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Smithsonian Miscellaneous Collections Volume 101
SMITHSONIAN MISCELLANEOUS COLLECTIONS VOLUME 101. NUMBER 15 FIFTH CONTRIBUTION TO NOMENCLATURE OF CAMBRIAN FOSSILS BY CHARLES E. RESSER Curator, Division of Stratigraphic Paleontology U. S. National Museum (Publication 3682) CITY OF WASHINGTON PUBLISHED BY THE SMITHSONIAN INSTITUTION MAY 22, 1942 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOLUME 101, NUMBER 15 FIFTH CONTRIBUTION TO NOMENCLATURE OF CAMBRIAN FOSSILS BY CHARLES E. RESSER Curator, Division of Stratigraphic Paleontology U. S. National Museum (Publication 3682) CITY OF WASHINGTON PUBLISHED BY THE SMITHSONIAN INSTITUTION MAY 12, 1942 Z?>i Botb QBafhtnore (prcee BALTIMORE, MD., U. S. A. ' FIFTH CONTRIBUTION TO NOMENCLATURE OF CAMBRIAN FOSSILS By CHARLES E. RESSER Curator, Division of Stratigraphic Paleonlolo<jy, U. S. National Museum This is the fifth in the series of papers designed to care for changes necessary in the names of Cambrian fossils. When the fourth paper was published it was hoped that further changes would be so few and so obvious that they could be incorporated in the Cambrian bibliographic summary, and would not be required to appear first in a separate paper. But even now it is impossible to gather all of the known errors for rectification in this paper. For example, correc- tion of some errors must await the opportunity to examine the speci- mens because the published illustrations, obviously showing incorrect generic determinations, are too poor to permit a proper understanding of the fossil. In the other instances where new generic designations are clearly indicated, erection of new genera should await the pub- lication of a paper with illustrations, because better-preserved speci- mens are in hand, or undescribed species portray the generic charac- teristics more fully and should therefore be chosen as the genotypes. -
An Exceptionally Preserved Arthropod Cardiovascular System from the Early Cambrian
ARTICLE Received 20 Dec 2013 | Accepted 4 Mar 2014 | Published 7 Apr 2014 DOI: 10.1038/ncomms4560 An exceptionally preserved arthropod cardiovascular system from the early Cambrian Xiaoya Ma1,2, Peiyun Cong1, Xianguang Hou1, Gregory D. Edgecombe2 & Nicholas J. Strausfeld3 The assumption that amongst internal organs of early arthropods only the digestive system withstands fossilization is challenged by the identification of brain and ganglia in early Cambrian fuxianhuiids and megacheirans from southwest China. Here we document in the 520-million-year-old Chengjiang arthropod Fuxianhuia protensa an exceptionally preserved bilaterally symmetrical organ system corresponding to the vascular system of extant arthropods. Preserved primarily as carbon, this system includes a broad dorsal vessel extending through the thorax to the brain where anastomosing branches overlap brain seg- ments and supply the eyes and antennae. The dorsal vessel provides segmentally paired branches to lateral vessels, an arthropod ground pattern character, and extends into the anterior part of the abdomen. The addition of its vascular system to documented digestive and nervous systems resolves the internal organization of F. protensa as the most completely understood of any Cambrian arthropod, emphasizing complexity that had evolved by the early Cambrian. 1 Yunnan Key Laboratory for Palaeobiology, Yunnan University, Kunming 650091, China. 2 Department of Earth Sciences, The Natural History Museum, Cromwell Road, London SW7 5BD, UK. 3 Department of Neuroscience and Center for Insect Science, University of Arizona, Tucson, Arizona 85721, USA. Correspondence and requests for materials should be addressed to X.H. (email: [email protected]) or to N.J.S. (email: fl[email protected]). -
A NEW METAZOAN from the MIDDLE CAMBRIAN of UTAH and the NATURE of the VETULICOLIA by DEREK E
