The Sciarid Fauna of the British Isles (Diptera: Sciaridae), Including Descriptions of Six New Species

Home , Ailladie, Aversley Wood, Beckenham Place Park, Begwary Brook, Bix Bottom, Boughton Fen

Blackwell Science, LtdOxford, UKZOJZoological Journal of the Linnean Society0024-4082The Lin- nean Society of London, 2006? 2006 146? 1147 Original Article

BRITISH SCIARID FAUNAF. MENZEL ET AL.

Zoological Journal of the Linnean Society, 2006, 146, 1–147. With 20 figures

The sciarid fauna of the British Isles (Diptera: Sciaridae), including descriptions of six new species

FRANK MENZEL1, JANE E. SMITH2* and PETER J. CHANDLER3

1Deutsches Entomologisches Institut, Leibniz-Zentrum für Agrarlandschafts- und Landnutzungsforschung (ZALF) e.V., Eberswalder Straße 84, 15374 Müncheberg, Germany 2Warwick HRI, Wellesbourne, Warwick CV35 9EF, UK 3606B Berryﬁeld Lane, Melksham, Wilts. SN12 6EL, UK

Received November 2004; accepted for publication March 2005

The results of a revision of the Sciaridae (Diptera: Nematocera) from the British Isles are presented, carried out as a preliminary to the preparation of a new Handbook for the identification of the British and Irish fauna of this family. A total fauna of 263 species is confirmed, including many species new to the British Isles: 111 new to Great Britain and 32 new to Ireland. Epidapus (Pseudoaptanogyna) echinatum Mohrig & Kozánek, 1992, hitherto known only from North Korea, is newly recorded from Europe. Six species are described as new to science: Bradysia austera Menzel & Heller sp. nov., Bradysia ismayi Menzel sp. nov., Bradysia nigrispina Menzel sp. nov., Corynoptera flavosignata Menzel & Heller sp. nov., Corynoptera uncata Menzel & Smith sp. nov. and Epidapus subgracilis Menzel & Mohrig sp. nov. The following new synonymies are proposed: Leptosciarella nigrosetosa (Freeman, 1990) = Leptosciarella truncatula Mohrig & Menzel, 1997; Sciara nursei Freeman, 1983 = Sciara ulrichi Menzel & Mohrig, 1998. Many misidentifications in the previous literature are corrected. Details of the collection data and location of specimens examined are provided under each species. The localities from which Sciaridae were collected in the British Isles are documented by modern county and grid references and the habitat indicated where known, to assist in assessing the ecological requirements of each species. © 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147.

ADDITIONAL KEYWORDS: Europe – faunistics – Nematocera – Palaearctic – systematics – taxonomy.

INTRODUCTION (formerly known as Mycetophiloidea) and were treated by many earlier authors as a subfamily of the The Sciaridae are mostly small, dark-coloured gnats Mycetophilidae. For this reason the English name fun- with a slender body, long legs and unmarked wings gus gnats, which was formerly applied to the whole with a characteristic venation. They have until group, has also been associated with them and has led recently remained one of the least well-known families to some confusion in the application of this name in of Diptera in the British fauna. This is perhaps sur- the literature. More recently, they have been termed prising given that the family includes a number of black fungus gnats or dark winged fungus gnats. The agricultural and horticultural pests. A firm taxonomic first of these terms is perhaps more appropriate as base is essential for the study of these pests. But their most species are predominantly black in colour, while study has not been popular among British dipterists, only some of the larger and more obvious species are both because of the perceived difficulty of identifica- particularly dark winged. The German name Trauer- tion and because of the rather uniform appearance of mücken (= mourning midges) also refers to their most members of the family. dark coloration and perhaps contrasts better with The Sciaridae are related to the Mycetophilidae and Pilzmücken (= fungus midges), the term used for other other families included in the superfamily Sciaroidea families of Sciaroidea. In addition, although some Sciaridae do develop in fungus fruiting bodies, this is by no means as frequent a life history as in the Myc- *Corresponding author. E-mail: [email protected] etophilidae, in which it is the dominant life history.

2 F. MENZEL ET AL.

Some Mycetophilidae are also mainly black but other regarded as a subfamily of Mycetophilidae and had types of coloration are more common in the family. this status in much of the earlier literature. The first Before the present study began, there was little idea systematic treatment was by Walker (1856), who of the size of the British fauna of Sciaridae owing to keyed 26 species placed in three genera. This was the relative neglect of the family by British dipterists. mainly followed in the first checklist of British Diptera Any estimate of the size of a fauna always has a direct (Verrall, 1888), which listed 27 species of Sciarinae in relationship with the number of specimens available four genera with few differences. A further early sys- for study by successive workers, which is itself a con- tematic account was by Theobald (1892), who included sequence of the number of active collectors and the only 14 species in four genera, but remarked that geographical coverage they achieved. K. G. V. Smith there were a large number of species of the genus Sci- (1974), describing the ways in which changes in ara found in England. His remark that Winnertz knowledge of the British Diptera fauna have taken (1867) had described 175 species in his monograph of place, commented that when a neglected family is the European species showed that he understood the studied a great change in the British list is to be scale involved. In his revised checklist Verrall (1901) expected. He particularly applied this to the leaf-min- listed 33 species in five genera, again under Sciarinae, ing Agromyzidae, of which knowledge had exploded in commenting on the Mycetophilidae as a whole that the 1950s and 1960s. ‘this family, though much improved, is still in a most For a little over 20 years there has been a Royal unfinished condition’ (p. 5). Verrall (1912) reinforced Entomological Society Handbook to the British species this when adding Trichosia morio (as Sciara longiven- of Sciaridae (Freeman, 1983b). The publication of such tris) to the British list, when he wrote ‘it is rather a a handbook can give the impression that a group is farce to add a single species of Sciara to our lists, when well known and this is reinforced when non-specialists at least 150 more species of this genus should occur in tend to conclude that their specimens belong to a spe- England’ (p. 20). cies included in the key. This assumption is, of course, There were no significant advances in knowledge often correct because the species most likely to be col- into the early part of the 20th century. Edwards lected are the common species that are well character- (1913b), in his earlier work on Mycetophilidae, did not ized and have long been known to be British. But it cover Sciaridae and possibly at that time preferred to has clearly led to many misidentifications in subse- regard them as a separate family as indicated in quent literature, as has been shown by the examina- Edwards (1922). However, they were again treated as tion of specimens in the present study. a subfamily by Edwards (1925a) when he provided the However, there was a 50% increase in the number of only systematic account of the entire group that has known British species in the 15 years from the publi- yet appeared in the British literature. He did then cation of the handbook to that of the latest checklist of acknowledge that they were far less well known than British Diptera (Chandler, 1998). This clearly demon- other groups of fungus gnats and recognized 45 spe- strated that knowledge of the British fauna was still cies in eight genera, most species still being retained at an early stage and the relative ease of finding fur- in the genus Sciara. One genus and species, Pnyxia ther species in new collections demonstrated that use scabiei (Hopkins), was then included by him in the of the existing handbook could not be a reliable means tribe Leiini. When he later (Edwards, 1941) provided of identifying British sciarids. Consequently, the an update on the British fauna he omitted the Scia- project to prepare a revised handbook was launched. ridae, stating that he was preparing a separate more Because of the large number of additional species detailed revision. Sadly this 1941 paper appeared that were found in British collections, including sev- posthumously due to Edwards’ premature death in eral new to science, it was considered that the present 1940, so his planned work on Sciaridae was never account of the material examined was a necessary pre- completed. liminary step to the preparation of a new handbook. Edwards was actively planning this sciarid revision With the addition of 111 species in the present paper in the 1930s and material was being gathered at the the fauna of the family in the British Isles is now con- Natural History Museum with this in mind. Ralph firmed to be 263, with 261 species found in Britain and Coe was Edwards’ assistant at that time and the 100 species in Ireland. extensive material collected by him in Scotland was presumably obtained for this purpose. Coe compiled the section on Sciaridae in the subsequent checklist HISTORY OF SCIARID STUDIES IN THE (Kloet & Hincks, 1945) and referred to the arrange- BRITISH ISLES ment following the known views of F. W. Edwards. An account of past work on the British fauna is nec- This checklist indicated a great increase in knowledge essary to put the present state of knowledge in per- over previously published work as it included 89 spe- spective. As mentioned, the Sciaridae were sometimes cies placed in only three genera, but with 12 subgen-

BRITISH SCIARID FAUNA 3 era of Sciara recognized. Much work on sorting attempt to revise fully the pinned collection and con- material to species was evidently done by Edwards sequently overlooked the presence of many species. In and there are many specimens in the BMNH collection addition to the species already recognized as distinct with genitalia preparations made by Edwards. These by Edwards, it was also found that most longer series are, as with preparations made by him in other fami- were mixed and there was a potential for recognizing lies of Nematocera, mounted in Canada Balsam on a a larger fauna at the time. strip of celluloid attached to the same pin as the The appearance of Freeman’s Handbook did, never- insect. A good number of the species newly added to theless, lead to an increase in interest in the family the British list in the present work were first recog- among British dipterists and further species rapidly nized in this way by Edwards, although not always came to notice in freshly collected material. Subse- determined by him. Freeman (1983b) generally quent papers (Freeman, 1987, 1990; Ashe & O’Connor, excluded these species from consideration, probably 1991; Hippa & Vilkamaa, 1994; Laurence, 1994; Lau- because there was no slide-mounted material rence & James, 1996; Mohrig & Menzel, 1997) added available. 25 further species. During this period knowledge of There was no further increase in knowledge of the the family in Europe was proceeding rapidly and British sciarid fauna in the next checklist of British many of the species already added or recognized here Diptera (Kloet & Hincks, 1976). The Sciaridae list for as new to the fauna were not distinguished at the time this was again based on the BMNH collection and was the handbook was written. compiled by Tony Hutson, who reduced the list to 85 This increase in knowledge of the family since 1983 named species plus seven unnamed species. However, was summarized by the account in the most recent the generic classification was revised following the checklist (Menzel, 1998), in which species were added monograph of the Finnish species by Tuomikoski that were known to be represented by British material (1960), which formed the basis for modern views on in collections but had not previously been published as generic definitions in the family; 17 genera were rec- British. The main sources of these additions were the ognized, several of them divided into subgenera, while private collections of Brian Laurence and Phil With- 12 species remained unplaced generically. In both the ers, who had both been actively gathering sciarids in 1945 and the 1976 lists the identity of several species the preceding decade. In the checklist there was a net was queried. gain of 29 species, bringing the total list to 153; one of The identification of the British species of the family these (Scatopsciara pusilliformis Mohrig & Mamaev) was still not practicable and this gap in the British lit- was later sunk in synonymy. The most significant erature remained until Paul Freeman embarked on aspect of the 1998 list was the large number of nomen- the preparation of a Royal Entomological Society clatural changes resulting from the examination of Handbook to the British species, published in 1983. In types, details of which were later published in the this work Freeman stated that it was based on the lay- monograph of the Palaearctic species by Menzel & out of species in the BMNH collection, ‘which is a Mohrig (2000). Apart from many generic changes, this record of Edwards’ work and views.’ However, he made also involved changes in specific names for about one- no suggestion that any manuscripts or notes on the third of the species included in the 1983 Handbook. family had been left by Edwards. Freeman (1983b) Little has hitherto been published on the British sta- used the 1976 checklist as a basis for the species tus of the species added in the checklist apart from the included and also followed the generic classification account of Bradysia lutaria (Winnertz) by White & proposed by Tuomikoski (1960). Although this classi- Smith (2000). Chandler et al. (2002) added a further fication has been modified and refined by later work in three species from both Britain and Ireland. Europe, it nevertheless provided a firm basis for fur- The Irish sciarid fauna has remained very poorly ther work on the British fauna. Freeman provided known even compared with that of Britain, although keys to 99 species in 18 genera, mentioning two addi- Haliday (1833) provided some early records and tional doubtful species so that 101 species were Walker (1856) summarized the distribution by indicat- claimed in the included checklist. ing national occurrence, citing 15 species from Ire- During the course of the present work, the BMNH land. This had only increased to 35 when occurrence in collection has been fully revised, resulting in correc- Ireland was similarly indicated in the 1998 checklist. tion of some records cited by Freeman (1983b). As indi- Recently, a further 37 species were added, with some cated above it was evident that Freeman had based deletions giving a net total of 68 species (Chandler his work principally on the slide-mounted collection. et al., 2002; Withers, 2002). He did make fresh slides from pinned specimens of There has been a recent increase in recording of the species where none was already available and cited smaller species, which are not readily obtained by net- some other distribution data from the pinned collecting and for which passive trapping techniques such tion. However, it was apparent that he made no as Malaise traps and water traps are more effective.

4 F. MENZEL ET AL.

The increased use of these techniques in recent years Museum of Ireland, Dublin (NMID) were revised. A has, to a large extent, been responsible for these addi- small amount of material from the Ulster Museum, tions, many of which are reported here for the first Belfast (UMB) was also identified. Examination of time. The wetland surveys carried out in Wales and material in the Hope Department of Entomology, East Anglia, mainly using water traps and the Mal- Oxford University Museum (UMO) was limited to spe- aise trap surveys of some ancient forest areas, such as cies that could be confirmed without preparation. Epping Forest and Burnham Beeches, have resulted Much work is still required on the collection there and in the capture of many interesting additions to the on other major museum collections. The private collec- British fauna. tions of several British dipterists have also been fully All of the collections examined in the course of the covered, including those of the authors and of Brian present work have added species not seen from other Laurence, Phil Withers, David Gibbs and Jonathan sources and the rate of additions to the known fauna Cole. has not diminished throughout the 3–4 years of this The Edinburgh collection mainly comprised the study. The substantial increase in number of species material reared from nests of birds and mammals by over the 1983 Handbook and the 1998 checklist and E. B. Basden. Basden had referred specimens to the fact that so many of these are still known from rel- Edwards for identification and some of those named by atively few records (individuals or localities) suggests Edwards had been deposited in BMNH but the bulk of that the actual number of species resident in the his material remained in his private collection, which British Isles cannot yet be accurately assessed. The went to NMS on his death. The revision of this mate- German fauna of 370 species is still being added to rial has been particularly useful from the biological from new collections, with frequent description of new point of view and it is indicated where present identi- species. It is therefore expected that examination of fications differ from those published by Basden (1933). further collections and coverage of more habitats and Much of the data presented here is based on regions will add further species. Another reason why recently collected material. Frank Menzel and Jane we think that there must be more species in the Smith have both collected sciarids in several parts of British Isles is because there are several species miss- England in recent years and their slide-mounted ing from our list that are widespread throughout material is held at their respective institutions (DEI, Europe and usually occur frequently (e.g. Bradysia HRI). This material was collected from areas with meigeni (Rübsaamen, 1894); B. procera (Winnertz, several suitable localities in Devon, Gloucestershire, 1868); Corynoptera boletiphaga (Lengersdorf, 1940); Leicestershire, Herefordshire, Oxfordshire, Shrop- C. tridentata Hondru, 1968; Ctenosciara lutea shire, Staffordshire, Warwickshire, West Midlands (Meigen, 1804); Epidapus detriticola (Kratochvil, and Worcestershire to improve the data for a new 1936); Phytosciara macrotricha (Lengersdorf, 1926); handbook. Scatopsciara curviforceps (Bukowski & Lengersdorf, Frank Menzel has determined in entirety the slide- 1936) and Sciara analis Schiner, 1864). A total British mounted collection of Brian Laurence, which will now fauna in excess of 300 species now seems within reach. be deposited at UMO apart from some type material However, the characterization of species in the new that will be placed at BMNH as indicated in his Handbook under preparation will, we hope, ensure papers. Frank Menzel also checked some slide- that any species subsequently found but not yet mounted specimens from the private collection of included can be detected more reliably than has hith- Patrick Roper and a large part of the slide-mounted erto been practicable for British workers. collection of Phil Withers, including much of the Welsh wetland material. He also determined and mounted on slides the alcohol-preserved material obtained by THE COLLECTIONS EXAMINED M. de Courcy Williams at the Gearagh, County Cork It was necessary to examine material of all species and this material will be deposited at DEI and NMID. previously recorded from the British Isles, and, as Frank Menzel also similarly processed samples mentioned above, the collection of the Natural History recently obtained by J. W. Ismay & B. Schulten at Museum, London (BMNH) was most important in this Chigwell Row Wood LNR and on Salisbury Plain (DEI, respect. This resulted in a number of corrections and UMO). also some taxonomic and nomenclatural changes, Most of the material from glasshouses, mushroom which are indicated under the relevant species. cultivation and plant compost was collected by E. S. The examination of other museum collections has Binns, P. A. Buxton, B. Gurney, N. W. Hussey, W. J. been more limited, mainly because they comprise Parr, J. E. Smith, E. R. Speyer and P. F. White. Many mostly older collections of dry pinned material. How- of the older specimens from the literature (e.g. Binns ever, the entire collections of the National Museums and Buxton) were found in the collection of HRI. of Scotland, Edinburgh (NMS) and the National Newer material came from farms and nurseries

BRITISH SCIARID FAUNA 5 throughout the UK. All of these specimens were of 14 987 specimens were checked and recorded. These revised by Jane Smith and Frank Menzel and are in comprised 12 529 males, 2419 females, five pupae and the collections of DEI and HRI. 34 larvae. The females, pupae and/or larvae (including The collections of P. J. Chandler, D. J. Gibbs and J. reared material) were only included in the species list H. Cole were determined by Peter Chandler but all when they were positively identified to species. Cor- critical material, including examples of all species rect identifications were made by comparison against additional to the British fauna, was referred to Frank specimens from the authors’ collections when neces- Menzel for verification. These collections are mainly sary. The results established that the sciarid fauna of pinned, but often male genitalia, and in the case of the British Isles has, to date, a total of 263 species. the first two collections many whole specimens, are These are included in the following genera: Bradysia mounted in DMHF (dimethyl hydantoin formalde- (74 species), Bradysiopsis (1), Camptochaeta (5), hyde) on glass cover slips glued to card, the specimen Claustropyga (3), Corynoptera (55), Cratyna (20), usually placed above a round hole in the card formed Ctenosciara (1), Epidapus (8), Hyperlasion (1), Leptos- using a hole punch. Some material from P. J. Chan- ciarella (16), Lycoriella (14), Phytosciara (7), Pnyxia dler’s collection has been deposited at NMS and (1), Pseudolycoriella (6), Scatopsciara (22), Schwenck- NMID. feldina (1), Sciara (6), Scythropochroa (2), Trichoda- A full list of the collectors whose names appear on pus (1), Trichosia (11), Xylosciara (6) and Zygoneura labels is given below. Many of these relate to museum (2). specimens or to material referred for identification to The nomenclature and sytematics used are based on one of us or to Brian Laurence or Phil Withers. For the revision by Menzel & Mohrig (2000) and Menzel & example, material from the East Anglian wetland sur- Heller (2005). The morphology of the adults and the vey (collected by A. Foster and D. Procter) was initially terminology (naming and description of characteris- determined by Brian Laurence and that from the tics of Sciaridae, technical terms) comply with Menzel Welsh wetland survey (collected by D. Boyce, P. & Mohrig (1997b, 2000). Holmes and D. Reed) was dealt with by Phil Withers. The Malaise trap material from Epping Forest and COLLECTIONS AND ABBREVIATIONS Burnham Beeches, for which J. Dagley and J. W. Ismay This investigation included British Sciaridae from the were, respectively, the collectors, was similarly pro- following collections: cessed by Peter Chandler. In all cases the data given for individual specimens includes indication of both BMNH The Natural History Museum, London, the collector and the collection in which the specimens England, Great Britain (loans via Dr B. R. are deposited. Pitkin, B. C. Townsend, J. Chainey, N. Wyatt) A list is also given of all localities from which spec- DEI Deutsches Entomologisches Institut, ZALF imens have been examined. Because of the way in e.V., Müncheberg (former Eberswalde), which these data were initially assembled it proved Germany (incl. private collection Dr Frank most practical to group these under the latest county Menzel, Eberswalde) boundaries, with some flexibility where counties have HRI Horticulture Research International, Welles- recently been divided into several unitary authorities bourne, England, Great Britain (e.g. Avon and Berkshire are retained as entities for NMID National Museum of Ireland, Dublin, Ireland convenience). Where there has been a change in the (loan via Dr M. de Courcy Williams, Dr J. P. county assignment of a locality the previous county O’Connor) name is indicated to assist in relating to the informa- NMS National Museums of Scotland, Edinburgh, tion on data labels. For each locality the precise loca- Scotland, Great Britain (loan via Dr G. E. tion and habitat have been identified as far as Rotheray) practicable. In the case of recently collected specimens PBLN Private collection Brian R. Laurence, such information has been obtained from the collec- Northumbria, England, Great Britain [for- tors where it was not already stated on the label. For merly School of Biological Sciences, Univer- older museum material data given on labels is often sity of East Anglia, Norwich] (loan via B. R. not very precise and in these cases grid references are Laurence to be deposited in UMO) only given or habitats suggested where these could be PDGB Private collection David J. Gibbs, Bristol reliably estimated. (loan via D. J. Gibbs) PJCB Private collection Jonathan H. Cole, Bramp- ton, Cambridgeshire, Great Britain (loan via MATERIAL AND METHODS J. H. Cole) Sciarid specimens from throughout the British Isles PPCM Private collection Peter J. Chandler, Melk- were examined and identified to species level – a total sham, England, Great Britain

6 F. MENZEL ET AL.

PPRS Private collection Patrick Roper, Sedle- Hale-Carpenter, H. M. Hallett, A. H. Hamm, K. M. scombe, East Sussex, England, Great Britain Harris, D. J. de C. Henshaw, B. M. Hobby, W. G. Hoff, (loan via P. Roper) P. Holmes, E. Holmes-Smith, M. Howe, E. Hurrell, C. PPWM Private collection Phil Withers, Montée du P. Hurst, N. W. Hussey, A. M. Hutson, J. W. Ismay, D. Cimetière, Sainte Euphémie, France (loan J. Jackson, E. A. Jarzembowski, F. Jenkinson, H. P. via P. Withers) Jones, M. Jones, T. J. Jordan, D. Keilin, D. K. Kevan, J. UMB Ulster Museum, Belfast, Northern Ireland J. F. X. King, D. L. Lack, R. P. Lane, Latham (initials (loan via R. Nash) unknown), B. R. Laurence, B. Levey, A. M. Low, M. Luff, UMO Oxford University Museum of Natural His- I. MacGowan, I. F. G. McLean, J. R. Malloch, G. Mans- tory, Oxford, Oxfordshire, Great Britain bridge, F. Menzel, R. K. Merriﬁeld, H. W. Miles, C. ZFMK Zoologisches Forschungsinstitut und Moreby, C. Morley, J. Mousley, R. Nash, R. Newstead, Museum Alexander Koenig, Bonn, Germany C. Nicholson, G. E. Nixon, C. G. Nurse, J. P. O’Connor, (loan via Dr H. Ulrich) M. O’Connor, J. A. Owen, L. Parmenter, W. J. Parr, M. Pavett, K. Paviour-Smith, E. C. Pelham-Clinton, J. F. In addition, three collections need to be mentioned Perkins, I. Perry, A. Piffard, C. G. Piffard, A. R. Plant, that have no Sciaridae material from the British Isles. C. W. Plant, C. M. Port, K. Porter, D. Procter, C. H. W. The type material from newly described species, col- Pugh, M. N. Pugh, B. Rands, C. Raper, D. Reed, J. C. lected in other European countries, are held in these Reid, O. W. Richards, N. Riddiford, Robinson (initials collections: unknown), Rochford (initials unknown), P. Roper, G. E. NKME Naturkundemuseum der Stadt Erfurt, Rotheray, N. C. Rothschild, W. M. Russell, B. Schulten, Erfurt, Germany (loan via M. Hartmann) M. J. Service, D. Sharp, M. R. Shaw, I. Sims, P. Skid- PKHH Private collection Kai Heller, Heikendorf more, J. E. Smith, K. G. V. Smith, M. C. D. Speight, E. (Germany) (loan via K. Heller) R. Speyer, G. M. Spooner, W. H. Staunton, P. Stephens PWMP Private collection Prof. Dr Werner Mohrig, [also written as P. Stevens], G. H. H. Stokes, M. Storey, Poseritz, OT Puddemin, Germany (loan via A. E. Stubbs, D. Sumner, T. H. Taylor, F. V. Theobald, I. Prof. Dr W. Mohrig) Thomas, J. Topham, B. C. Townsend, R. W. J. Uffen, R. I. Vane-Wright, C. R. Vardy, G. H. Verrall, F. Walker, T. de G. (Lord) Walsingham, A. Warne, J. Warner, J. War- COLLECTORS wick, C. O. Waterhouse, A. R. Waterston, J. Waterston, As collectors’ initials are omitted in the list of data J. H. Watson, M. A. Watson, J. A. Webb, W. Wesche, B. (except in cases where there is more than one collector Wheeler, P. F. White, O. H. Wild, C. B. Williams, S. A. with the same surname), they are given below: Williams, P. Withers, J. H. Wood, B. Woodcock, N. P. D. M. Ackland, F. C. Adams, A. Al-Amidi, P. Ashe, E. Wyatt, P. Wycherley, J. W. Yerbury. C. M. d’Assis-Fonseca, E. W. Aubrook, E. E. Austen, M. A. Austin, T. Bainbridge-Fletcher, C. J. G. Baker, Barn- HABITAT DESCRIPTIONS ston (initials unknown), E. B. Basden, E. G. Bayford, A. Beaumont, B. P. Beirne, H. E. J. Biggs, E. S. Binns, N. Bog: colloquial term for ombrotrophic mire (see under L. Birkett, R. Blackith, K. G. Blair, K. P. Bland, M. Bos- mire). They receive nutrients only from precipita- ton, P. A. Bottomley, D. Boyce, E. S. Bradford, R. C. Bra- tion (oligotrophic) so are poorly fed. Blanket bog: dley, C. T. H. Bradshaw, H. Britten, J. Brock, G. Brown, large area of peat. Raised bog: dome-shaped peat J. Buller, E. Burtt, H. Bury, P. A. Buxton, A. Calvert, A. bog that has developed in a former lake on a peat fen Cant, A. E. J. Carter, D. J. Carter, J. E. Chainey, S. J. once the mineral-rich water supply has been cut off. Chambers, P. J. Chandler, C. A. Cheetham, G. A. Clout, Breckland heath: dry, sandy grass heath of Breckland, R. L. Coe, M. Cohen, J. H. Cole, L. W. Cole, M. J. Collier, in East Anglia, with thin patterned sands overlying J. E. Collin, F. Collins, C. N. Colyer, J. Cooter, J. Coul- chalk. Some Breckland heaths are a mix of acidic son, M. de Courcy Williams, J. Cowley, P. S. Cranston, and chalk-dependent plants, often growing within R. Crossley, J. Dagley, C. W. Dale, J. P. Dear, R. H. L. inches of each other. Disney, W. L. Distant, H. StJ. K. Donisthorpe, C. M. Carr: an un-managed fen where small shrubs and Drake, Dudgeon (initials unknown), A. B. Duncan, R. trees start to grow, leading to the natural creation of Dunne, C. E. Dyte, K. M. Edelsten, F. W. Edwards, C. carr woodland. Elbourn, W. Elstow, P. Entwistle, R. E. Evans, M. Eyre, Fen: the ﬁrst stage in the natural change, or succession, J. W. Fidler, A. Foster, A. Fox-Wilson, P. Freeman, J. C. from open water to woodland. Waterlogged areas of F. Fryer, R. S. George, D. J. Gibbs, D. R. Giffard, P. A. land dominated by reeds, rushes and sedges, fens Goddard, A. Godfrey, J. Good, J. G. Gordon, A. Gordon- form a complicated system of plant and animal com- Smith, W. R. Grant, E. E. Green, L. W. Grensted, P. H. munities with a mineral-rich water supply in part at Grimshaw, B. Gurney, D. Hackett, J. N. Halbert, G. D. least from the groundwater table (geotrophic).

BRITISH SCIARID FAUNA 7

Grikes: the gaps between limestone rocks formed by mon mallow, Malva sylvestris; foxglove, Digitalis water erosion. purpurea; hair grass, Deschampsia species; heather, Hanger: a wood growing on the side of a hill. Calluna vulgaris; hoary ragwort, Senecio erucifolius; Heath: wide open landscape dominated by heathers, hogweed, Heracleum sphondylium; horsetails, Equise- gorse and grasses with occasional trees such as pine tum species; ivy, Hedera helix; ivy broomrape, and silver birch. The soil is usually sandy and thus Orobanche hederae; marsh marigold, Caltha palustris; acid, dry in summer and prone to fires. Wet heaths moss campion, Silene acaulis; nettle, Urtica dioica; occur where underlying rocks or clay prevent water purple moor grass, Molinia cearulea; ragwort, Senecio drainage so the water table is high; dry heaths are jacobaea; reed, Phragmites; reed mace, Typha; free draining and humid heaths are between the rhubarb, Rheum rhaponticum; royal fern, Osmunda; two. rushes, Juncus species; sea kale, Crambe maritima; Mire: a wetland that supports vegetation that is peat sedges, Carex, Cladium, Trichophorum; thistle, Cir- forming. Basin mire is a type of fen formed in a sium species; guelder rose, Viburnum lantana; water shallow hollow left by glacial drift; ombrotrophic dropwort, Oenanthe species; wild parsnip, Pastinaca mire has its water and nutrient supply directly from sativa; wood anemone, Anemone nemorosa. rainfall. Fungi: Agaricus campestris, Agrocybe praecox, Aman- Moor: open, semi-natural habitat with dwarf shrub ita muscaria, A. rubescens, Auricularia mesenterica, heaths often intermixing with farmland. It includes Bjerkandera adusta, Calocybe gambosa, Collybia dry and wet heaths, blanket and raised bogs, valley dryophila, Grifola gigantea, Ganoderma applanatum, mire and rough grassland. Gymnopilus junonius, Hebeloma species, Hirneola Montane: mountainous. auricula-judae, Hypholoma fasciculare, H. velutinum, Tufa: a limestone deposit that forms on cliffs, caves Hypoxylon fragiforme, H. rubiginosum, Inonotus and quarry faces where limestone is naturally cuticularis, I. hispidus, I. nodulosus, I. radiatus, Kue- occurring. This rock can form very quickly if condi- hneromyces mutabilis, Lactarius species, Laetiporus tions are favourable. Plants, mosses and inverte- sulphureus, Leccinum scabrum, Lepiota cristata, Pez- brates often become encrusted in the calcite, and iza carnea, P. vesiculosa, Phaeolus schweinitzii, Pipto- preserved as fossils. porus betulinus, Pleurotus cornucopiae, P. ostreatus, Turlough: a flat-bottomed sinkhole in limestone areas Polyporus squamosus and other Polyporus species, that can be seasonally flooded. Russula alutacea, R. atropurpurea, R. pectinata, R. vesca, Sparassis crispa, Stereum sanguinolentum, Trametes versicolor. LATIN AND COMMON NAMES OF FLORA AND FAUNA ASSOCIATED WITH HABITATS Invertebrates: ants, Formica rufa, Lasius fuliginosus, L. niger, Myrmica ruginodis; beetles, Hylesinus frax- Trees: alder, Alnus glutinosa; ash, Fraxinus excelsior; ini, Ischnomera caerulea; slug, Testacella species. aspen, Populus tremula; beech, Fagus sylvatica; birch, Betula (pendula); blackthorn, Prunus spinosa; cork Birds: barn owl, Tyto alba; blackbird, Turdus merula; oak, Quercus suber; cypress, Chamaecyparis lawsoni- blue tit, Parus caeruleus; coot, Fulica atra; dunnock, ana; elder, Sambucus nigra; elm, Ulmus species; fir, Prunella modularis; house sparrow, Passer domesti- Abies species; hawthorn, Crataegus monogyna; hazel, cus; linnet, Carduelis cannabina; little owl, Athene Corylus avellana; holly, Ilex aquifolium; hornbeam, noctua; magpie, Pica pica; moorhen, Gallinula chloro- Carpinus betulus; horse chestnut, Aesculus hippoc- pus; robin, Erithacus rubecula; rook, Corvus frugile- astanum; large-leaved lime, Tilia platyphyllos; lime, gus; sand martin, Riparia riparia; song thrush, Tilia species; maple, Acer campestre; mountain ash, Turdus philomelos. Sorbus aucuparia; pedunculate oak, Quercus robur; Mammals: brown rat, Rattus norvegicus; common pine, Pinus species; poplar, Populus nigra; sea buck- shrew, Sorex araneus; dormouse, Muscardinus avella- thorn, Hippophae rhamnoides; spruce, Picea abies; narius; field mouse, Apodemus sylvaticus; grey sycamore, Acer pseudoplatanus; wild olive, Olea euro- squirrel, Sciurus carolinensis; hedgehog, Erinaceus pea; willow, Salix species. europaeus; mole, Talpa europaea; rabbit, Oryctolagus Other plants and flowers: agrimony, Eupatorium; bil- cuniculus. berry, Vaccinium myrtillus; black bog rush, Schoenus nigricans; bluebell, Hyacinthoides nonscripta; bog- LOCALITIES bean, Menyanthes trifoliata; bog myrtle, Myrica gale; bramble, Rubus fruticosus; butterbur, Petasites hybri- There have been many changes in county names and dus; buttercup, Ranunculus species; cabbage thistle, borders since 1954 and there is considerable confusion Cirsium oleraceum; cinquefoil, Potentilla species; comin some areas between postal addresses and counties.

Where a change in boundary had occurred, former Luton Airport (TL1120) calcareous grassland on counties have been included and are given in []. Where steep bank near runway available, both details of the habitat and the approx- Maulden Wood (TL0739) mixed wood on Lower imate Ordnance Survey Landranger grid references – Greensand in ( ) – have been given. In many cases the specimens Old Warden (TL1343) reared from fungi were collected too long ago for this information to be Sandy (TL1749) available. Sandy Warren (TL1948) broad-leaved woodland on Lower Greensand Sharpenhoe, woods at south end (TL0629) broad- ENGLAND leaved woodland AVON (the former county now comprising the unitary Sharpenhoe, The Clappers (TL0630) beech authorities of Bath and North East Somerset, North woodland Somerset, City of Bristol and South Gloucestershire) Shefford (TL1438) Blaise Wood [Gloucestershire] (ST5577) broad- Tempsford (TL1654) leaved woodland BERKSHIRE [= Berks.] Bristol, Redfield [Gloucestershire] (ST6173) garden Aldermoors (SU7774) damp broad-leaved woodland of D. J. Gibbs Ashley Hill (SU8281) mixed woodland Bristol, Trooper’s Hill [Gloucestershire] (ST6273) Barkham, reared from fungus acid grassland and heath Binfield (SU8471) Brown’s Folly [Somerset] (ST7965) calcareous Bucklebury Common (SU52-55/69–70) broad- broad-leaved woodland leaved woodland Burledge Hill [Somerset] (ST5959) neutral Burghfield Common (SU6567) broad-leaved grassland woodland Cleeve Heronry [Somerset] (ST4666) calcareous Calcot (SU6671) floodplain meadow broad-leaved woodland California Country Park (SU7865) carr and mixed Coombe Dingle [Gloucestershire] (ST5577) broad- drier woodland leaved woodland Carbins Wood (SU5668) broad-leaved woodland Emerson’s Green [Gloucestershire] (ST6776) pas- Cock Marsh (SU8886) water meadows ture, hedgerow Dinton Pastures Country Park (SU77–78/71–72) Folly Farm [Somerset] (ST6060) neutral broad- carr woodland (Sandford Copse), wooded bank of leaved woodland River Loddon, water meadows and gravel pits, Gordano, Walton Moor [Somerset] (ST4372) herb- old hedges (Mortimer’s Meadow) rich grassland Hambleden Weir (SU7884) reared from nests Hengrove Park [Somerset] (ST5968) wasteland and Hungerford (SU3368) scrub Lodge Wood (SU7873) broad-leaved woodland Leigh Wood [Somerset] (ST5573) calcareous broad- Maidenhead (SU88) mushroom farm leaved woodland Newbury (SU46) reared from nests Lord’s Wood [Somerset] (ST6363) mixed woodland Pearman’s Copse (SU7369) broad-leaved woodland Middle Hope [Somerset] (ST3366) calcareous Pine Wood (SU8365) acid mixed woodland grassland Pinkney’s Green (SU6682) reared from nests Monk Wood [Somerset] (ST7570) calcareous broad- Searles Farm (SU67) carr woodland by gravel pit leaved woodland lakes (SU6970), carr by river Kennet (SU6871) Nailsea [Somerset] (ST4770) moorland Silwood Park (SU9468) mixed woodland Purn Hill [Somerset] (ST3357) calcareous grassland Temple (SU8384) reared from nests Tickenham Hill [Somerset] (ST4472) calcareous Thatcham (SU5166) reed (Phragmites) bed broad-leaved woodland and grassland Theale (SU6469-6570) carr woodland and ash Weston Big Wood [Somerset] (ST4574) calcareous coppice broad-leaved woodland Theale, Sheffield Bottom (SU6469) broad-leaved BEDFORDSHIRE [= Beds.] woodland between lakes and canal Begwary Brook (TL1656) marsh, rough grassland Theale Lake (SU6870) partly wooded lake and scrub margin Biggleswade (TL1944) Warren Row (SU8180) reared from nests Bromham (TL0151) reared from fungi Wildmoor Heath (SU8257) heathland and mixed Cardington (TL0847) woodland Flitwick Moor (TL0435) valley mire with acid and Windsor Forest (SU9274-9373) ancient oak and neutral carr beech forest

Windsor Great Park (SU97) parkland with ancient Badger Wood (TL3259) broad-leaved woodland oaks Bedford Purlieus [Northamptonshire] (TF0300– Wokefield Common (SU6566) broad-leaved wood- TL0499) broad-leaved woodland with stream land and heathland Beech, Devil’s Ditch (TL66) calcareous grassland BUCKINGHAMSHIRE [= Bucks.] Bone Mill Hollow (TF0301) Beaconsfield (SU99) reared from nests Brampton, Lenton Close [Huntingdonshire] Black Park (TQ0184) mixed woodland (TL2169) house and garden of J. H. Cole Bockmer End (SU8287) reared from nests Brampton, Ouse flood meadow [Huntingdonshire] Boveney and Boveney Lock (SU9477) wooded banks (TL2169) rough grazing meadow and ditches of river Brampton, Portholme Meadow [Huntingdonshire] Brockhurst Wood (SU9684) broad-leaved woodland (TL2370) ditch with old willow Burnham and East Burnham (SU98) reared from Brampton Wood [Huntingdonshire] (TL1870) nests ancient woodland on clay Burnham Beeches (SU9485, SU9584, SU9585) Cambridge (TL4559) ancient beech woodland, bog, heath Chippenham Fen (TL6468-6568) fen and carr Cliveden (SU9083) broad-leaved woodland woodland Collum Green, reared from nests Earith Gravel Pits [Huntingdonshire] (TL3976) Dancer’s End (SP9009) broad-leaved woodland scrub and rough grassland by pits Dropmore (SU9387) reared from nests Gamlingay Wood (TL2453) ancient broad-leaved Farnham Common (SU9685) reared from nests woodland on clay and sandy loam Farnham Royal (SU9683) reared from nests Godmanchester, Osier Lake [Huntingdonshire] Fawley (SU7687) reared from nests (TL2672) carr, marsh and pond George Green (TQ0081) reared from nests Grafham Water, Savage’s Creek [Huntingdon- Gerrard’s Cross (SU9987) reared from nests shire] (TL1369) stream through marsh into Great Missenden (SP9102) mushroom farm reservoir Greenlands (SU7886) reared from nests Hayley Wood (TL2953) ancient broad-leaved Hambleden Great Wood (SU7987) broad-leaved woodland woodland Hemingford [Huntingdonshire] (TL2871) Iver Heath (TQ0383) reared Hinchingbrooke Country Park [Huntingdonshire] Ivinghoe (SP9416) calcareous grassland (TL2171) mixed woodland Medmenham (SU8184) reared from nests Holme Fen [Huntingdonshire] (TL2089) scrub and Milton Keynes Sphagnum bog Old Slade Wood (TQ0378) broad-leaved woodland Holme Fen, Denton Covert [Huntingdonshire] Seer Green (SU9792) reared from nests (TL1989) dry birch woodland on peat Slough Kirtling (TL6957) Stoke Common (SU9885) broad-leaved woodland Lady’s Wood [Huntingdonshire] (TL2482) fragment Stoke Court Park (SU9783) woodland and parkland of ancient woodland Stoke Poges (SU9883) broad-leaved woodland Little Paxton Pits [Huntingdonshire] (TL1963) Stoke Poges, Hollybush Hill (SU9984) broad-leaved rough grassland and scrub woodland Monkfield Wood (TL3259) broad-leaved woodland Stoke Wood (SU9785) broad-leaved woodland Monk’s Wood [Huntingdonshire] (TL1979-80) Taplow (SU9283) ancient woodland on clay The Coombe (SP9615–9715) broad-leaved woodland Newmarket (TL6162) Turville Heath (SU7491) broad-leaved woodland Oakland Wood (TL3159) broad-leaved woodland and grassland Perry West Wood [Huntingdonshire] (TL1366) Wexham Place Wood (SU9984) broad-leaved broad-leaved woodland woodland Raveley Wood [Huntingdonshire] (TL2481) ancient CAMBRIDGESHIRE [= Cambs.] woodland on clay Abbot’s Ripton, Bevills Wood [Huntingdonshire] St. Ives, Holt Island [Huntingdonshire] (TL3171) (TL2079) mixed woodland on clay willow carr Alconbury [Huntingdonshire] (TL1874) ditch along Shepreth (TL3948) reared from fungus meadow hedge Sutton Fen (TL4078) fen Archers Wood [Huntingdonshire] (TL1781) broad- Sutton Wash (TL4178) grazing marsh on Ouse leaved woodland on clay Washes Aversley Wood (TL1782) broad-leaved woodland The Oaks (TL3159) broad-leaved woodland Babraham (TL5150) Upware (TL5470)

