Project : and Aphodiinae West Africa fauna

To contact :

Philippe MORETTO 5, rue Gimelli F.83000 Toulon FRANCE @ : [email protected] Scarabaeinae and Aphodiinae West Africa fauna

Africa, the continent of the Scarab , will soon benefit from the work of a new task force. The Group Catharsius has just assembled from a group of dedicated individuals interest- ed in developing an identification tool for Africa's yet to be better known Scarab fauna.

Recent collecting efforts in West Africa and taxonomic studies by members of Catharsius including Yves Cambefort, Philippe Moretto, Jean-François Josso, Patrick Prévost, Olivier Montreuil, Patrice Bordat, Frank Krell and François Génier, have generated a large quantity of specimens with precise distributional and ecological data. This material, in addition to material held in museums, will serve as the basis for a published synthesis of current taxonom- ical, distributional and ecological knowledge of Scarabaeinae and Aphodiinae inhabiting West Africa. The group aims to resolve most taxonomic problems for the area through the study of Scarabaeinae and Aphodiinae throughout the entire Afrotropical realm.

The multi-year project will begin by data basing current holdings of Scarabaeinae and Aphodiinae in order to identify areas of West Africa most in need of sampling by modern tech- niques. Catharsius members and interested partners will then target these areas in a series of col- lecting expeditions. Computerization of the newly acquired specimens will be performed accord- ing to standards developed by ScarabNet, with the goal of making the pooled data available to the entire ScarabNet community. Ultimately, the project will result in a monographic publication describing all known taxa, providing identification keys, and illustrating distributional and eco- logical data.

To contact :

Philippe MORETTO 5, rue Gimelli F.83000 Toulon FRANCE @ : [email protected] Yves Cambefort @ : [email protected]

Yves Cambefort has been the first to study West African dung ecology and biogeography in the field (Ivory Coast, etc.), and his 1984 dissertation was the first significant work pub- lished on the subject. The book he coedited in 1991 with Ilkka Hanski was also a landmark in the natural history of dung beetles. Since a few years, although he has been more involved in cultural entomology of beetles in general, he has maintained nevertheless his interest in African dung beetles, group of which he is presently one of the oldest specialists.

A few papers :

Cambefort Y. 1971. Scarabaeinae récoltés dans le Fouta Djalon (Guinée). Bulletin de l'Institut fondamental d'Afrique noire, A, 33 (2) : 420-424. Cambefort Y. 1972. Scarabaeinae récoltés en Mauritanie par J.-L. Amiet. Nouvelle Revue d'Entomologie, 2 (3) : 243- 251. Cambefort Y. 1975. Nouveaux Onthophagides du Sénégal (Col. ). Nouvelle Revue d'Entomologie, 5 (2): 187-189. Cambefort Y. 1981. Coléoptères Scarabaeinae nouveaux ou peu connus de Lamto (Côte d'Ivoire). Bulletin de la Société entomologique de France, 85 (9-10) : 262-265. Paulian R. & Cambefort Y. 1982. Coléoptères Lamellicornes coprophages du parc national du Niokolo Koba. Dans « Recherches scientifiques dans les parcs nationaux du Sénégal, XI ». Mémoires de l'Institut fondamental d'Afrique noire, 92 : 159-163. Cambefort Y. 1983. Nouveaux Scarabaeidae s. str. de Côte-d'Ivoire et de pays voisins (Coleoptera). Revue française d'Entomologie, (N.S.), 5 (1) : 7-9. Cambefort Y. 1984. Étude écologique des Coléoptères Scarabaeidae s. str. de Côte d'Ivoire. Travaux des Chercheurs de Lamto (Côte-d'Ivoire), n° 3. Paris : École Normale Supérieure, viii + 294 + 12 p. Hanski I. & Cambefort Y., éditeurs, 1991. Dung Beetle Ecology. Princeton : Princeton University Press, xiv + 481 p. Cambefort Y. & Nguyen-Phung Th. 1996. On the genus Copris Müller, 1764. Definition and phylogenetic survey of the Afrotropical species-groups (Coleoptera : Scarabaeidae). Journal of African Zoology, 110 (4) : 271-289. Cambefort Y. 1997. Food choice and environment occupancy in Afrotropical dung beetles : A phylogenetic study of two examples (Coleoptera, Scarabaeidae). Dans Grandcolas Ph., éditeur, « The origin of biodiversity in : Phylogenetic tests of evolutionary scenarios ». Mémoires du Muséum national d'Histoire naturelle, 173 : 125-134. Duringer P., Brunet M., Cambefort Y., Beauvilain A., MacKaye H.T., Vignaud P. & Schuster M. 2000. Des boules de bousiers fossiles et leurs terriers dans les sites à Australopithèques du Pliocène tchadien. Bulletin de la Société géologique de France, 171 (2) : 259-269. Cambefort Y. & Bordat P. 2004. Coléoptères Scarabaeidae s. str., Aphodiidae et Ceratocanthidae du mont Nimba et des régions limitrophes. Dans Lamotte M. & Roy R., éditeurs, « Le peuplement du mont Nimba ». Mémoires du Muséum national d'Histoire naturelle, 190 : 551-580. Patrice Bordat @ : [email protected]

