Tribe ANTHAXIINI Gory & Laporte, 1839

ANTHAXIA Eschscholtz, 1829 nitidula (Linnaeus, 1758)

NO RECORDS

quadripunctata (Linnaeus, 1758)

NO RECORDS

Tribe MELANOPHILINI Bedel, 1921

MELANOPHILA Eschscholtz, 1829 acuminata (De Geer, 1774)

NO RECORDS

Atlas of Yorkshire Coleoptera - part 7 - Eucinetidae to Byrrhidae page 19

Subfamily AGRILINAE Laporte, 1835

Tribe AGRILINI Laporte, 1835

AGRILUS Curtis, 1825

Subgenus AGRILUS Curtis, 1825 viridis (Linnaeus, 1758)

Agrilus (Agrilus) v iridis (Linnaeus, 1758)

3 WATSONIAN (0,2,0,0,0) RedList_pre94-VU. There are 2 YORKSHIRE two records of this species on the YNU 1 database, both very old and probably 0 NY NZ NZ OV SD SE SE TA doubtful: York (SE65) 1830, JF 9 Stephens; York (SE65) date unknown, 8 "Cook" (name given in YNU paper 7 6 archive), on oak. These two records 5 may refer to the same collecting event. 4 No vouchers have been traced. The 3 only authenticated British records are 2 from the SE of England and the New 1 Forest. SD SE SE TA 0 SJ SK SK TF 9 8 post 31/12/1999 Map produced for Yorkshire Naturalists' Union by DMAP 17/2/2017 pre 31/12/1999 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4

Subgenus ANAMBUS Thomson, C.G., 1864 angustulus (Illiger, 1803)

Agrilus (Anambus) angustulus (Illiger, 1803)

3 WATSONIAN (3,2,1,0,0) Notable B. Very scarce with 2 YORKSHIRE only six known county records on our 1 database. Where habitat is mentioned 0 NY NZ NZ OV SD SE SE TA in the records this is oak. Records are 9 from old growth areas rather than 8 specifically old forest (except 7 Helmsley). Specimens should be looked 6 5 at carefully to avoid confusion with 4 laticornis, which is more common and 3 widespread. See Levey (1977). 2 1

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Atlas of Yorkshire Coleoptera - part 7 - Eucinetidae to Byrrhidae page 20

biguttatus (Fabricius, 1777)

NO RECORDS

cuprescens (Ménétries, 1832)

NO RECORDS

cyanescens Ratzeburg, 1837

NO RECORDS

laticornis (Illiger, 1803)

Agrilus (Anambus) laticornis (Illiger, 1803)

3 WATSONIAN (2,7,8,2,0) Notable B. The commonest 2 YORKSHIRE Agrilus species in Yorkshire with 19 1 records. Recorded from oak and hazel 0 NY NZ NZ OV SD SE SE TA in ancient woodland sites and in old 9 growth areas.(vi-viii). 8 7 6 5 4 3 2 1

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sinuatus (Olivier, 1790)

NO RECORDS

sulcicollis Lacordaire, 1835

NO RECORDS

Atlas of Yorkshire Coleoptera - part 7 - Eucinetidae to Byrrhidae page 21

Tribe APHANISTICINI Jacquelin du Val, 1859

APHANISTICUS Latreille, 1829 emarginatus (Olivier, 1790)

NO RECORDS

pusillus (Olivier, 1790)

Aphanisticus pusillus (Oliv ier, 1790)

3 WATSONIAN (0,1,0,0,0) Notable B. Very rare in 2 YORKSHIRE Yorkshire with a single record on the 1 YNU database: Scarborough area 0 NY NZ NZ OV SD SE SE TA (TA08), c1900, RL. No habitat details 9 are given with this record and no 8 voucher has been traced by this author. 7 The species is associated with various 6 5 sedges. 4 3 2 1

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Tribe TRACHEINI Laporte, 1835

TRACHYS Fabricius, 1801 minuta (Linnaeus, 1758)

Trachys minuta (Linnaeus, 1758)

3 WATSONIAN (2,0,0,0,0) RedList_pre94-VU. Very rare 2 YORKSHIRE with only two records on the YNU 1 database, both of which may refer to 0 NY NZ NZ OV SD SE SE TA the same collecting event: Brough 9 (SE9527), date unknown, GBW; Brough 8 (SE9527), 1903, TS. The species is 7 6 known to be associated with Salix and 5 Carpinus species. 4 3 2 1

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scrobiculatus Kiesenwetter, 1857

