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Prebiological Evolution and the Metabolic Origins of Life
Prebiological Evolution and the Andrew J. Pratt* Metabolic Origins of Life University of Canterbury Keywords Abiogenesis, origin of life, metabolism, hydrothermal, iron Abstract The chemoton model of cells posits three subsystems: metabolism, compartmentalization, and information. A specific model for the prebiological evolution of a reproducing system with rudimentary versions of these three interdependent subsystems is presented. This is based on the initial emergence and reproduction of autocatalytic networks in hydrothermal microcompartments containing iron sulfide. The driving force for life was catalysis of the dissipation of the intrinsic redox gradient of the planet. The codependence of life on iron and phosphate provides chemical constraints on the ordering of prebiological evolution. The initial protometabolism was based on positive feedback loops associated with in situ carbon fixation in which the initial protometabolites modified the catalytic capacity and mobility of metal-based catalysts, especially iron-sulfur centers. A number of selection mechanisms, including catalytic efficiency and specificity, hydrolytic stability, and selective solubilization, are proposed as key determinants for autocatalytic reproduction exploited in protometabolic evolution. This evolutionary process led from autocatalytic networks within preexisting compartments to discrete, reproducing, mobile vesicular protocells with the capacity to use soluble sugar phosphates and hence the opportunity to develop nucleic acids. Fidelity of information transfer in the reproduction of these increasingly complex autocatalytic networks is a key selection pressure in prebiological evolution that eventually leads to the selection of nucleic acids as a digital information subsystem and hence the emergence of fully functional chemotons capable of Darwinian evolution. 1 Introduction: Chemoton Subsystems and Evolutionary Pathways Living cells are autocatalytic entities that harness redox energy via the selective catalysis of biochemical transformations. -
Geological Timeline
Geological Timeline In this pack you will find information and activities to help your class grasp the concept of geological time, just how old our planet is, and just how young we, as a species, are. Planet Earth is 4,600 million years old. We all know this is very old indeed, but big numbers like this are always difficult to get your head around. The activities in this pack will help your class to make visual representations of the age of the Earth to help them get to grips with the timescales involved. Important EvEnts In thE Earth’s hIstory 4600 mya (million years ago) – Planet Earth formed. Dust left over from the birth of the sun clumped together to form planet Earth. The other planets in our solar system were also formed in this way at about the same time. 4500 mya – Earth’s core and crust formed. Dense metals sank to the centre of the Earth and formed the core, while the outside layer cooled and solidified to form the Earth’s crust. 4400 mya – The Earth’s first oceans formed. Water vapour was released into the Earth’s atmosphere by volcanism. It then cooled, fell back down as rain, and formed the Earth’s first oceans. Some water may also have been brought to Earth by comets and asteroids. 3850 mya – The first life appeared on Earth. It was very simple single-celled organisms. Exactly how life first arose is a mystery. 1500 mya – Oxygen began to accumulate in the Earth’s atmosphere. Oxygen is made by cyanobacteria (blue-green algae) as a product of photosynthesis. -
And Abiogenesis