Upwood Meadow [Huntingdonshire] (TL2582) Paramoor Wood (SW9748) broad-leaved woodland unimproved grassland and hedges Pentewan (SX0147) alder woodland Wansford Meadow [Northamptonshire] (TL0799) Penzance (SW4730) limestone grassland with ﬂush Redmoor (SX0761) heath grassland and scrub Warboys, Pingle Cutting [Huntingdonshire] Rosevear (SW7024) wooded cottage garden (TL3181) grassland and scrub Sheviock (SX3755) Warboys, Pingle Wood [Huntingdonshire] (TL3181) St. Erth (SW5535) ancient woodland on clay St. Ives (SW5038) mushroom farm Warboys, Wistow Wood [Huntingdonshire] (TL2982) Scilly Isles, Tresco (SV9015) ancient woodland on clay Tintagel (SX0488) cliff grassland Warboys Wood [Huntingdonshire] (TL3082) broad- Tregardock Cliff (SX0483) leaved woodland on clay Tresillian (SW8546) broad-leaved woodland Welches Dam (TL4786) within Ouse Washes bird Valency Valley (SX1091-1190) broad-leaved hide woodland Wennington Wood [Huntingdonshire] (TL2478) CUMBRIA broad-leaved woodland on clay Bleathwaite Coppice (SD2995) broad-leaved Wicken Fen (TL5570) fen meadows and ditches woodland Woodditton Wood (TL6658) broad-leaved woodland Blelham Bog [Lancashire] (NY3600) Woodwalton Fen [= Wood Walton; Huntingdonshire] Brigsteer Park [Westmorland] (SD4887) (TL2283-84) fen and alder carr Burton Wood CHESHIRE Clifton [Cumberland] (NY5326) Arden Hall (SJ7683) Dent (SD5187) Bollin Valley (SJ78) Dun Fell [Cumberland] (NY7532) Chester Fish House Moss [Lancashire] (SD3382) Cotterill Clough (SJ8083) Gait Barrows [Westmorland] (SD4877) broad- Delamere (Forest) (SJ5471) broad-leaved woodland leaved woodland on limestone pavement and conifer plantations Grange-over-Sands [Lancashire] (SD4077) Longdendale [most of this valley is in Derbyshire] Great Ellerside [Lancashire] (SD3586) broad- (SK0499) leaved woodland Poulton (SJ4059) High Dam [Lancashire] (SD3688) Rostherne (SJ7483) Holker Moss [Lancashire] (SD3678) Warrington (SJ6088) Humphrey Head [Lancashire] (SD4075) CORNWALL Ickenthwaite [Lancashire] (SD3289) Bodmin (SX0768) Jonny Wood [Cumberland] (NY2514) broad-leaved Caerhays Castle (SW9741) woodland Caerkief Farm and Wood (SW7752, SW7852) Kendal (SD4695) mushroom farm Coombe, woods to north (SS2112-2211) broad- Lake Windermere [Westmorland] (SD3998) leaved woodland with stream Lancelot Clark Storth Nature Reserve [Westmor- Cottage Meadow, Redlake land] (SD5447) open birch woodland on limestone Croft Pascoe Pool (SW7319) Lodore [Cumberland] (NY2618) Devichoys Wood (SW7738) broad-leaved woodland Meathop [Westmorland] (SD4481) Glendurgan (SW7727) Moorthwaite (NY5151) Goonvrea (SW7150) Nichol’s Moss [Lancashire] (SD4382) Gurnard’s Head (SW4338) Roudsea Wood [Lancashire] (SD3382) broad-leaved Gweek Wood (SW7026) coastal broad-leaved woodland woodland Rydal Water [Westmorland] (NY3506) Hayle Kimbro Pool (SW6917) wooded margin Sandscale Haws [Lancashire] (SD1875) coastal Helston (SW6627) sand dunes Kennack Cove (SW7316) coastal broad-leaved Skinner’s Pastures [Lancashire] (SD3487) broad- woodland leaved woodland Land’s End (SW3425) vegetable detritus on cliff top Stock Ghyll [Westmorland] (NY3906) with salty spray Swindale Beck (NY5416) Lelant (SW5437) Thursby (NY3050) mushroom farm Maen Porth (SW7829) Tilberthwaite Ghyll [Lancashire] (NY2801) Mevagissey (SX0145) Ullswater, Glencoyne Wood [Cumberland] (NY3818) Nanjizal Cove (SW3523) Walney [Cumberland] (SD1867)

Way Beck [Westmorland] (SD4085) mixed woodland Ivybridge (SX6356) broad-leaved woodland Whitbarrow Scar [Westmorland] (SD4387) grass- Kismeldon Meadow (SS3416) land and scrub on limestone escarpment Knap Mill (SX7147) shaded riverbank Windermere (SD4199) Lifton Park (SX3985) Witherslack [Westmorland] (SD4385-87) Linton (SS2419) DERBYSHIRE Littlehempston (SX8162) Chatsworth Deer Park (SK2670) mixed woodland Lundy Island (SS1345) and parkland Manaton, Becky Woods (SX7580) wet woodland on Chee Dale (SK1273) granite, alder, birch Dovedale (SK1451) Newton Abbot (SX8772) Goyt Valley [Cheshire] (SK0177) Peppercombe (SS3823) stream in cliff gully Lathkill Dale (SK2066) Plymouth (SX4857) Longdendale [part in Cheshire] (SK0398) Plymstock (SX5153) Matlock (SK3060) Powler’s Piece (SS3718) Miller’s Dale (SK1473) Putsborough (SS4541) DEVON Rockford, River East Lyn (SS7547) wooded river Avon Valley Wood gorge Aylesbeare (Common) (SY0590) Riversmead (SS5935) Beaford Bridge (SS5414) Rousdon (SY2992) Beesands (SX8241) Salcombe (SX7439) Berry (SS9817) Seaton (SY2490) Berry Pomeroy Castle (SX8361) Sidmouth (SY1288) Bovey Valley (SX4595) broad-leaved woodland Slapton Ley Nature Reserve (SX8245) mixed vege- Bovey Valley, Rudge Wood (SX7879) broad-leaved tation by coastal lake woodland Spreacombe Chapel Wood (SS4841) broad-leaved Brendon (SS7647) woodland Brownsham (SS2925) broad-leaved woodland Stoke Gabriel [under S. Devon; = South Devon] Buckfastleigh (SX7466) (SX8457) Budshead Wood Tamerton Folliatt [= Foliot] (SX4761) Canonteign Falls (SX8382) ancient wet deciduous Torcross (SX8242) woodland, ferns Twitchen (SS6541) Clovelly (SS3124) mixed woodland Welcombe Valley (SS2217) broad-leaved woodland Cod Wood (SX7888) Whiddon Deer Park (SS5639) parkland Crownhill Park (SX45) Windbury (Wood) (SS2826) broad-leaved Dart Head (SX7767) woodland Dartmeet (SX6773) riverbank woodland with Wistmans Wood (SX6176) stunted oak woodland on ﬂushes moorland Dartmeet, Fernworthy Reservoir (SX6785) Woolacombe (SS4644) Dartmoor, Bush Down (SX6983) moorland Yarner Wood (SX7778) broad-leaved woodland Dartmouth (SX8751) Yelverton (SX5268) Dawlish Warren (SX9878) ﬁxed dune grassland and DORSET scrub Affpuddle Heath (SY8092) conifer plantation Dean Wood (SX7164) Arne (SY9788) dry heath and oak woodland, adja- Dendles Wood (SX6161) broad-leaved woodland cent to saltmarsh Dowlands Cliff (SY2990) Aunt Mary’s Bottom (ST5402) carr with chalk Edge Barton (SY1889) stream Ellerton Wood, broad-leaved woodland Badbury Rings (ST9603) calcareous grassland, Exeter (SX9292) woodland and seepages Fingle Bridge (SX7489) broad-leaved woodland Binnegar Quarry (SY8788) sandy exposures and Five Oaks heath Golden Park (SS2421) Chesil Beach (SY6182) coastal shingle Harpford Common (SY0791) Corfe Common (SY9680) grassland and scrub with Haytor (SX7677) moorland on granite stream Hembury Wood (SX7268) broad-leaved woodland Dorchester (SY6991) Holne (SX7169) woods by River Dart Ferndown (SU0701) Ilfracombe (SX7069) woodland Glanvilles Wootton (ST6808)

Godlingstone Heath (SZ0182) sandy heathland and Wadhurst Place (TQ6432) bog Withyham (TQ5036) Hartcliff Combe (ST8009) carr woodland with chalk ESSEX springs Belfairs (TQ8187) broad-leaved woodland Hod Hill (ST8510) calcareous grassland and scrub Chigwell Row, Chigwell Row Wood (TQ4692–93) Holt Forest (SU0406) ancient and secondary broad-leaved woodland Kingston Wood (SY9579) calcareous broad-leaved with sycamore, poplar, oak, hornbeam, hazel, woodland holly, ivy, bramble and a small patch of heath Long Crichel (ST9811) (heather, Juncus, Deschampsia) Piddletrenthide (SY7100) Clacton-on-Sea (TM1715) Portland (SY6975) Epping Forest (TQ39–43/82–98, TL43–44/00) Rempstone Heath (SY9985) ancient broad-leaved woodland and grassland; Ringstead (SY7682) (Monument Field (TQ4195), ﬁeld partly sur- Slepe Heath (SY9485) dry heathland, grassland rounded by woodland; Epping Thicks (TL4400), and scrub ancient beech woodland) Stoborough Heath (SY9485) dry heathland, grass- Leigh-on-Sea (TQ8285) salt marsh land and scrub Mersea (TM0513) Stokeford Heath (SY8788) sandy heathland Nazeing (TL4207) Sutton Common (SU0510) Thundersley Glen (TQ7987) broad-leaved woodland The Spittles (SY3593) coastal clay cliffs with broad- West Mersea (TM0112) leaved woodland GLOUCESTERSHIRE [= Gloucs.] Verwood (SU1108) Ashwell Grove (ST5699) broad-leaved woodland Winfrith (SY8187) Ban-y-Gor Wood (ST5496) calcareous broad-leaved Woolcombe Meadow and Wood (SY5595) acid grass- woodland land and wet woodland Chalford (SO9003) Yetminster (ST6011) mushroom farm Chedworth (SP0512) DURHAM Cheltenham (SP0220) mushroom farm Cassop Vale (NZ3438) calcareous grassland and Collinpark Wood (SO7427) acid broad-leaved mining subsidence lake woodland Castle Eden Dene (NZ4138) broad-leaved woodland Crickley Hills Country Park (SO9316) and stream in limestone gorge Daneway (SO9403) broad-leaved woodland High Force (NY8828) vicinity of waterfall Dowdeswell Wood (SO9920) broad-leaved woodland Nesbitt Dene (NZ4537) broad-leaved woodland and Forest of Dean, ancient broad-leaved forest stream in limestone gorge Hencliff Wood [Gloucestershire] (ST6371) acid Scargill (NZ0510) broad-leaved woodland by stream broad-leaved woodland Teesdale (NY9426) Hobb’s Quarry (SO6919) calcareous broad-leaved Wemmergill Hall (NY8922) woodland EAST RIDING OF YORKSHIRE (formerly included in Lancaut Nature Reserve (ST5496) calcareous Humberside) broad-leaved woodland Allerthorpe (SE7847) wooded heathland Longborough (SP1830) Barmby Moor (SE74) Perrott’s Brook (SP0106) broad-leaved woodland EAST SUSSEX Randwick (SO8206) broad-leaved woodland Crowborough (TQ5130) Rodborough Forest (SO8505) Duddleswell (TQ4627) Shortwood (SO8308) calcareous broad-leaved Frant (TQ6036) woodland Hailsham (TQ5909) mushroom farm Stroat (ST5797) Hastings, Hastings Castle (TQ8209) grassland on Witcombe Wood (SO9113) broad-leaved woodland maritime, sandstone cliff top Wotton-under-Edge (ST7593) Hellingly (TQ5812) GREATER LONDON Hurtwood, Peasmarsh (TQ8823) Acton [Middlesex] (TQ2182) Old Lodge Warren (TQ5531) Addington [Surrey] (TQ3764) Patcham (TQ3009) Beckenham Place Park [Kent] (TQ3870) Rye Harbour (TQ9419) shingle beach Beddington [Surrey] (TQ2965) Seaford (TV4899) Bromley [Kent] (TQ4167) former garden of P. J. Sedlescombe (TQ7819) garden of P. Roper, grass- Chandler land at woodland fringe Bromley, Bickley [Kent] (TQ4369)

Bromley, Brook Wood [Kent] (TQ4166) broad-leaved Stanmore (Common) [Middlesex] (TQ1593) broad- woodland leaved woodland Bromley, Fisher’s Wood [Kent] (TQ4067) broad- Teddington [Surrey] (TQ1671) leaved woodland Thornton Heath [Surrey] (TQ3268) Buckingham Palace Garden [Middlesex] Tulse Hill [Middlesex] (TQ3273) (TQ2879) formal garden with wooded fringes Uxbridge [Middlesex] (TQ0684) and lake Wimbledon [Surrey] (TQ2571) Bushy Park [Middlesex] (TQ1469) Wood Green [Middlesex] (TQ3191) Charing Cross [Middlesex] (TQ3181) GREATER MANCHESTER Chingford [Essex] (TQ3994) Denton (SJ9295) Chislehurst, Pond Wood [Kent] (TQ4369) broad- Manchester, Fallowfield (SJ8694) leaved woodland with pond Trafford, Sale (SJ7992) Clapham [Surrey] (TQ2975) Warburton (SJ7090) Coppets Wood, N12 Whitefield (SD8006) Coulsdon [Surrey] (TQ3158) Withington [Cheshire] (SJ8593) Cripplegate [Middlesex] (TQ3281) HAMPSHIRE [= Hants.] Crofton Heath [Kent] (TQ4366) broad-leaved Abbot’s Wood (SU8140) broad-leaved woodland woodland Alice Holt Forest (SU7942-43) broad-leaved wood- Cudham [Kent] (TQ4560) land and conifer plantations Downe, Cuckoo Wood [Kent] (TQ4462) calcareous Ampfield (SU4024) broad-leaved woodland Basingstoke (SU6351) East Ham [Essex] (TQ4383) Bishop’s Waltham, The Moors (SU5617) wet carr East Ham, St. Mary’s Nature Reserve [Essex] woodland (TQ4383) Bolderwood (SU2409) Enfield Wash Bournemouth [formerly Dorset] (SZ1091) garden Hampstead Heath [Middlesex] (TQ2786) Bransgore (SZ1998) Harlesden [Middlesex] (TQ2283) Chilbolton Common (SU3839) water meadows Harrow [Middlesex] (TQ1488) East Tisted (SU7133) Highgate Wood [Middlesex] (TQ2888) broad-leaved Farley Down(s) (SU3929) calcareous grassland woodland Fleet Pond (SU8254) pond with wooded margins Kew Gardens [Surrey] (TQ1876) botanical gardens and marsh Leaves Green [Kent] (TQ4162) Greywell Fen (SU7251) calcareous valley, alder carr London (TQ3081) woodland and sedge beds by River Whitewater Merton [Surrey] (TQ2569) Hayling Island (SU7201) Mitcham Common [Surrey] (TQ2968) broad-leaved Headley Common (SU5263) woodland Hengistbury Head (SZ1891) New Barnet (TQ2696) Hook Common (SU7254) New Malden [Surrey] (TQ2168) King’s Somborne (SU3631) Northwood [Middlesex] (TQ0991) Leckford Estate (SU36–37/36–38) carr woodland Norwood [Surrey] (TQ1378) and water meadows by River Test Old Coulsdon [Surrey] (TQ3157) Lyndhurst (SU2908) Pett’s Wood [Kent] (TQ4569) house and garden of P. New Forest [= N. Forest] (SU20-SU30) broad-leaved Freeman woodland Pinner [Middlesex] (TQ1290) New Forest, Bank (SU2908) Putney [Middlesex] (TQ2475) New Forest, Beaulieu and Beaulieu River (SU3903) Richmond Park [Surrey] (TQ2072) ancient park- New Forest, Brockenhurst (SU2903-3003) broad- land and broad-leaved woodland leaved woodland by stream Scadbury Park [Kent] (TQ4570) broad-leaved New Forest, Brockenhurst Wood (SU3100) broad- woodland leaved woodland Scratch Wood [Middlesex] (TQ2094) broad-leaved New Forest, Burley (SU2103) woodland New Forest, Cadnam (SU2915) broad-leaved Selsdon [Surrey] (TQ3562) woodland South Kensington, Natural History Museum [Mid- New Forest, Denny Wood (SU3306) ancient broad- dlesex] (TQ2679) wildlife garden leaved woodland Sparrow Wood [Kent] (TQ4366) broad-leaved New Forest, Eyeworth Wood (SU2215) ancient woodland broad-leaved woodland

New Forest, Fritham (SU2414) Lower Lye, Sned Wood (SO4167) conifer plantation New Forest, Great Huntly Bank (SU2705–06) with broad-leaved fringe, river side ancient broad-leaved woodland Moccas Park (SO3342) broad-leaved woodland and New Forest, Hatchett’s Pond (SU3601) sparse mar- parkland ginal vegetation around pond Olchon Valley (SO3131) New Forest, Ipley Manor (SU3706) Paradize Wood (SX6365) broad-leaved woodland New Forest, Linwood (SU1809) broad-leaved Stoke Wood (SO6040) broad-leaved woodland woodland Symond’s Yat (SO5616) broad-leaved woodland New Forest, Lyndhurst Road (SU20) Tarrington (SO6140) New Forest, Mark Ash Wood (SU2407) ancient The Flits (SO3741) wet pasture and hedges broad-leaved woodland Westhide (Wood) (SO5844) New Forest, Matley Bog (SU3307) bog within broad- West Wood, Bagpiper’s Tump (SO5736) mixed leaved woodland woodland New Forest, Oakley Inclosure (SU5751) broad- HERTFORDSHIRE [= Herts.] leaved woodland Aldbury Common (SP9711) broad-leaved woodland New Forest, Puttles Bridge (SU2702) broad-leaved Ashridge Estate (SU9912) broad-leaved woodland woodland by stream Ayot Green New Forest, Rhinefields (Inclosure) (SU2504) Baldock (TL2535) broad-leaved woodland Bayford (TL3109) New Forest, Sowley Pond (SZ3897) Bishop’s Stortford (TL4820) garden of C. W. Plant New Forest, Wood Crates (SU2508-2608) ancient Bramfield (TL2915) broad-leaved woodland Bushey Heath (TQ1594) Nightingale Wood (SU3818) Cheshunt (TL3602) Oakshott (SU7428) Clothall (TL2832) Odiham Common (SU7752) broad-leaved woodland Felden (TL0506) Ovington (SU5631) Frithsden Beeches (TL0109) ancient pollarded Pamber Forest (SU6161) broad-leaved woodland beech woodland with stream and marsh Great Offley (TL1426) broad-leaved woodland Pennington (SZ3295) Harpenden (TL1314) Ports Down (SU6406) calcareous grassland Hatfield Forest (TL2308) broad-leaved woodland Ridley Wood (SU2006) broad-leaved woodland Hertford, Bayfordbury Estate (TL3100) meadow Rotherfield Park (SU6931) broad-leaved hedge woodland Hitchin (TL1829) Rotherfield Park, Stonybrow Wood (SU6830) broad- Hitch Wood (TL1923) broad-leaved woodland leaved woodland Hoddesdon (TL3709) Sandbanks (SZ0588) King’s Walden (TL1524) Selborne, The Lythes (SU7434) broad-leaved wood- Knebworth (TL2520) land by stream Lea Valley (TL1616) Selborne Common and Hanger (SU7333) beech Letchworth (TL2233) woodland on dry slope Offley Holes (TL1729) Silchester Common (SU6262) partly wooded heath- Oughton Head (TL1730) land and bog Radwell (TL2336) Sladen (SU4255) Rickmansworth (TQ0694) Swaythling (SU4315) Rothamsted (TL1314) grounds of horticultural Totton (SU4416) research station Waggoner’s Wells (SU8534) broad-leaved woodland Rothamsted, Knott Wood (TL1314) with ponds and stream Royston (TL3641) Weston Common (SU6944) Symondshyde [as Simonshyde] (TL1910) West Walk (SU6013) Tewin (TL2715) Whitmoor Vale (SU8536) broad-leaved woodland Therfield Heath (TL2339) Winnall Moor (SU4830) rough pasture with ditches Todd’s Green (TL2226) HEREFORDSHIRE Tring (SP9312) Coldborough Park (SO6429) Wallington (TL2934) Credenhill Park Wood (SO4645) Watford (TL1197) Fishpool Valley (SO4566) Welwyn (TL2316) Haugh Wood (SO5935) broad-leaved woodland Welwyn, Sherrard’s Wood (TL2316)

Westbury Wood (TL1526) LEICESTERSHIRE [= Leics.] Weston (TL2630) Newtown Linford, Lea Meadows (SK5011) wet ISLE OF MAN meadow with mixed woodland with alder, birch, Peel (SC2584) hazel, spruce and fir ISLE OF WIGHT Ravenstone, Kelham Bridge (SK4114) wet meadow Apes Down (SZ4587) calcareous grassland with reed beds around pools Bembridge (SZ6588) Stockerston, Bolt Wood (SP8497) ancient broad- Blackgang (SZ4975) coastal landslip leaved woodland with ash, pedunculate oak, Brighstone Forest (SZ4284) conifer plantation hazel and field maple Corfheath Firs (SZ4490) mixed woodland Whetstone, Narborough Bog (SP5497) wet meadow Rowridge Valley (SZ4586) broad-leaved woodland with reed bed on peat and broad-leaved woodland and calcareous grassland with pedunculate oak, ash, elder and willow Walter’s Copse (SZ4390) broad-leaved woodland LINCOLNSHIRE [= Lincs.] KENT Colsterworth (SK9224) broad-leaved woodland Ashford (TR0549) mushroom farm Kirton (TF3038) Barming, Oaken Wood (TQ7155) broad-leaved Nettleton, Long Wood (TA1000) woodland Stamford (TF0308) Bearsted (TQ8056) Welton Wood (TF0179) broad-leaved woodland Bedgebury Pinetum (TQ7233) arboretum of conifers MERSEYSIDE Blean Wood (TQ0860) broad-leaved woodland Freshfield (SD2908) Chattenden Wood (TQ7571) broad-leaved woodland Liverpool (SJ3591) Cobham (TQ6768) NORFOLK Darenth (Wood) (TQ5771) broad-leaved woodland Barton Broad (TG3622) Dartford (TQ5374) Boston Common Dover (TR3242) Boughton Fen (TF7002) Dunton Green (TQ5258) Brancaster (TF7644) sedge bed Eynsford (TQ5465) Bridgeham, Thetford Chase (TL8884) Farningham Wood (TQ5368) broad-leaved Bure Marshes (TG3617) carr woodland and fen woodland Catfield (TG36-37/20–21) carr woodland and fen Folkestone (TR2267) Costessey (TG1711) Hothfield Bog (TQ9645) bog with wooded fringes Dickleburgh (TM1782) former garden of P. Withers, Knole Park (TQ5453) parkland and woodland with reclaimed agricultural land ancient beech East Harling (TM0087) glacial pool Mereworth Wood (TQ6455) broad-leaved woodland East Walton Common (TF7416) broad-leaved Orlestone Wood (TQ0035) mixed woodland woodland Sevenoaks (TQ5355) Felbrigg [Great] Wood (TG1946) broad-leaved Sevenoaks, One Tree Hill (TQ5653) broad-leaved woodland woodland Felthorpe Wood (TG1416) broad-leaved woodland Shoreham, Dunstall Wood (TQ5361) broad-leaved Foulden (TL7700) broad-leaved woodland woodland Fowl Mere (TL8789) Stodmarsh (TR2160) Foxley Wood (TG0623) Tunbridge Wells (TQ5839) Hickling [Hickling Reed Swamp; Hickling Stubb West Malling (TQ6857) Mill; Hickling Broad] (TG4120/4321) sedge bed, Westerham, Hosey Common (TQ4553) heathland alder carr and mixed woodland Holt Country Park (TG0939) Wye (TR0647) Honingham (TG1011) Wye Downs (TR0847) calcareous grassland Honingham, Warren Wood (TG1111) LANCASHIRE [= Lancs.] Honingham Fen Colne (SD8840) Horning Ferry (TG3417) Lower Brock Mill (SD5443) broad-leaved woodland How Hill (TG3719) damp broad-leaved woodland, Kirkham, Fairfield Experimental Horticulture peat hollows and fen Station (SD4332) Hunstanton Martin Mere (SD4214) lake and wildfowl reserve Lenwade Pits (TG1018) Preston (SD4432) mushroom farm Lolly Moor (TF9910) fen Winmarleigh Moss (SD4447) partly wooded Lopham Fen (TM0480) wetland in alder carr heathland Middle Harling (TL9986) wetland fen

Mills Marsh (TG3615) wet meadow Cotterdale (SD8492) riverbank with alders Narford Hall (TF7613) Coverham (SE1086) Norwich (TM4497, TG2108) mushroom farm Cowbeck, birch carr Norwich, The Loke (TG2309) malaise trap in former Crag Wood (SD9656) garden of B. R. Laurence Derwent Ings, water meadows and hedges Old Buckenham (Fen) (TM0491-92) open fen with Drifﬁeld,The Keld (TA0157) broad-leaved woodland wooded fringes Duncombe Park (SE6082) broad-leaved woodland Pashford (TL7383) poor fen Easington Beck (NZ7516) broad-leaved woodland Pulham St. Mary (TM2089) broad-leaved woodland Forge Valley (SE9887) ancient broad-leaved wood- Quidenham (TM0388) land with seepages Reedham (TG4202) sedge bed Fyningdale Moor (SE8698) Roydon Common (TF6822) broad-leaved woodland, Goathland (NZ8402) marsh Gunnerside (SD9598-9199) broad-leaved woodland Scarning (TF9512) wetland fen Harrogate, Birk Crag Wood (SE2955) Sea Mere (TG0300-01) damp meadow Hawnby, Low Pasture (SE5389) Snetterton (TL9990) decaying vegetation by race Hawnby, River Rye (SE5388) broad-leaved track woodland Snettisham Beach (TF6934) sea shore Hole of Horcum (SE8594) Southrepps Common (TG2635) Phragmites bed and Ingleborough (SD7475) marsh Ingleton (SD6973) Stanford battle area (TL8695) Janet’s Foss Wood (SD9163) Strumpshaw (TG3406) damp meadow Jeffry Bog (SE7576) Surlingham, Wheatfen (TG3306-3205) carr wood- Langton Hall (SE3095) broad-leaved woodland land and fen Malham Tarn (SD8966) broad-leaved woodland, bog Sutton Broad (TG3723) carr woodland and fen and lakeshore Thetford Forest (TL8884) Marrick, Colt Park Wood (SE0899) Thompson Common (TL9395-96) broad-leaved Monk Fryston (SE5129) broad-leaved woodland by woodland and grassland with pingo pools streams Thurgarton (TG1835) Mulgrave Wood (NZ8411) broad-leaved woodland Upton Broad and Upton Fen (TG3813) broad-leaved North Grimston (SE8568) woodland and fen Park End (NZ7009) Wayland Wood (TL9299) broad-leaved woodland Pen-y-Ghent (SD8474) Welbourne (TG0610) Raindale (SE8092) West Harling (TL98) water meadow by River Thet Rake Beck (NZ0805) wooded stream gorge West Harling Common (TL9585) conifer plantation Ravensworth (NZ1408) with streams and water meadow Rillington (SE8675) Weybourne Dunes (TG1143) Rosedale Moor (SE7299) Woodbastwick (TG3316-3416) carr woodland and Selby (SE6030) mushroom farm fen Slapston Holm Wood (SE0198) broad-leaved NORTHAMPTONSHIRE [= Northants.] woodland Buckingham Thick Copse (SP7043) broad-leaved Tup Hag Wood (SE5867) broad-leaved woodland woodland NOTTINGHAMSHIRE [= Notts.] NORTHUMBERLAND Bulwell Hall Park (SK5445) Berwick-upon-Tweed [= Berwick] (NT0053) garden Nottingham (SK5742) of B. R. Laurence Radcliffe on Trent (SK6539) Newburn (NT9347) Sherwood Forest (SK66) broad-leaved woodland Nine Nicks of Thirlwall (NY6667) with oaks NORTH YORKSHIRE [= N. Yorkshire] Strelley (SK5041) Ashberry Pastures (SE5684) broad-leaved wood- OXFORDSHIRE [= Oxon] land and meadows Aston Rowant (SU7296-99) calcareous grassland Austwick (Moss) (SD7768) raised mire Bagley Wood [Berkshire] (SP5102) broad-leaved Aysgarth Force (SE0089) waterfall with mossy woodland rocks Bayswater (SP5408) Bishop Wood (SE5532-5633) Bix Bottom (SU7288) broad-leaved woodland and Bolton Wood calcareous grassland Clapham (SD7569) Brasenose (SP5605)

Charlbury (SP3519) Snailbeach (SJ3803) Cothill [Berkshire] (SU4599) carr woodland and fen Trefonen Marsh (SJ2426) shaded stream in pasture Ditchley (SP4021) grazed by cattle Frilford Heath [Berkshire] (SU4498) Whale Moss Goring Heath (SU6579) SOMERSET Goring-on-Thames (SU6081) village garden, for- Burrington Combe (ST4758) stream in rocky gully merly of J. H. Cole Castle Neroche (ST2715) Goring-on-Thames, Coneyberry Wood (SU6181) Cogley Wood (ST7035) broad-leaved woodland beech woodland on chalk Dunster (SS9943) broad-leaved woodland Hartslock (SU6278) calcareous grassland adjacent East Harptree Combe (ST5554) broad-leaved wood- to hanger woodland land by stream Headington Hill and Wick (SP5306) Ebbor Gorge and Ebbor Rocks (ST5248) broad- Hill End [Berkshire] (SP4606) broad-leaved wood- leaved woodland land and wooded stream Edford Wood (ST6748) broad-leaved woodland Hogley (Bog) [= Lye Valley] (ST6564) Great Elm (ST7449) Little Wittenham Wood [Berkshire] (SU5692) Ivy Thorn Hill (ST4734) calcareous broad-leaved broad-leaved woodland woodland and grassland Lye Hill Loxley Wood (ST4037) broad-leaved woodland Marston (SP5208) reared from nests Pawlett Levels (ST3143) grazing marshes Nettlebed (SU7185) Priddy (ST5251) conifer plantation Oxey Mead (SP4810) water meadows and bank of Shapwick Heath (ST4340) grassland and scrub on River Thames with trees peat Oxford, Milham Ford School (SP5307) school Vallis Vale (ST7549) broad-leaved woodland grounds with old poplars West Town (ST5335) Oxford, Oxford University Museum and Parks Yeovil (ST5617) (SP5106) SOUTH YORKSHIRE Sarsgrove Wood (SP3024) broad-leaved woodland Barnsley (SE3506) Shotover (Country Park) (SP5606) broad-leaved Deffer Wood (SE2709) woodland Doncaster (SE5803) Spartum Fen (SP6501) carr woodland Doncaster, Hatfield Moor (SE7006) Stokenchurch (SU7696) Doncaster, Pot Riding Wood (SE5200) Stonor (SU78) Doncaster, Thorne Moors (SE7316) Tubney [Berkshire] (SU4398-4400) broad-leaved Howell Wood (SE4410) woodland Sheffield (SK3688) Waterperry Wood (SP6009) woodland including Sheffield, Ewden Valley (SK2597) conifer plantations STAFFORDSHIRE [= Staffs.] Weston Green Fen (SP5219) partly wooded fen Alvecote Pools (SK2504) alder carr and wetlands Weston-on-the-Green (SP5318) formed by mining subsidence Whitecross Green Wood (SP6614) broad-leaved Aqualate Mere, Decoy Wood (SJ7720) broad-leaved woodland woodland Wormsley [Park] (SU7394) Burton [Shropshire] (SJ4425) mushroom farm Wychwood [Forest] (SP3316) ancient broad-leaved Cannock Chase (SJ9818-20) mixed woodland woodland Colwich Park (SK0121) Wytham Woods [Berkshire] (SP4608) broad-leaved Consall Wood (SJ9849-9947) broad-leaved woodland woodland SHROPSHIRE [= Salop] Cop Mere (SJ8030) broad-leaved woodland with Bromfield, River Teme gulley Church Stretton (SO4694) Crowgutter Wood (SK0350) broad-leaved woodland Duxmoor Dingle (SO4477) Gibriddings Wood (SK0344) broad-leaved woodland Holly Bank (SO5695) calcareous broad-leaved Grindon Wood (SK0944) broad-leaved woodland woodland Hatherton (SJ9510) Leebotwood (SO4798) stream side with willow, Loynton Moss (SJ7825) alder and oak adjacent to oat field Lum Pool (SK0660) Llanmynach Hills (SJ2621) grassland and scrub in Manifold Valley (SK0952) broad-leaved woodland old limestone quarry SUFFOLK Oswestry (SJ2929) Barton Mills (TL7173)

Belstead House (TM3651) Chobham Common [Long Arm carr; Gracious Pond] Benacre Broad (TM5282) (SU9666, SU9863, respectively) Bradfield Wood (TL9357) broad-leaved woodland Cosford Mill (SU9138) carr woodland Butley Thicks (TM3651) broad-leaved woodland Devil’s Punchbowl (SU8936) broad-leaved woodland East Bergholt (TM0735) by stream Felixstowe (TM2738) mushroom farm Esher, West End Common (TQ1262-63) broad- Flatford Mill (TM0733) leaved woodland Hoxne (TM1877) river margin Felbridge (TQ3639) Ipswich (TM0838) mushroom farm Gomshall, Hackhurst Farm (TQ0947) Lakenheath [= Laken Heath] (TL7182) damp Hackhurst Down (TQ0948) meadow Hackhurst Wood (TQ1047) broad-leaved Kessingland (TM5286) woodland Market Weston Fen (TL9878) fen with sedge Horsell Common (TQ0060) heathland and pine pools woodland Mildenhall (TL7174) Horsley (TQ0954) Minsmere (TM4666) mixed damp woodland Limpsfield Common (TQ4152) Monk’s Soham (TM2165) Mayford (SU9956) reared from nests Occold (TM1570) decaying vegetation Nutcombe Down (SU8834) broad-leaved woodland Orford (TM4249) Oxshott (Heath) (TQ1361) heathland and pine RAF (Royal Air Force base) Barnham (TL8779) woodland heath grassland and mixed woodland with Puttenham (SU9347) stream Ranmore Common (TQ1451) broad-leaved Redgrave Lake (TM0576) lake margin woodland Reydon Wood (TM4778) damp broad-leaved Ripley (TQ0556) woodland Runnymede Meadow (TQ0072) water meadows and Santon Downham (TL8187) ponds Scotland Fen (TM3547) poplar plantation and fen, Send (TQ0255) oak woodland Staffhurst Wood (TQ4148) broad-leaved woodland Shrubland Park (TM1252) Thundry Meadow (SU89-90/44) water meadows and Snape Maltings (TM3957) carr woodland Stallode Wash (TL6884) marsh land Thursley Common (SU9041) heathland, bog and Staverton Thicks (TM3650) ancient broad-leaved mixed woodland woodland West Horsley, Sheep Leas (TQ0851) calcareous Thelnetham Fen (TM0178) fen grassland and woodland Thornham Magna (TM1071) damp broad-leaved White Downs (TQ1249) woodland Wisley (TQ0659) Tuddenham Fen (TL7472) wet heathland and birch Wisley Common (TQ0758) heathland and mixed woodland woodland Tunstall Common (TM3754) Witley Common (SU9240) Walberswick (TM4673-74) saltmarsh and reed beds Woking (TQ0058) Wangford (TL7584) Breckland heath Wotton (TQ1247) Wortham Ling (TM0879) heath Virginia Water (SU9968) SURREY TYNE & WEAR Albury, Blackheath (TQ0346) Derwent Valley, Lockhaugh Bank Wood Ashtead (Common) (TQ1759) heath [= Lockhaugh Wood] (NZ1759) broad-leaved Blackheath, Albury (TQ0346) woodland Bookham Common (TQ1256) broad-leaved WARWICKSHIRE [= Warks.] woodland Bishops Tachbrook, Oakley Wood (SP3059) ancient Boundless Copse (SU8936-9036) broad-leaved and woodland, partly coniferous mixed woodland Brandon, New Close Wood (SP4076) broad-leaved Box Hill (TQ1852) broad-leaved woodland woodland with pedunculate oak, birch and hazel Byfleet (TQ0661) Clowes Wood (SP1074) broad-leaved woodland Charterhouse Alder Holt (SU9544) carr woodland Coventry, Coombe Abbey Country Park [= Coombe by River Wey Abbey Park; West Midlands] (SP3979) broad- Chelsham (TQ3758) leaved woodland with ash, pedunculate oak, Chipstead (TQ2858) mountain ash and hornbeam

Coventry, Warwick University Campus, Tocil Wood Cuckfield (TQ3024) [West Midlands] (SP3075) broad-leaved wood- East Grinstead (TQ3038) land with pedunculate oak, mountain ash, maple Henfield (TQ2015) flood meadow ditch by River and hazel Adur Edge Hill (SO3748) deciduous woodland on steeply Inholm’s Copse (SU8526) broad-leaved woodland sloping sandstone Iping Common (SU8421) Farnborough Park (SP4349) wooded lake margin Littlehampton (TQ0403) horticultural research sta- Kingsbury Water Park (SP2096) parkland, wooded tion and mushroom farm lake margin, reed beds Parham (TQ0613) birch woodland Leamington Spa (SP3266) in house plant compost Pulborough (TQ1117) mushroom farm Leamington Spa, Newbold Comyn Park (SP3365) Pulborough, Chesswood Nurseries (TQ0418) damp meadow, young mixed woodland Rewell Wood (SU9808) broad-leaved woodland Leamington Spa, Whitnash brook, Sydenham Rogate (SU8023) meadow (SP3364) stream-side with alder, willow, black- Three Bridges (TQ2837) thorn, elder and hawthorn Tilgate Forest Leamington Spa, Willes Meadow [in part under Worthing (TQ1403) Newbold Comyn Park] (SP3266) meadow flooded Worthing, Clapham (TQ0906) in winter WEST YORKSHIRE Moreton-in-Marsh, Long Spinney (SP3135) wet Bradford (SE1633) woodland on limestone Leeds (SE23) Moreton-in-Marsh, Whichford Wood (SP3034) Leeds, Farnley (SE2147) ancient and plantation woodland Wakefield (SE2817) mushroom farm Moreton-in-Marsh, Wolford Wood (SP2433) ancient Wakefield, Wentbridge (SE4817) deciduous woodland, wet in part WILTSHIRE Oversley Wood (SP1056) broad-leaved woodland Lackham Park (ST91–92/69–70) broad-leaved Studley, Rough Hill Wood (SP0564) ancient wood- woodland land, mainly oak, ground cover of wood anemones Littlecote Park (SU2969) and bluebells Melksham (ST8962) in house of P. J. Chandler Ufton Bridge (SP3762) disused railway cutting Odstock (SU1324) Ufton Fields (SP3761) broad-leaved woodland, pools Salisbury Plain Training Area East (ST94–95) and grassland unimproved calcareous grassland Wellesbourne (SP2757) mushroom farm Savernake Forest (SU20–22/65–67) ancient beech Wellesbourne, Horticulture Research International woodland [= HRI] (SP2756) laboratory culture WORCESTERSHIRE Whitacre Heath (SP2193) wetland, willow carr, Bell Heath (SO97) birch, reed beds, old sand and gravel workings Evesham (SP0645) mushroom farm WEST MIDLANDS Hunthouse Woods (SO77) broad-leaved woodland Birmingham [Warwickshire] (SP08) Redditch, Lion Wood (SP0467) oak woodland on Birmingham, Edgbaston [Warwickshire] (SP0683) acid soil with heather and bilberry Birmingham, Sutton Park [Warwickshire] Shrawley Wood (SO8065) broad-leaved woodland (SP0998) mixed woodland, wetlands, heath and Wyre Forest (SO73–76/73–77) ancient broad-leaved parkland woodland Coventry [Warwickshire] (SP37) Coventry, Bubbenhall, Ryton Wood (SP3872) ancient woodland with pedunculate oak, lime, NORTHERN IRELAND hazel and birch ANTRIM Coventry, Ryton-on-Dunsmore, Brandon Marsh Glenariff (D2120) broad-leaved woodland in gorge (SP3875) willow carr, birch, ash and wetland, Lennymore Bay (J1175) former gravel workings CITY OF BELFAST Solihull, Baddesley Clinton, Hay Wood (SP2171) Belfast (J36) riverside with trees mixed woodland, mainly coniferous, fir, cypress, DOWN pedunculate oak, birch Crawfordsburn Country Park (J4682) mixed Sutton Coldfield [Warwickshire] (SP1296) woodland WEST SUSSEX Holywood (J37-47) Ambersham Common (SU9019-9120) heathland Newcastle (J33) and mixed woodland Rostrevor (J1817) broad-leaved woodland