[Palaeontology, Vol. 48, Part 4, 2005, pp. 681–686] RAPID COMMUNICATION A NEW METAZOAN FROM THE MIDDLE CAMBRIAN OF UTAH AND THE NATURE OF THE VETULICOLIA by DEREK E. G. BRIGGS*, BRUCES.LIEBERMAN , SUSAN L. HALGEDAHLà and RICHARD D. JARRARDà *Department of Geology and Geophysics, Yale University, PO Box 208109, New Haven, CT 06520-8109, USA; e-mail: [email protected] Department of Geology, University of Kansas, 1475 Jayhawk Boulevard, 120 Lindley Hall, Lawrence, KS 66045, USA àDepartment of Geology and Geophysics, University of Utah, 135 S. 1460 East, Salt Lake City, UT 84112, USA Typescript received 22 November 2004; accepted in revised form 24 March 2005 Abstract: A new metazoan, Skeemella clavula gen. et sp. arthropodan in character. The similarity of this fossil nov., is described from the Middle Cambrian Pierson Cove to vetulicolians throws hypotheses of their deuterostome Formation of the Drum Mountains, Utah, USA. Skeemella affinity into question and highlights their problematic sta- is similar to vetulicolians, but differs from other examples tus. of this group in the relative proportions of the anterior and posterior sections, the large number of divisions, and Key words: vetulicolian, Cambrian, deuterostome, arthro- the elongate bifid termination. The posterior section is pod. The very name vetulicolian conjures up creatures from species Banffia confusa, to erect a new class of stem-group another planet. These extraordinary fossils, with a head arthropods, the Vetulicolida. They argued that Banffia shield-like anterior and narrow segmented trunk-like pos- constricta from the Burgess Shale belongs to the same terior, are the latest Cambrian group to be accorded phy- class, extending its range to North America. -
Fossils from South China Redefine the Ancestral Euarthropod Body Plan Cédric Aria1 , Fangchen Zhao1, Han Zeng1, Jin Guo2 and Maoyan Zhu1,3*
Aria et al. BMC Evolutionary Biology (2020) 20:4 https://doi.org/10.1186/s12862-019-1560-7 RESEARCH ARTICLE Open Access Fossils from South China redefine the ancestral euarthropod body plan Cédric Aria1 , Fangchen Zhao1, Han Zeng1, Jin Guo2 and Maoyan Zhu1,3* Abstract Background: Early Cambrian Lagerstätten from China have greatly enriched our perspective on the early evolution of animals, particularly arthropods. However, recent studies have shown that many of these early fossil arthropods were more derived than previously thought, casting uncertainty on the ancestral euarthropod body plan. In addition, evidence from fossilized neural tissues conflicts with external morphology, in particular regarding the homology of the frontalmost appendage. Results: Here we redescribe the multisegmented megacheirans Fortiforceps and Jianfengia and describe Sklerolibyon maomima gen. et sp. nov., which we place in Jianfengiidae, fam. nov. (in Megacheira, emended). We find that jianfengiids show high morphological diversity among megacheirans, both in trunk ornamentation and head anatomy, which encompasses from 2 to 4 post-frontal appendage pairs. These taxa are also characterized by elongate podomeres likely forming seven-segmented endopods, which were misinterpreted in their original descriptions. Plesiomorphic traits also clarify their connection with more ancestral taxa. The structure and position of the “great appendages” relative to likely sensory antero-medial protrusions, as well as the presence of optic peduncles and sclerites, point to an overall -
The Weeks Formation Konservat-Lagerstätte and the Evolutionary Transition of Cambrian Marine Life
Downloaded from http://jgs.lyellcollection.org/ by guest on October 1, 2021 Review focus Journal of the Geological Society Published Online First https://doi.org/10.1144/jgs2018-042 The Weeks Formation Konservat-Lagerstätte and the evolutionary transition of Cambrian marine life Rudy Lerosey-Aubril1*, Robert R. Gaines2, Thomas A. Hegna3, Javier Ortega-Hernández4,5, Peter Van Roy6, Carlo Kier7 & Enrico Bonino7 1 Palaeoscience Research Centre, School of Environmental and Rural Science, University of New England, Armidale, NSW 2351, Australia 2 Geology Department, Pomona College, Claremont, CA 91711, USA 3 Department of Geology, Western Illinois University, 113 Tillman Hall, 1 University Circle, Macomb, IL 61455, USA 4 Department of Zoology, University of Cambridge, Downing Street, Cambridge CB2 3EJ, UK 5 Museum of Comparative Zoology and Department of Organismic and Evolutionary Biology, Harvard University, 26 Oxford Street, Cambridge, MA 02138, USA 6 Department of Geology, Ghent University, Krijgslaan 281/S8, B-9000 Ghent, Belgium 7 Back to the Past Museum, Carretera Cancún, Puerto Morelos, Quintana Roo 77580, Mexico R.L.-A., 0000-0003-2256-1872; R.R.G., 0000-0002-3713-5764; T.A.H., 0000-0001-9067-8787; J.O.-H., 0000-0002- 6801-7373 * Correspondence: [email protected] Abstract: The Weeks Formation in Utah is the youngest (c. 499 Ma) and least studied Cambrian Lagerstätte of the western USA. It preserves a diverse, exceptionally preserved fauna that inhabited a relatively deep water environment at the offshore margin of a carbonate platform, resembling the setting of the underlying Wheeler and Marjum formations. However, the Weeks fauna differs significantly in composition from the other remarkable biotas of the Cambrian Series 3 of Utah, suggesting a significant Guzhangian faunal restructuring. -
