The Occurrence of Gigantic Marsupials in Tasmania
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SUPPLEMENTARY INFORMATION for a New Family of Diprotodontian Marsupials from the Latest Oligocene of Australia and the Evolution
Title A new family of diprotodontian marsupials from the latest Oligocene of Australia and the evolution of wombats, koalas, and their relatives (Vombatiformes) Authors Beck, RMD; Louys, J; Brewer, Philippa; Archer, M; Black, KH; Tedford, RH Date Submitted 2020-10-13 SUPPLEMENTARY INFORMATION FOR A new family of diprotodontian marsupials from the latest Oligocene of Australia and the evolution of wombats, koalas, and their relatives (Vombatiformes) Robin M. D. Beck1,2*, Julien Louys3, Philippa Brewer4, Michael Archer2, Karen H. Black2, Richard H. Tedford5 (deceased) 1Ecosystems and Environment Research Centre, School of Science, Engineering and Environment, University of Salford, Manchester, UK 2PANGEA Research Centre, School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, New South Wales, Australia 3Australian Research Centre for Human Evolution, Environmental Futures Research Institute, Griffith University, Queensland, Australia 4Department of Earth Sciences, Natural History Museum, London, United Kingdom 5Division of Paleontology, American Museum of Natural History, New York, USA Correspondence and requests for materials should be addressed to R.M.D.B (email: [email protected]) This pdf includes: Supplementary figures Supplementary tables Comparative material Full description Relevance of Marada arcanum List of morphological characters Morphological matrix in NEXUS format Justification for body mass estimates References Figure S1. Rostrum of holotype and only known specimen of Mukupirna nambensis gen. et. sp. nov. (AMNH FM 102646) in ventromedial (a) and anteroventral (b) views. Abbreviations: C1a, upper canine alveolus; I1a, first upper incisor alveolus; I2a, second upper incisor alveolus; I1a, third upper incisor alveolus; P3, third upper premolar. Scale bar = 1 cm. -
Marsupial Lions & Methodological Omnivory
Marsupial Lions & Methodological Omnivory: Function, Success and Reconstruction in Paleobiology Penultimate Version, published in Biology & Philosophy Abstract Historical scientists frequently face incomplete data, and lack direct experimental access to their targets. This has led some philosophers and scientists to be pessimistic about the epistemic potential of the historical sciences. And yet, historical science often produces plausible, sophisticated hypotheses. I explain this capacity to generate knowledge in the face of apparent evidential scarcity by examining recent work on Thylacoleo carnifex, the ‘marsupial lion’. Here, we see two important methodological features. First, historical scientists are methodological omnivores, that is, they construct purpose-built epistemic tools tailored to generate evidence about highly specific targets. This allows them to produce multiple streams of independent evidence and thus maximize their epistemic reach. Second, investigative scaffolding: research proceeds in a piece-meal fashion, information only gaining evidential relevance once certain hypotheses are well supported. I illustrate scaffolding in a discussion of the nature of functional ascription in paleobiology. Frequently, different senses of ‘function’ are not discriminated during paleobiological investigation—something which can mar adaptationist investigations of extant organisms. However, I argue that, due to scaffolding, conflating senses of ‘function’ can be the right thing to do. Coarse grained functional hypotheses are required before it is clear what evidence could discriminate between more fine-grained ones. I draw on omnivory and scaffolding to argue that pessimists make a bad empirical bet. It is a bad idea to bet against the epistemic fortunes of such opportunistic and resourceful scientists, especially when we have reason to think we will systematically underestimate the amount of evidence ultimately available to them. -
THYLACOLEO CARNIFEX and the NARACOORTE CAVES Michael Curry, Liz Reed1,2 and Steve Bourne3