TYRONE Slade of Saggart (O0324) mixed woodland Moy (H8356) broad-leaved woodland Terenure (O1230) garden of P. Ashe GALWAY Newtown Lough (M4202) turlough REPUBLIC OF IRELAND KERRY CAVAN Castlegregory, Lough Gill (Q6014) sand dunes Bellananagh (H4000) lake-side Derrycunihy Cascade (V9280) broad-leaved wood- Virginia Woods (N5987) mixed woodland land by waterfall CLARE Drominahassig (V9566) broad-leaved woodland Ballynalacken Castle (M1000) small area of broad- Dromore Forest (V8267) broad-leaved woodland leaved woodland by castle Kenmare (V97) Burren, Ailladie [= Aill an Daill] (M0802) Killarney (V9586) mossy rocks by waterfall Burren, Caher River (M1509) Killarney, Castlerosse (V99) Burren, Fanore (M1307) green road Killarney, Galway’s Bridge (V9180) Burren, Formoyle (M1705) grikes Killarney, Kenmare Estate (V9490) Corker Pass (M3010) Killarney, Long Range Ennis (R2979) mixed broad-leaved woodland, Killarney, River Flesk mainly hazel Killarney, Ross Island (V9489, V9588) wooded pen- Lisdoonvarna [= Lisdoonvarna Spa] (R1397) insula in lake, damp broad-leaved woodland Lough Bunny (R3696) Killarney, Tomies Wood (V9188) broad-leaved Lough Luogh (R0593) woodland CORK Killarney, Torc Cascade [= Falls] (V9684) broad- Ballyvourney (W1977) damp broad-leaved leaved woodland by waterfall woodland Killarney, Upper Lake (V9282) wooded lake shore Castlefreke (W3335) saline-inﬂuenced woodland Killaun Bog (N1106) raised bog Clonpriest (X0573) saltmarsh Muckross Lake and House (V98) wooded lakeshore Coom Wood (W1273) wooded gully and grounds Dunamarc Falls (W0153) broad-leaved woodland by Muckross, Monk’s Wood (V9786) old beech waterfall woodland Gearagh (W2969) mature alluvial oak and ash Uragh Wood (V8363) oak woodland woodland with elm, willow and birch on braided KILDARE stream over limestone Ardscull Mote (S7397) broad-leaved woodland on Gearagh, Annahala bog (W3269) wet grassland and old castle mound fen subject to ﬂooding with streams, willow carr, Carton (N9637) mixed woodland Juncus, Typha and Osmunda over limestone Maynooth (S93) mushroom farm Glengarriff (State) Forest (V9057) oak woodland Newbridge Fen [= Pollardstown Fen] (N7615) Cla- Leap (W23) dium beds with tufa, open fen Leap, Myross Wood (W23) River Liffey, Victoria Bridge (N8419) riverbank Rahan (W6497) Rye Water (O0036) Riverstick, alder carr Straffan (N9326) Warren’s Court Wood, damp broad-leaved woodland KILKENNY Whites Marsh (W3738) salt marsh Ballielogue (S6731) DONEGAL Ballykeeffe Wood (S2151) broad-leaved woodland Inishowen Castlecomer (S5475) DUBLIN LAOIS Bohernabreena (O0922) vegetation beside reservoir Emo Park (N5307) mixed woodland Clondalkin (O0631) malaise trap in garden of J. P. LEITRIM O’Connor Dromahaire (G8032) mixed woodland Dublin City (O1530) LONGFORD Dublin, Kinsealy Research Centre (02142) horticul- Lough Derragh (N3979) tural research station MAYO Dublin, Phoenix Park (O19) parkland with old trees Brackloon Wood (M58) Howth (O23) broad-leaved woodland Castlebar Lough (M18) Malahide Castle (O2243) mixed woodland Lough Conn (G1606) broad-leaved woodland and River Liffey, Strawberry Beds (O9586) wooded lakeshore riverbank Rosserk Abbey (G2526) stream margin with trees

MONAGHAN SCOTLAND Carrickmacross, Nuremore Lake (H80) lakeside marsh and scrub ABERDEENSHIRE (formerly in Grampian Region) OFFALY Balmoral Forest (NO2387) conifer plantations Charleville Wood (N3222-3223) ancient broad- Ben Macdhui [= Ben Macdui] (NN99) high montane leaved woodland by lake habitat Mongan Bog (N0330) roadside ditch Braemar (NO19) ROSCOMMON Claybokie (NO0889) pine woodland Castlerea (M6780) broad-leaved woodland Den of Pitlurg Lough Key (G8403) mixed woodland Glen Gairn (NO3399) SLIGO Glen Lui (NO0690) Beltra Lough (G6031) wooded lakeshore Glen Quoich, Dubh Ghleann(NO0795) ancient pine Glencar Waterfall (G7643) woodland by woodland waterfall Glentanar (NO49) pine woodland Glen of Knocknarea (G6333) wooded gorge Linn of Dee (NO0689) pine woodland Templehouse Demesne (G6317) mixed woodland Loch Callater (NO1883) TIPPERARY Loch Davan (NJ4400) Ballina (R7172) Luibeg (NO0393) WATERFORD Mid Quoich (NO0992) ancient pine woodland Mahon Falls (S3009) Morrone Birkwood (NO1390-1491) birch woodland Passage East (S6811) hillside vegetation with calcareous seepages WESTMEATH Invercauld Bridge, Old Bridge of Dee (NO1892) Ballynafid Lake [= Lough] (N4161) fen wooded riverbank Glen Lough (N2866) wet meadows Upper Quoich (NO0995) ancient pine woodland Lough Slevin (N4556) mixed woodland ANGUS (formerly in Tayside) Scragh Bog (N4259) quaking bog Abernyte [Perthshire] (NO2631) stream along WEXFORD meadow hedge Ballyhighland (S8840) Braedownie (NO2875) Ballyteige (S9504) sand dunes Den of Alyth (NO2950) broad-leaved woodland by J.F. K ennedy Park (S7319) stream Lady’s Island Lake (T1007) Glen Doll (NO2576) Killoughrim (S9041) oak woodland ARGYLL & BUTE (formerly in Strathclyde) Oaklands (S7125) Arrochar [Dunbartonshire] (NN3004) Tintern (S7810) Barcaldine Forest [Argyllshire] (NM9742) WICKLOW Glasdrum [Argyllshire] (NM9945–NN0046) broad- Avoca Wood (T2079) broad-leaved woodland leaved woodland Blackditch Wood Glen Croe [Argyllshire] (NN2404) Blessington, shore of Lacken Reservoir (N8114) Lephinmore [Argyllshire] (NR99) open lake shore Port Appin [Argyllshire] (NM9045) Carrigower (O2508) broad-leaved woodland CITY OF EDINBURGH Derrybawn (T1495) broad-leaved woodland Blackford Hill (NT2570) Devil’s Glen (T29) mixed woodland in gorge Edinburgh (NT25) Glen of the Downs (O2611) broad-leaved CITY OF GLASGOW woodland Glasgow (NS6576) Milton of Campsie, mushroom Glendalough (T1196-1396) broad-leaved and mixed farm woodland by lakes DUMFRIES & GALLOWAY Glendalough State Forest (T1495) broad-leaved Caerlaverock [Dumfriesshire] (NY06) woodland by Avoca River Castlehill (NX8188) Knocksink Wood (O2117) broad-leaved woodland Castle Loch [Dumfriesshire] (NY0782–0881) broad- Murrough (including Black Ditch Wood O3103) leaved woodland saline-influenced woodland and dykes Corsemalzie [Kirkcudbright] (NX45) Powerscourt (O2116) parkland with old trees Newlaw Moss [Kirkcudbright] (NX7448) Sally Gap (O1311) montane habitat Stronord [Kirkcudbright] (NX4564) woodland in Stepaside (O1924) garden grounds of field centre The Quill (O2419) damp broad-leaved woodland Torrs [Wigtownshire] (NX5289) Wicklow town (T39) garden Tynron [Dumfriesshire] (NX89)

Wamphray Glen [Dumfriesshire] (NY1296) broad- Glen Coiltie [Inverness-shire] (NH4927) wet broad- leaved woodland leaved woodland EAST AYRSHIRE (formerly in Strathclyde) Glen Einich [Inverness-shire] Craufurdland Water [Ayrshire] (NS4641) Glen Glass [Ross & Cromarty] (NH5568) EAST LOTHIAN Glen More [Inverness-shire] (NH9809) ancient pine Gullane Bents (NT4784) dunes with sea buckthorn woodland scrub Glen Shiel [Ross & Cromarty] (NH0310) Humbie (NT4563) lush vegetation in wet meadow Glencoe [Argyllshire] (NN0958) alder woodland North Berwick (NT5585) mushroom farm Granish [Inverness-shire] (NH8914) birch FALKIRK woodland Limerigg (NS8570) wet heath Grantown-on-Spey [Morayshire] (NJ0326) pine FIFE woodland Cupar (NO3714) Insh Marshes [Inverness-shire] (NH7901) marshes HIGHLAND REGION by River Spey Ach na Bourin [Sutherland] (NC7158) Insh Marshes, Invertromie Wood [Inverness-shire] Arcan Mains [Ross & Cromarty] (NH5053) broad- (NH7899) aspen woodland leaved woodland Invermoriston [Inverness-shire] (NH4117) Aviemore [Inverness-shire] (NH8912) bank of Spey, Isle of Eigg, Galmsdale [Inverness-shire] (NM4686) birch woodland Kinrara [Inverness-shire] (NH8708) mixed Aviemore, Craigellachie [Inverness-shire] woodland (NH8812) birch woodland Kinrive Wood [Ross & Cromarty] (NH67) Aviemore, Spey bank [Inverness-shire] (NH9114) Lairig Ghru [Inverness-shire] (NH9503-9603) open Beinn Eighe [Ross & Cromarty] (NG9965) ﬂushes montane habitats in open mixed woodland Ledmore Wood [Sutherland] (NH6689) oak Belladrum Burn [Inverness-shire] (NH5138) woodland Bohuntine [Inverness-shire] (NN2983) Little Kildrummie [Nairn] (NH8653) by River Braelangwell [Ross & Cromarty] (NH6903) ancient Nairn pine woodland Loch Achilty [Ross & Cromarty] (NH4256) mixed Cairngorm [Inverness-shire] (NJ0004 and NH9806) woodland ancient pine woodland and open stream margin Loch Achnacloich [Ross & Cromarty] (NH6673) Cairngorm, Coire Raibeirt [Inverness-shire] Loch an Eilein [Inverness-shire] (NH8908) ancient (NJ0003) open montane habitat pine woodland Castletown [Caithness] (NH7442) Loch Assynt [Sutherland] (NC2024) Cawdor Wood [Nairn] (NH84) broad-leaved Loch Eye [Ross & Cromarty] (NH8480) alder woodland woodland Conon Island [Ross & Cromarty] (NH55) Loch Fleet [Sutherland] (NH8094) heathland and Corrimony [Inverness-shire] (NH3729) mixed scrub woodland above deep gorge Loch Garten [Inverness-shire] (NH91) ancient pine Culbin Forest and Sands [Nairn] (NH9057) dunes woodland and conifer plantations Loch Loy [Nairn] (NH9358) ancient pine Culrossie Wood [Ross & Cromarty] (NH7978) mixed woodland woodland, mainly pine Loch Meiklie [Inverness-shire] (NH4230) alder Dingwall [Ross & Cromarty] (NH55) woodland Drumguish [Inverness-shire] (NN79) Loch Morlich [Inverness-shire] (NH9709) ancient Drumnadrochit [Inverness-shire] (NH52) pine woodland Drumnadrochit, Urquhart Bay [Inverness-shire] Loch na Ba Ruaidhe [Inverness-shire] (NH4933) (NH5229) alder woodland Loch Pityoulish [Inverness-shire] (NH9113) Dulicht [Morayshire] (NJ0228) poplar woodland Loch Shieldaig [Ross & Cromarty] (NG87) broad- Easter Fearn [Ross & Cromarty] (NH6486) birch leaved woodland woodland Loch Ussie [Ross & Cromarty] (NH55) mixed Feshiebridge [= Feshie Bridge; Inverness-shire] woodland (NH80) wooded bank of River Feshie Lochanhully [Inverness-shire] (NH9123) birch Foxhole [Inverness-shire] (NH5240) broad-leaved woodland by stream woodland Migdale Loch and Wood [Sutherland] (NH6590) Glen Affric [Inverness-shire] (NH1923) ancient pine alder and birch carr by stream and pine wood- woodland land on higher ground

Moniack, Reelig Glen [Inverness-shire] (NH5543) Arran, Corriegills [Bute] (NS03) bogs and sea cliff mixed woodland Arran, Sannox [Bute] (NS04) broad-leaved Morvern, Glensanda [Argyllshire] (NM6654) woodland Nairn [Nairn] (NH85) Arran, Sliddery [Bute] (NR92) Nethy Bridge [Inverness-shire] (NJ0121) ORKNEY Pitmaduthy (Moss) [Ross & Cromarty] (NH7676) Eday (HY5531) Rosemarkie [Ross & Cromarty] (NH7358) broad- Kirkwall, Brandyquoi (HY4510) leaved woodland North Ronaldsay [= N. Ronaldsay] (HY7553) near Rothiemurchus [Inverness-shire] (NH9110) ancient shore of island pine woodland Orphir (HY3304) Rothiemurchus, by River Druie [Inverness-shire] Stronsay (HY6525) (NH9107) ancient pine woodland PERTH & KINROSS (formerly in Tayside) Skye, Moineach Mararulin [Inverness-shire] Balnaguard Burn (NN9451) broad-leaved woodland (NG4227) acid bog Beinn a’Chreachain (NN34) open montane habitats Spinningdale and Spinningdale Bog [Sutherland] Beinn Heasgarnich (MM43) open montane habitats (NH6690-6790) bog with stream Ben Lawers (NN6139) open montane habitats Strathglass (NH3836) Birks of Aberfeldy (NN8547) broad-leaved Strathglass, Cannich [Inverness-shire] (NH3331) woodland Strathrory [Ross & Cromarty] (NH67) Birnam Hill (NO0440) broad-leaved woodland Strathy Bog [Sutherland] (NC8365) bog and birch Blair Atholl (NN86) woodland Camghouran (NN5257) birch woodland Tarradale House [Ross & Cromarty] (NH5548) Glen Lyon, Cashlie (NN4841) wooded grounds and saltmarsh Comrie (NN7722) broad-leaved woodland, mainly The Mound (Mound Alderwood) [Sutherland] oak (NH7698) alder woodland Craig na Cailloch Torboll Wood [Sutherland] (NH7498) broad-leaved Dalnaglar Castle (NO1464, NO1465) woodland Dupplin Loch (NO0319) Udale Bay [Ross & Cromarty] (NH7065) alder Fungarth (NO0343) conifer plantations and stream woodland and saltmarsh Glen Eagles, Braco (NN8309) Victoria Falls [Ross & Cromarty] (NG8971) pine Glen Lednock (NN72) woodland Glendevon (NN90) MIDLOTHIAN Grandtully (NN9052) wet broad-leaved woodland Arniston (NT3259) Loch Earn (NN6423) Fanninghill Loch Loch Tummel (NN8159) Newbattle Abbey (NT3366) broad-leaved woodland Lochan an Daimh (NN7157) open marsh by pool, Pentland Hills, Carnkilton (NT1863) stream with alder Toxside Hill (NT2854) Lochan na Lairige (NN5940) MORAY (formerly in Grampian Region) Meall nan Tarmachan (NN5838) Auchindachy [Banffshire] Meikleour, North Wood (NO1540) pine plantation Ballindalloch [Banffshire] (NJ1836) Pass of Killiecrankie (NN9161) broad-leaved wood- Brodie Castle [Morayshire] (NH9757) mixed woodland in gorge land by lake Pitlochry (NO95) Convil Braes [Banffshire] Rannoch (NN4953) Falls of Tarnash [Banffshire] (NJ4449) Rannoch, Black Wood (NN5756) ancient pine Forres [Morayshire] (NJ05) woodland Glen of Drumloch [Banffshire] Rannoch, Carie (NN6257) conifer plantations Loch Park [Banffshire] (NJ3543) Rannoch, Loch Rannoch, north shore (NN5558) Logie [Morayshire] (NJ0050) broad-leaved woodland Tomintoul [Banffshire] (NJ11) Tummel Forest (NN8659) conifer plantations Spey Bay [Morayshire] (NJ3462) Vicars Bridge (NS9898) wooded riverbank NORTH AYRSHIRE (formerly in Strathclyde) Weem Castle Woods (NN8449) mixed woodland on Arran, Brodick [Bute] (NS03) woodland, possibly south-facing slope broad-leaved (in 1919) SCOTTISH BORDERS Arran, Catacol [Bute] (NR94) open glen with stream Ancrum, Ale Water [Roxburghshire] (NT6125) Arran, Correin Lochan [Bute] (NR9046) open lake wooded stream gully margin Gateheugh, by River Tweed

Newtown St. Boswell’s [Roxburghshire] (NT5731) WEST DUNBARTONSHIRE (formerly in broad-leaved woodland Strathclyde) Selkirk [Selkirk] (NT42) Bonhill [Dunbartonshire] (NS38) Yarrow [Roxburghshire] (NT32) WESTERN ISLES SHETLAND Barra (NF60) Bridge of Walls (HU2651) Barra, Craigston (NF6601) Fair Isle (HZ2271) Lewis, Barvas, sand dunes Gutcher Yell (HU5499) Lewis, Loch na Gainmhich (NB22-23) Kergord (HU4055) St. Kilda (NF09) Mid Yell (HU5190) Sand (HU3447) = Tresta (HU6090) WALES [ CYMRU] Unst (HP61) BRIDGEND Vidlin (HU4765) Porthcawl [Glamorgan] (SS8177) Weisdale (HU3852, HU3848) CARDIFF Yell (HU59) Pentyrch [Glamorgan] (ST1082) SOUTH AYRSHIRE (formerly in Strathclyde) CARMARTHEN (formerly in Dyfed) Culzean Castle [Ayrshire] (NS2209) mixed Afon Tywi (SN7538) woodland Beacon Bog (SN3516) basin mire, Molinia bog Failford, by River Ayr [Ayrshire] (NS4626) broad- Castell Cogan (SN3214) leaved woodland Carreg Cennen Castle (SN6619) SOUTH LANARKSHIRE (formerly in Strathclyde) Coed y Gelli (SN6019) broad-leaved woodland Chatelherault Country Park [Lanarkshire] Cors Farlais (SN6734) Juncus (NS7353) mixed woodland by river Dinefwr [= Dynevor] Park (SN6122) broad-leaved Gorge of Avon [Lanarkshire] (NS75) mixed wood- woodland and parkland land by river Ffrwd Fen (SN4102) Phragmites Lockhart Mill [Lanarkshire] (NS8745) grazing Llandeilo (SN6222) meadow and wooded riverbank Llanllwch (SN3618) unburnt raised bog Stonebyres, Walker Farm [Lanarkshire] (NS8544) Penallt (SN5318) STIRLING (formerly in Central Region) Pencarreg (SN5345) Aberfoyle (NN5101) Pont Aber, Sawdde Gorge (SN7322) broad-leaved Aberfoyle, Craigmore Wood [Perthshire] (NN5101) woodland broad-leaved woodland Pontarffinant Wood (SN5020) broad-leaved Balmaha [Dunbartonshire] (NS4291) woodland Beinn nan Eachan [Perthshire] (NN5738) open Rhôs Cruglas (SN6824) Juncus montane habitats Rhôs Pwllygawnen (SN2929-30) Juncus, Molinia Ben Chalum [= Ben Challum; Perthshire] (NN33) bog Callander [Perthshire] (NN6307) Stradey Wood (SN4801) broad-leaved woodland Crianlarich [Perthshire] (NN3825) Ynys Uchaf (SN4915) fen Glen Lochay [Perthshire] (NN5435) broad-leaved CEREDIGION [= Cardiganshire] (formerly in Dyfed) woodland Comin Esgair Maen (SN6564) valley fen Loch Chon [Perthshire] (NN4105) mixed woodland Cors Caron (SN6862/3/4 and SN7166) raised bog, and lakeshore marsh grazed river fen Loch Chon/Loch Ard [Perthshire] (NN4403) mixed Cors Caranod (SN5664) valley fen and wet woodland heathland Loch Katrine [Perthshire] (NN4907) mixed wood- Cors Fochno (SN6392) raised bog land on shore of loch Cors Gorsgoch (SN4850) valley and basin mire Loch Katrine Dam [Perthshire] (NN4906) birch Cors Llyn Farch a Llyn Fanod (SN5963) basin mire woodland Cwm Ystwyth (SN8587) watershed mire Lochan Maoil Dhuinne, Rowardennan Forest Figyn Blaen Brefi (SN5773) Molinia and Trichopho- [Stirlingshire] (NS3697) broad-leaved woodland rum bog with stream Gwaun Garthenor (SN6355) wet flood meadow Meall a’ Churain [Perthshire] (NN43) open mon- Gwaun Ystrad Caron (SN6759) floodplain mire tane habitats Llandre Meall Ghaordie [Perthshire] (NN53) open montane Llynoedd Ieuan (SN8081) blanket mire habitats Neuadd Fawr [= Neuadd-fawr] (SN5438)

Pentwd Marshes (SN1845) river fen with Coed y Rhygen [Merioneth] (SH6837) broad-leaved Glyceria woodland on lake shore Rhôs Cilcennin (SN5261) Cors Barfog [Merioneth] (SN6598) ungrazed basin Rhôs Llawr Cwrt (SN4249/4149) Juncus wet mire meadow, peat Cors Geirch (SH3037, SH3235) Schoenus nigricans, Rhôs Rhydd (SN5773) basin mire, peat Molinia bog Sphagnum and Phragmites, grazed Molinia and Salem (SN6785) valley fen Myrica Ynys Eidiol (SN6795) Phragmites, Molinia bog Cors Graianog (SH4945) Molinia bog, peat cutting, CONWY (formerly in Gwynedd) basin mire, grazed wet heath Bettws [= Betws-y-Coed; Caernarvon] (SH7955) Cors Gyfelog (SH4548, SH4647) Sphagnum, Carex Blaen y Wergloedd [Denbighshire] (SH9163) grazed limosa, Phragmites, floodplain fen grassland and fen Crawcwellt [Merioneth] (SH6930) grazed Molinia Bodnant [Caernarvon] (SH8072) bog, Carex rostrata Bwlch y Ddeufaen [Caernarvon] (SN7171) moor- Cwm Bychan [Merioneth] (SH6434) marginal vege- land flushes tation by upland lake Coed Dolgarrog [Caernarvon] (SH7766) broad- Cwm Crafnant [Caernarvon] (SH7460) leaved woodland Dolgellau [= Dolgelley; Merioneth] (SH7217) Migneint [Caernarvon] (SH 777447) blanket bog Glan y Wern [Merioneth] (SH6035) Nûg [Denbighshire] (SH9350) raised bog Hafod Garregog [under Caernarvon and Merio- Plas Islyn Bog (SH881501) poor fen neth] (SH6044) broad-leaved woodland (Levey DENBIGHSHIRE & Pavett), Molinia bog (Holmes, Boyce & Cefn Geirch, by Afon Alwen (SJ0146) Reed) Gatewen Marsh (SJ 318514) Typha latifolia Hermon Copper Bog (SH7325) thrift flush Gors Maen-llwyd (SH9858) blanket bog remnant Islau dref [Merioneth] (SH61) Hafod Wood (SJ0068) broad-leaved woodland Llanbedr [Merioneth] (SH5826) Llyn Creiniog (SH 926653) humid heath and grazed Llandecwyn [Merioneth] (SH6337) poor fen Llandwrog (SH4556) Llyn Fawnog (SH 852744) Carex rostrata Llyn Gwynant (SH6452) Llyn Tyn y Llyn (SH 846704) Carex rostrata Llyn Llennyrch (SH6537) Merddwr (SH 903498) grazed Eriophorum bog Llyn Pencraig (SH7758) Molinia and Phragmites FLINTSHIRE [formerly in Clwyd] Llyn Tyn y Mynydd (SH6659) wet heath Cefn-y-Bedd (SJ3156) Llyn Ystumllyn (SH5338, SH5539) floodplain fen Ffrith (SJ2855) (Sphagnum, Phragmites) GWYNEDD Pont Croesor [Merioneth] (SH5941) Aber Valley [Caernarvon] (SH6671) broad-leaved Rhôsgyll Fawr (SH4541) Molinia and Sphagnum woodland Trawsfynydd (SH7033) Molinia and Myrica Arthog Bog (SH6314) raised bog Vaynor Wood [Caernarvon] Bangor [Caernarvon] (SH5772) ISLE OF ANGLESEY [= Ynys Mon] (formerly in Brithdir-Isaf [Merioneth] (SH4751) Gwynedd) Bwlch y Dolenfawr (SJ1765) Aberffraw (SH3569) Bwlch y Dwr (SH9929) blanket bog Cors Bodeilio (SH5077) Cladium bed Cadair Idris [= Cader Idris] (SH7113) open mon- Cors Clegyrog (SH3889) acid basin mire tane habitats Cors Erddreiniog (SH4681, SH4783) fen with Cla- Chardeon (SH6417) grazed reed bed dium, wetland, Phragmites Coed Caeddafydd [Caernarvon] (SH6146) broad- Cors Goch (SH4581) Cladium bed leaved woodland Fedw Fawr (SH 602819) wet heath Coed Crafnant [Caernarvon] (SH6229) broad- Fferam Paradwys (SH4270) leaved woodland Gwenfro (SH5079) Glyceria, wet meadow, Coed Dol y Bebin (or Dolbebin) (SH6127) broad- Phragmites leaved woodland Llyn Hafodol (SH3989) floating fen Coed Gorswen [Caernarvon] (SH7571) broad-leaved Newborough Warren (SH4066) conifer woodland plantations Coed Tremadog [Caernarvon] (SH54) broad-leaved Red Wharf (SH5381) woodland Rhôs-y-gad (SH5078) calcareous fen Coed y Brenin [Merioneth] (SH7326) broad-leaved The Dingle (SH4576) woodland Tre Wilmot (SH2281) basin mire

MONMOUTHSHIRE (formerly Gwent) POWYS Black Cliff (ST5398) broad-leaved woodland on Aberithon Turbary (SO0157) brown moss, limestone cinquefoil Blorenge (SO2510) upland mire Afon Bran [Breconshire] Cleddon Bog (SO 509041) Molinia bog Blaen Cilieni (SN8938) Carex paniculata, Molinia Glyn Wood (SO5200) broad-leaved woodland heath Grondra Great Wood (ST4993) broad-leaved Blaen Glaswen [Montgomeryshire] (SJ 031301) woodland watershed blanket bog Langstone (ST3889) wet meadow Brechfa Pool (SO1137) Little Wenallt (SO5099) Cilkenny Dingle [Radnorshire] (SO1741) broad- Magor Marsh (ST4286) grazed Carex riparia leaved woodland Skenfrith (SO4520) wooded bank of River Monnow Colwyn Brook (SO0755) Carex acutiformis fen The British (SO2504) old mining spoil heaps with Cors Lawnt (SJ0412) valley mire developing vegetation and stream Cors-y-Llyn (SO0155) acidic mire Tregate Bridge (SO4716) willow and alder carr by Gors Goch (SN8963) blanket bog River Monnow Crickhowell [Breconshire] (SO2118) NEATH/PORT TALBOT Duthlas [Montgomeryshire] (SO2178) Cefn Grwhyd, Neath (SN7206) valley mire Grwyne Fechan [Breconshire] (SO2226) Gors Llwyn (SN8510, SN8511) fen (Phragmites, Illtyd Pools (SN9625) Juncus basin mire Carex), ombrotrophic mire Llanbrynmair Moor (SH9404) blanket bog Pant-y-Sais (SS7194) Phragmites, Molinia bog Llangammarch Wells [Breconshire] (SN9347) PEMBROKESHIRE (formerly in Dyfed) Llyn Coethlyn (SJ0114) basin mire, wet heath Aber Mawr (SM8834) Phragmites and cinquefoil Llyn Gwynllyn [Radnor] (SN 946692) Molinia Brynberian Moor (SN1234) Sphagnum and bog bog myrtle Rhôsgoch (SO1947) Juncus fen Castlemartin Corse (SR8999) meadow (Juncus, Ton Dingle [Montgomeryshire] (SO2690) broad- Iris) leaved woodland Cilast Isaf (SN2140) Y Gors [Breconshire] (SN 874298) ﬂoodplain mire Coed Tycanol (SN0936) broad-leaved woodland SWANSEA/ABERTAWE (formerly West Glamorgan) Cors Penally (SS1198) fen (Carex riparia, Cefn Bryn (SS 510909) humid heath Phragmites) Crymlyn Bog [Glamorgan] (SS6894-6994) damp Cwm Dewi (SN 010399) Carex riparia heath, Carex Dinas Head (SN0041) Fairwood Common (SS5792) Juncus Dowrog Common (SM7827) Carex valley fen Llanon (SN5408) Molinia bog, ungrazed heather Dyffryn Gwaun (SN0434) grazed meadow, Juncus Oxwich (SS5087) Phragmites fen, Molinia Rhôsili Down (SS4290) damp heath East Trewent Wood (SR9995) broad-leaved Welsh Moor (SS 926522) humid heath woodland VALE OF GLAMORGAN (formerly Mid Glamorgan) Freshwater East (SS0198) Penarth (ST1871) Grassholm (Island) (SM5909) rocky island with sea- St. Bride’s Major [Glamorgan] (SS8974) broad- bird colonies leaved woodland Kilgetty [= Killgety] (SN1307) mushroom farm WREXHAM (formerly in Clwyd) Llangloffan Fen (SM8931) grazed fen Bersham (SJ3149) Portheiddy Moor (SM 808314) grazed, sedge-rich Glyn Ceiriog [Denbighshire] (SJ2038) fen Llay [Denbighshire] (SJ3255) St. David’s Airﬁeld (SM7826) wet heath, Typha Sontley Marsh (SJ3347) fen Skokholm [Island] (SM7395) island with seabird Vicarage Bog (SJ3653) Sphagnum kettlehole colonies Skomer Island (SM7209) island with seabird colonies DESCRIPTION OF THE NEW SPECIES Stackpole Quay (SR9997) coastal cliffs BRADYSIA AUSTERA MENZEL & HELLER SP. NOV. The Ritec (SN1001) Phragmites (FIGS 1–3) Trefeiddan Moor (SM7325) basin mire, bogbean Waun Isaf (SN1429) damp heath Type locality: Lom, 2 km north-east of Libkovice Western Cleddau (SM9131) horsetails and Carex [south-west of Duchcov], 50°35′10″N 13°41′30″E paniculata fen (Czech Republic).

1 2 3

Figures 1–3. Bradysia austera Menzel & Heller sp. nov., male. Fig. 1. Hypopygium in ventral view. Fig. 2. Apical part of gonostyle in ventral view. Fig. 3. 4th ﬂagellomere.

Holotype: male, 280 m a.s.l., small pond with willows Other material: 1 male, England, Norfolk, Middle and rich reed beds, malaise trap, 14.v.−15.vi.1998, Harling (Great Britain), 28.vi.−12.ix.1988, Foster & Barták leg. Procter leg. (PBLN); 2 males, England, Norfolk, Reedham (Great Britain), 27.x.1988, Foster & Procter Paratypes: 1 male, same location, malaise trap, 25.vi.− leg. (PBLN); 2 males, Scotland, Perth & Kinross, Loch 23.vii.1998, Barták leg.; 1 male, Cork, Gearagh (Ire- Tummel, viii.1939, Edwards leg. (BMNH); 1 male, land), old oak-ash woodland on a stream with alder, Wales, Ceredigion, Cors Caranod (Great Britain), willow and birch, sweep net, 31.vi.1990, de Courcy 23.vii.1987, Holmes, Boyce & Reed leg. (PPWM); 1 Williams leg.; 1 male, Cork, Gearagh, Annahala bog male, Wales, Ceredigion, Cors Caron (Great Britain), (Ireland), damp meadow and bog with willow, Juncus, 15.vi.1987, Holmes, Boyce & Reed leg. (PPWM); 1 Typha and Osmunda, sweep net, 25.viii.1990, de male, Wales, Ceredigion, Gwaun Ystrad Caron (Great Courcy Williams leg.; 1 male, Sachsen, Leipzig (Ger- Britain), 26.vi.1987, Holmes, Boyce & Reed leg. many), nature reserve ‘Burgaue’ in north-west area of (PPWM); 1 male, Wales, Pembrokeshire, Castlemartin the town, 96 m a.s.l., riverine forest with pedunculate Corse (Great Britain), 14.x.1987, Holmes, Boyce & oak, ash, hornbeam, sycamore and maple, sweep net, Reed leg. (PPWM); 1 male, Wales, Swansea/Abertawe, 4.v.1993, Metzner leg.; 1 male, no. 865, Schleswig-Hol- Crymlyn Bog (Great Britain), 14.vi.1989, Holmes, stein, Kiel (Germany), garden in the University, light Boyce & Reed leg. (PPWM). trap, 23.vii.1994, Kolligs leg.; 1 male, 2 females, nos. 870, 876 and 962, Schleswig-Holstein, nature reserve Etymology: The name (latin: austera = dark) comes ‘Alte Sorge’ at Meggerdorf (Germany), lowland moor, from the dark bristles on the thorax and abdomen as damp fallow ground, photo-eclector, 2.v.−14.v.1994, well as the unicolourous dark brown base of the Heller leg.; 7 males, 4 females, nos. 862–864, 868–869, antenna (scape, pedicel, 1st and 2nd basal flagellom- 875 and 877, same locality data, 14.v.−1.vi.1994, eres). In contrast, the base of the antenna of the Heller leg.; 4 males, 1 female, nos. 835, 866–867 and closely related species Bradysia subaprica Mohrig & 873, same locality data, 15.vii.−15.viii.1994, Heller Krivosheina is distinctly pale yellow. leg.; 6 males, same locality data, 15.viii.−1.ix.1994, Description. Male. HEAD: eyebridge narrow, 2 facets Heller leg.; 2 males, no. 931, same locality data, 15.ix.− wide. Antennae moderately long and dark; basal seg- 1.x.1994, Heller leg.; 3 males, nos. 2694–2695, ments dark brown; 4th flagellomere (Fig. 3) 2.8–3.1 Schleswig-Holstein, Wankendorf (Germany), damp times as long as wide; setae coarse, dark and erect but fallow grassland, photo-eclector, 17.v.−31.v.1994, curved and almost as long as flagellomere width; basal Hingst leg.; 1 male, no. 2724, same locality data, part with rough surface structure and long sensilla; 13.vi.−27.vi.1995, Hingst leg. necks short and well differentiated; necks of flagellom- Location of types: holotype in the collection of DEI; eres 1–7(8) unicolourous dark brown, necks of paratypes in the collections of DEI (10 males, 1 flagellomeres 8(9)−13 whitish and bi-coloured. Prae- female), NMID (1 male) and PKHH (18 males, 6 frons strong and long with relatively dense, pale females). brown bristles; clypeus almost bare (with only 1–2

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 28 F. MENZEL ET AL. bristles). Palpi moderately long, 3-segmented and yel- Discussion. The new species belongs to the Bradysia lowish; basal segment slightly hump-backed with 2–4 fungicola-group. It is closely related to Bradysia sub- bristles (one distinctly longer than the others); sen- aprica Mohrig & Krivosheina, 1989, for which only the sory area simple, without distinct margin or pit; sen- type material from far eastern Russia is known to silla long, curved and pale; apical segment 0.7–0.9 date. In the past, the specimens examined from times as long as basal segment; 2nd segment longish- Europe were often wrongly identified as Bradysia sub- oval, about 2/3 as long as the apical segment with dis- aprica, so that there are also several incorrect litera- tinctly longer outer bristle. THORAX: dark brown with ture citations. Bradysia austera differs from Bradysia widespread lateral bright yellow colouration. Post- subaprica by the narrower, 2-facet-wide eyebridge, pronotum bare. Mesonotum with stronger, darker the unicolourous dark brown base to the antennae, brown setae (yellowish on faded specimens) and long, the slightly longer flagellomeres (length-width lateral and medial bristles. Katepisternum triangular, index = 2.8–3.1), the bi-coloured bands on the necks deeper than wide. Scutellum usually with 2 (rarely 3) from the 8th flagellomere onwards and a slightly very long marginal bristles among other setae. Legs wider apex on the gonostyles that is less narrow below long and strong; coxae and femora light brown to the spine group particularly on the inner side. The yellow; tibiae and tarsi darker due to dense setae; original description for Bradysia subaprica gave a anterior tibia with 2–5 spines among other setae wider, 3-facet-wide eyebridge, pale yellow on the base (sometimes weakly developed); anterior femur strong of the antennae as far as 2nd flagellomere, the 4th but not distinctly thickened; apex of anterior tibia flagellomere only 2.2 times as long as wide and a nar- with narrow comb of bristles in a single row (4–6 rower and more distinct point to the apex of the coarse bristles); middle and posterior tibial spurs slen- gonostyles. der and subequal; posterior tibia with dorsal row of spines; apex with a circlet of spines. Claws with fine teeth. Wings smoky-brown with well developed anal BRADYSIA ISMAYI MENZEL SP. NOV. area; posterior wing veins strong and, like the wing (FIGS 4–6) membrane, without macrotrichia; fork of M long- attenuated, weakly curved with M1 and M2 wide apart; Type locality: England, Essex, Chigwell Row, Chigwell stem of M slightly longer than fork of M; x = 1.1–1.4 y, Row Wood (Great Britain). both bare or y with 1–3 distal macrotrichia; stem of Holotype: male, 65–75 m a.s.l., heathland (heather) CuA = 0.5–0.7 x; R short = 0.4–0.5 R and joining c 1 with oak and holly, aerial trap, 11.vi.2002, Ismay & well before base of fork of M; R with only dorsal mac- 5 Schulten leg. rotrichia (very rarely with 1–2 ventral macrotrichia on apical part); C = 2/3–4/5 w. Halteres with short stalk, Paratypes: 1 male, same locality data, 20.vi.2002, pale yellow to light brown. ABDOMEN: brown; body Ismay & Schulten leg.; 2 males, England, Essex, Chig- setae strong, long and dark (setae pale only on ventral well Row, Chigwell Row Wood (Great Britain), 65– side of abdomen). Hypopygium (Fig. 1) compact, 75 m a.s.l., ancient woodland with oak, hazel and slightly wider than high and yellowish brown; ventral bramble, aerial traps, 23.v.−27.v.2002 and 18.vi.2002, base of genitalia without basal lobe or dense bristle Ismay & Schulten leg.; 1 male, England, Essex, group; ventral inner side of gonocoxites with short Epping Forest (Great Britain), ancient broad-leaved dark setae. Gonostyles slender, about 2.7 times as long woodland and grassland, Malaise trap, v.1999, Dagley as wide and flattened apically (Fig. 2); apex darker leg.; 3 males, England, Norfolk, Norwich, The Loke brown with darker, dense coarse bristles; apical tooth (Great Britain), garden, Malaise trap, 21.viii.− missing; apex rather flat, with distinct subapical spine 24.viii.1989, 28.iv.−4.v.1990 and 21.viii.1992, all Lau- group (4–5 curved, very closely aligned spines; all rence leg.; 1 male, England, Oxfordshire, Oxford, Mil- strong, dark brown and almost subequal). Tegmen ham Ford School (Great Britain), school grounds with slightly wider than high, brown and weakly sclero- old poplars, Malaise trap, 18.v.−22.v.2001, Webb leg.; 2 tized; apex flatly rounded; area with teeth large and males, Wales, Pembrokeshire, Coed Tycanol (Great circular with very coarse, long, single tipped teeth. Britain), mixed woodland, Malaise trap, 10.vi.− Aedeagus short with slender genital apodeme with 29.vi.1999, Levey & Pavett leg.; 1 male, Scotland, sclerotized, semicircular base. Body length: 1.8– Highland Region, Migdale Loch (Great Britain), alder 2.2 mm. and birch carr by stream surrounded by pine wood- Female. Antennal flagellomeres distinctly shorter; land on higher ground, sweep net, 31.v.2002, Chandler 4th flagellomere 2.0–2.2 times as long as wide; thorax leg.; 1 male, Wales, Gwynedd, Hafod Garregog (Great laterally also partly bright yellow as in male; wings Britain), broad-leaved woodland, Malaise trap, 18.ix.− slightly larger; all other characteristics as for males. 28.ix.1999, Levey & Pavett leg.; 1 male, no. 1891, Body length: 2.3–2.5 mm. Rhineland-Palatinate, Eifel, Daun district, Gönners-