Historical Development of the Distinction between Bio- and Abiogenesis. Robert B. Sheldon NASA/MSFC/NSSTC, 320 Sparkman Dr, Huntsville, AL, USA ABSTRACT Early greek philosophers laid the philosophical foundations of the distinction between bio and abiogenesis, when they debated organic and non-organic explanations for natural phenomena. Plato and Aristotle gave organic, or purpose-driven explanations for physical phenomena, whereas the materialist school of Democritus and Epicurus gave non-organic, or materialist explanations. These competing schools have alternated in popularity through history, with the present era dominated by epicurean schools of thought. Present controversies concerning evidence for exobiology and biogenesis have many aspects which reflect this millennial debate. Therefore this paper traces a selected history of this debate with some modern, 20th century developments due to quantum mechanics. It ¯nishes with an application of quantum information theory to several exobiology debates. Keywords: Biogenesis, Abiogenesis, Aristotle, Epicurus, Materialism, Information Theory 1. INTRODUCTION & ANCIENT HISTORY 1.1. Plato and Aristotle Both Plato and Aristotle believed that purpose was an essential ingredient in any scienti¯c explanation, or teleology in philosophical nomenclature. Therefore all explanations, said Aristotle, answer four basic questions: what is it made of, what does it represent, who made it, and why was it made, which have the nomenclature material, formal, e±cient and ¯nal causes.1 This aristotelean framework shaped the terms of the scienti¯c enquiry, invisibly directing greek science for over 500 years. For example, \organic" or \¯nal" causes were often deemed su±cient to explain natural phenomena, so that a rock fell when released from rest because it \desired" its own kind, the earth, over unlike elements such as air, water or ¯re. -
Calculating the Potato Radius of Asteroids Using the Height of Mt. Everest
Calculating the Potato Radius of Asteroids using the Height of Mt. Everest M. E. Caplan∗ Center for the Exploration of Energy and Matter, Indiana University, Bloomington, IN 47408 (Dated: November 16, 2015) Abstract At approximate radii of 200-300 km, asteroids transition from oblong `potato' shapes to spheres. This limit is known as the Potato Radius, and has been proposed as a classification for separating asteroids from dwarf planets. The Potato Radius can be calculated from first principles based on the elastic properties and gravity of the asteroid. Similarly, the tallest mountain that a planet can support is also known to be based on the elastic properties and gravity. In this work, a simple novel method of calculating the Potato Radius is presented using what is known about the maximum height of mountains and Newtonian gravity for a spherical body. This method does not assume any knowledge beyond high school level mechanics, and may be appropriate for students interested in applications of physics to astronomy. arXiv:1511.04297v1 [physics.ed-ph] 7 Nov 2015 1 I. INTRODUCTION Spacecraft are currently exploring asteroids and dwarf planets, such as the Near Earth Asteroid Rendezvous mission (NEAR) landing on Eros,1 the Dawn mission orbiting Ceres and Vesta,2 and the New Horizons flyby of Pluto and Charon.3 Additionally, the Mars Re- connaissance Orbiter (MRO) has observed the Martian moons Phobos and Deimos.4 These missions observe a remarkable variety of shapes for these bodies, shown in Fig. 1. Smaller asteroids have irregular shapes while dwarf planets (large asteroids) are nearly spherical. -
Ices on Mercury: Chemistry of Volatiles in Permanently Cold Areas of Mercury’S North Polar Region
Icarus 281 (2017) 19–31 Contents lists available at ScienceDirect Icarus journal homepage: www.elsevier.com/locate/icarus Ices on Mercury: Chemistry of volatiles in permanently cold areas of Mercury’s north polar region ∗ M.L. Delitsky a, , D.A. Paige b, M.A. Siegler c, E.R. Harju b,f, D. Schriver b, R.E. Johnson d, P. Travnicek e a California Specialty Engineering, Pasadena, CA b Dept of Earth, Planetary and Space Sciences, University of California, Los Angeles, CA c Planetary Science Institute, Tucson, AZ d Dept of Engineering Physics, University of Virginia, Charlottesville, VA e Space Sciences Laboratory, University of California, Berkeley, CA f Pasadena City College, Pasadena, CA a r t i c l e i n f o a b s t r a c t Article history: Observations by the MESSENGER spacecraft during its flyby and orbital observations of Mercury in 2008– Received 3 January 2016 2015 indicated the presence of cold icy materials hiding in permanently-shadowed craters