RESEARCH CATCHING the MARSUPIAL ‘LION’ by the TAIL: THYLACOLEO CARNIFEX and the NARACOORTE CAVES Michael Curry, Liz Reed1,2 and Steve Bourne3 1School of Physical Sciences, The University of Adelaide, Adelaide, SA, Australia; 2School of Biological Sciences, Flinders University, Bedford Park, SA, Australia; 3Naracoorte Lucindale Council, Naracoorte, SA, Australia. “Thylacoleo exemplifies the simplest and most effective dental machinery for predatory life and carnivorous diet known in the Mammalian class. It is the extreme modification, to this end, of the Diprotodont type of Marsupialia.” Owen (1866) Introduction defending Thylacoleo as “A very gentle beast, and of good conscience” (Macleay 1859). Macleay based his Of all the extinct Australian Pleistocene megafauna argument on Thylacoleo’s relationship with other species, Thylacoleo carnifex (the marsupial ‘lion’) has Diprotodont marsupials, most of which are herbivores. captured the imagination and interest of people more Gerard Krefft, Curator of the Australian Museum, was than any other. Perhaps it is the allure of its predatory almost equally as unimpressed with Thylacoleo’s habits, (Australia’s Pleistocene answer to T. rex); or the carnivory, opining that it “…was not much more intriguing notion that it used caves as dens (Lundelius, carnivorous than the Phalangers (possums) of present 1966 ). It is certainly an enigma and, as Owen (1866) time.” (Krefft, 1866). Owen, meanwhile, had received an suggested, an extreme and meat-eating version of the almost complete skull from the Darling Downs, in otherwise herbivorous diprotodont marsupials. Queensland and published a more detailed paper, Spectacular fossil finds over the past few decades have further describing the skull and teeth of Thylacoleo, put to rest much of the speculation regarding its habits acknowledging its diprotodont affiliation but more and morphology. -
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13 STUDIES IN TASMANIAN MAMMALS, LIVING AND EXTINCT. Number I. Nototherium mitchelli. (A Marsupial Rhinoceros.) lierium. mitchelli, Owen, British Association for Advancement of Science, Report 1844, p. ?Zygomaturus trilobus, De Vis, Proceedings Royal S of Queensland. 1888, Vol. V., pt. .1 p. 111. By I!. 11. Scott (Curator, Launceston Museum); and Clive E. Lord (Curator, Tasmanian Museum). (Received 3rd May, L920. Read 10th May, 1920.) The discovery at Smithton, during the present year, <• of a nearly complete i Nototherium mitchelli forms the occasion for a revision of many of our ideas - - markable marsupial animals, since 1 1 * I ntarv remains hitherto available for study have 1 to yield the sequence of evidence we now possess, be only— inf nded to place upon record the hat. Nototherium mitchelli was an extinct marsupial rhino< I that the four genera, Nototherium, Zygo- maturus, Euowenia, and Sthenomerus, with their - species, are accordingly under revision— and will later on be dealt with in detail. ma > Material 1 in ibids anything like speculation ; . hut it is within the mark to observe thai two groups of these animals have be instinctively felt (by all workers) to have existed, quite irrespective of sex ques- tion- —one a platyrhine and the other a latifrons type, and that it now appears that they were also a horned, and a hornless group, and that Nototherium mitchelli belonged to the former, or cerathine group, and that some other species constituted the acerathine group, in which the 14 STUDIES IN TASMANIAN MAMMALS, LIVING AND EXTINCT, weapons were reduced to very small things, or actually missing. -
A New Family of Diprotodontian Marsupials from the Latest Oligocene of Australia and the Evolution of Wombats, Koalas, and Their Relatives (Vombatiformes) Robin M
www.nature.com/scientificreports OPEN A new family of diprotodontian marsupials from the latest Oligocene of Australia and the evolution of wombats, koalas, and their relatives (Vombatiformes) Robin M. D. Beck1,2 ✉ , Julien Louys3, Philippa Brewer4, Michael Archer2, Karen H. Black2 & Richard H. Tedford5,6 We describe the partial cranium and skeleton of a new diprotodontian marsupial from the late Oligocene (~26–25 Ma) Namba Formation of South Australia. This is one of the oldest Australian marsupial fossils known from an associated skeleton and it reveals previously unsuspected morphological diversity within Vombatiformes, the clade that includes wombats (Vombatidae), koalas (Phascolarctidae) and several extinct families. Several aspects of the skull and teeth of the new taxon, which we refer to a new family, are intermediate between members of the fossil family Wynyardiidae and wombats. Its postcranial skeleton exhibits features associated with scratch-digging, but it is unlikely to have been a true burrower. Body mass estimates based on postcranial dimensions range between 143 and 171 kg, suggesting that it was ~5 times larger than living wombats. Phylogenetic analysis based on 79 craniodental and 20 postcranial characters places the new taxon as sister to vombatids, with which it forms the superfamily Vombatoidea as defned here. It suggests that the highly derived vombatids evolved from wynyardiid-like ancestors, and that scratch-digging adaptations evolved in vombatoids prior to the appearance of the ever-growing (hypselodont) molars that are a characteristic feature of all post-Miocene vombatids. Ancestral state reconstructions on our preferred phylogeny suggest that bunolophodont molars are plesiomorphic for vombatiforms, with full lophodonty (characteristic of diprotodontoids) evolving from a selenodont morphology that was retained by phascolarctids and ilariids, and wynyardiids and vombatoids retaining an intermediate selenolophodont condition. -