Figures 4–6. Bradysia ismayi Menzel sp. nov., male. Fig. 4. Hypopygium in ventral view. Fig. 5. Apical part of gonostyle in ventral view. Fig. 6. 4th flagellomere. dorf (Germany), grassland with fruit trees, Malaise curved (some setae closely aligned giving the impres- trap, 21.v.1994, Cölln leg. sion of a whorl); setae about 1/2 as long as flagellomere width; basal part with rough surface structure; necks Location of types: holotype in the collection of DEI; well differentiated and unicolourous dark brown. Face paratypes in the collections of DEI (3 males), HRI (1 with strong, long, dark brown bristles. Palpi long, 3- male), PBLN (3 males), PKHH (1 male) and PPCM (5 segmented and dark; basal segment hump-backed males). with 2–4 bristles (one distinctly longer than the oth- Other material: 1 male, England, Hampshire, Beau- ers); sensory area large with deep pit; sensilla short, lieu River (Great Britain), 22.v.1936, Edwards leg. dark and almost straight; 2nd and 3rd segments nar- (BMNH); 1 male, England, Hertfordshire, Letchworth row; apical segment 1.2 times as long as basal seg- (Great Britain), ix.1936, Edwards leg. (BMNH). ment; 2nd segment long-rod-like, =2/3–3/4 times as long as basal segment and with long outer bristle. Etymology: The new species was named after the dip- Thorax dark brown to black laterally with only a few terist Dr John Ismay (Oxford University Museum of paler areas. Postpronotum bare. Mesonotum with Natural History, Oxford) in recognition of his long ser- strong and coarse, dense, distinctly dark bristles as vice in collecting Diptera. He has contributed greatly well as several longer lateral and medial bristles. to the knowledge of the British sciarid fauna. Katepisternum triangular, deeper than wide. Scutel- Description. Male. HEAD: eyebridge 2–3 facets wide. lum with four longer marginal bristles among other Antennae short, strong and dark; basal segments dark setae. Legs long and strong; coxae, femora and tibiae brown; 4th flagellomere (Fig. 6) 2.0–2.2 times as long unicolourous dark brown; anterior tibia with spines as wide; setae of flagellomere dense, pale and erect but among other setae; anterior femur strong but not dis-

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 30 F. MENZEL ET AL. tinctly thickened; apex of anterior tibia with comb of Using the key in Menzel & Mohrig (2000), these two bristles in a single row (6–9 paler bristles extending to species could not be assigned to any of the species about middle of tibia); middle and posterior tibial group concepts as currently defined. At first glance, spurs slender and subequal; posterior tibia with dor- the Bradysia giraudii-group or the Bradysia mela- sal spine row; apex of posterior tibia with circlet of nura-group could be considered, but because of the spines. Claws without teeth. Wings distinctly brown absence of several characters they do not fit either of with well-developed anal area; posterior wing veins these species groups. In particular, R1 joining c before strong and, as membrane, without macrotrichia; fork the base of the fork of M, the narrow comb of bristles of M long-attenuated, base uniformly curved; M1 and on the anterior tibia, the conically narrowed tegmen,

M2 diverge in apical third; stem of M slightly shorter the pointed gonostyles with cheek-like swellings on than fork of M; x = 1.0–1.2 y; x bare and y with bristles their inner sides, as well as the sparse covering of (y on distal half or completely with macrotrichia); spines so that the apex of the gonostyles is not exten- stem of CuA = 0.6–0.7 x; R/R1-vein complex joining c sively surrounded by the group of spines: these before base of fork of M (with relatively long R and characters exclude the species from the Bradysia short R1); R1 = 0.6–0.7 R; R5 with macrotrichia on both melanura-group. On the other hand, these species sides (ventral macrotrichia on apex of wing as far as cannot be assigned to the Bradysia giraudii-group

R1-intersection, sometimes over entire R5); C = 2/3 w. because they do not have clubbed gonostyles with Halteres dark brown with short stalk. ABDOMEN: dark broadly rounded apices, nor a hook-like apical tooth, brown but paler than thorax; body setae strong, dense and no long spines on raised sockets. For this reason and dark. Hypopygium (Fig. 4) compact and slightly the Bradysia hortensis-group is established, to wider than high; setae distinctly coarse and dark which initially the two species Bradysia hortensis and brown to black; ventral base of genitalia without basal Bradysia ismayi are assigned. It is distinguished in lobe and without dense bristle group; ventral inner particular from the Bradysia giraudii-group by the side of gonocoxites with short, dark setae. Gonostyles following characters: antennal setae short, dense and slender, about 2.5 times as long as wide and pointed at ﬁne, at most 1/2 as long as the antennal width; apex of apex (Fig. 5); inner side in middle third (especially anterior tibia with a narrow comb of bristles (extend- dorsally) with cheek-like swelling; apex without apical ing at most to the middle of the tip); R5 with an exten- tooth and with 5–6 strong-short spines (arranged in sive covering of macrotrichia on both sides (at least two groups, one above the other; 2 + 3 or 2 + 4 spines). isolated macrotrichia extending from wing apex to the

Tegmen about as high as wide, strongly sclerotized tip of R1); gonostyles much narrower and pointed at and more tapered apically; basal structure strong, apex, apex without apical tooth and with only a group long and thick; area with teeth large and circular with of apical spines; spines short, strong, closely aligned coarse, single tipped teeth arranged in rows. Aedeagus and not displaced as far as the outer side of the gono- short with wide, semicircular, weakly sclerotized base. style; all spines on gonostyles on short sockets; middle Body length: 3.6–4.0 mm. of dorsal inner sides of gonostyles with cheek-like Female. Unknown. swelling; tegmen conically narrowed and narrowly rounded at apex. Discussion. The new species is very similar to Bradysia hortensis Heller, 2000, in its genital struc- BRADYSIA NIGRISPINA MENZEL SP. NOV. ture. However, it differs markedly in body length (FIGS 7–9) (Bradysia hortensis Heller, contrary to the original description, is only 2.7 mm) and in the distinctly Type locality: Hruby Jeseník, Velká Kotlina near Malá longer ﬂagellomeres (4th ﬂagellomere almost twice as Morávka (Czech Republic). long as in Bradysia hortensis). Additionally, Bradysia Holotype: male, 1185 m a.s.l., meadow by the source of ismayi has a very dark body (incl. genitalia) with the river Moravice (Daphne-Aceretum), pitfall trap, uniformly dark brown legs and halteres, large and 7.vi.1995, Frouz leg. strongly clouded wings, a higher tegmen with a smaller area of teeth and shorter teeth as well as a Paratypes: male, England, Warwickshire, Welles- more distinctly narrow apex to the gonostyles. While bourne (Great Britain), aerial trap outside mushroom the spines on the gonostyles of Bradysia hortensis are farm, 5.vii.−19.vii.2001, J.E. Smith leg.; 1 male, arranged in a circle and on a relatively broadly demar- England, Northumberland, ‘Berwick’ [= Berwick- cated and well-marked elevation, those of Bradysia upon-Tweed] (Great Britain), garden, Malaise trap, ismayi are in 2 narrow groups one over the other. In 24.v.−7.vi.1996, Laurence leg.; 2 males, same locality the new species the lobe-like elevation on which the data, water trap, viii.1999 and ix.1999, Laurence leg.; spines are set in B. hortensis is missing due to the 1 male, Norfolk, Norwich, The Loke (Great Britain), strongly pointed gonostyles. garden, 2.iii.1993, Laurence leg.; 1 male, Silkeborg

7 8 9

Figures 7–9. Bradysia nigrispina Menzel sp. nov., male. Fig. 7. Hypopygium in ventral view. Fig. 8. Gonostyle in ventral view. Fig. 9. 4th ﬂagellomere.

(Denmark), from Lengersdorf identified as ‘? triseri- elongated. Coxae and legs pale brown, anterior legs ata’, 16.v.1929, Nielsen leg. partly yellowish, anterior tibiae with several spines among other setae; apex of anterior tibia with comb Location of types: Holotype in the collection of DEI; of bristles in a single row (5–7 pale bristles); middle paratypes in the collections of DEI (1 male), HRI (1 and posterior tibiae with 2 subequal spurs; posterior male), PBLN (3 males) and ZFMK (1 male). tibia with dorsal row of spines; apex of posterior tibia Other material: 1 male, England, Hertfordshire, with circlet of spines. Claws without teeth. Wings Letchworth (Great Britain), vii.1918, Edwards leg. large and pale with well-developed anal area; poste- (BMNH); 2 males, England, Shropshire, Shrewsbury rior wing veins weak, very pale and, as the wing (Great Britain), 21.vii.1920, Edwards leg. (BMNH). membrane, without macrotrichia; stem of M longer than fork of M; fork of M long and narrow; M and M Etymology: The name ‘nigrispina’ describes the dis- 1 2 partially parallel, diverging towards wing margin; tinct black colour of the spines on the gonostyles that x = 1.1–1.9 y, both bare; stem of CuA = 1/2–4/5 x; is in distinct contrast to the pale brown colour of the R = 0.6–0.8 R, joining c well before base of fork of M; genitalia (latin: niger = black; spina = spine). 1 apical part of R5 with macrotrichia on both sides Description. Male. HEAD: eyebridge 2 facets wide. (with 5–9 ventral macrotrichia); C = 3/5 w. Halteres Antennae long and unicolourous brown; all flagellom- with short stalk, whitish-yellow. ABDOMEN: brown, eres about same length and slightly rough; 4th distinctly paler than thorax; body setae very fine, flagellomere (Fig. 9) 2.9–3.4 times as long as wide; whitish and moderately long. Hypopygium (Fig. 7) setae dense, erect, fine and pale, about as long as wider than high, pale brown, with fine, pale bristles; flagellomere width; basal part with long sensilla ventral base of gonocoxites without basal lobe or tuft among setae; necks short, well differentiated and uni- of setae; inner side v-shaped with sparse and short colourous brown. Face with very fine, pale yellow setae. Gonostyles (Fig. 8) long-attenuated, 2.8–3.0 setae. Palpi long, 3-segmented and pale; basal seg- times as long as wide; base distinctly tapered; inner ment hump-backed with 4–7 bristles (one distinctly side uniform, without emargination or excavation; longer than the others); sensory area on the basal apex on outer side with small, claw-like, backward segment large and dark with a deep pit; sensilla long set tooth; with 7–10 coarse, subequal and distinctly and slightly curved; apical segment very long and dark spines underneath; all spines curved inwards narrow, 1.3 times as long as basal segment; 2nd seg- and downwards, closely aligned and covering whole ment longish-oval and 0.6–0.7 times as long as apical apex; apical setae fine, short and very sparse. Tegmen segment. THORAX: dark brown, laterally yellowish. wider than high and trapezoidal; apex flattened and Postpronotum bare. Mesonotum with fine, sparse, membranous; area with teeth large and circular, with whitish setae and several strong, distinctly pale, long, single tipped teeth. Genital apodeme short, with lateral and medial bristles. Scutellum well differ- weakly sclerotized, funnel shaped base. Body length: entiated with 2 long, strong marginal bristles. 2.0–2.3 mm. Katepisternum triangular, deeper than wide, not Female. Unknown.

Discussion. The new species has all the characteristics gia, Thuringian Basin, Sieblebener Teich near Gotha of the Bradysia tilicola-group, in particular the dark, (Germany), broad-leaved woodland, sweep net, very deep sensory area on the basal segment of the 28.v.1988, Menzel leg.; 5 males, 3 females, nos. 1133– palpi, the fine comb of bristles on the anterior tibiae, 1136, Brandenburg, biosphere reserve ‘Schorfheide- the untoothed claws, the sparse-short hairs on the Chorin’, Altkünkendorf north-east of Joachimsthal mesonotum, the 2 strong marginal bristles of the (Germany), reeds, photo-eclector, 24.vi.−24.vii.1995, scutellum and the longish gonostyles with rounded Nötzold leg.; 1 male, Holibka, south-east of Bílina apex, with apical/subapical spine group and a reduced, (Czech Republic), 50°31′20″N 13°49′40″E, 360 m a.s.l., short hook-shaped tooth. The genitalia in Bradysia alder woodland on a small pond, yellow pan trap, nigrispina and Bradysia albochaeta Mohrig & 8.viii.−9.viii.1998, Barták leg.; 2 males, no. 400, Mamaev, 1985 are very similar. The distinguishing England, Berkshire, Dinton Pastures Country Park, characteristics in Bradysia nigrispina are the longer Mortimer’s Meadow (Great Britain), swampy wood- flagellomeres, the very pale halteres and the pale land, sweep net, 26.vi.1993, Chandler leg.; 2 males, brown hypopygium with distinct black-brown spines. nos. 59 and 61, England, Berkshire, Dinton Pastures The apex of the gonostyles covered with very dark Country Park, Sandford Copse (Great Britain), alder spines is very characteristic and stands in distinct woodland, sweep net, 3.vii.1994, Chandler leg.; 1 contrast to the general colour of the genitalia. The male, no. 381, England, Berkshire, Little Wittenham whitish, extremely fine and weak body setae are very Wood (Great Britain), broad-leaved woodland, sweep distinctive (especially on the mesonotum and scutel- net, 3.vii.1985, Chandler leg.; 6 males, 3 females, lum, coxae and femora, abdomen and hypopygium England, Berkshire, Reading, Searles Farm (Great including the stronger marginal bristles). Britain), carr woodland, sweep net, 26.vi.2003 and 22.vii.2003, all Chandler leg.; 10 males, England, Berkshire, Theale (Great Britain), carr woodland, CORYNOPTERA FLAVOSIGNATA MENZEL & HELLER sweep net, 7.vii.2003, Chandler leg.; 1 male, no. 380, SP. NOV. (FIGS 10–12) England, Cambridgeshire, Badger Wood (Great Brit- Type locality: Thuringia, nature reserve ‘Apfelstädter ain), broad-leaved woodland, Malaise trap, vi.1998, Ried’ near Apfelstädt (Germany). C.W. Plant leg.; 1 male, England, Cambridgeshire, Monkfield Wood (Great Britain), broad-leaved wood- Holotype: male, meadow with cabbage thistle, yellow land, Malaise trap, vi.1998, C.W. Plant leg.; 1 male, no. pan trap, 1.vii.1985, Weipert leg. 225, England, Durham, Castle Eden Dene (Great Brit- Paratypes: 3 males, same data; 2 males, Thuringia, ain), young mixed woodland, sweep net, 26.vi.1992, nature reserve ‘Apfelstädter Ried’ near Apfelstädt Chandler leg.; 1 male, no. 721, England, Somerset, (Germany), meadow with cabbage thistle, yellow pan Pawlett Levels (Great Britain), adjacent moorland trap, 13.vii.1985, Weipert leg.; 1 male, Thuringia, Thu- with old hedges, sweep net, 13.ix.1999, Gibbs leg.; 2 ringian Basin, Hanfsee near Schlotheim (Germany), males, England, Suffolk, Bradfield Wood (Great Brit- Barber trap, 3.vii.1989, Bellstedt leg.; 1 male, Thurin- ain), broad-leaved woodland, yellow pan trap, 6.vii.−

10 11 12

Figures 10–12. Corynoptera ﬂavosignata Menzel & Heller sp. nov., male. Fig. 10. Hypopygium in ventral view. Fig. 11. Apical part of gonostyle in ventral view. Fig. 12. 4th ﬂagellomere.

11.vii.1990, Laurence leg.; 1 male, no. 103, England, ial spurs on middle and posterior legs slender and Wiltshire, Lackham Wood (Great Britain), broad- subequal; posterior tibia with dorsal row of spines; leaved woodland, sweep net, 2.v.2002, Chandler leg.; 1 apex with circlet of spines. Claws without teeth. Wings female, England, Warwickshire, east of Coventry, well developed and slightly darkened (sometimes Coombe Abbey Country Park (Great Britain), slightly narrower and then with relatively weakly 52°24′53″N 01°25′12″W, 95 m a.s.l., broad-leaved developed anal area); posterior wing veins distinct woodland with ash, pedunculate oak, sycamore and and, as membrane, without macrotrichia; stem of M elder, sweep net, 28.viii.2003, Menzel leg.; 1 male, weak and longer than fork of M; fork of M short-curved England, Warwickshire, Leamington Spa, Newbold and wide apart; x = 0.8–1.1 y, both bare; stem of

Comyn Park (Great Britain), damp meadow, sweep CuA = 0.3–0.6 x (CuA1 and CuA2 sometimes converg- net, 15.vii.2001, J.E. Smith leg. ing directly before intersection); R1 short, = 0.5–0.7 R and joining c well before base of fork of M; R and R Location of types: holotype in the collection of DEI; 1 5 with only dorsal macrotrichia (rarely with additional paratypes in the collections of DEI (5 males, 1 female), 1–2 ventral macrotrichia in first half of R ); C = 3/5 w. HRI (12 males, 2 females), NKME (1 male), PBLN (2 5 Halteres with short stalk, brown and with 1–2 rows of males), PDGB (1 male), PKHH (3 males, 3 females), small bristles. ABDOMEN: bi-coloured with dark brown PPCM (14 males, 1 female) and PWMP (4 males). dorsal and pale yellow underside; body setae sparse Other material: 1 male, England, Hertfordshire, and short, fine and pale. Hypopygium (Fig. 10) about Letchworth (Great Britain), v.1918, Edwards leg. as high as wide and yellow; ventral base of genitalia (BMNH); 1 male, England, Oxfordshire, Charlbury without basal lobe and without basal bristle group; (Great Britain), 4.vii.1936, Edwards leg. (BMNH). inner side of gonocoxites with wide emargination and short, fine setae. Gonostyles strong, about twice as Etymology: The name ‘flavosignata’ refers to the yel- long as wide with coarse and sparse apical setae low colour of the basal segments of the antennae (Fig. 11); apex distinctly pointed, with 3 equally (latin: flavus = yellow; signata = marked). strong, long, evenly curved, closely grouped spines Description. Male. HEAD: eyebridge complete, 2–3 directly below apex (always 2 + 1); apical tooth, whip- facets wide. Basal segments of antenna distinctly lash-like hair and distinctly long bristles missing; whitish-yellow colour; flagellomeres long, rough and inner side of upper half slightly emarginated. Tegmen unicolourous brown (also 1st flagellomere dark); 4th distinctly wider than high, membranous with widely flagellomere (Fig. 12) 3.0–3.6 times as long as wide; rounded apex; area with teeth small, wider than high basal part without sensilla; setae on basal part long, with long, single tipped teeth. Aedeagus very short coarse, erect, slightly longer than flagellomere width; and narrow, with wide, funnel shaped base. Body necks short, unicolourous brown and well differenti- length: 1.7–2.2 mm. ated. Face with sparse, fine, pale bristles. Palpi short, Female. Flagellomeres distinctly shorter; 4th 3-segmented and pale yellow; basal segment short, flagellomere 1.8–2.1 times as long as wide; all slightly hump-backed and with a long outer bristle; flagellomeres with sparse setae that are distinctly sensory area on edge of segment and with margin; sen- shorter than flagellomere width. Palpi 3-segmented, silla fine and slightly curved; 2nd palpal segment with longer apical and 2nd segments than males; 2nd short-ovoid to almost spherical, about 0.7–1.0 times as segment 2/3–3/4 times as long as apical segment; sen- long as the narrower apical segment and with several sory area with larger, stronger margin; sensilla longer bristles (one outer bristle distinctly longer); 2nd and and more distinctly curved; patch of bristles on apex of 3rd palpal segments together only slightly longer than anterior tibia with weak, but distinctly curved margin; basal segment. THORAX: yellow-brown, coxae and legs wings larger and wider, with well developed anal area; whitish-yellow; thorax laterally always with much stem of CuA = 0.2–0.6 x; R1 = 0.6 R; C = 3/5–2/3 w; paler areas (especially katepisternum and ventral all other characters as in males. Body length: areas of metanotum, often whitish-yellow). Postprono- 1.9–2.3 mm. tum bare. Mesonotum with very sparse, pale setae and slightly longer and stronger lateral and medial bris- Discussion. The new species definitely belongs to tles. Scutellum with 2 long and strong, pale, marginal the Corynoptera membranigera-group falling between bristles among other fine setae. Katepisternum trian- Corynoptera membranigera (Kieffer, 1903) and gular, deeper than wide. Legs long and slender; ante- Corynoptera saccata Tuomikoski, 1960. Over the past rior tibia with fine (but distinct) spines among other few years, some collection data for Corynoptera fla- setae; anterior femur strong and not distinctly thick- vosignata were incorrectly published under the name ened; apex of anterior tibia with wide patch of bristles Corynoptera trispina Tuomikoski [a synonym of almost in a single row without basal carina and with- Corynoptera membranigera (Kieffer)]. The wrong out or with very weakly developed curved margin; tib- identification was based on a small amount of mate-

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 34 F. MENZEL ET AL. rial from Thuringia (Germany), which led Menzel & Holotype: male, broad-leaved woodland with peduncu- Mohrig (2000: 231) to describe the base of the anten- late oak, birch, hazel and large-leaved lime, sweep net, nae of Corynoptera membranigera as ‘more rarely 25.viii.2002, Menzel leg. distinctly yellow to whitish’. The new species is Paratypes: 13 males, same data, 25.viii.2002, Menzel distinguished from Corynoptera membranigera and leg.; 2 males, same data, 25.viii.2002, J.E. Smith leg.; Corynoptera saccata particularly by the pale base of 2 males, England, Leicestershire, nature reserve the antennae, of which only the scape and pedicel are ‘Kelham Bridges’ south of Ravenstone [south-west distinctly yellow. In addition to this characteristic, of Coalville] (Great Britain), 52°42′17″N 01°23′59″W, Corynoptera ﬂavosignata has thickened and pointed 124 m a.s.l., area around bank of pond in damp gonostyles with 3 long and strong, very closely aligned meadow, sweep net, 24.viii.2003, Menzel leg.; 1 male, spines on the apex (2 spines above and one spine no. 398, England, Greater London, Highgate Woods, directly beneath). All spines are shorter and more mixed woodland, Malaise trap, 1998–99, Hackett leg.; dense than in Corynoptera membranigera and are 1 male, no. 313, England, Buckinghamshire, Burnham distinctly longer than in Corynoptera saccata. Also Beeches, Moat (Great Britain), ancient beech wood- the upper half of the inner side of the gonostyles of land and earthworks, Malaise trap, 8.vii.−22.vii.1996, Corynoptera membranigera is more strongly emargin- Ismay leg.; 1 male, no. 202, England, Buckingham- ated and the 3–4 spines stand one above the other and shire, Burnham Beeches, Pumpkin Hill (Great Brit- are relatively widely isolated from each other in the ain), beech woodland, Malaise trap, 22.vii.−5.viii.1996, upper third. There are similarities to Corynoptera Ismay leg. hemiacantha Mohrig & Menzel, 1992 in the thickened gonostyles, the weak emargination of the inner side Location of types: holotype in the collection of DEI; and the typical arrangement of the spines, but the paratypes in collections of DEI (12 males), HRI (5 basal segments of the antennae of Corynoptera hemi- males) and PPCM (3 males). acantha are dark. Also in Corynoptera hemiacantha Other material: 3 males, Wales, Isle of Anglesey, Fedw the gonostyles are wider, the inner side is more dis- Fawr (Great Britain), 13.vii.1988, Holmes, Boyce & tinctly arched and the apex distinctly pointed. Reed leg. (PPWM); 1 male, Wales, Powys, Y Gors (Great Britain), 17.vii.1989, Holmes, Boyce & Reed leg. (PPWM); 1 male, Wales, Swansea/Abertawe, CORYNOPTERA UNCATA MENZEL & SMITH SP. NOV. Welsh Moor (Great Britain), 14.vi.1989, Holmes, (FIGS 13–15) Boyce & Reed leg. (PPWM). Type locality: England, West Midlands, nature reserve Etymology: The name was chosen because of the very ‘Ryton Wood’ near Bubbenhall [south-east of Coven- strongly curved gonostyles, characteristic of the spe- try] (Great Britain). cies (latin: uncata = curved).

13 14 15

Figures 13–15. Corynoptera uncata Menzel & Smith sp. nov., male. Fig. 13. Hypopygium in ventral view. Fig. 14. Api- cal part of gonostyle in ventral view. Fig. 15. 4th ﬂagellomere.

Description. Male. HEAD: eyebridge wide with 3–4 fac- Discussion. The new species belongs to the C. subtilis- ets. Antennae very long and slender, unicolourous group and is close to Corynoptera sedula Mohrig & Kri- brown; basal antennal segments dark; 4th flagellom- vosheina, 1985. Both species have long flagellomeres ere (Fig. 15) 3.6–4.4 times as long as wide; setae on and strong gonostyles with an inwardly curved, flagellomeres very long, coarse and erect, about as pointed apex. On the narrowly round apex there is a long or slightly longer than flagellomere width; necks strong tooth and several slender spines. The gonostyles uniformly dark and well differentiated; basal part of Corynoptera sedula are relatively slender (almost without sensilla. Palpi yellow and 3-segmented; basal the same width from the base to the apex), below the segment slender to weakly hump-backed, about as apical tooth there are always 3 spines and the inner long as slender apical segment with an outer bristle side of the gonocoxites has relatively short bristles. It but without deepened sensory pit; sensilla fine; 2nd has the eyebridge three facets wide and has very fine, segment short-oval and about 3/5 as long as apical seg- pale body hairs and unicolourous brown palpi, legs and ment. THORAX: dark brown, laterally with only a few halteres. By contrast, Corynoptera uncata has a longer paler, yellowish areas in front of base of wing. Post- body, yellow to light brown palpi and legs, sooty black pronotum bare. Mesonotum with sparse, coarse, dark halteres, dark and coarse body hairs and the eyebridge setae and long-strong lateral and medial bristles. to 4 facets wide. The genitalia of the new species has Coxae and legs yellow to pale brown (in distinct con- thicker gonostyles (much wider in the lower half and trast to dark thorax and abdomen); legs very long and with less deep emargination on the inner side at the slender; anterior legs with spines among other setae; upper half than Corynoptera sedula), always with 5 apex of anterior tibia with patch of bristles almost in a hyaline spines on the apex and distinctly long bristles single row, without basal carina but with curved mar- on the inner side of the gonocoxites. gin (all bristles dark, coarse and spine-like); spurs on middle and posterior tibiae slender and subequal; posterior tibia with dorsal row of spines; apex with EPIDAPUS (EPIDAPUS) SUBGRACILIS MENZEL & circlet of spines. Claws without teeth. Wings large and MOHRIG SP. NOV. (FIGS 16–20) brownish; posterior wing veins strong and as mem- Type locality: Holibka [south-east of Bílina], brane without macrotrichia; stem of M weak and 50°31′20″N 13°49′40″E (Czech Republic). slightly shorter or equal to fork of M; fork of M long and evenly curved (M1 and M2 almost parallel); Holotype: male, 450 m a.s.l., stony steppe, yellow pan x = 0.8–1.1 y, both bare; stem of CuA short, = 0.3–0.5 trap, 24.viii.−23.ix.1998, Barták leg. x; R and R with only dorsal macrotrichia (more 1 5 Paratypes: 1 male, no. 233, England, Essex, Epping rarely R with 1–3 ventral macrotrichia in first half); 5 Forest, Monument Field (Great Britain), meadow R = 0.6–0.7 R and joining c well before base of fork of 1 near ancient broad-leaved woodland, Malaise trap, M; R joining c at level of middle of fork of M; C = 2/3 5 29.iii.1999, Dagley leg.; 1 male, Vitosha Mountains w. Halteres dark brown, with short stalk and with (Bulgaria), sweep net, 9.ix.1989, Dimitrova leg.; 2 1–2 rows of bristles. ABDOMEN: dark brown; body males, Andalusia, Jimena de la Frontera (Spain), setae coarse, moderately long and dark. Hypopygium valley with stream, damp cork oak woodland with (Fig. 13) about as high as wide, uniformly dark brown bramble, wild olive and grasses, yellow pan trap, and without basal lobe or ventral tuft of setae; ventral iii.–iv.1995, Wilden leg. inner side of gonocoxites with long setae and long bristles crossed over tegmen. Gonostyles about twice as Location of types: holotype in the collection of DEI; long as wide, distinctly thickened and narrowly paratypes in the collections of DEI (2 males), PPCM (1 rounded apically (Fig. 14); apex slightly pointed with male) and PWMP (1 male). coarse and sparse bristles and with a strong, dark api- Etymology: The name ‘subgracilis’ was chosen to show cal tooth; group of spines below tooth subapically (5 that the new species is closely related to Epidapus pale, hyaline spines about as long as or slightly longer gracilis (Walker, 1848) (latin: sub = near, almost, a lit- than tooth); middle of inner side of gonostyles with tle as; gracilis = slender, graceful). cheek-like swelling, upper half distinctly emarginated and without long whiplash-like hair; longer bristles Description. Male. HEAD: head capsule roundish, eye- only present near group of spines ventrally. Tegmen bridge complete, 2 facets wide. Ocelli present. Anten- dark brown, strongly sclerotized and distinctly wider nae long, unicolourous brown and slightly rough; than high; apex of tegmen evenly rounded; area with basal antennal segments dark; 1st flagellomere teeth circular, with fine, single tipped teeth. Aedeagus elongated, = 3.9–4.7 times as long as wide; 4th short, with funnel shaped, scerotized base. Body flagellomere (Fig. 18) 3.6–4.2 times as long as wide; length: 1.9–2.5 mm. setae on flagellomeres coarse, bristle-like about 1.5 Female. Unknown. times as long as flagellomere width; neck long, = 1/3–

16 17 18

19 20

Figures 16–20. Epidapus subgracilis Menzel & Mohrig sp. nov., male. Fig. 16. Hypopygium in ventral view. Fig. 17. Apical part of gonostyle in ventral view. Fig. 18. 4th ﬂagellomere. Fig. 19. Palpus in lateral view. Fig. 20. Haltere.

2/5 times as long as basal part and strongly differen- erect; anterior tibia without spines among other tiated; basal part with long bristle-like setae among bristles; apex of anterior tibia with patch of slightly isolated, long sensilla. Face with moderately long, denser bristles without margin; all tibial spurs fine, coarse, dark bristles. Palpi (Fig. 19) one-segmented, slender and pencil-like; both spurs on middle and pos- short-ovoid swollen and brownish; basal segment with terior tibiae shortened so that they are almost sub- 5–7 coarse bristles, almost all on underside; sensory equal; posterior tibia with long and erect, coarse row of area large, towards edge and apex of segment, simple bristles which merge into other small fine spines in and without margin; sensilla long and straight (not apical third; apex of posterior tibia without dense cir- curved). THORAX: coxae and legs unicolourous brown clet of spines (with only a few coarse and long bristles to dark brown. Postpronotum bare. Mesonotum and some isolated, fine spines). Claws reduced in slightly arched; bristles coarse and dark with some width, without teeth. Wings slightly brownish with longer, slightly stronger bristles on lateral and medial wedge shaped, reduced anal area; posterior wing veins areas. Scutellum well differentiated with 2–3 longer weak and as membrane without macrotrichia; stem of marginal bristles among other finer bristles. Katepis- M pale and slightly shorter than wide fork of M (in ternum flat and elongate wedge-shaped. Coxae and Spanish specimen distinctly shorter and wider fork of legs slender and elongated; bristles on underside of M); fork of M short-triangular and wide apart (M1 and anterior and posterior femora not distinctly long and M2 curved and strongly diverging); x = 1.3–1.5 y, both

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 BRITISH SCIARID FAUNA 37 bare; stem of CuA short, = 0.4–0.5 x (in 2 paratypes THE BRITISH SPECIES not visible because overlaid by other body parts); R 1 In the British checklist, Menzel (1998) listed several very short, = 0.6–0.7 R and joining c well before base of species that were taken from unchecked literature or fork of M; R and R with dorsal macrotrichia only (R 1 5 1 were incorrect due to wrong interpretation of the lit- with 3–5 macrotrichia and R with 11–14 macrotri- 5 erature. Because of revised material, the following cor- chia); due to R meeting costa early, C very long, = 3/4 5 rections need to be made to Menzel (1998): w. Halteres (Fig. 20) brown, with relatively short stalk and elongated capitulum; capitulum with 6–9 bristles; Bradysia amoena (Winnertz, 1867) = Bradysia tilicola = haltere-capitulum index 1.8–2.0. ABDOMEN: dark (Loew, 1850) [synonymy by Menzel & Heller (2005)] brown; setae of body sparse and fine, relatively short Bradysia brunnipes (Meigen, 1804) = Bradysia pal- and dark. Hypopygium (Fig. 16) wider than high, with lipes (Fabricius, 1787) [synonymy by Heller (2004)] coarse, dark, erect bristles; ventral base of genitalia Bradysia arcana Menzel & Mohrig, 1998 without basal lobe or tuft of setae; inner side of gono- [misidentification] = Bradysia fenestralis (Zetter- coxites widely semicircular with short to moderately stedt, 1838) long setae. Gonostyles longish and pointed; main part Bradysia rufescens (Zetterstedt 1852) of gonostyles 2.2–2.6 times as long as wide; outer side [misidentification] = Bradysia rectinervis Frey, 1948 evenly arched and distinctly rounded apically; with Corynoptera acanthostyla Tuomikoski, 1960 coarse, moderately long apical tooth as elongation of [misidentification] = Claustropyga brevichaeta inner side; apical tooth about 1/3 as long as body of (Mohrig & Antonova, 1978) gonostyles, undivided and pointing inwards and down- Corynoptera dentiforceps (Bukowski & Lengersdorf, wards; apex with lighter, coarse and longer setae; 1936) [misidentification] = Corynoptera recur- inner side of gonostyle, under apical tooth, with 3 hya- vispina Freeman, 1987 line spines which extend to middle of gonostyle Corynoptera inexpectata Tuomikoski, 1960 [recte inex- (Fig. 17); several longer bristles on inner side; long spectata, misidentification] = Corynoptera tetracha- whiplash-like hair missing. Tegmen wide (about 1.3– eta Tuomikoski, 1960 1.5 times as wide as high), membranous and with very Corynoptera subtilis (Lengersdorf, 1929) short basal processes; apex widely rounded; area of [misidentification] = Corynoptera irmgardis (Leng- teeth very small and round with fine-single tipped ersdorf, 1930) teeth. Aedeagus short with wider, weakly sclerotized Leptosciarella truncata (Tuomikoski, 1960) [misiden- base. Body length: 1.0–1.4 mm. tification and incorrect synonymy with nigrosetosa Female. Unknown. (Freeman, 1990)] = Leptosciarella nigrosetosa (Free- man, 1990) restit. (reinstated) [= Leptosciarella Discussion. The new species is placed in the subgenus truncatula Mohrig & Menzel, 1997 syn. nov.] Epidapus s.s. in the Epidapus atomarius-group. This Lycoriella venosa (Staeger, 1840) [misinterpretation of species group is distinguished in particular by trian- the literature (O’Connor & Ashe, 1991)] = Lycoriella gular pointed gonostyles together with a strong and inflata (Winnertz, 1867) coarse tooth and hyaline spines on the apex. Epidapus Sciara flavomarginata Mohrig & Mamaev, 1982 [mis- subgracilis is similar to Epidapus schillei (Börner, identification and incorrectly synomymized from 1903) in the short body, the eyebridge two facets wide, Sciara nursei to Sciara flavomarginata in Menzel the short R1 and the weak bristles on the halteres (1998) and Menzel & Mohrig (2000)] = Sciara nursei (only 6–9 macrotrichia on the capitulum of the hal- Freeman, 1983 restit. [= Sciara ulrichi Menzel & tere). However, because of the longer apical tooth on Mohrig, 1998 syn. nov.] the gonostyles and the coarser bristles on the mesonotum and hypopygium it is also close to Epidapus gracilis (Walker, 1848), but the latter is distinctly larger. BRADYSIA WINNERTZ, 1867 Epidapus gracilis is distinguished from the new spe- BRADYSIA ALBANENSIS (LENGERSDORF, 1926) cies by a wider eyebridge three facets wide, a longer R 1 Literature: Freeman (1983b): 35 [in part]; Menzel and stronger bristles on the halteres (10–18 macrotri- (1998): 20. chia). Epidapus subgracilis is distinguished by several characteristics that cannot be found in either of the Material examined: Beinn a′ Chreachain, Perth & other two species. These are the much shorter necks of Kinross, 1 male, 14.vi.1932, Edwards leg. (BMNH). the flagellomeres, the large, ovoid swollen basal seg- Meall a′ Churain, Stirling, 1 male, 6.vi.1932, Edwards ment of the palpi, the stronger shortened stalk of the leg. (BMNH). halteres and the high, rounded tegmen. Also the inner side of the more slender gonostyles has 3 hyaline Ecological notes: found on open montane habitats and spines in the distal half. at 900–1050 m a.s.l. vi.