in Mercury’s Revised 29 July 2016 north polar region. These icy condensed volatiles are thought to be composed of water ice and frozen Accepted 2 August 2016 organics that can persist over long geologic timescales and evolve under the influence of the Mercury Available online 4 August 2016 space environment. Polar ices never see solar photons because at such high latitudes, sunlight cannot Keywords: reach over the crater rims. The craters maintain a permanently cold environment for the ices to persist. Mercury surface ices magnetospheres However, the magnetosphere will supply a beam of ions and electrons that can reach the frozen volatiles radiolysis and induce ice chemistry. -
Modern Astronomical Optics - Observing Exoplanets 2
Modern Astronomical Optics - Observing Exoplanets 2. Brief Introduction to Exoplanets Olivier Guyon - [email protected] – Jim Burge, Phil Hinz WEBSITE: www.naoj.org/staff/guyon → Astronomical Optics Course » → Observing Exoplanets (2012) Definitions – types of exoplanets Planet (& exoplanet) definitions are recent, as, prior to discoveries of exoplanets around other stars and dwarf planets in our solar system, there was no need to discuss lower and upper limits of planet masses. Asteroid < dwarf planet < planet < brown dwarf < star Upper limit defined by its mass: < 13 Jupiter mass 1 Jupiter mass = 317 Earth mass = 1/1000 Sun mass Mass limit corresponds to deuterium limit: a planet is not sufficiently massive to start nuclear fusion reactions, of which deuterium burning is the easiest (lowest temperature) Lower limit recently defined (now excludes Pluto) for our solar system: has cleared the neighbourhood around its orbit Distinction between giant planets (massive, large, mostly gas) and rocky planets also applies to exoplanets Habitable planet: planet on which life as we know it (bacteria, planets or animals) could be sustained = rocky + surface temperature suitable for liquid water Formation Planet and stars form (nearly) together, within first few x10 Myr of system formation Gravitational collapse of gas + dust cloud Star is formed at center of disk Planets form in the protoplanetary disk Planet embrios form first Adaptive Optics image of Beta Pic Large embrios (> few Earth mass) can Shows planet + debris disk accrete large quantity of -
Viscosity of Gases References
VISCOSITY OF GASES Marcia L. Huber and Allan H. Harvey The following table gives the viscosity of some common gases generally less than 2% . Uncertainties for the viscosities of gases in as a function of temperature . Unless otherwise noted, the viscosity this table are generally less than 3%; uncertainty information on values refer to a pressure of 100 kPa (1 bar) . The notation P = 0 specific fluids can be found in the references . Viscosity is given in indicates that the low-pressure limiting value is given . The dif- units of μPa s; note that 1 μPa s = 10–5 poise . Substances are listed ference between the viscosity at 100 kPa and the limiting value is in the modified Hill order (see Introduction) . Viscosity in μPa s 100 K 200 K 300 K 400 K 500 K 600 K Ref. Air 7 .1 13 .3 18 .5 23 .1 27 .1 30 .8 1 Ar Argon (P = 0) 8 .1 15 .9 22 .7 28 .6 33 .9 38 .8 2, 3*, 4* BF3 Boron trifluoride 12 .3 17 .1 21 .7 26 .1 30 .2 5 ClH Hydrogen chloride 14 .6 19 .7 24 .3 5 F6S Sulfur hexafluoride (P = 0) 15 .3 19 .7 23 .8 27 .6 6 H2 Normal hydrogen (P = 0) 4 .1 6 .8 8 .9 10 .9 12 .8 14 .5 3*, 7 D2 Deuterium (P = 0) 5 .9 9 .6 12 .6 15 .4 17 .9 20 .3 8 H2O Water (P = 0) 9 .8 13 .4 17 .3 21 .4 9 D2O Deuterium oxide (P = 0) 10 .2 13 .7 17 .8 22 .0 10 H2S Hydrogen sulfide 12 .5 16 .9 21 .2 25 .4 11 H3N Ammonia 10 .2 14 .0 17 .9 21 .7 12 He Helium (P = 0) 9 .6 15 .1 19 .9 24 .3 28 .3 32 .2 13 Kr Krypton (P = 0) 17 .4 25 .5 32 .9 39 .6 45 .8 14 NO Nitric oxide 13 .8 19 .2 23 .8 28 .0 31 .9 5 N2 Nitrogen 7 .0 12 .9 17 .9 22 .2 26 .1 29 .6 1, 15* N2O Nitrous -
Measurement of Cp/Cv for Argon, Nitrogen, Carbon Dioxide and an Argon + Nitrogen Mixture