The Chinchilla Local Fauna: an Exceptionally Rich and Well-Preserved Pliocene Vertebrate Assemblage from Fluviatile Deposits of South-Eastern Queensland, Australia
The Chinchilla Local Fauna: An exceptionally rich and well-preserved Pliocene vertebrate assemblage from fluviatile deposits of south-eastern Queensland, Australia JULIEN LOUYS and GILBERT J. PRICE Louys, J. and Price, G.J. 2015. The Chinchilla Local Fauna: An exceptionally rich and well-preserved Pliocene verte- brate assemblage from fluviatile deposits of south-eastern Queensland, Australia. Acta Palaeontologica Polonica 60 (3): 551–572. The Chinchilla Sand is a formally defined stratigraphic sequence of Pliocene fluviatile deposits that comprise interbed- ded clay, sand, and conglomerate located in the western Darling Downs, south-east Queensland, Australia. Vertebrate fossils from the deposits are referred to as the Chinchilla Local Fauna. Despite over a century and a half of collection and study, uncertainties concerning the taxa in the Chinchilla Local Fauna continue, largely from the absence of stratigraph- ically controlled excavations, lost or destroyed specimens, and poorly documented provenance data. Here we present a detailed and updated study of the vertebrate fauna from this site. The Pliocene vertebrate assemblage is represented by at least 63 taxa in 31 families. The Chinchilla Local Fauna is Australia’s largest, richest and best preserved Pliocene ver- tebrate locality, and is eminently suited for palaeoecological and palaeoenvironmental investigations of the late Pliocene. Key words: Mammalia, Marsupialia, Pliocene, Australia, Queensland, Darling Downs. Julien Louys [[email protected]], Department of Archaeology and Natural History, School of Culture, History, and Languages, ANU College of Asia and the Pacific, The Australian National University, Canberra, 0200, Australia. Gilbert J. Price [[email protected]], School of Earth Sciences, The University of Queensland, Brisbane, Queensland, 4072, Australia. -
Systematic and Palaeobiological Implications of Postcranial Morphology in the Diprotodontidae (Marsupialia)
Systematic and palaeobiological implications of postcranial morphology in the Diprotodontidae (Marsupialia) Aaron B. Camens School of Earth and Environmental Sciences Discipline of Ecology and Evolutionary Biology The University of Adelaide South Australia A thesis submitted in partial fulfilment of the degree of Doctor of Philosophy at the University of Adelaide February 2010 II Declaration This work contains no material which has been accepted for the award of any other degree or diploma in any university or other tertiary institution to Aaron Camens and, to the best of my knowledge and belief, contains no material previously published or written by another person, except where due reference has been made in the text. I give consent to this copy of my thesis when deposited in the University Library, being made available for loan and photocopying, subject to the provisions of the Copyright Act 1968. The author acknowledges that copyright of published works contained within this thesis (as listed below) resides with the copyright holder(s) of those works. I also give permission for the digital version of my thesis to be made available on the web, via the University’s digital research repository, the Library catalogue, the Australasian Digital Theses Program (ADTP) and also through web search engines, unless permission has been granted by the University to restrict access for a period of time. Publications in this thesis include: Camens, A. B. and Wells, R.T. 2009. Diprotodontid footprints from the Pliocene of Central Australia. Journal of Vertebrate Paleontology 29: 863-869. Copyright held by Taylor and Francis. Camens, A. B. and Wells, R.T. -