BRADYSIA ALPICOLA (WINNERTZ, 1867) don, 1 male, 6.vi.1992, Chandler leg. (PPCM). Slough, Synonyms: caliginosa (Winnertz, 1867); concolor Berkshire, 1 male, 26.vi.1942, Richards leg. (BMNH). − (Beling, 1873); egens (Winnertz, 1867); meridiana Stanford, Norfolk, 1 male, 27.iii. 10.iv.1988, Collier (Lengersdorf, 1926); moreensis (Lengersdorf, 1926); leg. (PPWM). Tomintoul, Moray, 11 males, 12 females, − mutabilis (Lengersdorf, 1926); obscura (Winnertz, 8.viii. 11.viii.1937, Coe leg. (BMNH). 1867); rogenhoferi (Winnertz, 1867). Ecological notes: very common in broad-leaved wood- Literature: Freeman (1983b): 36 [as Bradysia morio, land; also found in aspen woodland; wooded lakeshore; misidentification]; Menzel (1998): 20. mossy rocks by waterfall; wooded riverside; raised bog and bog by lakeshore; calcareous grassland and min- Material examined: Aversley Wood, Cambridgeshire, 1 ing subsidence lake; dunes with sea buckthorn scrub. male, 24.iv.1988, J.H. Cole leg. (PJCB). Ballina, Tip- iii–ix. perary, 1 male, 27.v.1984, J.P. O’Connor leg. (NMID). Bank, New Forest, Hampshire, 1 male, 1 female, vi.1939, Collin leg. (BMNH). Belfast, City of Belfast, 1 BRADYSIA ANGUSTATA TUOMIKOSKI, 1960 male, 17.v.1978, Nash leg. (UMB). Berwick-upon- New to British list. Tweed, Northumberland, 1 male, v.1999; 1 male, vii.1999, all Laurence leg. (PBLN). Brandyquoi, Kirk- Material examined: Great Ellerside, Cumbria, 1 male, wall, 1 male, v.–vi.2001, Laurence leg. (PBLN). Cassop 15.v.1999, Chandler leg. (PPCM). Vale, Durham, 1 male, 13.viii.1986, J.H. Cole leg. Ecological notes: found in broad-leaved woodland. v. (PJCB). Colt Park Wood, Marrick, North Yorkshire, 1 male, 27.viii.−30.viii.1981, Disney leg. (PBLN). Convil Braes, Highland Region, 1 male, 11.viii.1937, Coe leg. BRADYSIA ANGUSTOOCULARIS MOHRIG & (BMNH). Dingwall, Highland Region, 1 female, KRIVOSHEINA, 1989 5.viii.1909; 1 male, 1 female, 16.viii.1909; 1 female, Synonym: luteocoxa Mohrig & Krivosheina, 1989. 26.viii.1909; 1 male, 30.viii.1909, all King leg. (BMNH). Dowlands Cliff, Devon, 20 males, 3 females, New to British list. 9.vi.1937, Edwards leg. (BMNH). Dunton Green, Material examined: Letchworth, Hertfordshire, 1 Kent, 1 female, 1.v.1936, no collector detail (BMNH). male, v.1918, Edwards leg. (BMNH). The Loke, Nor- East Walton Common, Norfolk, 1 male, 24.iv.1989, wich, Norfolk, 1 male, 26.iv.1987; 1 male, 8.v.1987, 1 Stubbs leg. (PPCM). Edge Barton, Devon, 1 male, male, 2.v.−6.v.1992, all Laurence leg. (PBLN). 6.vi.1937, Edwards leg. (BMNH). Forest of Dean, Gloucestershire, 1 female, 27.v.1962, K.G.V. Smith leg. Ecological notes: found in garden. iv–v. (BMNH). Glen Lochay, Stirling, 1 male, 8.vi.1932, Edwards leg. (BMNH). Glendevon, Perth & Kinross, 1 male, 15.viii.1973, Dear leg. (BMNH). Gorge of Avon, BRADYSIA APRICA (WINNERTZ, 1867) South Lanarkshire, 1 male, 20.vi.1909, King leg. Synonym: ingrata var. varicornis (Strobl, 1910). (BMNH). Gullane Bents, East Lothian, 2 males, 1 Literature: Freeman (1983b): 38 [in part; not female, 19.vii.1990, Chandler leg. (NMS, PPCM). fig. 148, = Bradysia subalpina Frey (1948); Laurence Hatherton, Staffordshire, 2 males, 25.viii.1983, Wyatt (1994): 110 [in part] and 118 [in part as Bradysia nit- leg. (BMNH). Invertromie Wood, Insh Marshes, High- idicollis, misidentification]; Menzel (1998): 20. land Region, 1 male, 30.v.2002, Chandler leg. (PPCM). Killarney, Kerry, 1 male, 11.ix.1981, J.P. O’Connor leg. Material examined: Ballynafid Lake, Westmeath, 1 (NMID). Killaun Bog, Offaly, 1 male, ix.2000, Withers male, 10.v.1989, Withers leg. (PPWM). Brancaster, leg. (PPWM). Letchworth, Hertfordshire, 1 male, Norfolk, 9 males, 2.ix.1988, Foster & Procter leg. 1.vii.1934, Edwards leg. (BMNH). Loch Park, Moray, (PBLN). Burnham Beeches, Buckinghamshire, 3 1 female, 12.viii.−14.viii.1937, Coe leg. (BMNH). males, 10.v.−21.vi.1995; 1 male, 13.v.1998, all Ismay Malham Tarn, North Yorkshire, 2 males, 26.vi.− leg. (PPCM). Cothill, Oxfordshire, 6 males, 5.v.1934, 28.vi.1982, Disney leg. (PPWM). Mark Ash Wood, New Edwards leg. (BMNH); 1 male, 3.vi.1990, Chandler Forest, Hampshire, 1 male, 4.vi.1989, Chandler leg. leg. (PPCM). Dickleburgh, Norfolk, 1 male, 21.iv.1989, (PPCM). Moccas Park, Herefordshire, 6 males, Withers leg. (PPWM). Frilford Heath, Oxfordshire, 1 10.vi.2002, Godfrey leg. (HRI, PPCM). Muckross Lake, male, 25.iv.−25.v.1991, Porter leg. (PPCM). Glanvilles Kerry, 1 male, 8.v.1981, Chandler leg. (PPCM). Nairn, Wootton, Dorset, 2 males, viii.1939, Edwards leg. Highland Region, 2 males, 10.vi.1904, no collector (BMNH). Greywell Fen, Hampshire, 1 male, detail (BMNH); 3 males, 1 female, 10.viii.1904, King 11.viii.1999, Chandler leg. (PPCM). Hogley, Oxford- leg. (BMNH). Portland, Dorset, 2 males, 7.vi.1967, shire, 1 male, 1.v.1915, Hamm leg. (BMNH). How Hill, Goddard leg. (PPCM). Richmond Park, Greater Lon- Norfolk, 1 male, 8.iv.1989; 1 male, 1.x.1989 (reared),

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 BRITISH SCIARID FAUNA 39 all Withers leg. (PPWM). Leckford Estate, Hampshire, Ecological notes: found in broad-leaved woodland. viii. 1 male, 4.v.1974, S.A. Williams leg. (PPCM); 1 male, 18.v.1985, Chandler leg. (PPCM). Llangloffan Fen, Pembrokeshire, 1 male, 23.vi.1987, Holmes, Boyce & BRADYSIA AUSTERA MENZEL & HELLER SP. NOV. Reed leg. (PPWM). Lough Gill, Castlegregory, Kerry, 1 New to British and Irish list. male, 13.vii.1990, Withers leg. (PPWM). Mills Marsh, Material examined: Annahala bog, Gearagh, Cork, 1 Norfolk, 1 male, 11.viii.1988, Foster & Procter leg. male, 25.viii.1990, de Courcy Williams leg. [paratype (PBLN). Pass of Killiecrankie, Perth & Kinross, 1 of Bradysia austera] (DEI). Castlemartin Corse, Pem- male, 19.vi.1976, Chandler leg. (PPCM). Portheiddy brokeshire, 1 male, 14.x.1987, Holmes, Boyce & Reed Moor, Pembrokeshire, 1 male, 21.vii.1987, Holmes, leg. (PPWM). Cors Caron, Ceredigion, 1 male, Boyce & Reed leg. (PPWM). Rostherne, Cheshire, 1 15.vi.1987, Holmes, Boyce & Reed leg. (PPWM). Cors male, 30.vii.1932, Britten leg. (BMNH). Stanford, Nor- Caranod, Ceredigion, 1 male, 23.vii.1987, Holmes, folk, 1 male, 27.iii.−10.iv.1988, Collier leg. (PPWM). Boyce & Reed leg. (PPWM). Crymlyn Bog, Swansea/ West Harling Common, Norfolk, 1 male, 30.iii.1991, Abertawe, 1 male, 14.vi.1989, Holmes, Boyce & Reed Withers leg. (PPWM). Wheatfen Broad, Norfolk, 1 leg. (PPWM). Gearagh, Cork, 1 male, 31.vi.1990, de male, 8.x.1988; 1 male, 25.vii.1989, all Withers leg. Courcy Williams leg. [paratype of Bradysia austera] (PPWM). Whites Marsh, Cork, 1 male, 12.v.1989, (NMID). Gwaun Ystrad Caron, Ceredigion, 1 male, Withers leg. (PPWM). Wicken Fen, Cambridgeshire, 1 26.vi.1987, Holmes, Boyce & Reed leg. (PPWM). Llyn male, 12.v.1990, Withers leg. (PPWM). Wortham Ling, Ystumllyn, Gwynedd, 1 male, 8.x.1988, Holmes, Boyce Suffolk, 1 male, 10.v.1987, Withers leg. (PPWM). & Reed leg. [paratype of Bradysia austera] (PPWM). Ecological notes: found in broad-leaved and carr wood- Loch Tummel, Perth & Kinross, 2 males, viii.1939, land; conifer plantation; very common on fens; sedge Edwards leg. (BMNH). Middle Harling, Norfolk, 1 beds; common on wet meadows; sand dunes; salt marsh; male, 28.vi.−12.ix.1988, Foster & Procter leg. (PBLN). garden on reclaimed agricultural land. Reared from the Reedham, Norfolk, 2 males, 27.x.1988, Foster & fungus Leccinum scabrum and mole nest. iii–x. Procter leg. (PBLN). Ecological notes: found in damp broad-leaved and BRADYSIA ARCANA MENZEL & MOHRIG, 1998 mixed woodland; wet heathland; wetlands including fens, water meadows, floodplain mire and bogs. vi–x. Literature: Donisthorpe (1927): 138 [in part as Sciara sp.? (3)]; Menzel & Mohrig (1998): 355; Menzel & Mohrig (2000): 151. BRADYSIA BICOLOR (MEIGEN, 1818) New to British list. Synonyms: abdominalis (Lehmann, 1824); bicolor var. alpestris (Lengersdorf, 1926); bore (Walker, 1848); Material examined: Oxshott, Surrey, 1 male, rufiventris (Macquart, 1834). [26].iv.1901, Donisthorpe leg. (BMNH). Literature: Haliday (1833): 150 [as Sciara]; Walker Ecological notes: found in a site comprising heathland (1848): 105 [as Sciara rufiventris]; Edwards (1915a): and pine woodland; collected with the ant Lasius fulig- 164; Edwards (1925a): 538 [all as Sciara]; Bury (1920): inosus. iv. 250 [as Sciara rufiventris]; Brindle (1963): 83 [as Sci- Taxonomic note: In contradiction to the original ara]; Disney (1976): 92; Freeman (1983b): 34; Menzel description by Menzel & Mohrig (1998) and the ver- (1998): 20. sion in Menzel & Mohrig (2000) the apical spine group Material examined: Aviemore, Highland Region, 1 on the holotype has 10 spines [not 6–7]; they partially male, 11.viii.1903, King leg. (BMNH). Bagley Wood, overlap because of their close grouping. Also the spe- Oxfordshire, 3 males, 27.v.1916, Hamm leg. (UMO). cies has a hook-like tooth above the apical spine group Barton Mills, Suffolk, 1 male, 24.v.1916; 3 males, that is set back on the dorsal side of the gonostyle. In 20.v.1920; 1 female, 26.v.1920; 2 males, 29.v.1935; 2 the holotype the small tooth is only obvious on the left males, 30.v.1935, all Collin leg. (UMO). Beinn Eighe, gonostyle. Bradysia fenestralis (Zetterstedt) does not Highland Region, 1 female, 1953, Richards leg. have this tooth structure. (NMS). Ben Lawers, Perth & Kinross, 1 male, 1 female, 10.vii.1988, Chandler leg. (NMS). Berry BRADYSIA ASCENDA RUDZINSKI, 1994 Pomeroy, Devon, 1 female, 11.x.1980, Chandler leg. (PPCM). Bonhill, West Dunbartonshire, 1 male, New to British list. 8.viii.1908, Malloch leg. (NMS). Braedownie, Angus, Material examined: Blaise Wood, Avon, 1 male, 1 male, 24.vii.1937, Coe leg. (BMNH). Bwlch y 4.viii.1985, Chandler leg. (PPCM). Dolenfawr, Gwynedd, 1 male, 5.vii.1987, Mousley

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 40 F. MENZEL ET AL. leg. (NMS). Cambridge, Cambridgeshire, 1 male, Chandler leg. (HRI). Woodwalton Fen, Cam- 25.v.1904, Jenkinson leg. (BMNH). Camghouran, bridgeshire, 1 female, vi.1939; 1 male, 24.v.1940; 1 Perth & Kinross, 1 male, 21.vii.1990, Chandler leg. male, 7.vi.1940, all Edwards leg. (BMNH). No locality (PPCM). Cairngorm, Highland Region, 1 female, details [? Tullymore Park or Mountains of Mourne], 5.viii.1972, Chandler leg. (PPCM). Cannich, Strath- Down, 3 females, no date, Haliday leg. (NMID). glass, Highland Region, 1 male, 17.vii.1936, Collin leg. Ecological notes: widespread in broad-leaved (incl. (UMO). Carnkilton, Pentland Hills, Midlothian, 1 calcareous and neutral) and ancient pine woodland; female, vii.1924, Wild leg. (NMS). Church Stretton, flushes in open mixed woodland; fen and alder carr; Shropshire, 1 female, 26.viii.1913, Bury leg. (BMNH). bog; marginal vegetation by upland lake; river bank, Comrie, Perth & Kinross, 1 male, 11.vii.1901, A.E.J. pools and grassland; lakeshore; open montane habi- Carter leg. (UMO). Coombe, Cornwall, 1 female, tats. v–x. 16.vi.1989, Chandler leg. (NMS). Coombe Dingle, Avon, 1 male, 22.v.−30.v.1948, d’Assis-Fonseca leg. (PPCM). Cwm Bychan, Gwynedd, 1 female, BRADYSIA BREVISPINA TUOMIKOSKI, 1960 12.vii.1976, Chandler leg. (PPCM). Dalnaglar Castle, New to British list. Perth & Kinross, 1 male, 3.viii.1979, Chandler leg. (NMS). Daneway, Gloucestershire, 1 female, Material examined: Aviemore, Highland Region, 2 12.x.1979, Chandler leg. (PPCM). Dun Fell, Cumbria, males, v.1934, Edwards leg. (BMNH). Bradfield Wood, 1 male, 19.vii.1976, Coulson leg. (PPCM). Fanninghill Suffolk, 1 male, 16.viii.−22.viii.1991, Laurence leg. Loch, Midlothian, 1 male, 18.vii.1990, Chandler leg. (PBLN). Brancaster, Norfolk, 1 male, 2.ix.1988, Foster (PPCM). Folly Farm, Avon, 1 male, 23.x.2002, Gibbs & Procter leg. (PBLN). Buckingham Palace Garden, leg. (PDGB). Forres, Moray, 1 male, 2 females, Greater London, 2 males, 25.iii.−11.iv.1996, D.J. 3.viii.1904, King leg. (BMNH). Glen Lui, Aberdeen- Carter & C.W. Plant leg. (PPCM). Burnham Beeches, shire, 1 female, 12.vii.1977, Chandler leg. (NMS). Buckinghamshire, 1 male, 15.iv.1913, Edwards leg. Glen More, Highland Region, 1 male, 6.viii.1972, (BMNH); 1 male, 30.viii.−15.ix.1995; 18 males, 1.vii.− Chandler leg. (PPCM). Goyt Valley, Derbyshire, 1 26.ix.1996, all Ismay leg. (PPCM). Catfield, Norfolk, 1 male, 18.vii.−20.vii.1939, Edwards leg. (BMNH). male, 12.x.1988, Foster & Procter leg. (PBLN). Chig- Hobb’s Quarry, Gloucestershire, 1 male, 4.ix.2002, well Row Wood, Chigwell Row, Essex, 1 male, 1 female, Gibbs leg. (PDGB). Lancaut Nature Reserve, Glouces- 18.vi.2002; 5 females, 28.vi.2002; 1 female, 15.ix.2002; tershire, 1 female, 5.ix.2002, Gibbs leg. (PDGB). Llan- 2 females, 23.ix.2002; 3 females, 26.ix.2002; 7 females, gammarch Wells, Powys, 1 male, 27.vii.1913, Yerbury 8.x.2002; 2 males, 15 females, 16.x.2002; 2 females, leg. (BMNH). Loch Callater, Aberdeenshire, 8 males, 2 22.x.2002, all Ismay & Schulten leg. (DEI, UMO). females, 28.vii.−29.vii.1937, Coe leg. (BMNH). Mid Coed Tycanol, Pembrokeshire, 1 male, 10.vi.− Quoich, Strathdee, Aberdeenshire, 1 male, 22.ix.2000, 29.vi.1999; 1 male, 29.viii.−31.x.1999, all Levey & Godfrey leg. (HRI). Mildenhall, Suffolk, 1 male, Pavett leg. (PPCM). Denny Wood, New Forest, Hamp- 22.v.1909, Collin leg. (UMO). Monk’s Wood, Cam- shire, 1 male, 17.iv.1954, Colyer leg. (BMNH). Epping bridgeshire, 1 female, 31.v.1970, Chandler leg. Forest, Essex, 2 males, 24.v.-vii.1999, Dagley leg. (PPCM). Nethy Bridge, Highland Region, 1 male, (PPCM). Hafod Garregog, Gwynedd, 1 male, 14.vii.− 12.vii.1906; 1 male, 1 female, 27.vii.1906; 1 female, 8.ix.1999, Levey & Pavett leg. (PPCM). Lea Meadows, 3.vii.1911; 1 female, 20.vii.1913; 1 female, 9.viii.1918; Newtown Linford, Leicestershire, 1 male, 24.viii.2003, 1 male, 4.viii.1919; 1 female, 10.viii.1919, all King leg. Menzel leg. (DEI). Poulton, Cheshire, 1 male, (BMNH, NMS). New Forest, Hampshire, 1 male, 26.iv.1953, Colyer leg. (BMNH). Rhôs-y-gad, Isle of 19.v.1902; 1 male, 29.v.1902; 1 male, 30.v.1910, all Anglesey 1 male, 5.x.1988, Holmes, Boyce & Reed leg. Adams leg. (BMNH). Rannoch, Perth & Kinross, 1 (PPWM). Rough Hill Wood, Studley, Warwickshire, 4 male, 30.vii.1917; 1 male, 31.vii.1917; 1 female, males, 1 female, 10.iv.2003, J.E. Smith leg. (HRI). Sal- 20.viii.1917, all King leg. (BMNH); 1 male, isbury Plain Training Area, Warminster, Wiltshire, 2 21.vii.1990, Chandler leg. (NMS). Rothiemurchus, by males, 3.vii.2003, Ismay & Schulten leg. (DEI, UMO). River Druie, Highland Region, 1 male, 18.viii.1986, Sedlescombe, East Sussex, 1 male, 28.ix.1996, Roper Chandler leg. (NMS). Stoke Wood, Herefordshire, 1 leg. (PPRS). Selsdon, Greater London, 1 male, female, 13.x.1906, Wood leg. (UMO). Strathy Bog, 25.iv.1937, Coe leg. (BMNH). Sutton Park, Birming- Highland Region, 1 male, 31.vii.1972, Chandler ham, West Midlands, 1 male, 4 females, 7.iv.2003, J.E. leg. (PPCM). Tarrington, Herefordshire, 2 females, Smith leg. (HRI). Teddington, Greater London, 1 male, 31.v.1918, Yerbury leg. (UMO). Ufton Fields, Warwick- 1 female, 13.iv.1964, Harris leg. (BMNH). The Loke, shire, 1 female, 9.vi.1991, Chandler leg. (NMS). Vallis Norwich, Norfolk, 1 male, 10.viii.1987, Laurence leg. Vale, Somerset, 1 female, 2.vii.1985, Chandler leg. (PBLN). Woodwalton Fen, Cambridgeshire, 1 male, (PPCM). Windsor Forest, Berkshire, 1 male, 2.ix.1995, 24.v.1940, Edwards leg. (BMNH).

Ecological notes: common in broad-leaved, mixed and 8.v.1987; 1 male, 18.vi.1987; 1 male, 5.vii.1987; 3 carr woodland; ancient woodland with oak, hazel and males, 10.viii.1987; 2 males, 15.ix.1987; 1 male, 24.x.− bramble; heathland; wetlands including wet meadow, 25.x.1987; 1 male, 5.v.1988; 1 male, 18.ix.−20.ix.1988; calcareous fen, sedge bed and river bank; grassland at 1 male, 5.vi.−13.vi.1989; 1 male, 15.ix.−28.ix.1989; 1 woodland fringe; unimproved calcareous grassland; male, 27.vii.1992; 1 male, 27.viii.−1.ix.1992; 1 male, parkland; gardens. iii–x. 24.ix.−3.x.1992; 3 males, 29.ix.−3.x.1992, all Laurence leg. (PBLN). Turville Heath, Buckinghamshire, 2 males, 20.vii.2002, Chandler leg. (PPCM). No locality BRADYSIA BULBOSTYLA MOHRIG & MENZEL, 1990 details, Norfolk, 2 males, 1983, no collector details Literature: Freeman (1987): 202 [as Bradysia amoena, (PPWM). misidentification]; Chandler (2001): 200 [as Bradysia Ecological notes: very common in broad-leaved wood- arcana, misidentification]. land; ancient woodland with oak, hazel and bramble; New to British and Irish list. heathland; grikes; water meadows by river bank with trees; gardens; in stick insect cage. Reared from nests Material examined: Buckingham Palace Garden, of blackbird, thrush, robin. iii–x. Greater London, 1 male, 11.vi.−1.viii.1996; 1 male, 27.iii.−24.iv.1997; 1 male, 19.vi.−18.vii.1997; 1 male, 18.vii.−13.viii.1997, all D.J. Carter & C.W. Plant leg. BRADYSIA CINERASCENS (GRZEGORZEK, 1884) (PPCM). Buckingham Thick Copse, Northampton- shire, 1 male, 8.v.1992, Warne leg. (PPCM). Burnham Synonyms: interstincta (Grzegorzek, 1884); lanicauda Beeches, Buckinghamshire, 1 male, 10.v.−23.v.1995; 1 Tuomikoski, 1960. − male, 31.viii. 13.ix.1996, all Ismay leg. (PPCM). Cas- Literature: Chandler, O’Connor & Ashe (1996): 258 [as tle Neroche, Somerset, 1 male, 12.v.1936, Edwards leg. Bradysia confinis, misidentification]. (BMNH). Chigwell Row Wood, Chigwell Row, Essex, 2 males, 1 female, 13.v.−27.v.2002; 1 male, 1 female, New to British and Irish list. 20.v.2002; 2 males, 11.vi.2002; 28 males, 1 female, Material examined: Arne, Dorset, 1 male, 17.v.− 18.vi.2002; 41 males, 3 females, 28.vi.2002; 4 males, 5.vi.1992, C.W. Plant leg. (PPCM). Bank, New Forest, 15.ix.2002; 3 males, 1 female, 23.ix.2002; 3 males, Hampshire, 1 male, 17.v.1953, Colyer leg. (BMNH). 26.ix.2002; 12 males, 8.x.2002; 6 males, 3 females, Bellananagh, Cavan, 1 male, 3.viii.1981, J.P. & 16.x.2002; 2 males, 2 females, 22.x.2002, all Ismay & M.O’Connor leg. (NMID). Bohuntine, Highland Schulten leg. (DEI, UMO). East Burnham, Bucking- Region, 1 male, 20.vii.1997, Chandler leg. (PPCM). hamshire, 1 male, 15.vi.1931 (reared), Basden leg. Boundless Copse, Surrey, 1 male, 26.vi.2002, Chandler (BMNH). Farnham Royal, Buckinghamshire, 1 male, leg. (PPCM). Camghouran, Perth & Kinross, 1 male, 11.v.1931; 1 male, 8.iv.1932 (all reared), all Basden leg. 31.viii.1987, Chandler leg. (NMS). Hafod Garregog, − (BMNH, NMS). Formoyle, Clare, 1 male, 17.ix. Gwynedd, 6 males, 30.vi.−14.vii.1999; 5 males, 14.vii.− 25.ix.1991, Green leg. (PPCM). Gearagh, Cork, 1 male, 8.ix.1999, all Levey & Pavett leg. (HRI, PPCM). Islau 2.iv.1990, de Courcy Williams leg. (NMID). Gibrid- dref, Gwynedd, 1 male, 10.x.1975, Chandler leg. dings Wood, Staffordshire, 1 male, 14.x.1997, Chan- (PPCM). Glen Affric, Highland Region, 1 male, dler leg. (PPCM). Hartslock, Oxfordshire, 1 male, 1.ix.1984, Chandler leg. (PPCM). Monk Wood, Avon, 1 6.v.2000; 1 male, 14.v.2000, all Raper leg. (PDGB). male, 22.ix.2000, Gibbs leg. (PDGB). Purn Hill, Avon, Hitchin, Hertfordshire, 1 male, ix.1917, Edwards leg. 1 male, 24.v.2002, Gibbs leg. (PDGB). Scotland Fen, (BMNH). Knebworth, Hertfordshire, 1 male, x.1922; 1 Suffolk, 1 male, 19.x.1996, Chandler leg. (PPCM). male, 29.iv.1923, all Edwards leg. (BMNH). Lackham Staverton Thicks, Suffolk, 2 males, 19.v.1996, Chan- Park, Wiltshire, 1 male, 2.vii.2002, Chandler leg. dler leg. (PPCM). Tunstall Common, Suffolk, 1 male, (PPCM). Lancaut, Gloucestershire, 1 male, 5.ix.2002, 19.x.1996, Chandler leg. (PPCM). Gibbs leg. (PDGB). Letchworth, Hertfordshire, 1 male, v.1918, Edwards leg. (BMNH). Oxey Mead, Oxford- Ecological notes: widespread in broad-leaved wood- shire, 1 male, 15.ix.1999, Chandler leg. (PPCM). land (including calcareous); also found in mixed and Oxford, Oxfordshire, 1 male, 10.iv.1926; 1 male, ancient pine woodland; fen; dry heath; calcareous 12.vii.1926 (all reared), all Hamm leg. (BMNH). grassland. v–x. Oxford University Parks, Oxford, Oxfordshire, 2 males, 8.iii.−6.iv.1926 (reared), Hamm leg. (BMNH). Pett’s Wood, Greater London, 1 male, x.1984, Freeman BRADYSIA CONFINIS (WINNERTZ, 1867) leg. (BMNH). Temple, Berkshire, 6 males, 21.iv.− Synonyms: myrtilli (Winnertz, 1867); nigrescens (Win- 30.iv.1934 (reared), Basden leg. (BMNH, NMS). The nertz, 1869); occulta (Winnertz, 1867); sororcula (Win- Loke, Norwich, Norfolk, 1 male, 26.iv.1987; 1 male, nertz, 1867); tarda (Winnertz, 1867).

Literature: Edwards (1925a): 538 [as Sciara]; Free- Literature: Edwards (1925a): 540 [in part as Sciara man (1983b): 35; Chandler (2001): 200; Menzel (1998): praecox, misidentiﬁcation]; Poinar & Doncaster 20. (1965): 73; Binns (1981): 417; Freeman (1983b): 36 [all as Bradysia paupera]; Dunne & O’Connor (1989): 64 Material examined: Alice Holt Forest, Hampshire, 2 [as Bradysia aprica, misidentiﬁcation]; Buxton (1993): males, 10.v.1984, Chandler leg. (PPCM). Bagley 23 [as Bradysia paupera]; Menzel (1998): 20; White, Wood, Oxfordshire, 3 males, 18.v.1936, Edwards leg. Smith & Menzel (2000): 208; Menzel, Smith & Colauto (BMNH). Blean Wood, Kent, 2 males, 19.v.1973, Par- (2003): 451. menter leg. (BMNH). Brampton Wood, Cam- bridgeshire, 1 male, 13.v.1989, J.H. Cole leg. (PJCB). New to Irish list. Brockhurst Wood, Buckinghamshire, 2 males, 2 Material examined: Berwick-upon-Tweed, Northum- females, 5.v.1935, Basden leg. (NMS). Buckingham berland, 1 male, vi.1999; 2 males, ix.1999, all Lau- Palace Garden, Greater London, 1 male, 24.iv.− rence leg. (PBLN). Cheshunt, Hertfordshire, 1 male, 15.v.1997, D.J. Carter & C.W. Plant leg. (PPCM). Cas- 19.v.1939, Speyer leg. (HRI). Evesham, Worcester- tle Neroche, Somerset, 4 males, 12.v.1936, Edwards shire, 1 male, 1996, J.E. Smith leg. (HRI). Glasshouse leg. (BMNH). Charleville Wood, Offaly, 1 male, Crops Research Station, Littlehampton, West Sussex, 27.v.1984, Chandler leg. (PPCM). Clovelly, Devon, 1 1 male, 2 females, 3 larvae, xi.1952–ii.1953; 2 males, 3 male, 8.v.1936, Edwards leg. (BMNH). Corrimony, larvae, i.1953; 2 larvae, 19.i.1953; 5 males, 20.vi.1956; Highland Region, 1 male, 28.v.2002, Chandler leg. 3 males, 10.ix.1956; 4 males, 17.ix.1956; 1 male, (PPCM). Darenth, Kent, 3 males, 5 females, 3.v.1898, 24.ix.1956; 1 male, 1.x.1956; 3 males, 15.x.1956; 1 Grant leg. (BMNH). Epping Forest, Essex, 5 males, male, 23.x.1956; 1 male, 5.xi.1956; 1 male, 2 females, 26.iv.−2.v.1999; 1 male, v.1999, all Dagley leg. (HRI, x.1963, all Parr leg. (HRI); 1 female, 9.i.1964; 1 male, PPCM). Farley Down, Hampshire, 3 males, 20.v.1936, 23.iv.1965; 1 male, 1 female, 5.vii.1965; 2 males, Edwards leg. (BMNH). Fungarth, Perth & Kinross, 1 22.x.1965; 4 males, 1 female, no date, all Gurney leg. male, 30.v.1998, Chandler leg. (PPCM). Hook Com- (HRI); 5 males, 3.ix.1966; 4 males, 1 female, no date, mon, Hampshire, 3 males, 9.v.1976, Chandler leg. all Hussey leg. (HRI); 2 males, 24.ix.1975; 9 males, 2 (PPCM). Knebworth, Hertfordshire, 3 males, 4.v.1923; females, 1975; 5 males, 3 females, 1977; 1 male, 1 male, 6.v.1923; 1 male, 13.v.1923, all Edwards leg. 1.ii.1980; 6 males, 3.v.1982; 29 males, 2 females, no (BMNH). Ledmore Wood, Highland Region, 1 male, date, all Binns leg. (HRI). Kinsealy Research Centre, 28.v.1994, Chandler leg. (PPCM). Loch Achilty, High- Dublin, 2 males, 2 females, 24.ii.−25.ii.1986; 35 males, land Region, 1 male, 28.v.2002, Chandler leg. (PPCM). 15.ix.−26.x.1986 (reared), all Dunne leg. (NMID). Lea Lolly Moor, Norfolk, 1 male, 23.v.1979, Irwin leg. Valley, Hertfordshire, 1 male, 1 female, 20.iii.1965; 1 (PPWM). New Forest, Hampshire, 1 male, 16.v.1968, male, 20.viii.1965, all Butters leg. (HRI). Leamington Parmenter leg. (BMNH). Newmarket, Cambridge- Spa, Warwickshire, 3 males, 1998, J.E. Smith leg. shire, 1 male, 29.iv.1889, Verrall leg. (BMNH). Nure- (HRI). Mid Yell, Shetland, 1 male, ix.1990, Laurence more Lake, Carrickmacross, Monaghan, 2 males, leg. (PBLN). Nazeing, Essex, 7 males, no date, Stevens 20.v.1976, J.H. Cole leg. (PJCB). Savernake Forest, leg. (HRI). Sedlescombe, East Sussex, 1 male, xi.2001, Wiltshire, 1 male, 11.iv.−2.v.1990, Porter leg. (PPCM); Roper leg. (PPRS). Stanford, Norfolk, 1 male, 10.iv.− 3 males, 3.v.1993, Chandler leg. (PPCM). Sedlescombe, 24.iv.1988, Collier leg. (PPWM). Ravensworth, North East Sussex, 1 male, 2.v.1997; 1 male, 3.v.1997, all Yorkshire, 2 males, 5.v.1981, Port leg. (BMNH). The Roper leg. (PPRS). Staffhurst Wood, Surrey, 1 male, Loke, Norwich, Norfolk, 1 male, I.-iii.1990, Laurence 18.v.1991, Chandler leg. (PPCM). Stoke Wood, Buck- leg. (PBLN). Wellesbourne, Warwickshire, 1 male, inghamshire, 1 male, 2 females, 29.iv.1943, Richards 27.i.1998; 5 males, 27.vii.1998; 1 male, 4.viii.2000; 1 leg. (BMNH). Stokeford Heath, Dorset, 1 male, male, xi.2001, all J.E. Smith leg. (HRI, PPCM). No 7.v.2001, Gibbs leg. (PDGB). Windsor Forest, Berk- locality details, England, 11 males, 10 females, 1986, shire, 2 males, 10.v.1939, Edwards leg. (BMNH). White leg. (HRI). Ecological notes: very common in broad-leaved and Ecological notes: common in glasshouses and in house mixed woodland and conifer plantations; sandy heath- plant compost; also found in gardens; grassland at land; lakeside marsh with scrub; fen; formal garden woodland fringe; found damaging cucumbers in pots with wooded fringes and lake; grassland (including [Cheshunt record, Edwards (1925a)]; mushroom farm calcareous). iv–v. in old compost. Reared in laboratory culture. i–xii.

BRADYSIA DIFFORMIS FREY, 1948 BRADYSIA DISTINCTA (STAEGER, 1840) Synonyms: agrestis Sasakawa, 1978; paupera Tuomi- Synonyms: egregia (Beling, 1873); fastuosa (Winnertz, koski, 1960. 1867); insignis (Winnertz, 1867).

Literature: Edwards (1925a): 539 [as Sciara insignis]; (1983a, b) correctly identified in part the extremely Freeman (1983b): 36 [as Bradysia insignis]; Menzel rare species B. fenestralis (Zetterstedt) [nec B. arcana (1998): 20. Menzel & Mohrig, 1998]. A male from BMNH [details: Temple, Berkshire, 23.iv.1932, Basden leg.] was used Material examined: Baldock, Hertfordshire, 1 female, for Fig. 135 in Freeman (1983b). At least with this vi.1918, Edwards leg. (BMNH). Cors Erddreiniog, Isle specimen there is no doubt that it is B. fenestralis and of Anglesey, 2 males, 27.vii.1988; 3 males, 6.x.1988, all not B. arcana. There is another male from BMNH, Holmes, Boyce & Reed leg. (PPWM). Crag Wood, cited in Freeman (1983b: 36) [labelled ‘Kent, Cob- North Yorkshire, 1 male, 18.vi.1920, Cheetham leg. ham’], but the identification is unconfirmed as the (BMNH). Fowl Mere, Norfolk, 1 male, 1 female, genitalia preparation is black. vi.1939, Edwards leg. (BMNH). Ivybridge, Devon, 1 male, 18.v.1893, Yerbury leg. (BMNH). Thompson Common, Norfolk, 1 male, 7.vi.1986, Withers leg. BRADYSIA FLAVIPILA TUOMIKOSKI, 1960 (PPWM). Trawsfynydd, Gwynedd, 1 male, 5.viii.1988, New to British and Irish list. Holmes, Boyce & Reed leg. (PPWM). No locality details, 2 males, 1 female, no date [19th century], Dale Material examined: Annahala bog, Gearagh, Cork, leg. (BMNH). 1 male, 22.iii.1990; 1 male, 15.vi.1990; 1 male, 24.ix.1990, all de Courcy Williams leg. (DEI, NMID). Ecological notes: found in broad-leaved woodland; Brandyquoi, Kirkwall, Orkney, 1 male, 28.v.− wetlands with Cladium, Phragmites, Molinia and 8.vi.2001; 2 males, v.-.vi.2001, all Laurence leg. Myrica; grassland with pingo pools. v–viii, x. (PBLN). Cadair Idris, Gwynedd, 1 male, 13.xi.1993, Laurence leg. (PBLN). Eday, Orkney, 1 male, 1.vi.− 6.vi.1988, Laurence leg. (PBLN). Gearagh, Cork, 1 BRADYSIA DOLOSA LAURENCE, 1994 male, 5.viii.1990, de Courcy Williams leg. (DEI). Literature: Laurence (1994): 112; Menzel (1998): 20; Lundy Island, Devon, 1 male, 1 female, 11.vii.1972, Withers (2002): 114. Hutson leg. (BMNH); 1 male, 20.vii.−27.vii.1972, Lane Material examined: Catfield, Norfolk, 7 [not 8] males, leg. (BMNH). Narborough Bog, Whetstone, Leicester- 12.x.1988, Foster & Procter leg. [holotype and shire, 1 male, 24.viii.2003, J.E. Smith leg. (HRI). − paratypes of Bradysia dolosa] (DEI, PBLN). Foulden, North Ronaldsay, Orkney, 1 male, 4.ix. 6.ix.1989; 2 Norfolk, 12 males, 10.vi.−8.vii.1988, Foster & Procter males, 9.ix.1989, 2 males, ix.1989, all Laurence leg. leg. [paratypes of Bradysia dolosa] (PBLN). Hickling, (PBLN). Salisbury Plain Training Area, Warminster, Norfolk, 1 male, 22.viii.1988, Foster & Procter leg. Wiltshire, 11 males, 4.vii.2003, Ismay & Schulten leg. [paratype of Bradysia dolosa] (PBLN). Killaun Bog, (DEI, UMO). The Loke, Norwich, Norfolk, 1 male, Offaly, 1 male, ix.2000, Withers leg. (PPWM). Middle 4.vii.1987; Laurence leg. (PBLN). Tomintoul, Moray, − Harling, Norfolk, 1 male, 28.vi.−12.vii.1988, Foster & 88 males, 11 females, 8.viii. 11.viii.1937, Coe leg. − Procter leg. [paratype of Bradysia dolosa] (PBLN). (BMNH). Unst, Shetland, 6 males, 30.v. 2.vi.1997, Laurence leg. (PBLN). Vidlin, Shetland, 1 male, Ecological notes: found in broad-leaved and carr wood- 14.vi.−16.vi.1992, Laurence leg. (PBLN). Weisdale, land; wetlands including fen, sedge bed and raised Shetland, 4 males, viii.1996, Laurence leg. (PBLN). bog. vi–x. Wicklow town, Wicklow, 1 male, 19.x.−22.x.1998, Lau- rence leg. (PBLN). Willes Meadow, Leamington Spa, Warwickshire, 1 male, 27.viii.2002, Menzel leg. (DEI). BRADYSIA FENESTRALIS (ZETTERSTEDT, 1838) No locality details, ? Wales, 1 male, ix.1988, Eyre & Synonym: frigida (Winnertz, 1867). Luff leg. (PPWM). Literature: Freeman (1983a): 168; Freeman (1983b): Ecological notes: found in damp broad-leaved wood- 36; Menzel (1998): 20 [as Bradysia arcana, misidenti- land; wet meadow with reed bed on peat; bog; meadow fication]; Menzel & Mohrig (1998): 354; Menzel & flooded in winter; unimproved calcareous grassland; Mohrig (2000): 153; Menzel & Heller (2005): 350. open montane habitats; near shore; gardens. iii, v–xi. Material examined: Temple, Berkshire, 1 male, 23.iv.1932, leg. Basden (BMNH). The Loke, Norwich, BRADYSIA FUNGICOLA (WINNERTZ, 1867) Norfolk, 1 male, 2.v.−6.v.1992, Laurence leg. (DEI). Synonyms: fera (Winnertz, 1867); hercyniae (Win- Ecological notes: found in garden. Reared from narcis- nertz, 1869); incana (Strobl, 1910); ingrata (Winnertz, sus bulbs and nests of moorhen (Freeman, 1983a). iv–v. 1867); sylvicola (Winnertz, 1869). Taxonomic note: Contrary to the literature study by Literature: Menzel (1998): 20; Chandler et al. (2002): Menzel & Mohrig (2000: 151 and 154), Freeman 122.

Material examined: Alice Holt Forest, Hampshire, 1 (PPCM). Lea Meadows, Newtown Linford, Leicester- male, 23.vii.1999, Chandler leg. (PPCM). Alvecote shire, 3 females, 24.viii.2003, Menzel leg. (DEI). Leck- Pools, Staffordshire, 4 males, 13.x.2001, J.E. Smith ford Estate, Hampshire, 1 male, 5.v.1984; 1 male, leg. (HRI). Annahala bog, Gearagh, Cork, 2 males, 1.ix.1999, all Chandler leg. (PPCM). Loch Achilty, 1.vi.1990, de Courcy Williams leg. (DEI, NMID). Highland Region, 1 male, 28.v.2002, Chandler leg. Ashridge Estate, Hertfordshire, 2 males, 8.vii.1999, (PPCM). Loch Loy, Highland Region, 1 male, Chandler leg. (PPCM). Aviemore, Highland Region, 1 14.vii.1984, Chandler leg. (PPCM). Monk’s Wood, male, v.1934, Edwards leg. (BMNH). Bix Bottom, Muckross, Kerry, 1 male, 4.v.1981, Chandler leg. Oxfordshire, 2 males, 23.ix.2000, Raper leg. (PDGB). (PPCM). Narborough Bog, Whetstone, Leicestershire, Bolt Wood, Stockerston, Leicestershire, 1 male, 1 1 male, 24.viii.2003, Menzel leg. (DEI). New Close female, 24.viii.2003, Menzel leg. (DEI). Boundless Wood, Brandon, Warwickshire, 1 male, 28.viii.2003, Copse, Surrey, 1 male, 26.vii.2002, Chandler leg. Menzel leg. (DEI). New Forest, Hampshire, 1 male, (PPCM). Brancaster, Norfolk, 1 male, 1.vii.− 17.x.1937, Edwards leg. (BMNH). Newbattle Abbey, 15.vii.1988, Foster & Procter leg. (PBLN). Burnham Midlothian, 3 males, 6.vi.1994 (reared), Rotheray leg. Beeches, Buckinghamshire, 1 male, 15.ix.−4.x.1995; 1 (NMS). Nuremore Lake, Carrickmacross, Monaghan, male, 22.vii.−5.viii.1996; 1 male, 17.v.−25.v.1999, all 2 males, 20.v.1976, J.H. Cole leg. (PJCB). Paradize Ismay leg. (PPCM). Caerhays Castle, Cornwall, 1 Wood. Herefordshire, 1 male, 9.x.1998, Chandler leg. male, 2.vii.1988, Chandler leg. (PPCM). California (PPCM). Redgrave Lake, Suffolk, 1 male, 24.v.1987, Country Park, Berkshire, 1 male, 27.vii.1996, Chan- Withers leg. (PPWM). Rhôs Rydd, Ceredigion, 1 male, dler leg. (PPCM). Castle Eden Dene, Durham, 1 male, 25.ix.1986, Holmes, Boyce & Reed leg. (PPWM). Ring- 14.viii.1990, J.H. Cole leg. (PJCB). Charleville Wood, stead, Dorset, 1 male, viii.1939, Edwards leg. Offaly, 1 male, 19.ix.1999, Chandler leg. (PPCM). (BMNH). Rosevear, Cornwall, 1 male, 6.vii.1983, Chester, Cheshire, 1 male, 27.ii.1953, Gordon-Smith Chandler leg. (PPCM). Rotherfield Park, Hampshire, leg. (BMNH). Chippenham Fen, Cambridgeshire, 1 1 male, 3.vii.2002; 1 male, 2.x.2002, all Chandler male, 2.vii.1988, Chandler leg. (PPCM). Clovelly, leg. (PPCM). Savernake Forest, Wiltshire, 1 male, Devon, 1 male, 8.v.1936, Coe leg. (BMNH). Collinpark 3.v.1993, Chandler leg. (PPCM). Scargill, Durham, 1 Wood, Gloucestershire, 1 male, 3.ix.2002, Gibbs leg. male, 5.viii.1979, Chandler leg. (PPCM). Sedlescombe, (PDGB). Coombe Abbey Country Park, Coventry, War- East Sussex, 1 male, 11.ix.1999, Roper leg. (PPRS). wickshire, 1 male, 28.viii.2003, J.E. Smith leg. (HRI). Sidmouth, Devon, 1 male, 10.v.1936, Edwards leg. Cuckfield, West Sussex, 1 male, 15.i.1969 (reared), (BMNH). Spartum Fen, Oxfordshire, 1 male, Lane leg. (BMNH). Cuckoo Wood, Downe, Greater 15.x.1999, Chandler leg. (PPCM). Spey Bay, Moray, 1 London, 1 male, 20.vii.1985, Chandler leg. (PPCM). male, 2.vii.1936, Coe leg. (BMNH). Stanford, Norfolk, Culbin Sands, Highland Region, 1 male, 5.vii.− 1 male, 27.iii.−10.iv.1988, Collier leg. (PPWM). Stony- 7.vii.1936, Coe leg. (BMNH). Dinton Pastures Country brow Wood, Rotherfield Park, Hampshire, 1 male, Park, Berkshire, 1 male, 20.xi.1992, Chandler leg. 2.x.2002, Chandler leg. (PPCM). The Loke, Norwich, (PPCM). Ditchley, Oxfordshire, 1 male, v.1937, Norfolk, 1 male, 26.iv.1987; 1 male, 15.v.−18.v.1992, all Edwards leg. (BMNH). Dowdeswell Wood, Gloucester- Laurence leg. (PBLN). The Spittles, Dorset, 1 male, shire, 1 male, 16.ix.2001, Gibbs leg. (PDGB). Epping 2.vii.1998, Perry leg. (PPCM). Torboll Wood, Highland Forest, Essex, 10 males, vii.–ix.1998; 1 male, 1999, all Region, 1 male, 26.v.2002, Chandler leg. (PPCM). Dagley leg. (PPCM). Felbrigg Wood, Norfolk, 1 male, Udale Bay, Highland Region, 1 male, 29.v.2002, 10.vii.1993, Chandler leg. (PPCM). Folly Farm, Avon, Chandler leg. (PPCM). Uragh Wood, Kerry, 1 male, 1 male, 25.vii.2001, Gibbs leg. (PDGB). Frithsden 8.vii.1990, Withers leg. (PPWM). West End Common, Beeches, Hertfordshire, 1 male, 16.x.1997, Chandler Esher, Surrey, 1 male, 19.v.2002, Chandler leg. leg. (PPCM). Gearagh, Cork, 1 male, 7.iv.1990; 1 male, (PPCM). Wheatfen Broad, Norfolk, 1 male, 8.x.1988, 11.v.1990; 1 male, 12.vi.1990; 1 male, 30.vi.1990; 1 Withers leg. (PPWM). Wicken Fen, Cambridgeshire, 1 male, 5.viii.1990; 1 male, 8.ix.1990; 1 male, 26.ix.1990, male, 12.v.1990, Withers leg. (PPWM). Woodbastwick, all de Courcy Williams leg. (DEI, NMID). Glen Coiltie, Norfolk, 1 male, viii.1932, Edwards leg. (BMNH). Highland Region, 1 male, 28.v.2002, Chandler leg. Wychwood, Oxfordshire, 1 male, v.1937, Edwards leg. (PPCM). Goring-on-Thames, Oxfordshire, 1 male, (BMNH). Wytham Woods, Wytham, Oxfordshire, 1 19.iv.1962, J.H. Cole leg. (PJCB). Grange-over-Sands, male, 21.viii.2002, Menzel leg. (DEI). No locality Cumbria, 1 male, 20.iii.1996, Birkett leg. (PPCM). details, Wales, 1 male, no date, Holmes, Boyce & Reed Highgate Wood, Greater London, 1 male, 1998–99, leg. (PBLN). Hackett leg. (PPCM). Hitch Wood, Hertfordshire, 1 male, 26.ix.1937, Edwards leg. (BMNH). Killaun Bog, Ecological notes: very common in broad-leaved and Offaly, 1 male, ix.2000, Withers leg. (PPWM). Lack- carr woodland; conifer plantations; ancient pine wood- ham Park, Wiltshire, 1 male, 2.vii.2002, Chandler leg. land; wetlands including water meadows, sedge bed,

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 BRITISH SCIARID FAUNA 45 wooded river banks and lake shores, bogs; grassland (BMNH); 1 male, 4.vi.1977, J.H. Cole leg. (PJCB). No (including calcareous); gardens; old hedges; salt- locality details, Devon, 2 males, 4 females, iv.1942, marsh; dunes; coastal clay cliffs. Reared from fallen Perkins leg. (BMNH). beech trunk and rotten wood. ii–xi. Ecological notes: common in broad-leaved and carr woodland; mixed drier woodland; fen; Cladium beds BRADYSIA GIRAUDII (EGGER, 1862) with tufa; herb-rich grassland; grassland with pingo pools; water meadows and gravel pits; wooded river Synonyms: clavigera (Lengersdorf, 1926); nemorum bank; gardens; meadow hedge. Reared from dead stem (Winnertz, 1867). of foxglove, decaying pith of old thistle stems, dead Literature: Freeman (1983b): 36; Chandler et al. stem of agrimony, stem of old nettles, stem of common (1996): 259 [all as Bradysia giraudi (Schiner) [recte mallow, thistle stem, rotten wood. iv–x. giraudii (Egger)]; Menzel (1998): 20.