Measurement of Cp/Cv for Argon, Nitrogen, Carbon Dioxide and an Argon + Nitrogen Mixture Stephen Lucas 05/11/10 Measurement of Cp/Cv for Argon, Nitrogen, Carbon Dioxide and an Argon + Nitrogen Mixture Stephen Lucas With laboratory partner: Christopher Richards University College London 5th November 2010 Abstract: The ratio of specific heats, γ, at constant pressure, Cp and constant volume, Cv, have been determined by measuring the oscillation frequency when a ball bearing undergoes simple harmonic motion due to the gravitational and pressure forces acting upon it. The γ value is an important gas property as it relates the microscopic properties of the molecules on a macroscopic scale. In this experiment values of γ were determined for input gases: CO2, Ar, N2, and an Ar + N2 mixture in the ratio 0.51:0.49. These were found to be: 1.1652 ± 0.0003, 1.4353 ± 0.0003, 1.2377 ± 0.0001and 1.3587 ± 0.0002 respectively. The small uncertainties in γ suggest a precise procedure while the discrepancy between experimental and accepted values indicates inaccuracy. Systematic errors are suggested; however it was noted that an average discrepancy of 0.18 between accepted and experimental values occurred. If this difference is accounted for, it can be seen that we measure lower vibrational contributions to γ at room temperature than those predicted by the equipartition principle. It can be therefore deduced that the classical idea of all modes contributing to γ is incorrect and there is actually a „freezing out‟ of vibrational modes at lower temperatures. I. Introduction II. Method The primary objective of this experiment was to determine the ratio of specific heats, γ, for gaseous Ar, N2, CO2 and an Ar + N2 mixture. -
Bepicolombo - a Mission to Mercury
BEPICOLOMBO - A MISSION TO MERCURY ∗ R. Jehn , J. Schoenmaekers, D. Garc´ıa and P. Ferri European Space Operations Centre, ESA/ESOC, Darmstadt, Germany ABSTRACT BepiColombo is a cornerstone mission of the ESA Science Programme, to be launched towards Mercury in July 2014. After a journey of nearly 6 years two probes, the Magneto- spheric Orbiter (JAXA) and the Planetary Orbiter (ESA) will be separated and injected into their target orbits. The interplanetary trajectory includes flybys at the Earth, Venus (twice) and Mercury (four times), as well as several thrust arcs provided by the solar electric propulsion module. At the end of the transfer a gravitational capture at the weak stability boundary is performed exploiting the Sun gravity. In case of a failure of the orbit insertion burn, the spacecraft will stay for a few revolutions in the weakly captured orbit. The arrival conditions are chosen such that backup orbit insertion manoeuvres can be performed one, four or five orbits later with trajectory correction manoeuvres of less than 15 m/s to compensate the Sun perturbations. Only in case that no manoeuvre can be performed within 64 days (5 orbits) after the nominal orbit insertion the spacecraft will leave Mercury and the mission will be lost. The baseline trajectory has been designed taking into account all operational constraints: 90-day commissioning phase without any thrust; 30-day coast arcs before each flyby (to allow for precise navigation); 7-day coast arcs after each flyby; 60-day coast arc before orbit insertion; Solar aspect angle constraints and minimum flyby altitudes (300 km at Earth and Venus, 200 km at Mercury). -
The Search for Another Earth – Part II
GENERAL ARTICLE The Search for Another Earth – Part II Sujan Sengupta In the first part, we discussed the various methods for the detection of planets outside the solar system known as the exoplanets. In this part, we will describe various kinds of exoplanets. The habitable planets discovered so far and the present status of our search for a habitable planet similar to the Earth will also be discussed. Sujan Sengupta is an 1. Introduction astrophysicist at Indian Institute of Astrophysics, Bengaluru. He works on the The first confirmed exoplanet around a solar type of star, 51 Pe- detection, characterisation 1 gasi b was discovered in 1995 using the radial velocity method. and habitability of extra-solar Subsequently, a large number of exoplanets were discovered by planets and extra-solar this method, and a few were discovered using transit and gravi- moons. tational lensing methods. Ground-based telescopes