NO RECORDS

subglaber Rey, 1891

NO RECORDS

troglodytes Gyllenhal in Schönherr, 1817

Trachys troglodytes Gyllenhal in Schoenherr, 1817

3 WATSONIAN (1,1,0,0,0) We have two records in the 2 YORKSHIRE YNU database: Scarborough area 1 (TA08), c1900, RL; Hornsea (TA24), 0 NY NZ NZ OV SD SE SE TA undated, WWF. The latter record is 9 mentioned in Fowler 4:73. 8 7 6 5 4 3 2 1

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Superfamily Latreille, 1804

29. Family BYRRHIDAE, 1804

Subfamily Latreille, 1804

SIMPLOCARIA Stephens, 1829 maculosa Erichson, 1847

Simplocaria maculosa Erichson, 1847

3 WATSONIAN (1,0,0,0,0) The YNU has a single record 2 YORKSHIRE on its database: Kelfield (SE5938), 1 14/5/1956, JP, swept on banks of R. 0 NY NZ NZ OV SD SE SE TA Ouse. This was the first recent British 9 specimen, see Naturalist 1960:17. The 8 specimen is deposited in Leeds City 7 Museum. The only other British 6 5 specimens were located by Colin 4 Johnson in the Manchester Museum. 3 These had been collected by W G 2 Blatch at Bewdley, Worcs, in the 1890's, 1 and were found among a series of S. 0 SD SE SE TA SJ SK SK TF semistriata. 9 8 post 31/12/1999 Map produced for Yorkshire Naturalists' Union by DMAP 17/2/2017 pre 31/12/1999 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4

semistriata (Fabricius, 1794)

Simplocaria semistriata (Fabricius, 1794)

3 WATSONIAN (11,12,44,14,1 ) Fairly common and 2 YORKSHIRE very widely distributed. Swept around 1 pond margins, in saltmarshes, in flood 0 NY NZ NZ OV SD SE SE TA refuse and in piles of cut reeds. The 9 species is associated with the roots of 8 plants and in which the larvae 7 6 develop. (ii-xii, peak ix). 5 4 3 2 1

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MORYCHUS Erichson, 1846 aeneus (Fabricius, 1775)

Morychus aeneus (Fabricius, 1775)

3 WATSONIAN (1,0,0,0,2) A little known species and 2 YORKSHIRE very rare in the county with only three 1 records known: Great Langton 0 NY NZ NZ OV SD SE SE TA (SE2996), 23/4/1984, PJH, by grubbing 9 under stones on R. Swale bank; Great 8 Langton (SE2896), 7/7/2005, AG, on 7 river bank in ground litter: Filey 6 5 (TA1281), 14/12/1980, MLD. 4 Apparently a northern species which 3 has affinities with riverine habitats. 2 1

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CYTILUS Erichson, 1846 sericeus (Forster, 1771)

Cytilus sericeus (Forster, 1771)

3 WATSONIAN (3,16,66,16,5) Widespread and fairly 2 YORKSHIRE common but mainly recorded in the 1 centre of the county. In pitfalls in wet 0 NY NZ NZ OV SD SE SE TA grassland and on peat moors, in ground 9 litter in wet meadows and at lake and 8 pond edges. The larvae are root feeders 7 mainly on grasses. (ii-xii, peak vi). 6 5 4 3 2 1

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BYRRHUS Linnaeus, 1767 arietinus Steffahny, 1842

Byrrhus arietinus Steffahny, 1842

3 WATSONIAN (0,1,6,1,0) Notable B. Scarce in the 2 YORKSHIRE county with eight records only. In 1 ground litter at edges of a reservoir, on 0 NY NZ NZ OV SD SE SE TA high moorland and at margins of damp 9 woodland. The larvae are root feeders. 8 7 6 5 4 3 2 1

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fasciatus (Forster, 1770)

Byrrhus fasciatus (Forster, 1770)

3 WATSONIAN (2,7,16,0,3) Widespread but rarely 2 YORKSHIRE recorded post-2000. The larvae are root 1 feeders. NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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pilula (Linnaeus, 1758)

Byrrhus pilula (Linnaeus, 1758)

3 WATSONIAN (8,17,91,26,6) Widespread and the 2 YORKSHIRE commonest of Yorkshire's Byrrhus 1 species. Again, the larvae develop in 0 NY NZ NZ OV SD SE SE TA the roots of grasses and mosses. 9 8 7 6 5 4 3 2 1

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pustulatus (Forster, 1770)

Byrrhus pustulatus (Forster, 1770)

3 WATSONIAN (3,5,17,1,1) Widespread but infrequent, 2 YORKSHIRE and mainly recorded from the eastern 1 half of the county. The larvae develop 0 NY NZ NZ OV SD SE SE TA in plant roots. 9 8 7 6 5 4 3 2 1