Material examined: Bayfordbury Estate, Hertford, BRADYSIA HELLERI MENZEL & MOHRIG, 1998 Hertfordshire, 1 male, 10.vi.1975, J.H. Cole leg. Literature: Laurence (1994): 118; Laurence (1996): 87 (PJCB). Bickley, Bromley, Greater London, 2 males, [all as Bradysia brunnipes, misidentification]. 12.v.1978 (reared), Dear leg. (BMNH). Bishop’s Stort- ford, Hertfordshire, 1 male, vii.1993, C.W. Plant leg. New to British list. (HRI). Buckingham Palace Garden, Greater London, 1 Material examined: The Loke, Norwich, Norfolk, 2 male, 23.v.−6.vi.1998, D.J. Carter & C.W. Plant leg. males, 8.v.1987; 1 male, 15.vi.−19.vi.1987, all Lau- (PPCM). Byfleet, Surrey, 1 male, 14.v.1932, Coe leg. rence leg. (PBLN). (BMNH). California Country Park, Berkshire, 1 male, 8.vii.1998, Chandler leg. (PPCM). Cothill, Oxford- Ecological notes: found in garden. v–vi. shire, 1 male, 7.v.1972, Chandler leg. (PPCM). Dinton Pastures Country Park, Berkshire, 1 male, 9.vii.1994, Chandler leg. (PPCM). Ennis, Clare, 1 male, 30.v.1984, BRADYSIA HEYDEMANNI (LENGERSDORF, 1955) J.P. O’Connor leg. (NMID). Failford, by River Ayr, New to British list. South Ayrshire, 1 male, 3.vii.1995, Chandler leg. Material examined: Burnham Beeches, Buckingham- (PPCM). Goring-on-Thames, Oxfordshire, 1 male, shire, 2 males, 7.vi.−21.vi.1995; 10 males, 26.v.− 15.v.1962, J.H. Cole leg. (PJCB). Hogley, Oxfordshire, 26.xi.1996, all Ismay leg. (HRI, PPCM). Cadair Idris, 1 male, 25.iv.1924 (reared), Hamm leg. (UMO). Lack- Gwynedd, 1 male, 4.ix.1993, Laurence leg. (PBLN). ham Park, Wiltshire, 1 male, 2.viii.2002, Chandler leg. Epping Forest, Essex, 1 male, x.1998, Dagley leg. (PPCM). Leckford Estate, Hampshire, 1 male, (PPCM). Stanford battle area, Norfolk, 1 male, 27.iii.− 6.vii.1991, Chandler leg. (PPCM). Mark Ash Wood, 10.iv.1988, Withers leg. (PPWM). New Forest, Hampshire, 1 male, 2.vi.1984, Chandler leg. (NMS). Newbridge Fen, Kildare, 1 male, 8.viii.− Ecological notes: found in ancient broad-leaved wood- 19.viii.1998, Good leg. (PPWM). Oversley, Warwick- land and grassland; open montane habitats. iii–xi. shire, 1 male, ix.1969; 1 male, 4 females, x.1969 (reared), all Evans leg. (BMNH). Rostherne, Cheshire, 1 female, 29.v.1932, Britten leg. (BMNH). Runnymede BRADYSIA HILARIFORMIS TUOMIKOSKI, 1960 Meadow, Surrey, 1 male, 2.vii.1990, Chandler leg. Literature: Freeman (1983b): 38 [as Bradysia hilaris, (NMS). Scadbury Park, Greater London, 3 males, misidentification]; Menzel (1998): 20. 21.vii.1975, Chandler leg. (PPCM). Seaton, Devon, 1 New to Irish list. male, v.1937 (reared), Blair leg. (BMNH). The Loke, Norwich, Norfolk, 1 male, 12.vii.1987; 1 male, 15.v.− Material examined: Annahala bog, Gearagh, Cork, 1 18.v.1992, all Laurence leg. (PBLN). Theale, Berk- male, 1.vi.1990, de Courcy Williams leg. (NMID). shire, 1 male, 7.vii.2003, Chandler leg. (HRI). Thomp- Brownsham, Devon, 1 male, 18.v.1987, Chandler son Common, Norfolk, 1 male, 11.vii.1997, Chandler leg. (PPCM). Ditchley, Oxfordshire, 1 male, v.1937, leg. (PPCM). Toxside Hill, Midlothian, 1 male, 4 Edwards leg. (BMNH). Falls of Tarnash, Moray, 1 females, 1.xi.−9.xi.2001 (reared), Bland leg. (NMS). male, 1.vii.1936, Coe leg. (BMNH). Gearagh, Cork, 2 Walton Moor, Gordano, Avon, 3 males, 6 females, males, 30.vi.1990, de Courcy Williams leg. (DEI). Glen 20.iv.1952, d’Assis-Fonseca leg. (NMS, PPCM). Wel- Coiltie, Highland Region, 1 male, 20.v.2002, Chandler bourne, Norfolk, 1 male, 18.viii.1983 (reared), Evans leg. (PPCM). Lackham Park, Wiltshire, 4 males, leg. (PPWM). Windbury Wood, Devon, 1 male, 20.ix.2002, Chandler leg. (PPCM). Leckford, Hamp- 18.v.1980, Chandler leg. (NMS). Woodwalton Fen, shire, 1 male, 25.vii.1998, Chandler leg. (PPCM). Mal- Cambridgeshire, 1 male, 24.v.1940, Edwards leg. ham Tarn, North Yorkshire, 1 male, 3.vi.−5.vi.1982,

Disney leg. (PPWM). Savernake Forest, Wiltshire, 1 13.vii.1989; 4 males, 3.x.1989, all Holmes, Boyce & male, 3.v.1993, Chandler leg. (PPCM). Tre Wilmot, Isle Reed leg. (PPWM). Llyn Coethlyn, Powys, 2 males, of Anglesey, 1 male, 14.vi.1988, Holmes, Boyce & Reed 26.x.1988, Holmes, Boyce & Reed leg. (PPWM). Llyn leg. (PPWM). Wychwood, Oxfordshire, 1 male, v.1937, Creiniog, Denbighshire, 1 male, 6.vii.1988; 1 male, Edwards leg. (BMNH). Wytham Woods, Wytham, 13.x.1988; 1 male, 27.x.1988, all Holmes, Boyce & Oxfordshire, 5 males, 2 females, 21.viii.2002, Menzel Reed leg. (PPWM). Llyn Tyn y Mynydd, Gwynedd, 1 & J.E. Smith leg. (DEI, HRI). male, 20.vii.1988; 3 males, 18.viii.1988, all Holmes, Boyce & Reed leg. (PPWM). Merddwr, Denbighshire, 1 Ecological notes: widespread in broad-leaved wood- male, 3.viii.1988, Holmes, Boyce & Reed leg. (PPWM). land; also found in bogs; lakeshore; basin mire. v–ix. Magor Marsh, Monmouthshire, 1 male, 28.ix.1988, Holmes, Boyce & Reed leg. (PPWM). Nûg, Conwy, 1 BRADYSIA HILDAE HELLER, 2000 male, 12.x.1988, Holmes, Boyce & Reed leg. (PPWM). Rhôs Cruglas, Carmarthen, 1 male, 3.x.1989, Holmes, New to British list. Boyce & Reed leg. (PPWM). Rhôs LLawr Cwrt, Cere- Material examined: Bromley, Greater London, 1 male, digion, 3 males, 6.x.1987, Holmes, Boyce & Reed leg. 11.vii.1953, Biggs leg. (NMS). Cadair Idris, Gwynedd, (PPWM). Rhôsili Down, Swansea/Abertawe, 2 males, 1 male, 22.x.1993, Laurence leg. (PBLN). Epping 12.vii.1989; 1 male, 3.x.1989, all Holmes, Boyce & Forest, Essex, 1 male, iv.-v.1999, Dagley leg. (PPCM). Reed leg. (PPWM). Trawsfynydd, Gwynedd, 1 male, Letchworth, Hertfordshire, 1 male, iii.1936; 1 male, 7.vii.1988, Holmes, Boyce & Reed leg. (PPWM). vi.1936, all Edwards leg. (BMNH). Ecological notes: found in broad-leaved woodland; Ecological notes: found in ancient broad-leaved wood- common on wetlands in Wales including damp heathland; grassland; open montane habitats. iii–vii, x. land, fens, bogs, valley and basin mires; grazed meadow. vi–x.

BRADYSIA HOLSATICA HELLER, 2004 New to British list. BRADYSIA INUSITATA TUOMIKOSKI, 1960 Literature: Freeman (1983a): 169; Freeman (1983b): Material examined: Blaen y Wergloedd, Conwy, 1 34; Laurence (1997b): 228; Menzel (1998): 20. male, 4.viii.1988; 1 male, 13.x.1988, all Holmes, Boyce & Reed leg. (PPWM). Brynberian Moor, Pem- New to Irish list. brokeshire, 1 male, 12.x.1987, Holmes, Boyce & Reed Material examined: Balmaha, Stirling, 1 male, leg. (PPWM). Cefn Grwhyd, Neath/Port Talbot, 1 male, 5.vii.1909, King leg. (BMNH). Burnham Beeches, 12.vii.1989, Holmes, Boyce & Reed leg. (PPWM). Col- Buckinghamshire, 1 male, 10.v.−23.v.1995, Ismay leg. wyn Brook, Powys, 1 male, 18.vii.1989, Holmes, Boyce (PPCM). Crickhowell, Powys, 1 male, v.1937, Edwards & Reed leg. (PPWM). Comin Esgair Maen, Ceredigion, leg. (BMNH). Dickleburgh, Norfolk, 1 male, 5.viii.1989, 1 male, 15.x.1987, Holmes, Boyce & Reed leg. Withers leg. (PPWM). Eday, Orkney, 1 male, 30.v.− (PPWM). Coombe Abbey Country Park, Coventry, 31.v.1988; 9 males, 1.vi.−6.vi.1988, all Laurence leg. Warwickshire, 1 male, 28.viii.2003, Menzel leg. (DEI). (PBLN). Gearagh, Cork, 1 male, 10.vi.1990; 1 male, Cors Caron, Ceredigion, 1 male, 27.vii.1987, Holmes, 12.vi.1990, all de Courcy Williams leg. (DEI, NMID). Boyce & Reed leg. (PPWM). Cors Caranod, Ceredigion, Holt Island, St. Ives, Cambridgeshire, 1 male, 1 male, 15.vi.1987; 2 males, 1.x.1987, all Holmes, 13.ix.2002, J.H. Cole leg. (PJCB). Malham Tarn, North Boyce & Reed leg. (PPWM). Cors Erddreiniog, Isle of Yorkshire, 1 male, 2.vi.−3.vi.1982, no collector detail Anglesey, 2 males, 14.vii.1988, Holmes, Boyce & Reed (BMNH). Milham Ford School, Oxford, Oxfordshire, 1 leg. (PPWM). Cors Farlais, Carmarthen, 1 male, male, 18.v.2001, Webb leg. (PPCM). North Ronaldsay, 28.vi.1989, Holmes, Boyce & Reed leg. (PPWM). Cors Orkney, 1 male, 1.ix.1989, Laurence leg. (PBLN). Oak- Geirch, Gwynedd, 1 male, 28.vii.1988, Holmes, Boyce land Wood, Cambridgeshire, 1 male, 1.v.−31.v.1998, & Reed leg. (PPWM). Cors Graianog, Gwynedd, 1 C.W. Plant leg. (PPCM). Pawlett Levels, Somerset, 1 male, 28.vii.1988, Holmes, Boyce & Reed leg. (PPWM). male, 13.ix.1999, Gibbs leg. (PDGB). Stanford, Norfolk, Dyffryn Gwaun, Pembrokeshire, 1 male, 6.vii.1987; 1 1 male, 27.iii.−10.iv.1988, Collier leg. (PPWM). Stradey male, 20.vii.1987; 1 male, 26.ix.1987; 1 male, Wood, Carmarthen, 1 male, 13.x.1995, Chandler leg. 28.ix.1987, all Holmes, Boyce & Reed leg. (PPWM). (PPCM). Vidlin, Shetland, 4 males, 14.vi.−16.vi.1992, Fairwood Common, Swansea/Abertawe, 1 male, Laurence leg. (PBLN). Wicken Fen, Cambridgeshire, 1 4.x.1989, Holmes, Boyce & Reed leg. (PPWM). Illtyd male, 25.ix.1990, J.H. Cole leg. (PJCB). Pools, Powys, 1 male, 5.vii.1989; 1 male, 19.vii.1989; 1 male, 11.x.1989, all Holmes, Boyce & Reed Ecological notes: found in broad-leaved woodland; wil- leg. (PPWM). Llanon, Swansea/Abertawe, 1 male, low carr; bog and lakeshore; fen meadows and ditches;

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 BRITISH SCIARID FAUNA 47 grazing marshes; school grounds with old poplars; BRADYSIA LAPPONICA (LENGERSDORF, 1926) near shore; gardens. iii–x. Synonyms: nigerrima (Lengersdorf, 1940); pseu- dopraecox Frey, 1948; quinquedentata (Lengersdorf, 1936). BRADYSIA ISMAYI MENZEL SP. NOV. Literature: Menzel (1998): 20. New to British list. Material examined: Cadair Idris, Gwynedd, 1 male, Material examined: Beaulieu River, Hampshire, 1 1.viii.1993; 1 male, 16.viii.1993, all Laurence leg. male, 22.v.1936, Edwards leg. (BMNH). Coed Tycanol, (PBLN). Coed y Gelli, Carmarthen, 2 males, Pembrokeshire, 2 males, 10.vi.−29.vi.1999, Levey & 12.x.1995, McLean leg. (PPCM). Cotterdale, North Pavett leg. [paratypes of Bradysia ismayi] (HRI, Yorkshire, 1 male, 22.ix.1986, J.H. Cole leg. (PJCB). PPCM). Chigwell Row Wood, Chigwell Row, Essex, 1 Hafod Garregog, Gwynedd, 1 male, 16.vi.−30.vi.1999, male, 23.v.−27.v.2002; 1 male, 11.vi.2002; 1 male, Levey & Pavett leg. (HRI). Loch Callater, Aberdeen- 18.vi.2002; 1 male, 20.vi.2002, all Ismay & Schulten shire, 1 male, 22.vii.1938, Coe leg. (BMNH). Loch leg. [holotype and paratypes of Bradysia ismayi] Tummel, Perthshire, 1 male, 8.vii.1933, Collin leg. (DEI). Epping Forest, Essex, 1 male, v.1999, Dagley (BMNH). Loch Ussie, Highland Region, 1 male, leg. [paratype of Bradysia ismayi] (PPCM). Hafod 18.vi.1976, Chandler leg. (PPCM). Strathrory, High- Garregog, Gwynedd, 1 male, 18.ix.−28.ix.1999, Levey land Region, 1 male, 15.vi.1976, Chandler leg. & Pavett leg. [paratype of Bradysia ismayi] (PPCM). (PPCM). Letchworth, Hertfordshire, 1 male, ix.1936, Edwards leg. (BMNH). Migdale Loch, Highland Region, 1 male, Ecological notes: found in broad-leaved and mixed 31.v.2002, Chandler leg. [paratype of Bradysia ismayi] woodland; riverbank with alders; open montane hab- (PPCM). Milham Ford School, Oxford, Oxfordshire, 1 itats. v–x. male, 18.v.−22.v.2001, Webb leg. [paratype of Bradysia ismayi] (PPCM). The Loke, Norwich, Norfolk, 1 male, 21.viii.−24.viii.1989; 1 male, 28.iv.−4.v.1990; 1 male, BRADYSIA LEMBKEI MOHRIG & MENZEL, 1990 21.viii.1992, all Laurence leg. [paratypes of Bradysia Literature: Freeman (1983b): 35 [in part as Bradysia ismayi] (PBLN). albanensis, misidentification]. Ecological notes: found in broad-leaved woodland; New to British list. ancient woodland with oak, hazel and bramble; pine woodland on higher ground; school grounds with old Material examined: Glen of Drumloch, Moray, 1 male, poplars; alder and birch carr by stream; heathland 16.viii.1937, Coe leg. (BMNH). Harpenden, Hertford- (heather) with oak and holly; garden. v–vi, viii–ix. shire, 1 male, 7.x.1934, C.B. Williams leg. (BMNH). Shefford, Bedfordshire, 1 male, x.1917, Edwards leg. (BMNH). The Loke, Norwich, Norfolk, 1 male, BRADYSIA KASSEBEERI HELLER, 1998 26.iv.1987, Laurence leg. (PBLN). Tomintoul, Moray, 1 − Literature: Laurence (1994): 110 [in part as Bradysia male, 8.viii. 11.viii.1937, Coe leg. (BMNH). Tresco, aprica and Bradysia fungicola, misidentification]. Scilly Isles, Cornwall, 1 male, 6.x.1970, Hutson leg. (BMNH). New to British list. Ecological notes: found in garden and at 335 m a.s.l. iv, Material examined: Cadair Idris, Gwynedd, 6 males, viii, x. 13.x.−19.xii.1988, Laurence leg. (DEI). Catfield, Nor- folk, 3 males, 29.ix.−12.x.1988; 19 males, 13.x.− 12.xii.1988, all Foster & Procter leg. (PBLN). Dowrog BRADYSIA LEPTOPTERA TUOMIKOSKI, 1960 Common, Pembrokeshire, 1 male, 13.x.1987, Holmes, Literature: Chandler et al. (2002): 122 [in part Boyce & Reed leg. (PPWM). Gatewen Marsh, Den- Bradysia normalis, misidentification]. bighshire, 4 males, 3.viii.1988, Holmes, Boyce & Reed leg. (PPWM). Reedham, Norfolk, 1 male, 21.x.1988, Material examined: Ailladie, Clare, 1 male, 24.v.1985, Foster & Procter leg. (PBLN). Sutton, Norfolk, 1 male, J.P. & M.A. O’Connor leg. (NMID). California Country 9.vi.−23.vi.1988, Foster & Procter leg. (PBLN). Park, Berkshire, 1 male, 19.v.1998, Chandler leg. Thompson Common, Norfolk, 1 male, 13.ix.− (PPCM). North Ronaldsay, Orkney, 1 male, 9.ix.− 27.ix.1988, Foster & Procter leg. (PBLN). 10.ix.1989, Laurence leg. (PBLN). Tomintoul, Moray, 1 male, 8.vii.−11.vii.1937, Coe leg. (BMNH). Ecological notes: found in carr woodland and fen including Carex, Typha latifolia beds; open montane Ecological notes: found in carr and mixed drier wood- habitats. vi, viii–xii. land; sea shore on island. v, vii, ix.

BRADYSIA LEUCOPEZA MOHRIG & MAMAEV, 1989 Ecological notes: widespread in broad-leaved wood- Literature: Menzel (1998): 20. land; also found in mixed woodland; alder and birch carr by stream; pine woodland on higher ground; gar- Material examined: Cors Goch, Isle of Anglesey, 1 den. v–ix. male, 28.vii.1988, Holmes, Boyce & Reed leg. (PPWM). Gors Llwyn, Neath/Port Talbot, 1 male, 28.vi.1989, Holmes, Boyce & Reed leg. (PPWM). Illtyd Pools, BRADYSIA LONGICUBITALIS (LENGERSDORF, 1924) Powys, 1 male, 19.vii.1989, Holmes, Boyce & Reed leg. Synonym: cinereovittata Frey, 1948. (PPWM). Literature: Edwards (1938): 201 [in part as Sciara Ecological notes: found on wetlands including mire (Neosciara) brunnipes, misidentiﬁcation]; Freeman with Juncus, fens with Phragmites and Carex, Cla- (1983a): 168 [as Bradysia cinereovittatum (recte dium bed. vi, vii. cinereovittata)]; Freeman (1983b): 35 [as Bradysia cinereovittata]; Menzel (1998): 20.

BRADYSIA LOBATA HONDRU, 1968 Material examined: Craigmore Woods, Aberfoyle, New to British and Irish list. Stirling, 1 male, 22.vi.1999, Chandler leg. (NMS). Crymlyn Bog, Swansea/Abertawe, 1 male, 19.vi.1906, Material examined: Chilbolton Common, Hampshire, Yerbury leg. (BMNH). Duddleswell, East Sussex, 1 1 male, 6.vii.1991, Chandler leg. (PPCM). Gearagh, male, 9.x.1976, Chandler leg. (PPCM). Fungarth, Cork, 1 male, 7.iv.1990, de Courcy Williams leg. Perth & Kinross, 1 male, 30.v.1998, Chandler leg. (DEI). Glyn Ceiriog, Wrexham, 1 male, 26.vi.− (PPCM). Harpenden, Hertfordshire, 14 males, 1934, 7.vii.1995, Brock leg. (PPCM). Moy, Tyrone, 1 male, C.B. Williams leg. (BMNH). Loch Chon/Loch Ard, 24.ix.−30.ix.1984, Boston leg. (PPCM). Woolcombe Stirling, 1 male, 22.vi.1999, Chandler leg. (PPCM). Meadow, Dorset, 1 male, 3.vii.1998, Chandler leg. Migdale Wood, Highland Region, 1 male, 31.v.2002, (PPCM). Chandler leg. (PPCM). Rhôs Rydd, Ceredigion, 1 male, Ecological notes: found in damp broad-leaved wood- 25.ix.1986, Holmes, Boyce & Reed leg. (PPWM). land; water meadows; acid grassland. iv, vi–vii, ix. Sherrard’s Wood, Welwyn, Hertfordshire, 1 male, 22.x.1936, Edwards leg. (BMNH). Ecological notes: found in broad-leaved and mixed BRADYSIA LOBULIFERA FREY, 1948 woodland; pine woodland on higher ground; conifer Literature: Edwards (1938): 201 [as Sciara (Neosciara) plantations with stream; alder and birch carr by pullula, misidentification]; Freeman (1983b): 35; Lau- stream; damp heath with Carex. v–vi, x. rence (1997b): 228 [all as Bradysia pullula, misidentification]; Menzel (1998): 20 [in part as Bradysia rufescens, misidentification]. BRADYSIA LONGISTYLIA M OHRIG & KRIVOSHEINA, 1982 Synonym: longistyla Mohrig & Krivosheina, 1989. Material examined: Birnam Hill, Perth & Kinross, 1 male, 13.vii.1991, Chandler leg. (PPCM). Brandyquoi, New to British list. Orkney, 1 male, v.-vi.2001, Laurence leg. {PBLN). Material examined: Burnham Beeches, Buckingham- Crickhowell, Powys, 1 male, v.1937, Edwards leg. shire, 1 male, 9.x.−26.xi.1996, Ismay leg. (PPCM). (BMNH). Failford, by River Ayr, South Ayrshire, 1 Lackham Park, Wiltshire, 1 male, 14.x.2002, Chandler male, 3.vii.1995, Chandler leg. (PPCM). Falls of Tar- leg. (PPCM). Sedlescombe, East Sussex, 1 male, nash, Moray, 1 male, 1.vii.1936; 4 males, 9.vii.1936, all xi.2001, Roper leg. (PPRS). The Loke, Norwich, Coe leg. (BMNH). Glen Coiltie, Highland Region, 1 Norfolk, 1 male, 8.v.1987; 4 males, 15.ix.1987, all male, 23.vi.1999; 1 male, 28.v.2002, all Chandler leg. Laurence leg. (PBLN). Upton Fen, Norfolk, 1 male, (PPCM). Lackham Park, Wiltshire, 1 male, 20.ix.2002, 11.x.2002, Chandler leg. (PPCM). West End Common, Chandler leg. (PPCM). Migdale Wood, Highland Esher, Surrey, 1 male, 19.x.2001, Chandler leg. Region, 1 male, 26.vi.1999, Chandler leg. (PPCM). (PPCM). Reelig Glen, Moniack, Highland Region, 3 males, 23.vi.1999, Chandler leg. (PPCM). Teesdale, Durham, Ecological notes: widespread in ancient beech and 1 male, 22.vi.1924, Edwards leg. (BMNH). The Loke, other broad-leaved woodland; grassland at woodland Norwich, Norfolk, 1 male, 18.vii.1981, Laurence fringe; fen; garden. v, ix–xi. leg. (PBLN). Tomintoul, Moray, 1 male, 8.viii.− 11.viii.1937, Coe leg. (BMNH). Wytham Woods, Wytham, Oxfordshire, 1 male, 2 females, 21.viii.2002, BRADYSIA LUCICHAETA MOHRIG & KRIVOSHEINA, 1989 J.E. Smith leg. (HRI). New to British list.

Material examined: North Ronaldsay, Orkney, 1 male, Material examined: Bix Bottom, Oxfordshire, 1 male, 4.ix.−6.ix.1989, Laurence leg. (DEI). 6.vii.2002, Chandler leg. (PPCM). Ecological notes: taken in a water trap on the shore of Ecological notes: found in site comprising broad- an island. ix. leaved woodland and calcareous grassland. vii.

BRADYSIA LUTARIA (WINNERTZ, 1869) BRADYSIA NERVOSA (MEIGEN, 1818) Synonym: postbicolor Mohrig & Krivosheina, 1983. Synonyms: fucata (Meigen, 1818); variabilis (Zetter- Literature: Laurence (1994): 118; Laurence (1996): 87 stedt, 1838). [all as Bradysia brunnipes, misidentification]; Menzel Literature: ? Walker (1848): 107 and 108 [as Sciara (1998): 20; White et al. (2000): 208; White & Smith fucata and Sciara nervosa]; ? Walker (1856): 54 [as (2000): 165. Sciara fucata]; ? Edwards (1938): 201 [as Sciara Material examined: Cors Erddreiniog, Isle of Anglesey, (Neosciara)]; Freeman (1983b): 37; Menzel (1998): 20. 2 males, 20.x.1988, Holmes, Boyce & Reed leg. Material examined: Bucklebury Common, Berkshire, (PPWM).Cuckfield, West Sussex, 4 males, 16.i.1969 1 male, 27.v.1990, Chandler leg. (PPCM). Colster- (reared), Lane leg. (BMNH). Dyffryn Gwaun, Pem- worth, Lincolnshire, 1 male, 3.vi.1987, Chandler brokeshire, 1 male, 20.vii.1987, Holmes, Boyce & Reed leg. (PPCM). Corrimony, Highland Region, 1 male, leg. (PPWM). Glasshouse Crops Research Station, 28.v.2002, Chandler leg (PPCM). Craigellachie, Avi- Littlehampton, West Sussex, 2 males, 13.viii.1956; 2 emore, Highland Region, 1 male, 30.v.2002, Chandler males, 15.viii.1956, all Hussey leg. (HRI); 1 male, leg. (PPCM). Decoy Wood, Aqualate Mere, Stafford- 26.iv.1965; 1 female, 26.v.1965, all Gurney leg. (HRI). shire, 1 male, 23.v.1994, Chandler leg. (PPCM). Dubh Lephinmore, Argyll & Bute, 1 female, 14.iv.1951; 2 Ghleann, Glen Quoich, Aberdeenshire, 3 males, females, 20.iv.1951 (all reared), all Pelham-Clinton 15.vi.2003, Chandler leg. (PPCM). Epping Thicks, leg. (NMS). Monkfield Wood, Cambridgeshire, 1 male, Epping Forest, Essex, 2 males, 14.vi.−20.vi.1999, Dag- vi.1998, C.W. Plant leg. (PPCM). Rhôs-y-gad, Isle of ley leg. (HRI). Frithsden Beeches, Hertfordshire, 1 Anglesey, 1 male, 27.vii.1988, Holmes, Boyce & Reed male, 6.vi.1997, Chandler leg. (PPCM). High Force, leg. (PPWM). Rothamsted, Harpenden, Hertfordshire, Durham, 1 male, 27.vi.−28.vi.1987, Disney leg. 10 males, 1950, Laurence leg. (PBLN). The Loke, (BMNH). Migdale Wood, Highland Region, 1 male, Norwich, Norfolk, 1 male, 23.iv.1987; 1 male, 8.v.1987; 11.vi.1984; 2 males, 31.v.2002, all Chandler leg. 7 males, 18.vi.1987; 1 male, 4.vii.1987; 2 males, (PPCM, PPWM). Roydon Common, Norfolk, 1 male, − − 5.vii.1987; 1 male, 25.vi. 19.vi.1988; 1 male, 15.ix. 5.vi.1993, Chandler leg. (PPCM). Selborne Common, 28.ix.1989, all Laurence leg. (PBLN). Waun Isaf, Hampshire, 1 male, 28.v.1988, Chandler leg. (PPCM). Pembrokeshire, 1 male, 20.vii.1987, Holmes, Boyce & Staffhurst Wood, Surrey, 1 male, 19.v.1991, Chandler Reed leg. (PPWM). Wellesbourne, Warwickshire, 7 leg. (NMS). Windsor Forest, Berkshire, 1 male, males, 26.vi.1997, White leg. (DEI); 8 males, 29 30.v.1097; 1 male, 15.v.1988, all Chandler leg. (NMS). females, 16.x.2000; 2 males, 11 females, xi.2000; 8 males, 29.xi.2001, all J.E. Smith leg. (DEI, HRI, Ecological notes: widespread in broad-leaved and carr PPCM). Yarrow, Scottish Borders, 4 males, 2 females, woodland; mixed and pine woodland; by waterfall. v– 20.iv.1951 (reared), Pelham-Clinton leg. (NMS). No vi. locality details, England, 3 females, 14.x. [no year]; 2 males, 1 female, 16.x.1950, all no collector details BRADYSIA NIGRISPINA MENZEL SP. NOV. (PBLN); 1 male, 1 female, 2 larvae, no date, Speyer leg. (HRI). Literature: Freeman (1983b): 37 [in part as Bradysia regularis, misidentification]. Ecological notes: found in broad-leaved woodland; damp heathland; wetlands including fens, reed beds New to British list. and water meadows; garden; grounds of horticultural Material examined: Berwick-upon-Tweed, Northum- research station; in glasshouses on cucumber roots. berland, 1 male, 24.v.−7.vi.1996; 1 male, viii.1999; 1 Reared from rotten wood, dung, mushroom farm and male, ix.1999, all Laurence leg. [paratypes of Bradysia laboratory cultures. iv-ix. nigrispina] (DEI, PBLN). Letchworth, Hertfordshire, 1 male, vii.1918, Edwards leg. (BMNH). Shrewsbury, BRADYSIA MOESTA FREY, 1948 Shropshire, 2 males, 21.vii.1920, Edwards leg. (BMNH). The Loke, Norwich, Norfolk, 1 male, Synonym: albosetosa Frey, 1948. 2.iii.1993, leg. Laurence [paratype of Bradysia New to British list. nigrispina] (PBLN). Wellesbourne, Warwickshire, 1

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 50 F. MENZEL ET AL. male, 5.vii.−19.vii.2001, J.E. Smith leg. [paratypes of Angus, 2 males, 24.vii.1937, Coe leg. (BMNH). Brae- Bradysia nigrispina] (HRI). mar, Aberdeenshire, 1 male, 19.vii.−30.vii.1938, Coe leg. (BMNH). Brancaster, Norfolk, 1 male, 16.ix.− Ecological notes: found in gardens; outside mushroom 8.x.1988, Foster & Procter leg. (PBLN). Brockenhurst, farm. iii, v–viii, ix. Hampshire, 1 male, 10.ix.1977, Chandler leg. (PPCM). Brynberian Moor, Pembrokeshire 1 male, 6.vii.1987, Holmes, Boyce & Reed leg. (PPWM). Burley, Hamp- BRADYSIA NITIDICOLLIS (MEIGEN, 1818) shire, 1 male, 30.vii.1953, Colyer leg. (BMNH). Burn- Synonyms: alacris (Winnertz, 1867); albicans (Win- ham Beeches, Buckinghamshire, 1 male, 4.x.1999, nertz, 1867); aprilina (Meigen, 1818); atroparva Frey, Chandler leg. (NMS). Cadair Idris, Gwynedd, 2 1948; fenestrata (Meigen, 1818); inornata (Winnertz, males, 19.vi.1993; 3 males, 3.vii.1993; 1 male, 1867); reforma Rudzinski, 1993; scatopsoides (Meigen, 13.vii.1993; 1 male, 17.vii.1993; 1 male, 1.viii.1993; 4 1818); tenella (Winnertz, 1867); trichoptera (Lengers- males, 4 ix.1993; 8 males, 22.x.1993; 12 males, dorf, 1926). 13.xi.1993, all Laurence leg. (PBLN). Caerkief Farm, Cornwall, 1 male, 3.vii.1983, Chandler leg. (PPCM). Literature: ? Walker (1848): 107 and 108; ? Walker Cannock Chase, Staffordshire, 1 male, 21.x.1983, (1856): 52, 54 and 55 [as Sciara nitidicollis, Sciara Chandler leg. (NMS). Carton, Kildare, 1 male, fenestrata, Sciara aprilina and Sciara scatopsoides]; 29.iv.1987, J.P. O’Connor leg. (NMID). Castlemartin Edwards (1925a): 540 [as Sciara albinervis, misiden- Corse, Pembrokeshire, 2 males, 22.vii.1987; 2 males, tification]; Edwards & Collin (1932): 264; Edwards 14.x.1989, all Holmes, Boyce & Reed leg. (PPWM). (1934b): 34 [all as Sciara]; Edwards (1938): 201 [as Catfield, Norfolk, 1 male, 12.x.−13.xii.1988, Foster & Sciara (Neosciara)]; Freeman (1983b): 38 [in part as Procter leg. (PBLN). Cefn Bryn, Swansea/Abertawe, 1 Bradysia aprica, misidentification]; Laurence (1994): male, 28.vi.1989, Holmes, Boyce & Reed leg. (PPWM). 113; Chandler et al. (1996): 259; Laurence (1997a): 85; Cefn Geirch, Denbighshire, 1 male, 15.x.1994, Chan- Laurence (1996): 85; Laurence (1997b): 228; Laurence dler leg. (PPCM). Cefn Grwhyd, Neath/Port Talbot, 4 & James (1997): 166; Menzel (1998): 20. males, 12.vii.1989; 1 male, 4.x.1989, all Holmes, Boyce Material examined: Ailladie, Clare, 1 male, 24.v.1985, & Reed leg. (PPWM). Chigwell Row Wood, Chigwell J.P. & M.A. O’Connor leg. (NMID). Ambersham Com- Row, Essex, 1 male, 4 females, 28.vi.2002; 4 males, 3 mon, West Sussex, 1 male, 9.vi.1979, Chandler leg. females, 15.ix.2002; 1 male, 7 females, 3.x.2002; 2 (PPCM). Annahala bog, Gearagh, Cork, 4 males, males, 2 females, 8.x.2002; 3 males, 8 females, 30.iii.1990; 2 males, 1 female, 6.iv.1990; 6 males, 1 16.x.2002; 2 males, 3 females, 22.x.2002, all Ismay & female, 10.viii.1990; 4 males, 1 female, 25.viii.1990; 8 Schulten leg. (DEI, UMO). Chilbolton Common, males, 1 female, 24.ix.1990, all de Courcy Williams leg. Hampshire, 2 males, 6.vii.1991, Chandler leg. (NMS, (DEI, NMID). Ashberry Pastures, North Yorkshire, 1 PPCM). Chippenham Fen, Cambridgeshire, 1 male, male, 12.x.1990, Chandler leg. (NMS). Ashtead, Sur- 2.vii.1988, Chandler leg. (NMS). Clothall, Hertford- rey, 1 male, 8.v.1937, Coe leg. (BMNH). Austwick, shire, 1 male, 4.v.1918, Edwards leg. (BMNH). Clov- North Yorkshire, 1 male, 10.x.1930; 1 male, v.1943, elly, Devon, 2 males, 8.v.1936, Edwards leg. (BMNH). all Cheetham leg. (BMNH). Aversley Wood, Cam- Collinpark Wood, Gloucestershire, 1 male, 3.ix.2002, bridgeshire, 1 male, 24.iv.1988, J.H. Cole leg. (PJCB). Gibbs leg. (PDGB). Colne, Lancashire, 3 males, 7 Ballyvourney, Cork, 1 male, 10.vii.1990, Withers leg. females, 28.v.1927, Britten leg. (BMNH). Consall (PPWM). Balmoral Forest, Aberdeenshire, 9 males, Wood, Staffordshire, 1 male, 12.x.1997, Chandler leg. 1.viii.−4.viii.1937; 2 males, 19.vii.−30.vii.1938, all Coe (PPCM). Convil Braes, Moray, 1 male, 11.viii.1937, Coe leg. (BMNH). Beaulieu River, Hampshire, 1 male, leg. (BMNH). Correin Lochan, Arran, North Ayrshire, 22.vi.1936, Edwards leg. (BMNH). Ben Lawers, Perth 3 males, 29.v.1919, Edwards leg. (BMNH). Corriegills, & Kinross, 1 male, 1.vi.1932, Edwards leg. (BMNH). Arran, North Ayrshire, 1 male, 2.vi.−4.vi.1919, Blean Wood, Kent, 1 male, 19.v.1973, Parmenter leg. Edwards leg. (BMNH). Cors Caron, Ceredigion, 1 (BMNH). Blaen y Wergloedd, Wrexham, 1 male, male, 29.vii.1987, Holmes, Boyce & Reed leg. (PPWM). 4.viii.1988; 1 male, 13.x.1988, all Holmes, Boyce & Cors Erddreiniog, Isle of Anglesey, 4 males, 6.x.1988, Reed leg. (PPWM). Bodnant, Conwy, 1 male, 2.v.1953, Holmes, Boyce & Reed leg. (PPWM). Cors Fochno, Colyer leg. (BMNH). Bonhill, West Dunbartonshire, 1 Ceredigion, 1 male, 1.ix.1989, Holmes, Boyce & Reed male, 28.vii.1908, Malloch leg. (NMS). Boveney Lock, leg. (PPWM). Cors Graianog, Gwynedd, 2 males, Buckinghamshire, 1 male, 18.iv.1981, Chandler leg. 28.vii.1988; 1 male, 7.x.1988, all Holmes, Boyce & Reed (PPCM). Bradfield Wood, Suffolk, 2 males, 6.vii.− leg. (PPWM). Cors-y-Llyn, Powys, 1 male, 11.vii.1989; 11.vii.1990; 1 male, 11.vii.−16.vii.1990; 5 males, 19.x.− 2 males, 10.x.1989, Holmes, Boyce & Reed leg. 16.x.1990; 2 males, 16.viii.−22.viii.1991; 4 males, (PPWM). Cothill, Oxfordshire, 1 male, 14.x.1999, 23.v.−30.v.2000, all Laurence leg. (PBLN). Braedownie, Chandler leg. (NMS). Craig na Cailloch, Perth & Kin-

