Southern Southwestern Madagascan Vegetation
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Plant Formations in the Southern and Southwestern Madagascan BioProvince Peter Martin Rhind Southern and Southwestern Madagascan Spiny Forest These forests are probably more accurately described as thorn-scrub since they normally grows to no more than about 6 m tall, but often include a scattering of emergent trees growing to about 10 m high. Otherwise there is no stratification and many stands are virtually impenetrable. The most distinctive elements are members of the near endemic family Didiereaceae and arborescent species of Euphorbia. The family includes four genera and twelve species - Didierea (D. madagascariensis and D. trollii), Alluaudia (six species), Alluaudiopsis (two species) and the montypic genus Decaryia (D. madagascariensis). Alluaudia ascendens and A. procera reach heights of 8m but most others are smaller than this. Of the Euphorbia species many are endemic and include several that have caducous leaves and green fleshy stems. One of the most abundant of these is the spiny endemic Euphorbia stenoclada (Euphorbiaceae), which can grow to a height of 10 m or more. The emergent trees include a few island endemics such as Adansonia fony, A. za (Malvaceae), Dicoma incana, D. carbonaria (Asteraceae) and Tetrapterocarpon geayi (Fabaceae). These forests are very rich in species but species composition varies greatly from place to place. Important woody plants often include the endemic or near endemics genera Megistostegium (Malvaceae) and Uncarina (Rubiaceae), the endemic species Rhigozum madagascariensis (Bignoniaceae), together with many endemic species of Acacia, Commiphora, Dichrostachys, Iphione, Cadaba, Sclerocarya, and the endemic palm Dypsis decaryi (Arecaceae). As expected, many of the taxa, such as Allaudia, Didierea, Mimosa and Pachypodium, have abundant spines. Lianas are numerous but rather small and include several endemic members of the Asclepiadaceae. Examples include species of Adenia, leafless species of Cissus, and species of the endemic genus Xerosicyos (Cucurbitaceae) with their thick fleshy leaves. The ground flora is usually sparse but may include tufts of the endemic grass Humbertochloa bambusiuscula (Poaceae) together with endemic succulent species of Aloe (such as the rare Aloe suzannae), Euphorbia, Senecio and Notonia. Further information required. References Chapman, V. J. 1977. Africa B - the remainder of Africa. In: Ecosystems of the World 1 - Wet Coastal Ecosystems. Ed. V. J. Chapman. Elsevier Scientific Publishing Company. Buam, D. A. 1996. The ecology and conservation of the baobabs of Madagascar. In: Ecology of tropical dry forests in Madagascar. Eds. J. U. Ganzhorn and P. J. Sorg. Primate Ecology 46: 311-327. Dorr, L. J. & Oliver, E. G. H. 1999. New taxa, names, and combinations in Erica (Ericaceae-Ericoideae) from Madagascar and the Comora Islands. Adansonia, ser. 3. 21(1): 75-91. Good, R. 1950. Madagascar and New Caledonia - a problem in plant geography. Blumea, 6: 470-490. Copyright © 2010 Peter Martin Rhind Goodman, S. M. 1996. A floral and faunal inventory of the eastern slopes of the Reserve Naturale Integrale d' Andringitra, Madagascar. Fieldana: Zoology, N. S. No. 85: 1-319. Goodman, S. M. & Benstead, J. P. (eds). 2003. The Natural History of Madagascar. The University of Chicago Press. Humbert, H. 1962. Flore de Madagascar et des Comores - Composees. Tomes 1-3. Typographie Firmin-Didot. Jolly, A., Oberle, P. & Albignac, R. 1984. Key Environments Madagascar. Pergamon Press. Menaut, J. C. 1983. The Vegetation of the African Savannas. In: Ecosystems of the World 13 - Tropical Savannas. Ed. F. Bourliere. Elsevier Scientific Publishing Company. Rakotonirina 1996. Composition and structure of the dry forests on sandy soils near Morondava. In: Ecology of tropical dry forests in Madagascar. Eds. J. U. Ganzhorn and P. J. Sorg. Primate Ecology 46: 311-327. Rauh, W. 1979. Problems of biological conservation in Madagascar. In: Plants and Islands. Ed. D. Bramwell. Academic Press. Rauh, W. 1986. Arid region of Madagascar. In: Ecosystems of the World 12B - Hot Deserts and Arid Shrubland, B. Eds. M. Evenari, I. Noy-Meir and D. W. Goodall. Elsevier. Renvoize, S. A. 1979. The origins of Indian Ocean Island floras. In: Plants and Islands. Ed. D. Bramwell. Academic Press. Schatz, G. E. 2001. Generic Tree Flora of Madagascar. Royal Botanic Gardens Kew and Missouri Botanical Gardens. White, F. 1983. The Vegetation of Africa. UNESCO. Wild, H. 1975. Phytogeography and Gondwanaland position of Madagascar. Boissiera, 24: 107-117. Copyright © 2010 Peter Martin Rhind .