were used for these discoveries and the search region was confined to about 300 light-years from the Earth. On December 27, 2006, the European Space Agency launched 1The movement of the star a space telescope called CoRoT (Convection, Rotation and plan- towards the observer due to etary Transits) and on March 6, 2009, NASA launched another the gravitational effect of the space telescope called Kepler2 to hunt for exoplanets. Conse- planet. See Sujan Sengupta, The Search for Another Earth, quently, the search extended to about 3000 light-years. Both Resonance, Vol.21, No.7, these telescopes used the transit method in order to detect exo- pp.641–652, 2016. planets. Although Kepler’s field of view was only 105 square de- grees along the Cygnus arm of the Milky Way Galaxy, it detected a whooping 2326 exoplanets out of a total 3493 discovered till 2Kepler Telescope has a pri- date. -
Saturn — from the Outside In
Saturn — From the Outside In Saturn — From the Outside In Questions, Answers, and Cool Things to Think About Discovering Saturn:The Real Lord of the Rings Saturn — From the Outside In Although no one has ever traveled ing from Saturn’s interior. As gases in from Saturn’s atmosphere to its core, Saturn’s interior warm up, they rise scientists do have an understanding until they reach a level where the tem- of what’s there, based on their knowl- perature is cold enough to freeze them edge of natural forces, chemistry, and into particles of solid ice. Icy ammonia mathematical models. If you were able forms the outermost layer of clouds, to go deep into Saturn, here’s what you which look yellow because ammonia re- trapped in the ammonia ice particles, First, you would enter Saturn’s up- add shades of brown and other col- per atmosphere, which has super-fast ors to the clouds. Methane and water winds. In fact, winds near Saturn’s freeze at higher temperatures, so they equator (the fat middle) can reach turn to ice farther down, below the am- speeds of 1,100 miles per hour. That is monia clouds. Hydrogen and helium rise almost four times as fast as the fast- even higher than the ammonia without est hurricane winds on Earth! These freezing at all. They remain gases above winds get their energy from heat ris- the cloud tops. Saturn — From the Outside In Warm gases are continually rising in Earth’s Layers Saturn’s atmosphere, while icy particles are continually falling back down to the lower depths, where they warm up, turn to gas and rise again. -
Inhibition of Premixed Carbon Monoxide-Hydrogen-Oxygen-Nitrogen Flames by Iron Pentacarbonylt
Inhibition of Premixed Carbon Monoxide-Hydrogen-Oxygen-Nitrogen Flames by Iron Pentacarbonylt MARC D. RUMMINGER+ AND GREGORY T. LINTERIS* Building and Fire Research Laboratory, National Institute of Standards and Technology, Gaithersburg, MD 20899, USA This paper presents measurements of the burning velocity of premixed CO-HrOrNz flames with and without the inhibitor Fe(CO)s over a range of initial Hz and Oz mole fractions. A numerical model is used to simulate the flame inhibition using a gas-phase chemical mechanism. For the uninhibited flames, predictions of burning velocity are excellent and for the inhibited flames, the qualitative agreement is good. The agreement depends strongly on the rate of the CO + OH <-> COz + H reaction and the rates of several key iron reactions in catalytic H- and O-atom scavenging cycles. Most of the chemical inhibition occurs through a catalytic cycle that converts o atoms into Oz molecules. This O-atom cycle is not important in methane flames. The H-atom cycle that causes most of the radical scavenging in the methane flames is also active in CO-Hz flames, but is of secondary importance. To vary the role of the H- and O-atom radical pools, the experiments and calculations are performed over a range of oxygen and hydrogen mole fraction. The degree of inhibition is shown to be related to the fraction of the net H- and O-atom destruction through the iron species catalytic cycles. The O-atom cycle saturates at a relatively low inhibitor mole fraction (-100 ppm), whereas the H-atom cycle saturates at a much higher inhibitor mole fraction (-400 ppm).