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 BRITISH SCIARID FAUNA 51 ross, 1 male, 5.vi.−8.vi.1932, Edwards leg. (BMNH). Boyce & Reed leg. (PPWM). Gwaun Ystrad Caron, Credenhill Park Wood, Herefordshire, 1 male, Ceredigion, 1 male, 4.ix.1986; 1 male, 11.ix.1986; 1 11.x.1998, Chandler leg. (NMS). Croft Pascoe Pool, male, 18.ix.1986, all Holmes, Boyce & Reed leg. Cornwall, 1 male, 2.x.1983, Chandler leg. (PPCM). (PPWM). Hartslock, Oxfordshire, 1 male, 6.v.2000, Crymlyn Bog, Swansea/Abertawe, 5 males, 5.x.1989, Raper leg. (PDGB). Hayle Kimbro Pool, Cornwall, 1 Holmes, Boyce & Reed leg. (PPWM). Culbin Forest, male, 4.x.1983, Chandler leg. (PPWM). Haytor, Devon, Highland Region, 1 male, 27.v.2002, Chandler leg. 8 males, 23.vi.2002, J.E. Smith leg. (HRI). Hengrove (PPCM). Culbin Sands, Highland Region, 3 males, Park, Avon, 1 male, 23.iv.2003, Gibbs leg. (PDGB). 5.vii.−7.vii.1936, Coe leg. (BMNH). Cwm Bychan, Hickling, Norfolk, 12 males, 23.vi.−7.vii.1988; 1 male, Gwynedd, 2 males, 12.vii.1976, J.H. Cole & Chandler vii.1988; 6 males, x.1988; 3 males, 1988; 8 males, leg. (PJCB, PPCM). Cwm Dewi, Pembrokeshire, 1 18.v.1989; 5 males, v.1989, all Foster & Procter leg. male, 28.ix.1987, Holmes, Boyce & Reed leg. (PPWM). (PBLN). Hickling Broad, Norfolk, 2 males, 1 female, Dartford, Kent, 1 male, 22.iv.1957, Parmenter leg. 17.ix.−19.ix.1932, Coe leg. (BMNH). Hinchingbrooke (BMNH). Dartmoor, Bush Down, Devon, 2 males, Country Park, Cambridgeshire, 1 male, 30.iv.1989, 28.v.1978, J.H. Cole leg. (PJCB). Dendles Wood, Devon, J.H. Cole leg. (PJCB). Hosey Common, Kent, 1 male, 1 male, 29.v.1978, J.H. Cole leg. (PJCB). Derrycunihy 29.x.1968, Chandler leg. (PPCM). How Hill, Ludham, Cascade, Kerry, 1 male, 16.x.1973, Chandler leg. Norfolk, 1 male, 24.vi.1990, Withers leg. (PPWM). Ill- (PPCM). Derwent Valley, Tyne & Wear, 1 male, 26.vii.− tyd Pools, Powys, 2 males, 11.x.1989, Holmes, Boyce & 13.viii.1990, Jessop leg. (PBLN). Devil’s Glen, Wicklow, Reed leg. (PPWM). Ingleborough, North Yorkshire, 1 1 male, 4.x.1980, Chandler leg. (PPCM). Dinton Pas- male, 13.v.1921, Edwards leg. (BMNH). Iping Com- tures Country Park, Berkshire, 1 male, 18.ix.1993; 2 mon, West Sussex, 2 males, 12.x.1984, Godfrey leg. males, 23.iv.1994; 1 male, 17.x.1998, all Chandler (PPCM). Ivybridge, Devon, 1 male, 11.x.1980, Chan- leg. (NMS, PPCM). Ditchley, Oxfordshire, 2 males, dler leg. (PPCM). Kenmare, Kerry, 1 male, 15.x.1973, v.1937, Edwards leg. (BMNH). Dromore Forest, Kerry, Chandler leg. (PPCM). Kessingland, Suffolk, 1 male, 1 male, 15.x.1973, Chandler leg. (PPCM). Dublin 22.iv.−25.iv.1933, Edwards leg. (BMNH). Killaun Bog, City, Dublin, 1 male, 13.v.1982, J.P. O’Connor leg. Offaly, 5 males, ix.2000, Withers leg. (PPWM). Laken- (NMID). Dyffryn Gwaun, Pembrokeshire, 1 male, heath, Suffolk, 2 males, 24.vi.−8.vii.1988, Foster & 20.vii.1987, Holmes, Boyce & Reed leg. (PPWM). Procter leg. (PBLN). Leckford Estate, Hampshire, Epping Forest, Essex, 1 male, 28.vi.2000; 1 male, 1 male, 3.x.1970, Stubbs leg. (PPCM); 1 male, 11.x.2000, all Chandler leg. (NMS). Fair Isle, Shetland, 30.iv.1977; 1 male, 12.iv.1981; 1 male, 27.vii.1985; 1 2 males, 21.vii.−3.viii.1966, Moreby leg. (BMNH); 3 male, 1.ix.1999, all Chandler leg. (NMS, PPCM). males, 31.viii.2000, Riddiford leg. (PBLN). Falls of Tar- Leebotwood, Shropshire, 2 females, 14.vii.2003, J.E. nash, Moray, 2 males, 9.vi.1936; 22 males, 9.vii.1936, Smith leg. (HRI). Lenwade Pits, Norfolk, 1 male, all Coe leg. (BMNH). Fferam Paradwys, Isle of Angle- 6.x.1980 (reared), Evans leg. (BMNH). Letchworth, sey, 1 male, 5.vii.1987, Chandler leg. (NMS). Fleet Hertfordshire, 2 males, vii.1939, Edwards leg. Pond, Hampshire, 1 male, 15.vii.1990, Chandler leg. (BMNH). Limerigg, Falkirk, 1 male, 18.vii.1990, Chan- (NMS). Gearagh, Cork, 3 females, 9.iii.1990; 3 males, dler leg. (PPCM). Linn of Dee, Aberdeenshire, 1 male, 31.iii.1990; 17 males, 7.iv.1990; 7 males, 20.iv.1990; 31.x.1998, Chandler leg. (NMS). Llanbedr, Gwynedd, 1 7 males, 7.vi.1990; 3 males, 9.vi.1990; 2 males, male, 16.x.1992, Chandler leg. (PPCM). Llandwrog, 10.vi.1990, all de Courcy Williams leg. (DEI, NMID). Gwynedd, 6 males, 9.vii.1914 [in part labelled as ‘? Glen Coiltie, Highland Region, 1 male, 28.v.2002, albinervis Winnertz’], Edwards leg. (BMNH, ZFMK). Chandler leg. (PPCM). Glendalough, Wicklow, 1 male, Llangloffan Fen, Pembrokeshire, 1 male, 29.ix.1987, 16.x.1973, Chandler leg. (PPCM). Glen Einich, High- Holmes, Boyce & Reed leg. (PPWM). Llanon, Swansea/ land Region, 1 male, 30.vi.1933, Coe leg. (BMNH). Abertawe, 2 males, 13.vii.1989, Holmes, Boyce & Reed Glen Lough, Westmeath, 1 male, 28.vi.1987, Chandler leg. (PPWM). Llyn Hafodol, Isle of Anglesey, 1 male, leg. (PPCM). Glen of Drumloch, Moray, 1 male, 11.vii.1987, Chandler leg. (NMS). Llyn Llennyrch, 16.viii.1937, Coe leg. (BMNH). Godlingstone Heath, Gwynedd, 2 males, 16.vii.1976, Chandler leg. (PPCM). Dorset, 1 male, 1.vii.1998, Chandler leg. (PPCM). Gors Llyn Pencraig, Gwynedd, 1 male, 27.x.1988, Holmes, Llwyn, Neath/Port Talbot, 1 male, 28.vi.1988; 2 males, Boyce & Reed leg. (PPWM). Llyn Tyn y Llyn, Den- 12.vii.1989, all Holmes, Boyce & Reed leg. (PPWM). bighshire, 1 male, 4.viii.1988; 5 males, 13.x.1988, all Grassholm, Pembrokeshire, 2 males, 25.viii.1934, Holmes, Boyce & Reed leg. (PPWM). Llyn Ystumllyn, Edwards leg. (BMNH). Great Huntly Bank, New For- Gwynedd, 1 male, 13.vii.1988; 1 male, 24.vii.1988; 1 est, Hampshire, 1 male, 29.vi.1989, Chandler leg. male, 5.x.1988; 3 males, 8.x.1988, all Holmes, Boyce & (NMS). Greywell Fen, Hampshire, 1 male, 15.vii.1990; Reed leg. (PPWM). Loch Callater, Aberdeenshire, 1 1 male, 7.vii.1991, all Chandler leg. (PPCM). Gwaun male, 28.vi.−29.vi.1937; 1 male, 28.vii.−29.vii.1937; Garthenor, Ceredigion, 1 male, 14.x.1986, Holmes, 4 males, 22.vii.1938, all Coe leg. (BMNH). Loch na

Ba Ruaidhe, Highland Region, 1 male, 13.vi.1984, River Druie, Highland Region, 1 male, 18.viii.1986, Chandler leg. (NMS). Loch Park, Moray, 4 males, Chandler leg. (PPCM). Ryton Wood, Bubbenhall, Cov- 12.viii.−14.viii.1937, Coe leg. (BMNH). Lockhaugh entry, West Midlands, 1 male, 25.viii.2002, Menzel leg. Bank Wood, Derwent Valley, Tyne & Wear, 2 males, (DEI). Salisbury Plain Training Area, Warminster, 24.vii.−13.viii.1990, Jessop leg. (PBLN). Lough Key, Wiltshire, 1 male, 3.vii.2003, Ismay & Schulten Roscommon, 1 male, 11.v.1970, Chandler leg. (PPCM). leg. (UMO). Sandscale Haws, Cumbria, 2 males, Lough Slevin, Westmeath, 1 male, 28.vi.1987, Chan- 18.vi.1999, Chandler leg. (NMS). Scadbury Park, dler leg. (PPCM). Lower Brock Mill, Lancashire, 2 Greater London, 1 male, 18.vi.1983, Chandler leg. males, 14.vi.1999, Chandler leg. (PPCM). Lundy (NMS). Scargill, Durham, 2 males, 2.vi.2002, Chandler Island, Devon, 1 male, 20.vii.−27.vii.1972, Lane leg. leg. (PPCM). Shapwick Heath, Somerset, 1 male, (BMNH). Maen Porth, Cornwall, 1 male, 5.x.1983, 7.ix.1930, Edwards leg. (BMNH); 1 male, 30.vi.1985, Chandler leg. (NMS). Mid Yell, Shetland, 1 male, J.H. Cole leg. (PJCB). Sidmouth, Devon, 1 male, 29.v.1987; 1 male, 5.ix.1990, all Laurence leg. (PBLN). 19.iv.1925, Edwards leg. (BMNH). Slepe Heath, Mills Marsh, Norfolk, 1 male, 27.vi.−11.vii.1988; 7 Dorset, 1 male, 1.vii.1998, Chandler leg. (NMS). males, 11.viii.1988, all Foster & Procter leg. (PBLN). Sned Wood, Lower Lye, Herefordshire, 2 females, Monument Field, Epping Forest, Essex, 5 males, 14.vii.2003, J.E. Smith leg. (HRI). Spartum Fen, 19.iv.−25.iv.1999, Dagley leg. (HRI). Nanjizal Cove, Oxfordshire, 1 male, 15.x.1999, Chandler leg. (PPCM). Cornwall, 1 male, 3.iv.1972, Chambers leg. (BMNH). St. Kilda, Western Isles, 1 male, 28.vii.1931, Lack leg. Newbold Comyn Park, Leamington Spa, Warwick- (BMNH); 1 male, 13.vii.−22.vii.1960, Giffard leg. shire, 67 males, 15 females, 29.vii.2001, J.E. Smith leg. (NMS); 1 male, 3.vi.−17.vi.1961, Russell leg. (NMS). (HRI). Newborough Warren, Isle of Anglesey, 1 male, Stoborough Heath, Dorset, 1 male, 1.vii.1998, Chan- 9.vii.1987, Chandler leg. (NMS). Newbridge Fen, Kil- dler leg. (NMS). Stradey Wood, Carmarthen, 1 male, dare, 1 male, 12.iv.1982, J.P. & M.A. O’Connor leg. 13.x.1995, McLean leg. (PPCM). Templehouse, Sligo, 1 (NMID); 1 male, 27.vi.1984, Chandler leg. (NMS); 1 male, 1.x.1977, Chandler leg. (PPCM). The Flits, Here- male, 16.vii.1990, Withers leg. (PPWM). New Malden, fordshire, 1 male, 11.x.1998, Chandler leg. (PPCM). Greater London, 1 male, 8.v.1932, Coe leg. (BMNH). The Loke, Norwich, Norfolk, 1 male, 26.iv.1987; 2 Nine Nicks of Thirlwall, Northumberland, 2 males, males, 4.v.1987; 5 males, 8.v.1987; 16 males, 9.v.1987; 1.viii.1923, Edwards leg. (BMNH). North Ronaldsay, 2 males, 15.ix.−28.ix.1989; 1 male, 27.vii.1992, all Lau- Orkney, 1 male, 4.iv.1989; 1 male, 12.ix.1989, all Lau- rence leg. (PBLN). The Ritec, Pembrokeshire, 1 male, rence leg. (PBLN). Nûg, Conwy, 1 male, 3.viii.1988, 8.vii.1987, Holmes, Boyce & Reed leg. (PPWM). Holmes, Boyce & Reed leg. (PPWM). Odiham Common, Thompson Common, Norfolk, 1 male, 11.vii.1993, Hampshire, 1 male, 7.vii.1984, Chandler leg. (PPCM). Chandler leg. (PPCM). Tomintoul, Moray, 26 males, Oswestry, Shropshire, 1 male, 10.v.1913, Colyer leg. 6.viii.−11.viii.1937; 3 males, 8.viii.−11.viii.1937, all (BMNH). Oxey Mead, Oxfordshire, 2 males, 15.x.1999, Coe leg. (BMNH). Torboll Wood, Highland Region, 1 Chandler leg. (PPCM). Oxford, Oxfordshire, 1 male, male, 18.x.1995, Chandler leg. (PPCM). Trawsfynydd, 10.v.1924, Hamm leg. (BMNH). Peel, Isle of Man, 4 Gwynedd, 1 male, 14.x.1988, Holmes, Boyce & Reed males, 1 female, 7.v.1988, Laurence leg. (PBLN). Pin- leg. (PPWM). Tresta, Shetland, 1 male, 10.vi.1993, gle Wood, Warboys, Cambridgeshire, 2 males, 2.v.1990, Laurence leg. (PBLN). Trooper’s Hill, Bristol, Avon, 1 J.H. Cole leg. (PJCB). Port Appin, Argyll & Bute, 1 male, 23.iv.2001, Gibbs leg. (PDGB). Tup Hag Wood, male, 30.ix.1951, Pelham-Clinton leg. (NMS). Power- North Yorkshire, 1 male, 12.x.1990, Chandler leg. scourt, Wicklow, 1 male, 1.x.1909, Halbert leg. (NMID). (NMS). Uragh Wood, Kerry, 3 males, 10.vii.1990, Redgrave Lake, Suffolk, 1 male, 24.v.1987, Withers Withers leg. (PPWM). Vidlin, Shetland, 1 male, leg. (PPWM). Reedham, Norfolk, 1 male, 11.vii.− 18.vi.−21.vi.1993, Laurence leg. (PBLN). Virginia 12.viii.1988; 1 male, 26.viii.−27.x.1988, all Foster & Water, Surrey, 1 male, 6.x.1974, Chandler leg. (NMS). Procter leg. (PBLN). Rhineﬁelds, New Forest, Hamp- Watford, Hertfordshire, 3 males, 21.x.1944, no shire, 1 male, 16.iv.1934, Colyer leg. (BMNH). Rhôs collector detail (BMNH). Waun Isaf, Pembrokeshire, 2 Llawr Cwrt, Ceredigion, 1 male, 15.vii.1987, Holmes, males, 20.vii.1987, Holmes, Boyce & Reed leg. Boyce & Reed leg. (PPWM). Rhôs Pwllygawnen, Car- (PPWM). Wellesbourne, Warwickshire, 10 males, 6.iv.− marthen, 2 males, 11.vii.1989; 1 male, 12.viii.1989, all 13.iv.2000; 1 male, 17.viii.−31.viii.2000; 10 males, 2 Holmes, Boyce & Reed leg. (PPWM). Rhôs-y-gad, Isle of females, 28.iii.−26.iv.2001; 19 males, 10.v.−24.v.2001; 3 Anglesey, 2 males, 5.x.1988, Holmes, Boyce & Reed leg. males, 13.ix.−4.x.2001, all J.E. Smith leg. (HRI). West (PPWM). Rhôsili Down, Swansea/Abertawe, 1 male, End Common, Esher, Surrey, 1 male, 16.vii.2002, 3.x.1989, Holmes, Boyce & Reed leg. (PPWM). Rhôs- Chandler leg. (PPCM). West Harling Common, goch, Powys, 1 male, 10.x.1989, Holmes, Boyce & Norfolk, 2 males, 20.x.1990; 1 male, 23.iv.1991, all Reed leg. (PPWM). Rosserk Abbey, Mayo, 2 males, Withers leg. (PPWM). West Walk, Hampshire, 1 male, 30.ix.1977, Chandler leg. (PPCM). Rothiemurchus, by 6.vi.1998, Chandler leg. (NMS). Whitacre Heath, War-

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 BRITISH SCIARID FAUNA 53 wickshire, 2 males, 13.x.2001, Chandler & J.E. Smith shire, 2 males, 2 females, 5.vi.−11.vi.1931 (reared); 2 leg. (HRI). Whitnash brook, Sydenham, Leamington males, 9.vi.1939, all Basden leg. (BMNH, NMS); 1 Spa, Warwickshire, 5 males, 4.ix.2003, J.E. Smith male, 9.x.1990, Chandler leg. (PPCM); 2 males, leg. (HRI). Wicken Fen, Cambridgeshire, 2 males, 30.viii.−15.ix.1995; 1 male, 5.viii.−31.viii.1996; 1 male, 8.vii.1989, Chandler leg. (NMS). Willes Meadow, 9.x.−26.xi.1996; 7 males, 26.vi.−7.vii.1999, all Ismay Leamington Spa, Warwickshire, 4 males, 5 females, leg. (HRI, PPCM). Cambridge, Cambridgeshire, 11 27.viii.2002, Menzel & J.E. Smith leg. (DEI, HRI). males, 3 females, 14.vi.−15.ix.1934, Basden leg. Windsor Forest, Berkshire, 1 male, 27.vii.1991, Chan- (NMS). Carie, Rannoch, Perth & Kinross, 1 male, dler leg. (PPCM). Windsor Great Park, Berkshire, 1 23.x.1990, Chandler leg. (PPCM). Carreg Cennen male, no date, Coe leg. (BMNH). Winnall Moor, Hamp- Castle, Carmarthen, 1 male, 14.x.1995, Chandler leg. shire, 6 males, 7.vii.1990, Chandler leg. (NMS, PPCM); (PPCM). Chatsworth Deer Park, Derbyshire, 1 male, 2 males, 7.vii.1990, J.H. Cole leg. (PJCB). Wood Crates, 11.x.1991, Chandler leg. (PPCM). Coed Tremadog, New Forest, Hampshire, 2 males, 8.vii.1990, Chandler Gwynedd, 1 male, 14.vii.1976, Chandler leg. (PPCM). leg. (NMS, PPCM). Wyre Forest, Worcestershire, 1 Cogley Wood, Somerset, 1 male, 19.x.1986, Chandler male, 14.x.1984, Chandler leg. (NMS). Yell, Shetland, leg. (PPCM). Coombe Abbey Country Park, Coventry, 2 males, 3.ix.−10.ix.1990, Laurence leg. (PBLN). No Warwickshire, 1 male, 28.viii.2003, J.E. Smith leg. locality details, Wicklow, 1 male, 19.x.−22.x.1998, Lau- (HRI). Cotterdale, North Yorkshire, 1 male, rence leg. (PBLN). 22.ix.1956, J.H. Cole leg. (PJCB). Dancer’s End, Buckinghamshire, 1 male, 8.iv.1984, Merrifield leg. Ecological notes: very common in all types of wood- (PPCM). Denton, Greater Manchester, 3 males, 7 land; wet and dry heathland; wetlands including females, 15.iv.1923 (reared), Newton leg. (BMNH). water meadows, fens, mires and bogs; dry grassland; Derwent Ings, North Yorkshire, 1 male, 4.viii.1989, old hedges; old sand and gravel workings; on rocky Chandler leg. (PPCM). Dinton Pastures Country Park, island with seabird colonies; coastal sand dunes; salt- Berkshire, 1 male, 21.viii.1993; 1 male, 7.x.1998, all marsh; gardens; in thousands on rhubarb flowers; Chandler leg. (PPCM). Dubh Ghleann, Glen Quoich, open montane habitats; near shore; in house. Reared Aberdeenshire, 2 males, 21.vii.2000, Godfrey leg. from the fungus Russula alutacea. iii–xi. (PPCM). Farnham Royal, Buckinghamshire, 5 Taxonomic note: All data under ‘Sciara? nitidicollis’ females, 13.v.−15.v.1931 (reared), Basden leg. (NMS). listed by Edwards (1925a) related to Lycoriella Foxhole, Highland Region, 1 male, 26.x.2000, Chan- ingenua [misidentification]. dler leg. (PPCM). Gearagh, Cork, 1 male, 22.v.1990, de Courcy Williams leg. (DEI). Glenariff, Antrim, 1 male, 12.xi.1986, Chandler leg. (PPCM). Goring-on-Thames, BRADYSIA NOCTURNA TUOMIKOSKI, 1960 Oxfordshire, 1 male, 29.x.1962, J.H. Cole leg. (PJCB). Greenlands, Buckinghamshire, 23 males, 12 females, Literature: Edwards (1938): 201 [in part as Sciara 13.v.−17.x.1933 (reared), Basden leg. (NMS). Hamble- (Neosciara) brunnipes, misidentification]; Chandler den Weir, Berkshire, 9 males, 1 female, 7.iv.−23.iv.1932 et al. (1996): 258 [in part as Bradysia brunnipes, mis- (reared), Basden leg. (NMS). Harpenden, Hertford- identification]; Chandler et al. (2002): 122. shire, 1 male, 7.v.1934; 1 male, 30.vii.1934; 1 male, Material examined: Alconbury, Cambridgeshire, 1 8.viii.1934, all C.B. Williams leg. (BMNH). Heming- male, 3.v.1976, J.H. Cole leg. (PJCB). Annahala bog, ford, Cambridgeshire, 1 male, ix.1938, Edwards leg. Gearagh, Cork, 1 male, 22.v.1990, de Courcy Williams (BMNH). Invertromie Wood, Insh Marshes, Highland leg. (NMID). Austwick, North Yorkshire, 1 male, Region, 1 male, 30.v.2002, Chandler leg. (PPCM). Loch 18.x.1930, Cheetham leg. (BMNH). Aversley Wood, Assynt, Highland Region, 1 male, 24.vii.1911, Yerbury Cambridgeshire, 1 male, 24.iv.1988, J.H. Cole leg. leg. (BMNH). Loch an Eilein, Highland Region, 1 (PJCB). Bagpiper’s Tump, West Wood, Herefordshire, male, 29.v.1973, Chandler leg. (PPCM). Loch Morlich, 1 male, 10.x.1998, Chandler leg. (PPCM). Baldock, Highland Region, 1 male, 14.vii.1936, Coe leg. Hertfordshire, 1 male, vii.1918, Edwards leg. (BMNH). Mahon Falls, Waterford, 1 male, 3.vii.1988, (BMNH). Ballyteige, Wexford, 1 male, 3.vi.1987, J.P. J.P. O’Connor leg. (NMID). Marston, Oxfordshire, 1 O’Connor leg. (NMID). Beinn Eighe, Highland Region, male, 25.xii.1932; 24 males, 14 females, 25.xii.1932– 1 male, 10.vi.1984, Chandler leg. (PPCM). Black Cliff, 19.v.1933 (all reared), all Basden leg. (BMNH, NMS). Monmouthshire, 1 male, 9.vii.1986, Chandler leg. Martin Mere, Lancashire, 1 male, 30.iv.1996, Under- (PPCM). Brampton, Cambridgeshire, 1 male, 8.v.1977; wood leg. (PPCM). Medmenham, Buckinghamshire, 1 1 male, 13.v.1989, all J.H. Cole leg. (PJCB). Bransgore, male, 15.vi.1931; 4 males, 1 female, 12.v.−19.v.1932 Hampshire, 1 male, 10.viii.1933 (reared), Basden leg. (all reared), all Basden leg. (BMNH, NMS). Old Buck- (NMS). Brodie Castle, Moray, 1 male, 23.x.2000, Chan- enham Fen, Norfolk, 1 male, 9.v.1990, Chandler leg. dler leg. (PPCM). Burnham Beeches, Buckingham- (PPCM). Old Lodge Warren, East Sussex, 1 male,

9.x.1996, Chandler leg. (PPCM). Pass of Killiecrankie, facing the Atlantic Ocean; in debris on shore of lake. Perth & Kinross, 1 male, 18.x.1971, Chandler leg. iii–iv, vi–vii. (PPCM). Penallt, Carmarthen, 1 male, 14.x.1995, Chandler leg. (PPCM). Peppercombe, Devon, 1 male, 14.vi.1984, J.H. Cole leg. (PJCB). Portholme Meadow, BRADYSIA OCELLARIS (COMSTOCK, 1882) Brampton, Cambridgeshire, 3 males, 3.vi.1976, J.H. Synonyms: garretti (Shaw, 1952); johannseni (Shaw, Cole leg. (PJCB). Radwell, Hertfordshire, 1 male, 1952); laffooni (Shaw, 1952); prothalliorum (de 22.v.1921, Edwards leg. (BMNH). Rotherfield Park, Meijere, 1946); rubicundula Frey, 1948; tritici (Coquil- Hampshire, 1 male, 3.vii.2002, Chandler leg. (PPCM). lett, 1895). Salisbury Plain Training Area, Warminster, Wiltshire, Literature: Edwards & Williams (1916): 259 [as Sciara 1 male, 3.vii.2003, Ismay & Schulten leg. (UMO). Scot- tritici]; Edwards (1925a): 539 [as Sciara pectoralis, land Fen, Suffolk, 1 male, 10.xi.1996, Chandler leg. misidentification]; Kloet & Hincks (1945): 350; (PPCM). Selsdon, Greater London, 1 male, 26.iv.1930; Kloet & Hincks (1976): 31; Menzel (1998): 23 [all 1 male 10.iv.1939, all Coe leg. (BMNH). Sheep Leas, as Bradysia radicum, misidentification]; Freeman West Horsley, Surrey, 1 male, 16.vi.1999, Chandler (1983b): 36 [as Bradysia tritici]; O’Connor & Ashe leg. (PPCM). Skenfrith, Monmouthshire, 1 male, (1991): 29 [as Bradysia tritici]; Menzel (1998): 20; 12.vi.1997, Chandler leg. (PPCM). Spartum Fen, Menzel et al. (2003): 454. Oxfordshire, 1 male, 1.vi.1991, Chandler leg. (PPCM). Stonybrow Wood, Rotherfield Park, Hampshire, 1 Material examined: Edgbaston, Birmingham, West male, 23.x.2002, Chandler leg. (PPCM). Temple, Berk- Midlands, 1 male, 13.ix.1922, Edwards leg. (BMNH). shire, 1 male, 22.iv.1932; 2 males, 5 females, 21.iv.− Glasshouse Crops Research Institute, Littlehampton, 22.iv.1934; 2 females, 7.iv.−16.vi.1934, all Basden Sussex, 1 male, no date, Binns leg. (HRI). Kew Gar- leg. (BMNH, NMS). Warren Row, Berkshire, 1 male, dens, Greater London, 2 males, no date, Edwards leg. 19.ii.1932; 3 males, 1 female, 21.ii.−19.iii.1932 (all (BMNH). London, Greater London, 1 female, no date, reared), all Basden leg. (BMNH, NMS). West Harling Baker leg. (BMNH). Merton, Greater London, 3 males, Common, Norfolk, 1 male, 28.x.1989, Withers leg. x.1911, C.B. Williams leg. (BMNH). Milton Keynes, (PPWM). Buckinghamshire, 7 males, 2 females, ii.1986, no collector detail (BMNH). Rillington, North Yorkshire, Ecological notes: common in all types of woodland; cal- 4 males, 4 females, 4.vi.1915 (reared), Staunton leg. careous grassland; wetlands including fen, bog, raised (BMNH). Terenure, Dublin City, Dublin, 1 male, mire, lake and wildfowl reserve, water meadows with 25.x.1989, Ashe leg. (NMID). Wellesbourne, Warwick- hedges and gravel pits; parkland; garden; sand dunes. shire, 4 males, 1.iv.1999, J.E. Smith leg. (HRI). Reared from nests of moles (several rearings), common shrew, red squirrel, rabbit and from old wasp’s Ecological notes: found in gardens; glasshouses, build- nest. ii–xi. ings, soil of pot plants. Reared from decaying roots of young orchids; primula seedlings. ii, iv, vi, ix–x.

BRADYSIA NORMALIS FREY, 1948 Literature: Freeman (1983a): 168; Freeman (1983b): BRADYSIA PALLIPES (FABRICIUS, 1787) 35; Freeman (1987): 197; Laurence (1994): 118; Men- Synonyms: agilis (Winnertz, 1867); brunnipes zel (1998): 20; Chandler et al. (2002): 122 [in part as (Meigen, 1804); conica (Grzegorzek, 1884); dispar Bradysia leptoptera, misidentification and locality (Winnertz, 1867); engadinica (Winnertz, 1867); fallax given in error as Lennymore Bog]. (Winnertz, 1867); kowarzii (Grzegorzek, 1884); laeta (Grzegorzek, 1884); luctuosa (Winnertz, 1867); New to Irish list. morbosa (Winnertz, 1867); picipes (Zetterstedt, 1838); Material examined: Barvas, Lewis, Western Isles, 9 rufipodex (Frey, 1945); rufipodex var. elysiaca (Frey, males, 2 females, 26.vi.1984, Laurence leg. (DEI, 1945); spreta (Winnertz, 1867); tristis (Winnertz, PBLN). Lennymore Bay, Antrim, Northern Ireland, 1867); umbratica (Zetterstedt, 1851). 1 male, 22.iv.1976, Reid leg. (PPCM). Loch Morlich, Literature: ? Walker (1848): 105 [as Sciara brunnipes Highland Region, 29 males, 14.vii.1936, Coe leg. and Sciara annulata, misidentification]; ? Walker (BMNH). Stanford, Norfolk, 1 male, 27.iii.−10.iv.1988, (1856): 52 [as Sciara brunnipes]; Edwards (1925a): Collier leg. (PPWM). Walberswick, Suffolk, 1 male, 538 [in part as Sciara brunnipes and Sciara annulata, 15.vi.−29.vi.1988, Foster & Procter leg. (PBLN). misidentification]; ? Edwards (1938): 201 [as Sciara Ecological notes: found in ancient pine woodland; salt- (Neosciara) umbratica]; Chandler (1978): 209 [as marsh and reed beds; males swarming in shelter of Bradysia umbratica, misidentification]; Skidmore, dunes, in sand among marram in an exposed situation Limbert & Eversham (1978): 76; Freeman (1983b): 34;

Skidmore (1991): 149; Laurence (1994): 118 [in part]; Ecological notes: common in broad-leaved and carr Laurence (1996): 87 [in part]; Laurence & James woodland; wetlands including fens, water meadows (1997): 166; Chandler et al. (1996): 258 [in part]; Men- and sedge beds; grazed meadow; grassland; gardens; zel (1998): 20; White et al. (2000): 207 [all as Bradysia open montane habitats; mushroom beds. Reared from brunnipes]; Heller (2004): 248 [brunnipes (Meigen) as cow and sheep dung. ii–xi. synonym to pallipes (Fabricius)].

Material examined: Ardscull Mote, Kildare, 1 male, BRADYSIA PECTORALIS (STAEGER, 1840) 23.vi.1975, Chandler leg. (PPCM). Beddington, Synonym: castanea Mohrig & Menzel, 1990. Greater London, 4 males, 14.iii.1935 (reared), Par- Literature: Freeman (1983b): 38; O’Connor & Ashe menter leg. (BMNH). Bone Mill Hollow, Cam- (1991): 29; Chandler et al. (1996): 259; Laurence bridgeshire, 1 male, 29.vi.1996, Drake leg. (NMS). (1994): 118 [all as Bradysia fungicola, misidentiﬁca- Cadair Idris, Gwynedd, 1 male, 13.xi.1993, Laurence tion]; Menzel (1998): 20; Chandler et al. (2002): 122. leg. (PBLN). Cors Erddreiniog, Isle of Anglesey, 1 male, 20.x.1988, Holmes, Boyce & Reed leg. (PPWM). Material examined: Aber Valley, Gwynedd, 1 male, Dyffryn Gwaun, Pembrokeshire, 1 male, 12.x.1987, 7.vii.1987, Chandler leg. (NMS). Archers Wood, Holmes, Boyce & Reed leg. (PPWM). Eday, Orkney, 1 Cambridgeshire, 1 male, 27.v.1990, J.H. Cole leg. male, 1.vi.1988, Laurence leg. (PBLN). Epping Forest, (PJCB). Ashberry Pastures, North Yorkshire, 1 male, Essex, 1 male, 29.iii.1999; 1 male, 20.ix.−10.x.1999, all 14.x.1990, Chandler leg. (NMS). Ballynalacken Cas- Dagley leg. (HRI, PPCM). Felden, Hertfordshire, 1 tle, Clare, 1 male, 19.v.1979, Chandler leg. (PPCM). male, 19.xi.1896, A. Piffard leg. (BMNH). Glen Gairn, Ban-y-Gor Wood, Gloucestershire, 1 male, 12.ix.2002, Aberdeenshire, 1 male, 17.vii.1997, Chandler leg. Gibbs leg. (PDGB). Bedford Purlieus, Cambridgeshire, (PPCM). Glendurgan, Cornwall, 1 male, 3.x.1983, 2 males, 14.v.1988, J.H. Cole leg. (PJCB). Beltra Chandler leg. (PPWM). Glyn Ceiriog, Wrexham, 2 Lough, Sligo, 1 male, 13.vi.1985, Chandler leg. males, 26.vi.−7.vii.1995, Brock leg. (HRI). Goonvrea, (PPCM). Birk Crag Wood, Harrogate, North Yorkshire, Cornwall, 1 male, 2.vii.−3.vii.1983, Chandler leg. 1 male, 16.vii.1977, Chandler leg. (NMS). Bolt Wood, (PPCM). Goring-on-Thames, Oxfordshire, 1 male, Stockerston, Leicestershire, 2 males, 24.viii.2003, 24.iv.1962; 1 male, 21.vi.1963, all J.H. Cole leg. Menzel leg. (DEI). Bolton Wood, North Yorkshire, 4 (PJCB). Harlesden, Greater London, 5 males, 20.vi. males, 14.ix.1931, Cheetham leg. (BMNH). Boundless 1894, Austen leg. (BMNH). Hickling, Norfolk, 3 males, Copse, Surrey, 1 male, 26.vii.2002; 1 male, 8.x.2002, no date, Foster & Procter leg. (PBLN). Ingleborough, all Chandler leg. (PPCM). Brancaster, Norfolk, 12 North Yorkshire, 2 males, 13.v.1921, Edwards leg. males, 2.ix.1988; 15 males, 2.ix.−16.ix.1988; 10 males, (BMNH). King’s Walden, Hertfordshire, 1 male, 28.x.1988, all Foster & Procter leg. (PBLN). Brandon 26.iv.1964, Colyer leg. (BMNH). Leckford Estate, Marsh, Ryton-on-Dunsmore, West Midlands, 1 male, Hampshire, 1 male, 4.x.1970, Stubbs leg. (PPCM). 22.viii.2002, Menzel leg. (DEI). Brasenose, Oxford- Letchworth, Hertfordshire, 1 male, viii.1917, Edwards shire, 1 male, 28.viii.1915, Hamm leg. (BMNH). Brig- leg. (BMNH). Llyn Ystumllyn, Gwynedd, 1 male, steer Park, Cumbria, 1 male, 14.vi.1999, Howe leg. 19.x.1988, Holmes, Boyce & Reed leg. (PPWM). (NMS). Brithdir-Isaf, Gwynedd, 3 males, 25.ix.1971; 1 Lopham Fen, Norfolk, 1 male, 22.ii.1990, Withers leg. male, 25.x.1971, all Hutson leg. (BMNH). Bucklebury (PPWM). Meall nan Tarmachan, Perth & Kinross, 8 Common, Berkshire, 1 male, 30.viii.1983; 1 male, males, 8 females, 2.x.−20.x.2002 (reared), Bland leg. 5.iii.1997 (reared), all Chandler leg. (PPCM). Burn- (NMS). Newlaw Moss, Dumfries & Galloway, 1 male, ham Beeches, Buckinghamshire, 1 male, 13.v.1990, 20.viii.1979, Chandler leg. (NMS). Newmarket, Cam- Chandler leg. (PPCM). California Country Park, Berk- bridgeshire, 1 male, 21.iii.1897, Verrall leg. (BMNH). shire, 2 males, 19.v.1998, Chandler leg. (NMS). Carie, Oxford University Parks, Oxford, Oxfordshire, 1 male, Rannoch, Perth & Kinross, 1 male, 29.v.1998, Chan- 1 female, 18.iv.1913; 1 male, 29.iv.1914, all Hamm leg. dler leg. (NMS). Castle Loch, Dumfries & Galloway, 1 (BMNH). Pont Aber, Carmarthen, 1 male, 12.x.1995, male, 16.ix.1989, Chandler leg. (PPCM). Castlerea, Chandler leg. (PPCM). Rhôs-y-gad, Isle of Anglesey, 1 Roscommon, 1 male, 28.ix.1977, Chandler leg. male, 5.x.1988, Holmes, Boyce & Reed leg. (PPWM). (PPCM). Chalford, Gloucestershire, 1 male, 12.x.1979, Taplow, Buckinghamshire, 1 male, 1 female, Chandler leg. (PPCM). Charleville Wood, Offaly, 1 26.iv.1968, Chandler leg. (PPCM). The Loke, Norwich, male, 27.v.1984; 4 males, 15.vi.1985; 2 males, Norfolk, 1 male, 20.iv.1987; 3 males, 26.iv.1987; 1 19.ix.1999, all Chandler leg. (NMS, PPCM, PPWM). male, 21.iv 1988, all Laurence leg. (PBLN). Thornham Chigwell Row Wood, Chigwell Row, Essex, 1 female, Magna, Suffolk, 1 male, 23.ii.1990, Withers leg. 8.x.2002; 1 female, 16.x.2002; 1 female, 22.x.2002, all (PPWM). No locality details, Kent, 3 males, vi.1928, Ismay & Schulten leg. (DEI, UMO). Chippenham Fen, Theobald leg. (BMNH). Cambridgeshire, 1 male, 21.x.1990, Chandler leg.