Non professional entomologist, Patrice BORDAT has been interested in the systematic of worldwide Aphodiinae, group in which he specialized for the past 25 years. Author and coauthor of more than 60 scientific publica- tions of which the majority are dealing with the Afrotropical fauna, he described over one hundred Aphodiinae species. Through the study of numerous types, primarily Afrotropical, he acquired a very good knowledge of the African fauna. Member of the Société Entomologique de France and the l'Association des Coléoptéristes de la Région Parisienne. Patrice Bordat is also an active publisher of two French journals: "Le Coléoptériste" and "La Nouvelle Revue d'Entomologie".

some publications:

BORDAT P., 2005 - Note Scientifique. Description du mâle de Haroldaphodius panticosus Bordat, 2003 (Coleoptera, , Aphodiidae). Nouvelle Revue d'Entomologie (N. S.), 22 (2) : 192-193. BORDAT P., 2005 - Nouvelles espèces d'Aphodiinae de la région afrotropicale (Coleoptera, Scarabaeoidea). Nouvelle Revue d'Entomologie (N. S.), 22 (1) : 35-61. BORDAT P. & DEGALLIER N., 2004 - Notes sur Rhyssemus plicatus (Germar). Le Coléoptériste [2003], 6 (3) : 207- 208. BORDAT P. & DELLACASA G., 1996 - (Aganocrossus) postpilosus Reitter, 1895, bona species (Coleoptera, Aphodiidae). Bollettino della Societa entomologica italiana, Genova, 128: 143-150. BORDAT P. & DELLACASA G., 1999 - Le type de Psammodius (Sicardia) pierrei Hollande & Thérond, 1998. Nouvelle Revue d'Entomologie (N. S.), 16 : 288. BORDAT P. & HOWDEN H. F., 1995 - Trois nouveaux genres, trois nouvelles espèces de Stereomerinae de Bornéo (Coleoptera, Aphodiidae). Bulletin de la Société entomologique de France, 100: 11-20. BORDAT P., CAMBEFORT Y. & DE MIRE P. B., 1991 - Coléoptères coprophages associés au Gorille de montagne dans la chaine des Volcans Virunga (Zaïre Oriental). Bulletin de la Société entomologique de France, 96: 77-85. BORDAT P., DELLACASA G. & DELLACASA M., 2000 - Révision du genre Macroretrus Péringuey, 1908 et description d'une nouvelle espèce. Macroretroides lumareti n. g., n. sp (Coleoptera, Scarabaeoidea, Aphodiidae). Nouvelle Revue d'Entomologie (N. S.), 17 : 107-122. CAMBEFORT Y & BORDAT P., 2003 - Coléoptères Scarabaeidae s. str., Aphodiidae et Ceratocanthidae du mont Nimba et des régions limitrophes, in Lamotte M. et Roy R. (eds), Le peuplement animal du mont Nimba (Guinée, Côte d'Ivoire, Liberia). Mémoires du Muséum national d'Histoire naturelle 190 : 551-580. DELLACASA G., BORDAT P. & DELLACASA M., 2001 - A revisional essay of world genus-group taxa of Aphodiinae. Memorie della Societa entomologica italiana, 79 (2000) : 1-482. DELLACASA M., DELLACASA G. & BORDAT P., 2002 - Aphodius davidkrali, a new species from China (Coleoptera, Aphodiidae). Nouvelle Revue d'Entomologie (N. S.), 19 (1) : 7-9. DELLACASA M., DELLACASA G. & BORDAT P., 2002 - Systematic redefinition of taxa belonging to genera Ahermodontus Baguena, 1930 and Ammoecius Mulsant, 1842, with description of the new genus Vladimirellus (Coleoptera; Aphodiidae). Acta Zoologica Academiae Scientiarum Hungaricae, 48 (4) : 269-316. DELLACASA M., DELLACASA G. & BORDAT P., 2004 - Aphodius davidkrali Dellacasa et al., 2002 = Aphodius (Phaeaphodius) plutenkoi Kral, 2002, nouvelle synonymie. Nouvelle Revue d'Entomologie (N. S.), [2003] 20 (3) : 279. MORETTO Ph. & BORDAT P., 2007 - Les Scarabéides coprophages du Parc National du Niokolo-Koba au Sénégal (Coleoptera, Scarabaeoidea). Nouvelle Revue d'Entomologie (N. S.), [2006] 23 (2) : 115-145. François Génier @ : [email protected]