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https://doi.org/10.24199/j.mmv.1973.34.03 9 May 1973 FOSSIL VERTEBRATE FAUNAS FROM THE LAKE VICTORIA REGION, S.W. NEW SOUTH WALES, AUSTRALIA By Larry G. Marshall Zoology Department, Monash University, Victoria* Abstract Fossil vertebrate localities and faunas in the Lake Victoria region of S.W. New South Wales, Australia, are described. The oldest fossil bearing deposit, the late Pliocene or early Pleistocene Moorna Formation and associated Chowilla Sand have yielded specimens of Neoceratodus sp., Emydura macquarrii, several species of small dasyurid, specimens of Glaucodon cf. G. ballaratensis, a species of Protemnodon which compares closely with P. cf. P. otibandus from the late Pliocene or early Pleistocene Chinchilla Sand in SE. Queensland, specimens of Lagostrophus cf L. fasciatus, species of Petrogale, Macropus, Osphranter, Sthenurus, Bettongia, Diprotodon, Lasiorhinus, a peramelid, at least two species of pseudomyine rodents and a species of Rattus cf. R. lutreolus. It is shown that the holotype of Zygomaturus victoriae (Owen) 1872 may have been collected from these sediments. The late Pliocene or early Pleistocene Blanchetown Clay has yielded species of Neoceratodus, Thylacoleo, Phas- colonus, Bettongia, Sthenurus, a diprotodontid, and a rodent. The late Pleistocene Rufus Formation has yielded species of Dasycercus, Sarcophilus, Thylacinus, Phascolonus, Lasiorhinus, Bettongia, Procoptodon, Onychogalea, Macropus, and Leporillus. A large species of macropod and a species of Phascolonus were collected from the late Pleistocene Monoman Formation. The lunette on the E. side of Lake Victoria has yielded a large, diverse fauna of late Pleistocene —Holocene age, that includes such extinct species as Protemnodon anak, P. brehus, Procoptodon goliah, Sthenurus andersoni, S. -
Thy Thylacoleo Is a Thylacine. Australian Archaeology, No. 80
In this issue... NUMBER 80 | JUNE 2015 • ‘Small, individually nondescript and • Mapping a millstone: The dynamics easily overlooked’: Contact beads of use-wear and residues on a Central from northwest Arnhem Land in an Australian seed-grinding implement | Indigenous-Macassan-European Mike Smith, Elspeth Hayes and hybrid economy | Daryl Wesley Birgitta Stephenson and Mirani Litster • Compliance-based archaeological • The palaeo-environmental history heritage management and place-based of Big Willum Swamp, Weipa: An participatory mapping for negotiated environmental context for the outcomes | David R. Guilfoyle and archaeological record | Janelle Myles B. Mitchell Stevenson, Sally Brockwell, • Attributes, preservation and Cassandra Rowe, Ulrike Proske management of dendroglyphs and Justin Shiner from the Wet Tropics rainforest of • A multidisciplinary investigation northeast Australia | Alice Buhrich, of a rock coating at Ngaut Ngaut Åsa Ferrier and Gordon Grimwade (Devon Downs), South Australia | • Mid-Holocene exploitation of marine Amy Roberts, Isobelle Campbell, molluscs in the lower Mid West, Allan Pring, Graham Bell, Alan Western Australia | Carly Monks, Watchman, Rachel S. Popelka-Filcoff, Bob Sheppard and Joe Dortch Claire E. Lenehan, Christopher T. Gibson, Natalie Franklin and the • The archaeology of Bindjarran Mannum Aboriginal Community rockshelter in Manilikarr Country, Association Inc. (MACAI) Kakadu National Park, Northern Territory | Denis Shine, Melissa • Thy Thylacoleo is a thylacine | Marshall, Duncan Wright, Tim David M. Welch Denham, Peter Hiscock, Geraldine NUMBER 80 • A fine-grained analysis of the Jacobsen and Sean-Paul Stephens macropod motif in the rock art of the • The Brremangurey pearl: A 2000 Sydney region, Australia | Alandra K. year old archaeological find from the Tasire and Iain Davidson coastal Kimberley, Western Australia | • Investigating standardisation in the Katherine Szabo, Brent Koppel, Mark | JUNE 2015 form of backed artefacts at two sites W. -
Rock Art Thematic Study
Rock Art Thematic Study Jo McDonald and Lucia Clayton 26 May 2016 Report to the Department of the Environment and the Australian Heritage Council Centre for Rock Art Research and Management, University of WA Rock Art Thematic Study Page ii Table of Contents 1 Introduction ....................................................................................................................................................... 1 2 Rock art overview ............................................................................................................................................... 2 2.1 Introduction to rock art ............................................................................................................................... 2 2.2 Regional overview of Australian Aboriginal rock art ..................................................................................... 3 2.2.1 Australian Capital Territory (ACT) ....................................................................................................................... 7 2.2.2 New South Wales ................................................................................................................................................ 7 2.2.3 Northern Territory ............................................................................................................................................. 14 2.2.4 Queensland ...................................................................................................................................................... -