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PORCINOLUS Mulsant & Rey, 1869 murinus (Fabricius, 1794)

Porcinolus murinus (Fabricius, 1794)

3 WATSONIAN (1,1,1,0,0) Notable B. Very scarce in 2 YORKSHIRE Yorkshire with only three known very 1 old records for the county: Castleton 0 NY NZ NZ OV SD SE SE TA (NZ60), 1879, HP; Allerthorpe Common 9 (SE7848), 19/6/1921, WJF; Doncaster 8 (SE50), 1900, EGB. A southern English 7 6 species with few records nationally. 5 4 3 2 1

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Subfamily SYNCALYPTINAE Mulsant & Rey, 1869

CHAETOPHORA Kirby & Spence, 1823 spinosa (Rossi, 1794)

NO RECORDS

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CURIMOPSIS Ganglbauer, 1902 maritima (Marsham, 1802)

Curimopsis maritima (Marsham, 1802)

3 WATSONIAN (2,0,0,0,0) Very rare in the county, with 2 YORKSHIRE only two records: Spurn (TA4012), 1 19/7/1948, WDH, at grass roots in 0 NY NZ NZ OV SD SE SE TA saltmarsh; Spurn (TA4012), 6/1950, 9 WDH, in saltmarsh. There are vouchers 8 for both records in Manchester 7 Museum British Coleoptera collection. 6 5 4 3 2 1

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nigrita (Palm, 1934)

Curimopsis nigrita (Palm, 1934)

3 WATSONIAN (0,0,20,0,0) UKBAP-2007. A byrrhid 2 YORKSHIRE with a very restricted range in the 1 county, known nationally only from 0 NY NZ NZ OV SD SE SE TA Thorne and Hatfield Moors and with a 9 single record from neighbouring Haxey 8 Grange Fen in North Lincolnshire. The 7 species is found in Sphagnum and 6 5 heather litter, and as adults and larvae 4 feed on the roots of mosses. (iv-x). 3 2 1

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setigera (Illiger, 1798)

Curimopsis setigera (Illiger, 1798)

3 WATSONIAN (1,0,0,0,0) Notable A. Very rare with a 2 YORKSHIRE single county record: Flixton Sand Pits 1 (TA0479), 21/5/1921, GBW, at roots of 0 NY NZ NZ OV SD SE SE TA grasses in a sand pit. 9 8 7 6 5 4 3 2 1

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References

Cooter, J., 1999. On ‘English’ names for British Coleoptera. Coleopterist 8 (1): 43-45.

Duff, A.G., (ed.), 2008 Checklist of the of the British Isles, 2008 edition. A.G. Duff publishing, Wells, Somerset.

Duff, A.G., (ed.), 2012 Checklist of Beetles of the British Isles, 2nd edition. Pemberley Books.

Fowler, W.W., 1887. Coleoptera of the British Islands, volume 4. Lovell Reeve, London.

Hyman, P.S., and Parsons, M.S., 1992. A review of the scarce and threatened Coleoptera of Great Britain, part 1. JNCC, Peterborough.

Hyman, P.S. and Parsons, M.S., 1994. A review of the scarce and threatened Coleoptera of Great Britain, part 2. JNCC, Peterborough.

Johnson, C.J., 1992. Ent. Rec. J. Var. 104:305-306, short notes on British species of Elodes.

Johnson, C.J., 1997. Clambus simsoni Blackburn (Col., Clambidae) new to Britain, with notes on its wider distribution. Ent. Mon. Mag. 133(1997):161-164.

Kevan, D.K., 1962. The British species of the genus Cyphon Paykull (Col., Helodidae), including three new to the British list. Ent. Mon. Mag. 98:114-121.

Key, R.S., 1999. More on ‘noddy names’ for British beetles. Coleopterist 8 (2): 94.

Levey, B., 1977. Handbooks for the Identification of British V part 1(b). Coleoptera: Buprestidae. Royal Entomological Society, London.

Lohse, G.A., 1979. In: Freude, H., et al. Die Kafer Mitteleuropas 6:255. Goecke & Evers, Krefeld.

Pope, R.D., 1977. Kloet & Hincks. A Check List of British Insects. Part 3: Coleoptera and Strepsiptera. Second revised edition. Handbks. Ident. Brit. Insects 11(3), pp. xiv+105. Royal Entomological Society, London.

Shirt, D.B., (ed), (1987). British Red Data Books: 2. Insects. Nature Conservancy Council, Peterborough.

Errors and omissions

The author would be pleased to receive feedback regarding any errors or omissions discovered by users of this Atlas.

R J Marsh FRES

YNU Coleoptera recorder 26 April 2017

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