François Génier is an amateur entomologist. He has been studying Scarabaeinae for more than 30 years. His interest for studying insects resulted in several scientific publications on Coleoptera and especially Scarabaeinae. Several expeditions in most Countries of America, South and West Africa gave him the opportunity to acquire in-depth knowledge of modern collecting techniques. F. Génier is a member of the Coleopterist Society, The Entomological Society of America and the Association des entomologistes amateurs du Québec.

Publications (Scarabaeidae)

Génier F. 1984. Malformation de la patte droite chez un Aphodius subterraneus L. (Coleoptera: Scarabaeidae: Aphodiinae). Fabreries 10(4): 75-76. Génier F. 1996. A revision of the Neotropical genus Ontherus Erichson. Memoirs of the Entomological Society of Canada 170: 1-169. Génier F. 1998. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae), Supplement 1. The Coleopterists Bulletin 52(3): 270-274. Génier F. & H.F. Howden. 1999. Two new Central American Latreille of the mirabilis Species Group (Coleoptera: Scarabaeidae, Scarabaeinae). The Coleopterists Bulletin 53(2): 130-144. Génier F. 2000a. Dichotomius comarapensis sp. nov., une nouvelle espèce bolivienne de scarabée brachyptère (Coleoptera: Scarabaeidae, Scarabaeinae). Fabreries 25: 25-31pp. Génier F. 2000b. A new North American Ateuchus (Weber) (Coleoptera: Scarabaeidae, Scarabaeinae). The Coleopterists Bulletin 54(3): 341-346. Génier F. 2001. Note sur les espèces de Deltochilum Eschscholtz décrites en 1939 par V. Balthasar (Coleoptera: Scarabaeidae, Scarabaeinae). Fabreries 26: 1-8. Smith A.T.B. & F. Génier 2001. Revision of the Genus Holocephalus Hope (Coleoptera: Scarabaeidae, Scarabaeinae, Coprini). The Canadian Entomologist 133: 777-791. Génier F.& F. Vaz-de-Mello. 2002. A Review of Arrow's Types of Trichillum and Pedaridium (Coleoptera: Scarabaeinae, Ateuchini) with description of two new species of Pedaridium. Acta Zoologica Cracoviana 45(3):185- 196. Génier F. 2003a. New Synonymy in the Genus Paragymnopleurus Shipp (Coleoptera: Scarabaeidae, Scarabaeinae). The Coleopterists Bulletin 57(2): 122 Génier F. & B. Kohlmann. 2003. Revision of the Genera Scatimus Erichson and Scatrichus gen. nov.(Coleoptera: Scarabaeidae: Scarabaeinae). Fabreries 28(2): 57-111. Génier F. 2003b. présumés néotropicaux : provenance erronées et nouvelle synonymie (Coleoptera : Scarabaeidae, Scarabaeinae). Fabreries 28(2) : 120-121. Howden, H.F. & F. Génier. 2004. Seven new species of Onthophagus Latreille from Mexico and the United States (Coleoptera: Scarabaeidae, Scarabaeinae). Fabreries 29(1) : 53-76. Génier F. & C. A. Medina. 2005. Onthophagus mirabilis Bates, Description of the Newly Discovered Female (Coleoptera: Scarabaeidae, Scarabaeinae). The Coleopterists Bulletin 58(4): 608-612. Vaz-de-Mello, F. & F. Génier. 2005. Lectotype designations, new synonymies, and new species in the genera Trichillum Harold and Pedaridium Harold (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 1038: 41-52.

Jean-François Josso @ : [email protected]

Born in 1961, member of Entomologic Society of France, Jean-François Josso studies African Scarabaeinae for 15 years and often visits Africa (Ivory Coast, Benin, Burkina Faso, Niger, Cameroon, Kenya, Tanzanie, Ethiopia, Malawi...). He has published some papers of taxonomy with Patrick Prevost and will soon be publishing other papers with Tristao Branco. He is a member of Association Magellanes, a french entomological rewiev.