Early Cambrian Fuxianhuiids from China Reveal Origin of the Gnathobasic Protopodite in Euarthropods
Early Cambrian fuxianhuiids from China reveal origin of the gnathobasic protopodite in euarthropods The Harvard community has made this article openly available. Please share how this access benefits you. Your story matters Citation Yang, Jie, Javier Ortega-Hernández, David A. Legg, Tian Lan, Jin-bo Hou, and Xi-guang Zhang. 2018. “Early Cambrian fuxianhuiids from China reveal origin of the gnathobasic protopodite in euarthropods.” Nature Communications 9 (1): 470. doi:10.1038/s41467-017-02754-z. http://dx.doi.org/10.1038/s41467-017-02754-z. Published Version doi:10.1038/s41467-017-02754-z Citable link http://nrs.harvard.edu/urn-3:HUL.InstRepos:35015071 Terms of Use This article was downloaded from Harvard University’s DASH repository, and is made available under the terms and conditions applicable to Other Posted Material, as set forth at http:// nrs.harvard.edu/urn-3:HUL.InstRepos:dash.current.terms-of- use#LAA ARTICLE DOI: 10.1038/s41467-017-02754-z OPEN Early Cambrian fuxianhuiids from China reveal origin of the gnathobasic protopodite in euarthropods Jie Yang1, Javier Ortega-Hernández 2,3, David A. Legg 4, Tian Lan5, Jin-bo Hou1 & Xi-guang Zhang 1 Euarthropods owe their evolutionary and ecological success to the morphological plasticity of their appendages. Although this variability is partly expressed in the specialization of the 1234567890():,; protopodite for a feeding function in the post-deutocerebral limbs, the origin of the former structure among Cambrian representatives remains uncertain. Here, we describe Alacaris mirabilis gen. et sp. nov. from the early Cambrian Xiaoshiba Lagerstätte in China, which reveals the proximal organization of fuxianhuiid appendages in exceptional detail. -
Early and Middle Cambrian Trilobites from Antarctica
Early and Middle Cambrian Trilobites From Antarctica GEOLOGICAL SURVEY PROFESSIONAL PAPER 456-D Early and Middle Cambrian Trilobites From Antarctica By ALLISON R. PALMER and COLIN G. GATEHOUSE CONTRIBUTIONS TO THE GEOLOGY OF ANTARCTICA GEOLOGICAL SURVEY PROFESSIONAL PAPER 456-D Bio stratigraphy and regional significance of nine trilobite faunules from Antarctic outcrops and moraines; 28 species representing 21 genera are described UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON : 1972 UNITED STATES DEPARTMENT OF THE INTERIOR ROGERS C. B. MORTON, Secretary GEOLOGICAL SURVEY V. E. McKelvey, Director Library of Congress catalog-card No. 73-190734 For sale by the Superintendent of Documents, U.S. Government Printing Office Washington, D.C. 20402 - Price 70 cents (paper cover) Stock Number 2401-2071 CONTENTS Page Page Abstract_ _ ________________________ Dl Physical stratigraphy______________________________ D6 I&troduction. _______________________ 1 Regional correlation within Antarctica ________________ 7 Biostratigraphy _____________________ 3 Systematic paleontology._____-_______-____-_-_-----_ 9 Early Cambrian faunules.________ 4 Summary of classification of Antarctic Early and Australaspis magnus faunule_ 4 Chorbusulina wilkesi faunule _ _ 5 Middle Cambrian trilobites. ___________________ 9 Chorbusulina subdita faunule _ _ 5 Agnostida__ _ _________-____-_--____-----__---_ 9 Early Middle Cambrian f aunules __ 5 Redlichiida. __-_--------------------------_---- 12 Xystridura mutilinia faunule- _ 5 Corynexochida._________--________-_-_---_----_ -
An Appraisal of the Great Basin Middle Cambrian Trilobites Described Before 1900