Nimbadon, a I:,Iew Genus and Three New Species Of
NIMBADON,A I:,IEWGENUS AND THREENEW SPECIESOF TERTIARY ZYGOMATURINES(MARSUPIALIA: DIPROTODONTIDAE) FROM NORTHERN AUSTRALIA,WITH A REASSESSMENTOF N'OI''ZOS S.J. HAND, M. ARC}IER,H. C,ODTHELP.T.H. RICH AND N.S. PLEDGE Hand, S.J.,Archer, M., Codthelp, H., Rich, T.H. & Pledge,N. S. 199306 3O:Nimbadon,a new genusand three new speciesofTeniary zygomaturines(Marsupialia: Diprotodontidae) from nonhem Australia, with a reassessmentof Neohelos.Memoirs of the Queenslnnd Museum 33(l\: 193-210.Brisbane. ISSN 0079-8835. Thrce new species of Oligo-Miocene zygomatu.ine dip.otodontids are described from northem Australia. All are small,l, plesiomorphicplesiomorDhicand appearto comprise a distinctive clade of zygomaturines.naEed here Nimbadon.Nimbalon. TlreThe clade is Dartlvpartly defined on the basis of the posleriorly iqclined P" parastyleand blade exrendingfrom the parametaconeto the lingual halfofhalfofthe the P'crown and then to lhe anterolinpualanterolingualcinpulum.cingulum. Two ofthe sDe.iesspe.ies areiue known from Oligo-Miocenelocalfaunas ofRiversleigh Slalion,nonhwestern Queensland. The third is from the middle Miocene Bullock Creek Local Faunaof Camfield Station. norlhwestern Nonhern Territory. Descriptionof additionalfossil materialreferable ro Neohelostrrarensrs Stinon, 1967,a middle Miocene zygomaturinefrom the Kutjamarpu lacal Faunaoi South Australia, enablesthe genus to be distin9tished frcm Nimbadon. The chronoslratigraphic significanceof the ncw zygomaturinesis consideredand found lo approximalelycorrespond to curent undeBtandingofthe relative agesof the dcpositsfrom which the specimenswere obtained. n Diprotodontidae, Z;gomaturinae, Ninbadon, Neohelos, Oligo-Miocene, Riversleigh, Bullock Creek local Fauna, Henk's Hollow lacal Fautv, Fig Tree lncal Suzonne Hand, M. Archer & H. Godthelp, School of Biologicol Science, UniversiD, of New South Wales, PO Box I, Kensinglon, New South llales 2033, Australia: T.H. -
A Tiny New Marsupial Lion (Marsupialia, Thylacoleonidae) from the Early Miocene of Australia
Palaeontologia Electronica palaeo-electronica.org A tiny new marsupial lion (Marsupialia, Thylacoleonidae) from the early Miocene of Australia Anna K. Gillespie, Michael Archer, and Suzanne J. Hand ABSTRACT Microleo attenboroughi, a new genus and species of diminutive marsupial lion (Marsupialia: Thylacoleonidae), is described from early Miocene freshwater limestones in the Riversleigh World Heritage Area, northwestern Queensland, Australia. A broken palate that retains incomplete cheektooth rows demonstrates that this new, very small marsupial lion possessed the elongate, trenchant P3 and predominantly subtriangular upper molars characteristic of thylacoleonids, while other features of the premolar sup- port its placement in a new genus. Phylogenetic analysis suggests that Microleo atten- boroughi is the sister taxon to all other thylacoleonids, and that Thylacoleonidae may lie outside Vombatomorphia as the sister taxon of all other wombat-like marsupials including koalas. However, given limited data about the cranial morphology of M. atten- boroughi, Thylacoleonidae is concluded here, conservatively, to be part of the vom- batomorphian clade. This new thylacoleonid brings to three the number of marsupial lion species that have been recovered from early Miocene deposits at Riversleigh and indicates a level of diversity previously not seen for this group. It is likely that the differ- ent size and morphology of the three sympatric taxa reflects niche partitioning and hence reduced competition. Thylacoleonids may have been the dominant arboreal predators of Cenozoic Australia. Anna K. Gillespie*, Palaeontology, Geobiology and Earth Archives (PANGEA) Research Centre, School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney 2052, Australia; [email protected], *corresponding author Michael Archer, Palaeontology, Geobiology and Earth Archives (PANGEA) Research Centre, School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney 2052, Australia; [email protected] Suzanne J.