Association Magellanes : www.magellanes.com

J.F. Josso - Pendjari reserve (Benin, 2005)

Publications :

J.F. Josso & P. Prévost - 2000- Révision du genre Diastellopalpus. Collection systématique Magellanes (Vol. 3). J.F. Josso & P. Prévost - 2001- Contribution à la connaissance des Scarabaeidae africains I . Les Cahiers Magellanes Hors-série n°2. J.F. Josso & P. Prévost - 2003- Contribution à la connaissance des Scarabaeidae africains II . Etude du genre Pedaria I. Les Cahiers Magellanes Hors-série n°7. J.F. Josso & P. Prévost - 2006- Contribution à la connaissance des Scarabaeidae africains III . Etude du genre Pedaria II. Les Cahiers Magellanes Hors-série n°14. J.F. Josso & P. Prévost - 2006- Description de cinq Onthophagini nouveaux pour l'Afrique de l'Ouest (Parc du W). Les Cahiers Magellanes Hors-série n°21. J.F. Josso & P. Prévost - (in preparation)- Révision du genre Pedaria. Collection systématique Magellanes.

Patrick Prévost @ : [email protected]

Patrick Prévost is interrested in African Scarabaeinae for 20 years. He has published several papers on this fauna with J.F. Josso and will be publishing other papers with Ph. Moretto. He is a founder member of “Association Magellanes”, a french entomological rewiev.

Association Magellanes : www.magellanes.com

Frank-Thorsten Krell @ : [email protected]

Frank Krell was born in Stuttgart in 1966, began collect- ing insects in 1976, and specialized in lamellicorns in 1981. In 1992 he received his Diploma in Biology from Tübingen University for his taxonomic revision and phylogenetic analysis of the dynastine genus Temnorhynchus. He received his PhD in 1996 with a study of the functional morphology of the copulatory appa- ratus of the cockchafer, Melolontha melolontha. Beginning in 1995, he has conducted studies on the biodiversity of Lamellicornia of the Ivory Coast and the influence of human land- use on dung beetle communities, first as a postdoctoral researcher with the University of Würzburg, then as Research Entomologist with The Natural History Museum (London) where he was the Head of the Coleoptera Division from 2005 to 2007. His current research as Curator of Entomology with the Denver Museum of Nature and Science emphasizes taxonomy, faunistics, ecology, F.T. Krell paleontology, and phylogeny of scarabaeoids. He is co-editor of (Scarabnet conference - Santa Cruz 2006) Systematic Entomology and a Commissioner of the ICZN.

Some publications :