An Appraisal of the Great Basin Middle Cambrian Trilobites Described Before 1900 By ALLISON R. PALMER A SHORTER CONTRIBUTION TO GENERAL GEOLOGY GEOLOGICAL SURVEY PROFESSIONAL PAPER 264-D Of the 2ty species described prior to I(?OO, 2/ are redescribed and 2C} refigured, and a new name is proposedfor I species UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON : 1954 UNITED STATES DEPARTMENT OF THE INTERIOR Douglas McKay, Secretary GEOLOGICAL SURVEY W. E. Wrather, Director For sale by the Superintendent of Documents, U. S. Government Printing Office Washington 25, D. C. - Price $1 (paper cover) CONTENTS Page Abstract..__________________________________ 55 Introduction ________________________________ 55 Original and present taxonomic names of species. 57 Stratigraphic distribution of species ____________ 57 Collection localities._________________________ 58 Systematic descriptions.______________________ 59 Literature cited____________________________ 82 Index __-_-__-__---_--______________________ 85 ILLUSTRATIONS [Plates 13-17 follow page 86] PLATE 13. Agnostidae and Dolichometopidae 14. Dorypygidae 15. Oryctocephalidae, Dorypygidae, Zacanthoididae, and Ptychoparioidea 16. Ptychoparioidea 17. Ptychoparioidea FIGUBE 3. Index map showing collecting localities____________________________ . Page 56 in A SHORTER CONTRIBUTION TO GENERAL GEOLOGY AN APPRAISAL OF THE GREAT BASIN MIDDLE CAMBRIAN TRILOBITES DESCRIBED BEFORE 1900 By ALLISON R. PALMER ABSTRACT the species and changes in their generic assignments All 29 species of Middle Cambrian trilobites -
Arthropod Pattern Theory and Cambrian Trilobites
Bijdragen tot de Dierkunde, 64 (4) 193-213 (1995) SPB Academie Publishing bv, The Hague Arthropod pattern theory and Cambrian trilobites Frederick A. Sundberg Research Associate, Invertebrate Paleontology Section, Los Angeles County Museum of Natural History, 900 Exposition Boulevard, Los Angeles, California 90007, USA Keywords: Arthropod pattern theory, Cambrian, trilobites, segment distributions 4 Abstract ou 6). La limite thorax/pygidium se trouve généralementau niveau du node 2 (duplomères 11—13) et du node 3 (duplomères les les 18—20) pour Corynexochides et respectivement pour Pty- An analysis of duplomere (= segment) distribution within the chopariides.Cette limite se trouve dans le champ 4 (duplomères cephalon,thorax, and pygidium of Cambrian trilobites was un- 21—n) dans le cas des Olenellides et des Redlichiides. L’extrémité dertaken to determine if the Arthropod Pattern Theory (APT) du corps se trouve généralementau niveau du node 3 chez les proposed by Schram & Emerson (1991) applies to Cambrian Corynexochides, et au niveau du champ 4 chez les Olenellides, trilobites. The boundary of the cephalon/thorax occurs within les Redlichiides et les Ptychopariides. D’autre part, les épines 1 4 the predicted duplomerenode (duplomeres or 6). The bound- macropleurales, qui pourraient indiquer l’emplacement des ary between the thorax and pygidium generally occurs within gonopores ou de l’anus, sont généralementsituées au niveau des node 2 (duplomeres 11—13) and node 3 (duplomeres 18—20) for duplomères pronostiqués. La limite prothorax/opisthothorax corynexochids and ptychopariids, respectively. This boundary des Olenellides est située dans le node 3 ou près de celui-ci. Ces occurs within field 4 (duplomeres21—n) for olenellids and red- résultats indiquent que nombre et distribution des duplomères lichiids. -
The Leanchoilia-Ottoia Fauna from the Middle Cambrian Burgess Shale of British Columbia
77 THE LEANCHOILIA-OTTOIA FAUNA FROM THE MIDDLE CAMBRIAN BURGESS SHALE OF BRITISH COLUMBIA. COLLINS, Desmond, Royal ontario Museum, 100 Queen's Park, Toronto, ontario M5S 2C6, CANADA The Leanchoilia-ottoia fauna from the Raymond quarry level of the Burgess Shale is different in both content and average size to the classic Marrel1a-Burgessia fauna excavated by Walcott from the Phyllopod bed just 20 m below. The animals most common in the fauna, Leanchoilia, Ottoia, Sidneyia and Vauxia, are typically 5 to 10 cm in length, whereas Phyllopod bed animals such as Marrella and Burgessia which make up half of this fauna are only 1 to 2 cm in length. This distinct difference also applies to the major predators, where large Anomalocaris and Hurdia dominate the Leanchoilia-ottoia fauna compared to the smaller Laggania in the Phyllopod bed fauna. Along with the different forms, there are elements common to both faunas, such as Choia, Helmetia, Olenoides, Ottoia, Sidneyia, Tuzoia, Vauxia and Waptia. New discoveries include a large jellyfish, a ctenophore, a "sea moth", a benthic sea-cucumber, Isoxys with eyes and appendages, tubular burrows containing commensal worms and the barnacle, Priscansermarinus, previously found in talus. The environment of burial of the two faunas also differs. Most of the Phyllopod bed animals occur within 3 to 6 cm thick bands, indicating transport from elsewhere. In contrast, many of the Leanchoilia-ottoia animals were buried in life position on the bedding planes, inclUding sessile forms such as the sponge, Chancelloria, rooted in the bedding surface and bent over in parallel.. -