KRELL, F.-T. 1999. Southern African dung beetles attracted by diplopod defensive secretions (Coleoptera: Scarabaeidae). African Entomology 7 (2): 287-288. Pretoria. KRELL, F.-T. 1999. Dung Beetles Unharmed by Army Ants in Tropical Africa (Coleoptera: Scarabaeidae; Hymenoptera: Formicidae: Dorylinae). Coleopterists Bulletin 53 (4): 325-328. Sacramento. KRELL, F.-T. & Cambefort, Y. 2000. Onthophagus clementianus Cambefort from Guinea, a junior synonym of Onthophagus stehliki Balthasar (Coleoptera: Scarabaeidae). Bulletin de la Société Entomologique de France 105 (4): 398. Paris. KRELL, F.-T. 2000. The fossil record of Mesozoic and Tertiary Scarabaeoidea (Coleoptera: ). Invertebrate Taxonomy 14 (6): 871-905. KRELL, F.-T. 2002. Reasons why impact factors don't work for taxonomy. Its long-term relevance, few specialists and lack of core journals put it outside ISI criteria. Nature 415: 957. London. KRELL, F.-T. & Krell-Westerwalbesloh, S. 2003. First record of the genus Eusaproecius Branco from West Africa (Col., Scarabaeidae, Coprinae). Entomologist's Monthly Magazine 139: 13-14. Wallingford. KRELL, F.-T., Krell-Westerwalbesloh, S., Weiß, I., Eggleton, P. & Linsenmair, K.E. 2003. Spatial separation of Afrotropical dung beetle guilds: a trade-off between competitive superiority and energetic constraints (Coleoptera: Scarabaeidae). Ecography 26 (2): 210-222. København. KRELL, F.-T. 2003. Onthophagus mankonoensis Balthasar from Côte d'Ivoire, a junior synonym of Onthophagus moc- querysi d'Orbigny (Coleoptera: Scarabaeidae). Casopis Národního Muzea, Rada Prirodovedná 172 (1-4): 51-54. Praha. KRELL, F.-T., Korb, J. & Walter, P. 2003. The beetle fauna of hyaena latrines: Coprocenoses consisting of necrophagous beetles (Coleoptera: Trogidae, Scarabaeidae). Tropical Zoology 16 (2): 145-152. Firenze. KRELL, F.-T. 2004. Parataxonomy vs. taxonomy in biodiversity studies - pitfalls and applicability of 'morphospecies' sorting. Biodiversity and Conservation 13 (4): 795-812. Dordrecht. Krell-Westerwalbesloh, S., KRELL, F.-T. & Linsenmair, K.E. 2004. Diel separation of Afrotropical dung beetle guilds - avoiding competition and neglecting resources. Journal of Natural History 38 (17): 2225-2249. Abingdon. Pfrommer, A. & KRELL, F.-T. 2004. Who Steals the Eggs? Coprophanaeus telamon (Erichson, 1847) Buries Decomposing Eggs in Western Amazonian Rain Forest (Coleoptera: Scarabaeidae). Coleopterists Bulletin 58 (1): 21-27. Sacramento. Schmitt, T., KRELL, F.-T. & Linsenmair, K.E. 2004. Quinone mixture as attractant for necrophagous dung beetles spe- cialized on dead millipedes. Journal of Chemical Ecology 30 (4): 731-740. Dordrecht. KRELL, F.-T., Mahiva, V.S., Kouakou, C., N'goran, P., Krell-Westerwalbesloh, S., Newman, D.H., Weiß, I. & Doumbia, M. 2005. Human influence on the dung fauna in Afrotropical grasslands (Insecta: Coleoptera). Pp. 133-139 in: Huber, B.A., Sinclair, B.J. & Lampe, K.-H. (eds): African Biodiversity: Molecules, Organisms, Ecosystems. Proceedings of the 5th International Symposium on Tropical Biology, Museum Koenig, Bonn. Springer, New York. Pdf on request.

Olivier Montreuil @ : [email protected]

Olivier Montreuil is a professional entomologist in charge of the management of the Scarabaeoidea collec- tion of the Muséum national d'Histoire naturelle, Paris. He has published, alone or in collaboration with other special- ists, several papers on the Melolohonthidae, Cetoniidae and Scarabaeidae of the palearctic region and Madagascar. He has done many collecting trips since 2001, mostly in Asia ((Lebanon, Iran, Nepal, Laos) but also in Madagascar. He is member of the Société entomologique de France since 1997.

O. Montreuil (Madagascar)

Some publications :