Ecdysis in a Stem-Group Euarthropod from the Early Cambrian of China Received: 2 November 2018 Jie Yang1,2, Javier Ortega-Hernández 3,4, Harriet B
www.nature.com/scientificreports OPEN Ecdysis in a stem-group euarthropod from the early Cambrian of China Received: 2 November 2018 Jie Yang1,2, Javier Ortega-Hernández 3,4, Harriet B. Drage5,6, Kun-sheng Du1,2 & Accepted: 20 March 2019 Xi-guang Zhang1,2 Published: xx xx xxxx Moulting is a fundamental component of the ecdysozoan life cycle, but the fossil record of this strategy is susceptible to preservation biases, making evidence of ecdysis in soft-bodied organisms extremely rare. Here, we report an exceptional specimen of the fuxianhuiid Alacaris mirabilis preserved in the act of moulting from the Cambrian (Stage 3) Xiaoshiba Lagerstätte, South China. The specimen displays a fattened and wrinkled head shield, inverted overlap of the trunk tergites over the head shield, and duplication of exoskeletal elements including the posterior body margins and telson. We interpret this fossil as a discarded exoskeleton overlying the carcass of an emerging individual. The moulting behaviour of A. mirabilis evokes that of decapods, in which the carapace is separated posteriorly and rotated forward from the body, forming a wide gape for the emerging individual. A. mirabilis illuminates the moult strategy of stem-group Euarthropoda, ofers the stratigraphically and phylogenetically earliest direct evidence of ecdysis within total-group Euarthropoda, and represents one of the oldest examples of this growth strategy in the evolution of Ecdysozoa. Te process of moulting consists of the periodical shedding (i.e. ecdysis) of the cuticular exoskeleton during growth that defnes members of Ecdysozoa1, a megadiverse animal group that includes worm-like organisms with radial mouthparts (Priapulida, Loricifera, Nematoida, Kinorhyncha), as well as more familiar forms with clawed paired appendages (Euarthropoda, Tardigrada, Onychophora). -
An Inventory of Trilobites from National Park Service Areas
Sullivan, R.M. and Lucas, S.G., eds., 2016, Fossil Record 5. New Mexico Museum of Natural History and Science Bulletin 74. 179 AN INVENTORY OF TRILOBITES FROM NATIONAL PARK SERVICE AREAS MEGAN R. NORR¹, VINCENT L. SANTUCCI1 and JUSTIN S. TWEET2 1National Park Service. 1201 Eye Street NW, Washington, D.C. 20005; -email: [email protected]; 2Tweet Paleo-Consulting. 9149 79th St. S. Cottage Grove. MN 55016; Abstract—Trilobites represent an extinct group of Paleozoic marine invertebrate fossils that have great scientific interest and public appeal. Trilobites exhibit wide taxonomic diversity and are contained within nine orders of the Class Trilobita. A wealth of scientific literature exists regarding trilobites, their morphology, biostratigraphy, indicators of paleoenvironments, behavior, and other research themes. An inventory of National Park Service areas reveals that fossilized remains of trilobites are documented from within at least 33 NPS units, including Death Valley National Park, Grand Canyon National Park, Yellowstone National Park, and Yukon-Charley Rivers National Preserve. More than 120 trilobite hototype specimens are known from National Park Service areas. INTRODUCTION Of the 262 National Park Service areas identified with paleontological resources, 33 of those units have documented trilobite fossils (Fig. 1). More than 120 holotype specimens of trilobites have been found within National Park Service (NPS) units. Once thriving during the Paleozoic Era (between ~520 and 250 million years ago) and becoming extinct at the end of the Permian Period, trilobites were prone to fossilization due to their hard exoskeletons and the sedimentary marine environments they inhabited. While parks such as Death Valley National Park and Yukon-Charley Rivers National Preserve have reported a great abundance of fossilized trilobites, many other national parks also contain a diverse trilobite fauna.