Boucher S., Montreuil O. & Le Restif A., 2003. Découverte de la femelle et nouvelles localisations de Osmoderma sikhotense Boucher, 2002 (Coleoptera : Cetoniidae) . Bulletin de la Société entomologique de France, 108(4) : 401-403. Montreuil O., 2004. Contribution à l'étude des Canthonini malgaches, 3e note : description de deux Apotolamprus Olsoufieff et mises au point taxonomiques et nomenclaturales (Coleoptera : Scarabaeidae). Revue française d'Entomologie (N.S.), 26(2) : 67-72. Montreuil O., Ghaioule D., Maatouf N., Rochat D. & Lumaret J.-P., 2004. Premières données biologiques sur Sphodroxia maroccana Ley (Coleoptera : Melolonthidae) et description de la femelle de ce nouveau ravageur forestier. Bulletin de la Société entomologique de France, 109(3) : 317-321. Montreuil O., 2005. Nouveaux Helictopleurus d'Orbigny, 1915, de Madagascar et révision du 'groupe semivirens' sensu Lebis, 1960 (Coleoptera : Scarabaeidae : Oniticellini). Zoosystema, 27(1) : 123-135. Montreuil O., 2005. Contribution à l'étude des Canthonini malgaches, 4e note : deux nouveaux Apotolamprus Olsoufieff, 1947 (Coleoptera : Scarabaeidae). Revue française d'Entomologie (N.S.), 27(1) : 1-4. Montreuil O., 2005. Contribution à l'étude du genre Haplidia Hope : Description de deux nouvelles espèces du Liban (Coleoptera, Melolonthinae). Bulletin de la Société entomologique de France, 110(2) : 129-131. Keith D. & Montreuil O., 2005. Révision des genres Otoclinius Brenske, 1896, Falsotoclinius n. gen. et Tanyproctoides Petrovitz, 1971 (Coleoptera : Melolonthidae : Pachydeminae). Annales de la Société entomologique de France, 40(3-4) [2004] : 213-225. Montreuil O., 2005. Contribution à l'étude du genre Helictopleurus d'Orbigny, 1915 (Coleoptera, Scarabaeidae). Bulletin de la Société entomologique de France, 110(4/5) : 373-376. Montreuil O., 2005. Contribution à l'étude des Canthonini malgaches, 5e note : description de nouveaux Aleiantus Olsoufieff, 1947 (Coleoptera : Scarabaeidae). Revue française d'Entomologie (N.S.), 27(4) : 153-160. Frolov A. & Montreuil O., 2006. Description of the male of the rare species Pseudorphnus hiboni Paulian, 1959, with notes on the genus Pseudorphnus Benderitter, 1913, endemic to Madagascar (Coleoptera, Scarabaeidae, Orphninae). Zootaxa, 1154: 27-33. Montreuil O., 2006. Contribution à l'étude des Canthonini malgaches, 6e note : description de nouvelles espèces du genre Arachnodes Westwood, 1847, et mises au point taxonomiques et nomenclaturales (Coleoptera : Scarabaeidae). Revue française d'Entomologie (N.S.), 28(3) : 97-110. Montreuil O., 2006. Nouvelles Eupotosia et Cetonischema d'Iran (Coleoptera : Cetoniidae). Coléoptères, 12(32): 427- 434. Montreuil O., 2006. Un nouveau Scarabaeus d'Iran. Bulletin de la Société entomologique de France, 111(4) : 445-447. Montreuil O., 2007. Nouveaux Helictopleurus d'Orbigny, 1915 (Coleoptera, Scarabaeidae). Bulletin de la Société ento- mologique de France, 112(1) : 79-84. Montreuil O. & Viljanen H., 2007. Contribution à l'étude des Canthonini de Madagascar, 7e note : mises au point tax- onomiques et nomenclaturales dans le genre Nanos Westwood, 1847, (Coleoptera : Scarabaeidae). Revue française d'Entomologie (N.S.), 29(1) : 1-10. Philippe Moretto @ : [email protected] Catharsius group coordinator

Philippe Moretto is a free lance entomologist studying African Scarabaeinae for the past 20 years, with special focus on Onthophagini. Philippe has acquired a good experience in the field through 15 trips in Africa, mostly in West Africa and constituted a large collection with more than 1500 species of african Scarabaeinae. He is member of the « Société entomologique de France » since 1975 and of the « Société d'Entomologie Africaine » since 1997. He has published, alone or in cooperation with other specialists, several papers on african Scarabaeinae.

Publications :

MORETTO P. (1998). “Kheper festivus (Harold) : un complexe d'espèces.” Entomologia Africana 3(2): 27-38. MORETTO P. (1999). “Diastellopalpus Lansberge nouveaux du bassin du Congo et de l'Angola.” Entomologia Africana 4(2): 33-40. MORETTO P. et NICOLAS J.L. (2001). « Description de 5 nouveaux Proagoderus Lansberge d'Afrique ». Entomologia Africana. VII: 28-37. NICOLAS J.L. et MORETTO P. (2002). « Description de deux nouveaux Scarabaeidae africains appartenant aux genres Allogymnopleurus Janssens et Scarabaeus Linné. » Entomologia Africana 7(2): 3-10. MORETTO P. (2004) “Descriptions de nouveaux Proagoderus Lansberge 1883 d'Afrique et remarques sur quelques espèces peu ou mal connues.” Entomologia Africana 9(1): 20-48. MORETTO P. & PREVOST P. Les « Onthophagus, Latreille, 1802 africains des 5ème et 6ème groupes de d'Orbigny. Essai de classification. Descriptions d'un genre nouveau et d'espèces nouvelles. » (sous presse) MORETTO P. & BORDAT P. « Les Scarabéides coprophages du Parc National du Niokolo Koba au Sénégal.» (sous presse) MORETTO P. « Les Onitis Fabricius, 1798 du XIIIème groupe. Espèces nouvelles et espèces affines » (sous presse)

Philippe Moretto is the Catharsius group coordinator

Philippe MORETTO 5, rue Gimelli F.83000 Toulon FRANCE @ : [email protected]