(NMS). Chobham Common, Surrey, 1 male, 8.ix.1974, (PJCB). Hickling, Norfolk, 1 male, v.1989, Foster & Chandler leg. (PPCM). Cilast Isaf, Pembrokeshire, 1 Procter leg. (PBLN). How Hill, Norfolk, 1 male, male, 7.x.1973, Chandler leg. (PPCM). Cleeve Her- 13.viii.1989, Withers leg. (PPWM). Knebworth, Hert- onry, Avon, 1 male, 11.ix.2000, Gibbs leg. (PDGB). fordshire, 1 male, vi.1922, Edwards leg. (BMNH). Coed Dolgarrog, Conwy, 1 male, 6.vii.1987, Chandler Lady’s Island Lake, Wexford, 1 male, 25.v.1987, J.P. leg. (NMS). Coombe Abbey Country Park, Coventry, O’Connor leg. (NMID). Lady’s Wood, Cambridgeshire, Warwickshire, 2 males, 28.viii.2003, Menzel leg. 1 male, 3.vi.1989, J.H. Cole leg. (PJCB). Lancaut (DEI). Cop Mere, Staffordshire, 1 male, 23.v.1994, Nature Reserve, Gloucestershire, 1 male, 5.ix.2002, Chandler leg. (PPCM). Cosford Mill, Surrey, 1 male, Gibbs leg. (PDGB). Lea Meadows, Newtown Linford, 15.ix.1973, Chandler leg. (PPCM). Cothill, Oxford- Leicestershire, 2 males, 1 female, 24.viii.2003, Menzel shire, 1 male, 14.vi.1986; 1 male, 3.vi.1990, all Chan- & J.E. Smith leg. (DEI, HRI). Leckford Estate, Hamp- dler leg. (NMS). Cotterdale, North Yorkshire, 1 male, shire, 1 male, 5.x.1995; 1 female, 23.ix.1997; 1 male, 27.ix.1986, J.H. Cole leg. (PJCB). Craigmore Wood, 12.x.1997; 2 males, 25.vii.1998, all Chandler leg. Aberfoyle, Stirling, 1 male, 21.vi.1999, Chandler (NMS, PPCM). Linton, Devon, 1 male, 16.vi.1989, leg. (PPCM). Crickhowell, Powys, 1 female, ix.1936, Chandler leg. (NMS). Loch Achilty, Highland Region, 1 Edwards leg. (BMNH). Culbin Forest, Highland male, 28.v.2002, Chandler leg. (PPCM). Logie, Moray, Region, 1 male, 27.v.2002, Chandler leg. (PPCM). 1 male, 1 female, 8.ix.1910, Jenkinson leg. (BMNH). Dartmeet, Devon, 2 males, 27.v.1978, J.H. Cole leg. Lough Slevin, Westmeath, 2 males, 28.vi.1987, Chan- (PJCB). Dowlands Cliff, Devon, 1 male, 9.vi.1937, dler leg. (NMS). Low Pasture, Hawnby, North York- Edwards leg. (BMNH). Dromore Forest, Kerry, 1 male, shire, 1 male, 12.x.1990, Chandler leg. (NMS). Maen 15.x.1973, Chandler leg. (PPCM). Ebbor Gorge, Som- Porth, Cornwall, 1 male, 5.x.1983, Chandler leg. erset, 1 male, 16.x.1986, Chandler leg. (NMS). Epping (PPCM). Matley Bog, New Forest, Hampshire, 1 male, Forest, Essex, 1 male, 3.v.−9.v.1999; 1 male, vi.1999, viii.1936, Edwards leg. (BMNH). Mulgrave Wood, all Dagley leg. (HRI, PPCM); 1 male, 25.v.1999, Chan- North Yorkshire, 6 males, 23.viii.−3.ix.1937, Edwards dler leg. (PPCM). Ewden Valley, Shefﬁeld, South York- leg. (BMNH). Newbridge Fen, Kildare, 1 male, 8.viii.− shire, 1 female, 21.ix.1971, Chandler leg. (NMS). 19.viii.1998; 1 male, 8.ix.−19.ix.2000, all Withers leg. Forest of Dean, Gloucestershire, 1 male, 27.v.1962, (PPWM). Oaklands, Wexford, 1 male, 29.v.1987, J.P. K.G.V. Smith leg. (BMNH). Galway’s Bridge, Killar- O’Connor leg. (NMID). Oakshott, Hampshire, 1 male, ney, Kerry, 1 male, 1 female, 9.ix.1981, J.P. & M.A. 14.vi.1979, Chandler leg. (PPCM). Olchon Valley, O’Connor leg. (NMID). Gatewen Marsh, Denbighshire, Herefordshire, 1 male, 21.v.1977, Chandler leg. 1 male, 12.x.1988, Holmes, Boyce & Reed leg. (PPCM). Richmond Park, Greater London, 1 male, (PPWM). Gearagh, Cork, 1 male, 1 female, 26.ix.1990, 6.vi.1992, Chandler leg. (NMS). Roudsea Wood, Cum- de Courcy Williams leg. (DEI, NMID). Glanvilles bria, 1 male, 16.vi.1999, Chandler leg. (PPCM). Roy- Wootton, Dorset, 1 male, viii.1939, Edwards leg. don Common, Norfolk, 1 male, 5.vi.1993, Chandler leg. (BMNH). Glen Coiltie, Highland Region, 1 male, (PPCM). Santon Downham, Suffolk, 1 male, viii.1935, 29.v.1994, Chandler leg. (PPCM). Glengarriff Forest, Edwards leg. (BMNH). Sladen, Hampshire, 1 male, Cork, 1 male, 4.vii.1985, J.P. & M.A. O’Connor leg. vi.1939, Edwards leg. (BMNH). Sherwood Forest, (NMID). Glen of the Downs, Wicklow, 1 male, Nottinghamshire, 1 male, 22.x.−24.x.1922, Edwards 27.vii.1981; 1 male, 3.x.1982, all J.P. O’Connor leg. leg. (BMNH). Spartum Fen, Oxfordshire, 1 male, (NMID). Goathland, North Yorkshire, 5 males, 1.vi.1991, Chandler leg. (PPCM). Stradey Wood, Car- 5.ix.1937, Edwards leg. (BMNH). Gorge of Avon, South marthen, 1 male, 13.x.1995, McLean leg. (PPCM). Lanarkshire, 1 female, 30.v.1909; 1 male, 18.viii.1909; Strawberry Beds, River Liffey, Dublin, 1 male, 1 male, 24.x.1910, all King leg. (BMNH). Great Eller- 26.ix.1999, Chandler leg. (NMS). Templehouse side, Cumbria, 1 male, 15.vi.1999, Chandler leg. Demesne, Sligo, 1 male, 1.x.1977, Chandler leg. (PPCM). Great Elm, Avon, 2 males, 16.x.1986, Chan- (PPCM). Thompson Common, Norfolk, 1 male, dler leg. (PPCM). Great Ofﬂey, Hertfordshire, 1 male, 1.vi.1985, Chandler leg. (NMS). Tickenham Hill, Avon, 29.ix.1968, Chandler leg. (NMS). Greywell Fen, 1 male, 30.v.2001, Gibbs leg. (PDGB). Tocil Wood, War- Hampshire, 1 male, 15.x.1989; 2 males, 11.viii.1999, wick University Campus, Coventry, Warwickshire, 4 all Chandler leg. (NMS, PPCM). Grindon Wood, males, 28.viii.2003, Menzel & J.E. Smith leg. (DEI, Staffordshire, 1 male 13.x.1997, Chandler leg. HRI). Tomies Wood, Killarney, Kerry, 1 female, (PPCM). Hafod Wood, Denbighshire, 1 male, 25.ix.1981, J.P. O’Connor leg. (NMID). West Walk, 24.v.1994, Chandler leg. (PPCM). Haugh Wood, Here- Hampshire, 2 males, 6.vi.1998, Chandler leg. (NMS). fordshire, 1 male, 12.x.1998, Chandler leg. (PPCM). Whale Moss, Shropshire, 1 male, 29.viii.1931, C.H.W. Hay Wood, Baddesley Clinton, Solihull, West Mid- Pugh leg. (BMNH). Whiddon Deer Park, Devon, 1 lands, 1 male, 25.viii.2002, Menzel leg. (DEI). Hem- male, 2.vi.2000, Chandler leg. (PPCM). Wicken Fen, bury Wood, Devon, 1 male, 29.v.1978, J.H. Cole leg. Cambridgeshire, 1 male, 21.v.1990; 1 male, 14.v.1992,

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 BRITISH SCIARID FAUNA 57 all J.H. Cole leg. (PJCB). Windsor Forest, Berkshire, 10.v.1934, Edwards leg. (BMNH). Ballynafid Lake, 1 male, 8.vi.1979; 6 males, 6.ix.1986; 2 males, Westmeath, 1 male, 10.v.1989, Withers leg. (PPWM). 13.vi.1987; 2 males, 8.viii.1987; 2 males, 27.v.1989; 1 Bayford, Hertfordshire, 1 male, 15.x.1931, Edwards male, 15.vi.1991; 1 male, 15.v.1994, all Chandler leg. leg. (BMNH). Beaford Bridge, Devon, 1 male, (NMS, PPCM, PPWM). Witcombe Wood, Gloucester- 14.vi.1989, Chandler leg. (NMS). Bix Bottom, Oxford- shire, 1 male, 2 females, 12.x.1979, Chandler leg. shire, 1 male, 23.ix.2000, Raper leg. (PDGB). Blean (NMS). Witherslack, Cumbria, 2 males, 15.vi.1999, Wood, Kent, 1 male, 19.v.1973, Parmenter leg. Chandler leg. (NMS). Woodwalton Fen, Cam- (BMNH). Bleathwaite Coppice, Cumbria, 1 male, bridgeshire, 4 males, 24.v.1940, Edwards leg. 8.x.2001, Gibbs leg. (PDGB). Bradfield Wood, Suffolk, (BMNH); 1 male, 4.vi.1977, J.H. Cole leg. (PJCB). 1 male, 9.iv.−16.iv.1990; 1 male, 16.viii.−22.viii.1991, Wytham Woods, Wytham, Oxfordshire, 1 male, all Laurence leg. (PBLN). Brendon, Devon, 2 males, 21.viii.2002, Menzel leg. (DEI). viii.1935, Edwards leg. (BMNH). Bridgeham, Thetford Chase, Norfolk, 1 male, 26.iv.1969, Cogan leg. Ecological notes: very common in broad-leaved and (BMNH). Brockenhurst, New Forest, Hampshire, 1 mixed woodland; conifer plantations; wooded river male, 23.v.1969, Hutson leg. (BMNH). Brockenhurst and lake sides; common in carr woodland; heathland; Wood, New Forest, Hampshire, 1 male, 8.vii.1990, wetlands including fen, bog, wet meadows, grassland Chandler leg. (PPCM). Brook Wood, Bromley, Greater with pingo pools, sedge beds Typha latifolia and London, 2 males, 3 females, 21.iv.1966, Chandler leg. marsh; former gravel workings; calcareous grassland; (PPCM). Buckingham Palace Garden, Greater Lon- ancient parkland; dunes. Reared from the fungus don, 1 male, 19.ii.−27.iii.1997, D.J. Carter & C.W. Laetiporus sulphureus. v–x. Plant leg. (NMS). Burnham Beeches, Buckingham- shire, 1 male, 5.vii.−20.vii.1995, Ismay leg. (PPCM); 1 male, 20.iv.1998; 1 male, 4.x.1999, all Chandler leg. BRADYSIA PERAFFINIS TUOMIKOSKI, 1960 (NMS, PPCM). Byfleet, Surrey, 2 males, 14.v.1932, New to British and Irish list. Coe leg. (BMNH). Cadair Idris, Gwynedd, 1 male, Material examined: Birks of Aberfeldy, Perth & Kin- 19.vi.1993; 1 male, 22.x.1993; 1 male, 13.xi.1993, all ross, 1 male, 23.vi.1999, Chandler leg. (PPCM). Col- Laurence leg. (PBLN). California Country Park, Berk- linpark Wood, Gloucestershire, 1 male, 3.ix.2002, shire, 1 male, 6.vii.1997, Chandler leg. (PPCM). Gibbs leg. (PPCM). Coombe Dingle, Avon, 1 male, Canonteign Falls, Devon, 1 female, 22.vi.2002, J.E. 18.vii.1987, Chandler leg. (PPCM). Gearagh, Cork, 1 Smith leg. (HRI). Castle Neroche, Somerset, 1 male, male, 2.iv.1990; 2 males, 26.iv.1990, all de Courcy Wil- 12.v.1936, Edwards leg. (BMNH). Cefn Grwhyd, liams leg. (DEI, NMID). Peppercombe, Devon, 1 male, Neath/Port Talbot, 1 male, 12.vii.1989, Holmes, Boyce 13.vii.2001, Chandler leg. (PPCM). & Reed leg. (PPWM). Charterhouse Alder Holt, Sur- rey, 1 male, 16.vii.1989, Chandler leg. (PPCM). Chat- Ecological notes: found in damp and acid broad-leaved elherault, South Lanarkshire, 1 male, 27.v.1994, woodland; by stream in cliff gully. iv, vi–vii, ix. Chandler leg. (PPCM). Chigwell Row Wood, Chigwell Row, Essex, 2 males, 28.vi.2002; 1 male, 16.x.2002, all Ismay & Schulten leg. (DEI, UMO). Clowes Wood, BRADYSIA PLACIDA (WINNERTZ, 1867) Warwickshire, 2 males, 19.iv.1987, Withers leg. Synonym: fimbricauda Tuomikoski, 1960. (PPWM). Coed Crafnant, Gwynedd, 1 male, 16.x.1992, Chandler leg. (NMS). Coed Dolbebin, Gwynedd, 1 Literature: Freeman (1983a): 169; Freeman (1983b): male, 16.x.1992, Chandler leg. (NMS). Coombe Abbey 37; O’Connor & Ashe (1991): 29; Laurence (1994): 118; Country Park, Coventry, Warwickshire, 1 male, Chandler et al. (1996): 259; Laurence (1996): 86; Lau- 28.viii.2003, J.E. Smith leg. (HRI). Craig na Cailloch, rence (1997a): 85; Laurence (1997b): 228; Laurence & Perth & Kinross, 1 male, 5.vi.−18.vi.1932, Edwards James (1997): 166 [all as Bradysia fimbricauda]; Men- leg. (BMNH). Crowgutter Wood, Staffordshire, 1 male, zel (1998): 20; Chandler (2001): 201. 12.x.1997, Chandler leg. (PPCM). Cuckoo Wood, Material examined: Allerthorpe, East Riding of Downe, Greater London, 1 male, 20.vii.1985, Chandler Yorkshire, 1 male, 18.ix.1923, Edwards leg. (BMNH). leg. (PPCM). Cwm Bychan, Gwynedd, 1 male, Ampfield, Hampshire, 1 male, 20.xi.1932, Edwards 16.x.1982, Chandler leg. (PPCM). Dickleburgh, Nor- leg. (BMNH). Annahala bog, Gearagh, Cork, 1 male, folk, 2 males, 7.i.−15.i.1989; 1 male, 29.i.−12.ii.1989, 6.iv.1990; 1 male, 23.vi.1990, all de Courcy Williams all Withers leg. (PPWM). Dinton Pastures Country leg. (DEI, NMID). Ashridge Estate, Hertfordshire, 1 Park, Berkshire, 1 male, 21.v.1995; 1 male, 17.x.1998, male, 8.vii.1999, Chandler leg. (NMS). Austwick, all Chandler leg. (PPCM). Duthlas, Powys, 1 male, North Yorkshire, 2 males, 26.v.−27.v.1934, Edwards 17.x.1987, McLean leg. (PPCM). Epping Forest, Essex, leg. (BMNH). Baldock, Hertfordshire, 1 male, 2 males, 19.iv.1999, Dagley leg. (NMS). Fritham, New

Forest, Hampshire, 2 males, 17.iv.1954, Colyer leg. say, Orkney, 1 male, 7.ix.1989, Laurence leg. (PBLN). (BMNH). Gearagh, Cork, 1 male, 2.iv.1990, de Courcy Oakley Wood, Bishops Tachbrook, Warwickshire, 1 Williams leg. (NMID). Gibriddings Wood, Stafford- male, 14.x.2001, Chandler leg. (HRI). Paramoor Wood, shire, 1 male, 14.x.1997, Chandler leg. (PPCM). Cornwall, 2 males, 7.vii.1983, Chandler leg. (PPCM). Glen Affric, Highland Region, 1 male, 25.vi.1999, Pashford, Norfolk, 1 male, 24.vi.−8.vii.1988, Foster & Chandler leg. (PPCM). Glendalough, Wicklow, 1 male, Procter leg. (PBLN). Pentewan, Cornwall, 2 males, 24.iv.1989, J.P. & M.A. O’Connor leg. (NMID). Glen- 7.vii.1983, Chandler leg. (NMS). Pine Wood, Berk- garriff, Cork, 1 male, 6.vii.1985, J.P. & M.A. O’Connor shire, 1 male, 10.ix.2001, Gibbs leg. (PDGB). Pond leg. (NMID). Glen Lochay, Stirling, 1 male, 5.vi.− Wood, Chislehurst, Greater London, 6 males, 18.vi.1932, Edwards leg. (BMNH). Greywell Fen, 20.iii.1966 (reared), Chandler leg. (PPCM). Powler’s Hampshire, 1 male, 2.viii.1998, Chandler leg. (PPCM). Piece, Devon, 1 male, 13.x.1988, Chandler leg. Grindon Wood, Staffordshire, 1 male, 13.x.1997, Chan- (PPCM). Pulham St. Mary, Norfolk, 1 male, 2.v.1988, dler leg. (PPCM). Hafod Garregog, Gwynedd, 1 male, Withers leg. (PPWM). Radwell, Hertfordshire, 2 13.x.1988, Holmes, Boyce & Reed leg. (PPWM). males, 21.x.1917, Edwards leg. (BMNH). Rahan, Cork, Hartslock, Oxfordshire, 1 male, 6.v.2000, Raper leg. 1 male, 6.vii.1989, J.P. O’Connor leg. (NMID). Rhine- (PDGB). Hay Wood, Baddesley Clinton, Solihull, West ﬁelds, New Forest, Hampshire, 3 males, 16.v.1956, Midlands, 1 male, 25.viii.2002, Menzel leg. (DEI). Colyer leg. (BMNH). Ringstead, Dorset, 1 male, Haytor, Devon, 1 male, 23.vi.2002, J.E. Smith leg. viii.1939, Edwards leg. (BMNH). Riversmead, Devon, (HRI). Hencliff Wood, Avon, 1 male, 14.ii.2001, Gibbs 1 male, 13.x.1988, M.N. Pugh leg. (NMS). Rosevear, leg. (PDGB). Hickling, Norfolk, 1 male, 1.vi.1988; 5 Cornwall, 1 male, 6.vii.1983, Chandler leg. (PPCM). males, x.1988, all Foster & Procter leg. (PBLN). Rough Hill Wood, Studley, Warwickshire, 1 male, Holker Moss, Cumbria, 2 males, 20.v.1938, Edwards 10.iv.2003, J.E. Smith leg. (HRI). Ryton Wood, leg. (BMNH). Humphrey Head, Cumbria, 1 male, Bubbenhall, Coventry, West Midlands, 3 males, 1.viii.−8.viii.1923, Edwards leg. (BMNH). Inholm’s 12.x.2001, Chandler & J.E. Smith leg. (HRI). Sand- Copse, West Sussex, 1 male, 20.vii.1974, Chandler leg. scale Haws, Cumbria, 1 male, 18.vi.1999, Chandler (NMS). Ipley Manor, New Forest, Hampshire, 1 male, leg. (NMS). Savernake Forest, Wiltshire, 2 males, 17.vii.1990, Chandler leg. (PPCM). Killaun Bog, 13.xi.1978, Chandler leg. (PPCM). Scadbury Park, Offaly, 2 males, ix.2000, Withers leg. (PPWM). Kneb- Greater London, 1 male, 3.xi.1984; 2 males, worth, Hertfordshire, 2 males, 10.x.1917, Edwards leg. 21.vii.1985, all Chandler leg. (NMS). Seer Green, (BMNH). Knole Park, Kent, 1 male, 12.xi.1967, Chan- Buckinghamshire, 1 male, 10.xii.1932; 7 males, dler leg. (NMS). Leckford Estate, Hampshire, 2 males, 13.xii.−21.xii.1932 (all reared), all Basden leg. 12.iv.1981; 2 males, 27.vii.1985; 1 male, 15.iii.1992, all (BMNH, NMS). Shapwick Heath, Somerset, 1 male, Chandler leg. (NMS, PPCM). Letchworth, Hertford- 5.vii.1985, Chandler leg. (NMS). Shefford, Bedford- shire, 2 males, vii.1915; 1 male, xi.1917, all Edwards shire, 14 males, x.1917 [in part labelled as ‘sp. no. 4’]; leg. (BMNH). Lion Wood, Redditch, Worcestershire, 2 7 males, 17.xi.1917, all Edwards leg. (BMNH, ZFMK). males, 10.iv.2003, J.E. Smith leg. (HRI). Loch Morlich, Shotover Country Park, Oxfordshire, 1 male, Highland Region, 1 male, 4.vii.1936, Coe leg. (BMNH). 17.x.1999, Chandler leg. (PPCM). Slade of Saggart, Lockhaugh Bank Wood, Derwent Valley, Tyne & Wear, Dublin, 1 male, 1.xi.1981, J.P. O’Connor leg. (NMID). 1 male, 7.vi.−23.vi.1990; 1 male, 24.vii.−13.viii.1990, Spey Bay, Moray, 1 male, 2.vii.1936, Coe leg. (BMNH). all Jessop leg. (PBLN). Lopham Fen, Norfolk, 1 male, Spinningdale, Highland Region, 2 males, 14.v.1994, 2.vii.1989, Withers leg. (PPWM). Lum Pool, Stafford- Entwistle leg. (NMS). Stanford, Norfolk, 1 male, shire, 1 male, 1.ix.1997, Chandler leg. (PPCM). Lundy 27.iii.−10.iv.1988, Collier leg. (PPWM). Sutton Park, Island, Devon, 6 males, 20.vii.−27.vii.1972, Lane leg. Birmingham, West Midlands, 8 males, 7.iv.2003, J.E. (BMNH). Mark Ash Wood, New Forest, Hampshire, Smith leg. (HRI). Thelnetham Fen, Suffolk, 1 male, 5 males, 4.vi.1989; 1 male, 29.vi.1989, all Chandler 20.vi.1989, Withers leg. (PPWM). Thornham Magna, leg. (NMS, PPCM). Market Weston Fen, Suffolk, 1 Suffolk, 1 male, 25.iv.1991, Withers leg. (PPWM). male, 18.iii.1990; 1 male, 12.v.1991, all Withers Thursley Common, Surrey, 1 male, 20.v.1966, Chan- leg. (PPWM). Minsmere, Suffolk, 1 male, 18.x.1996, dler leg. (PPCM). Tomies Wood, Killarney, Kerry, 1 McLean leg. (PPCM). Moccas Park, Herefordshire, 1 male, 15.ix.1981, J.P. O’Connor leg. (NMID). Tom- male, 11.x.1998, Chandler leg. (NMS). Monk’s Wood, intoul, Moray, 16 males, 8.viii.−11.viii.1937, Coe leg. Cambridgeshire, 1 male, 25.ix.1922, Edwards leg. (BMNH). Torc Falls, Killarney, Kerry, 1 male, (BMNH). Mulgrave Wood, North Yorkshire, 1 male, 11.vii.1990, Withers leg. (PPWM). University 23.viii.−3.ix.1937, Edwards leg. (BMNH). Murrough, Museum, Oxford, Oxfordshire, 1 male, 28.i.1921, Wicklow, 2 males, v.1989, Blackith leg. (PPWM). New- Hamm leg. (UMO). Uragh Wood, Kerry, 1 male, bold Comyn Park, Leamington Spa, Warwickshire, 1 8.vii.1990, Withers leg. (PPWM). Walberswick, Suf- male, 15.vii.2001, J.E. Smith leg. (HRI). North Ronald- folk, 2 males, 1.vi.−15.vi.1988, Foster & Procter leg.

(PBLN). Waterperry Wood, Oxfordshire, 1 male, shire, 2 males, 2 females, 28.viii.2003, Menzel & J.E. 5.v.1940, Edwards leg. (BMNH). Welcombe Valley, Smith leg. (DEI, HRI). Grafham Water, Savage’s Devon, 1 male, 15.x.1988, Chandler leg. (PPWM). Creek, Cambridgeshire, 1 male, 30.v.1988, J.H. Cole Wellesbourne, Warwickshire, 1 male, 5.viii.− leg. (PJCB). Greywell Fen, Hampshire, 3 males, 17.viii.2000, J.E. Smith leg. (HRI). West Harling Com- 29.viii.2002, Chandler leg. (HRI). Hartslock, Oxford- mon, Norfolk, 1 male, 25.v.1987; 1 male, 1.iv.1989, all shire, 1 male, 10.vii.1999, Chandler leg. (NMS); Withers leg. (PPWM). Weston, Hertfordshire, 2 males, 1 male, 14.v.2000, Raper leg. (PDGB). Hemingford, 4.x.1931, Edwards leg. (BMNH). West Town, Avon, 1 Cambridgeshire, 3 males, ix.1938, Edwards leg. male, 22.iii.1953, d’Assis-Fonseca leg. (NMS). Wheat- [paratypes of Bradysia edwardsi] (BMNH). Highgate fen Broad, Norfolk, 1 male, 8.x.1988, Withers leg. Wood, Greater London, 1 male, 1998–99, Hackett leg. (PPWM). Whichford Wood, Moreton-in-Marsh, War- (NMS). Lackham Park, Wiltshire, 1 male, 5.viii.2002; wickshire, 7 females, 11.x.2001, J.E. Smith leg. (HRI). 3 males, 20.ix.2002, all Chandler leg. (PPCM). Leck- Whitacre Heath, Warwickshire, 1 female, 13.x.2001, ford Estate, Hampshire, 1 male, 11.x.1999, Chandler J.E. Smith leg. (HRI). Windsor Forest, Berkshire, 1 leg. (PPCM).Letchworth, Hertfordshire, 1 male, 1 male, 10.vi.1939, Edwards leg. (BMNH); 4 males, female, vi.1917; 1 male, ix.1917, all Edwards leg. 1.v.1966; 1 male, 3.xi.1990, all Chandler leg. (NMS, [paratypes (males) of Bradysia edwardsi] (BMNH). PPCM). Woodwalton Fen, Cambridgeshire, 1 male, Lopham Fen, Norfolk, 1 male, 16.iv.1984, Withers leg. 24.v.1940, Edwards leg. (BMNH). Yarner Wood, (PPWM). Monkfield Wood, Cambridgeshire, 1 male, Devon, 1 male, 28.v.1978, Chandler leg. (NMS). vi.1998, C.W. Plant leg. (PPCM). Mulgrave Wood, North Yorkshire, 2 males, 23.viii.−3.ix.1937, Edwards Ecological notes: very common in all types of wood- leg. [paratypes of Bradysia edwardsi] (BMNH). Nar- land and carr; heathland; moorland; wetlands includ- borough Bog, Whetstone, Leicestershire, 1 male, ing fens, mires, bogs and water meadows; grassland; 24.viii.2003, Menzel leg. (DEI). Newbridge Fen, Kil- scrub; old hedges; old sand and gravel workings; mar- dare, 1 male, no date, Withers leg. (PPWM). Newtown ginal vegetation by upland lake; calcareous valley; Lough, Galway, 1 male, 3.v.1999, Withers leg. parkland; gardens; saltmarsh and reed beds; coastal (PPWM). North Ronaldsay, Orkney, 1 male, 8.ix.1989, sand dunes; open montane habitats. On ivy. Reared Laurence leg. (PBLN). Odstock, Wiltshire, 1 male, 1 from squirrel’s nest and rotten wood. I–x. female, 13.viii.1972, Cranston & Dear leg. (BMNH). Pingle Cutting, Warboys, Cambridgeshire, 1 male, 26.v.1985, J.H. Cole leg. (PJCB). Pingle Wood, War- BRADYSIA POLONICA (LENGERSDORF, 1928) boys, Cambridgeshire, 1 male, 2.v.1990, J.H. Cole leg. Synonyms: edwardsi Freeman, 1983; spinidensa (PJCB). Ringstead, Dorset, 1 male, viii.1939, Edwards Hondru, 1968. leg. (BMNH). Rotherfield Park, Hampshire, 2 males, 2.x.2002, Chandler leg. (PPCM). Ryton Wood, Bubben- Literature: Freeman (1983a): 166; Freeman (1983b): hall, Coventry, West Midlands, 1 male, 25.viii.2002, 35; Laurence (1997b): 228 [all as Bradysia edwardsi]; Menzel leg. (DEI). Salisbury Plain Training Area, Menzel (1998): 20; Menzel & Mohrig (2000): 132; Warminster, Wiltshire, 4 males, 29.v.2003; 1 male, Withers (2002): 114. 30.v.2003, all Ismay & Schulten leg. (DEI, UMO). Material examined: Badger Wood, Cambridgeshire, 1 Searles Farm, Reading, Berkshire, 1 male, 9.vii.2003, male, 1.iv.−3.iv.1998, C.W. Plant leg. (NMS). Bagley Chandler leg. (PPCM). Selsdon, Greater London, 1 Wood, Oxfordshire, 3 males, 20.v.−21.v.1933, Edwards male, 27.v.1934, Coe leg. (BMNH). St. Bride’s Major, leg. [holotype and paratypes of Bradysia edwardsi] Vale of Glamorgan, 1 male, 27.v.1968, Chandler leg. (BMNH); 1 male, 18.v.1936, Edwards leg. (BMNH). (PPCM). Theale, Berkshire, 1 male, 6.viii.2003, Chan- Bedford Purlieus, Cambridgeshire, 1 male, 19.v.1988, dler leg. (PPCM). Thornham Magna, Suffolk, 2 males, J.H. Cole leg. (PJCB). Brampton, Ouse flood meadow, 20.v.1989, Withers leg. (PPWM). Trawsfynydd, Cambridgeshire, 1 male, 23.v.1976, J.H. Cole leg. Gwynedd, 1 male, 5.viii.1988, Holmes, Boyce & Reed (PJCB). Brandon Marsh, Ryton-on-Dunsmore, West leg. (PPWM). Ufton Fields, Warwickshire, 1 male, Midlands, 35 males, 8 females, 22.viii.2002, Menzel & 9.vi.1991, Chandler leg. (PPCM). Warrington, J.E. Smith leg. (DEI, HRI). Cothill, Oxfordshire, 1 Cheshire, 1 male, 10.v.1934 (reared), Holmes-Smith male, 26.v.1936, Edwards leg. [paratype of Bradysia leg. [paratype of Bradysia edwardsi] (BMNH). Wel- edwardsi] (BMNH). Dinton Pastures Country Park, wyn, Hertfordshire, 1 male, vii.1934; 7 males, 21.vii.− Berkshire, 1 male, 24.viii.1995; 1 male, 9.vi.1996, all 24.vii.1934, all Edwards leg. [paratypes of Bradysia Chandler leg. (PPCM). Clonpriest, Cork, 1 male, edwardsi] (BMNH). Willes Meadow, Leamington Spa, 13.vii.1994, Withers leg. (PPWM). Gearagh, Cork, 1 Warwickshire, 2 males, 27.viii.2002, J.E. Smith leg. male, 2.iv.1990, de Courcy Williams leg. (NMID). (HRI). Whitnash brook, Sydenham, Leamington Spa, Coombe Abbey Country Park, Coventry, Warwick- Warwickshire, 2 males, 1 female, 4.ix.2003, J.E. Smith

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 60 F. MENZEL ET AL. leg. (HRI). Wytham Woods, Wytham, Oxfordshire, 49 male, 1.ix.1999, all Chandler leg. (NMS, PPCM). males, 27 females, 21.viii.2002, Menzel & J.E. Smith Letchworth, Hertfordshire, 2 males, vi.1937, Edwards leg. (DEI, HRI). leg. (BMNH). Lough Slevin, Westmeath, 1 male, 28.vi.1987, Chandler leg. (NMS). Nethy Bridge, High- Ecological notes: very common in broad-leaved and land Region, 1 male, 27.viii.1906, King leg. (BMNH). carr woodland; common in wetlands including fens, Old Bridge of Dee, Aberdeenshire, 1 male, 3.viii.1979, sedge beds, water meadows, stream sides and tur- Chandler leg. (PPCM). Scargill, Durham, 1 male, loughs; unimproved calcareous grassland; scrub; near 27.vii.1988, Chandler leg. (NMS). Tomintoul, Moray, shore; saltmarsh. iv–x. 17 males, 13 females, 8.viii.−11.viii.1937, Coe leg. (BMNH). Trawsfynydd, Gwynedd, 1 male, 5.viii.1988, Holmes, Boyce & Reed leg. (PBLN). Wheatfen Broad, BRADYSIA POSTRUFESCENS MOHRIG & MENZEL, 1990 Norfolk, 1 male, 13.vii.1986, Withers leg. (PPWM). New to British list. Woodwalton Fen, Cambridgeshire, 1 male, 4.vi.1977, Material examined: The Loke, Norwich, Norfolk, 1 J.H. Cole leg. (PJCB). Wyre Forest, Worcestershire, 1 male, 26.iv.1987; 1 male, 8.v.1987; 2 males, 9.v.1987; 1 male, 7.viii.1988, Chandler leg. (PPCM). male, 10.viii.1987; 2 males, 15.viii.1987; 2 males, Ecological notes: found in broad-leaved, mixed and 16.viii.1988, all Laurence leg. (DEI, PBLN). West Har- carr woodland; wetlands including fen, sedge beds, ling by River Thet, Norfolk, 1 male, 5.v.1986, Withers Molinia and Myrica, raised mire, marshes and water leg. (PPWM). meadows; garden. iv–ix. Ecological notes: found in water meadow by river; garden. iv–v, viii. BRADYSIA PRATINCOLA TUOMIKOSKI, 1960 Literature: Menzel (1998): 20; Withers (2002): 114. BRADYSIA PRAECOX (MEIGEN, 1818) Material examined: Dickleburgh, Norfolk, 1 male, Synonyms: albinervis (Winnertz, 1867); brevipalpis 26.iv.1979 (reared), Withers leg. (PPWM). Killaun Bog, (Winnertz, 1868); leclerqi (Lengersdorf, 1950); maci- Offaly, 4 males, ix.2000, Withers leg. (PPWM). Loch lenta (Winnertz, 1867); morosa (Winnertz, 1867); noc- Callater, Aberdeenshire, 1 male, 28.viii.−29.viii.1937, ticolor (Winnertz, 1867); simplex (Winnertz, 1867); Coe leg. (BMNH). Sontley Marsh, Wrexham, 1 male, simplex var. subsimplex (Lengersdorf, 1926); unicolor 3.viii.1988, Holmes, Boyce & Reed leg. (PPWM). (Winnertz, 1868). Ecological notes: found on fen; raised bog; garden on Literature: ? Walker (1848): 107; ? Walker (1856): 53 reclaimed agricultural land. Reared from elm log. iv, [all as Sciara]; Freeman (1983b): 38; Menzel (1998): viii–ix. 20. Material examined: Austwick, North Yorkshire, 2 BRADYSIA PSEUDOCAMPESTRIS MOHRIG, 1978 males, 1 female, 26.v.−27.v.1934, Edwards leg. New to British list. (BMNH). Aversley Wood, Cambridgeshire, 1 male, 24.iv.1988, J.H. Cole leg. (PJCB). Baldock, Hertford- Material examined: Salisbury Plain Training Area, shire, 1 male, v.1918; 1 male, 1 female, 10.v.1934, all Warminster, Wiltshire, 2 males, 4.vii.2003, Ismay & Edwards leg. (BMNH). Bank, New Forest, Hampshire, Schulten leg. (DEI, UMO). Stanford battle area, Nor- − 1 male, vi.1939 (reared), Collin leg. (BMNH). Bramp- folk, 3 males, 10.iv. 24.iv.1988, Withers leg. (PPWM). ton, Cambridgeshire, 1 male, 30.iv.1977; 1 male, Ecological notes: found on unimproved calcareous 1.vii.1987, all J.H. Cole leg. (PJCB). Carrigower, Wick- grassland. iv, vii. low, 1 male, 26.v.1984, Chandler leg. (PPCM). Chilbol- ton Common, Hampshire, 2 males, 6.vii.1991, Chandler leg. (PPCM). Chippenham Fen, Cam- BRADYSIA PSEUDODALMATINA MOHRIG & bridgeshire, 1 male, 2.vii.1988, Chandler leg. (NMS). RÖSCHMANN, 1993 Cothill, Oxfordshire, 1 male, 10 females, 5.v.1934, New to British list. Edwards leg. (BMNH). Greywell Fen, Hampshire, 5 Material examined: Cadair Idris, Gwynedd, 1 male, males, 3 females, 29.viii.2002, Chandler leg. (HRI, 4.ix.1993, Laurence leg. (PBLN). Hastings Castle, PPCM). Insh Marshes, Highland Region, 1 male, Hastings, East Sussex, 2 males, 20.xi.2001, Roper leg. 28.v.2002, Uffen leg. (PPCM). Leckford Estate, Hamp- (PPRS). shire, 1 female, 8.v.1976; 3 males, 2 females, 18.ix.− 19.ix.1976; 1 male, 21.ix.1977; 1 male, 23.ix.1977; 3 Ecological notes: found on grassland on maritime males, 2 females, 12.iv.1981; 1 male, 6.vii.1991; 1 sandstone cliff top; open montane habitats. ix, xi.

BRADYSIA RECTINERVIS FREY, 1948 land; fen with sedge pools; sedge bed; wetlands formed Literature: Menzel (1998): 20 [in part as Bradysia by mining subsidence; Juncus basin mire; ﬂoodplain rufescens, misidentiﬁcation]. mire; open montane habitats; decaying vegetation. v– xi. New to British and Irish list.

Material examined: Alvecote Pools, Staffordshire, 2 BRADYSIA REFLEXA TUOMIKOSKI, 1960 males, 13.x.2001, J.E. Smith leg. (HRI). Becky Woods, New to British list. Manaton, Devon, 1 male, 23.vi.2002, J.E. Smith leg. (HRI). Bradfield Wood, Suffolk, 2 males, 9.x.− Material examined: Manifold Valley, Staffordshire, 1 16.x.1990, Laurence leg. (PBLN). Brancaster, Nor- male, 15.vi.1938, Britten leg. (BMNH). folk, 1 male, 1.vii.−15.vii.1988, Foster & Procter leg. Ecological notes: found in broad-leaved woodland. vi. (PBLN). Burnham Beeches, Buckinghamshire, 5 males, 28.v.−1.vii.1996; 5 males, 26.ix.−26.xi.1996, all Ismay leg. (PPCM). Cadair Idris, Gwynedd, 29 males, BRADYSIA REGULARIS (LENGERSDORF, 1934) 19.vi.1993; 1 male, 3.vii.1993; 3 males, 17.vii.1993; 4 Synonym: subnervosa Frey, 1948. males, 1.viii.1993; 8 males, 16.viii.1993; 39 males, 4.ix.1993; 1 male, 22.x.1993; 1 male, 4.ix.1997, all Literature: Freeman (1983b): 37 [in part]; Laurence & Laurence leg. (DEI, PBLN). Canonteign Falls, Devon, James (1997): 166; Menzel (1998): 20. 1 male, 22.vi.2002, J.E. Smith leg. (HRI). Chigwell Material examined: Cadair Idris, Gwynedd, 2 males, Row Wood, Chigwell Row, Essex, 1 male, 15.ix.2002, 4.ix.1993; 1 male, 22.x.1993, all Laurence leg. (PBLN). Ismay & Schulten leg. (DEI). Coed Tycanol, Pem- Cors Caranod, Ceredigion, 1 male, 10.ix.1986; 1 male, brokeshire, 1 male, 2.vi.−10.vi.1999; 1 male, 20.vi.− 17.ix.1986; 1 male, 24.ix.1986; 1 male, 1.x.1986,all 29.vi.1999; 1 male, 29.vi.−7.ix.1999, all Levey & Holmes, Boyce & Reed leg. (PPWM). Cors Geirch, Pavett leg. (PPCM). Gearagh, Cork, 1 male, Gwynedd, 1 male, 28.vii.1988; 1 male, 21.x.1988, all 26.ix.1990, de Courcy Williams leg. (DEI). Gwaun Holmes, Boyce & Reed leg. (PPWM). Cors Graianog, Ystrad Caron, Ceredigion, 1 male, 18.ix.1986, Gwynedd, 1 male, 28.vii.1988; 1 male, 4.viii.1988, all Holmes, Boyce & Reed leg. (PPWM). Hafod Garregog, Holmes, Boyce & Reed leg. (PPWM). Cors Llyn Farch Gwynedd, 1 male, 16.vi.−30.vi.1999, Levey & Pavett a Llyn Fanod, Ceredigion, 1 male, 10.ix.1986, Holmes, leg. (PPCM). Illtyd Pools, Powys, 1 male, 11.ix.1989, Boyce & Reed leg. (PPWM). Culrossie Wood, Highland Holmes, Boyce & Reed leg. (PPWM). Lackham Park, Region, 1 male, 24.x.2000, Chandler leg. (PPCM). Wiltshire, 1 male, 14.x.2002, Chandler leg. (PPCM). Hickling Broad, Norfolk, 1 male, viii.1937, Edwards Loch Callater, Aberdeenshire, 1 male, 27.vi.1938, Coe leg. (BMNH). Rhôsgyll Fawr, Gwynedd, 1 male, leg. (BMNH). Loch Park, Moray, 15 males, 12.viii.− 14.vii.1988, Holmes, Boyce & Reed leg. (PPWM). 14.viii.1937, Coe leg. (BMNH). Mark Ash Wood, New Forest, Hampshire, 1 male, 2.vi.1984, Chandler leg. Ecological notes: found in mixed woodland, mainly (PPCM). Market Weston Fen, Suffolk, 1 male, pine; alder carr; grazed wet heath; wetlands including 12.v.1991, Withers leg. (PPWM). Mid Yell, Shetland, valley fen, bog, sedge bed, basin mire with Schoenus 1 male, ix.1990, Laurence leg. (PBLN). Murrough, nigricans, Sphagnum and Phragmites and grazed Wicklow, 1 male, 8.v.1989, Withers leg. (PPWM). New Molinia and Myrica; open montane habitats. vii–x. Forest, Hampshire, 1 male, 21.ix.1909, Adams leg. Taxonomic note: Highland Region record in Freeman (BMNH). Rhôsili Down, Swansea/Abertawe, 1 male, (1983b) was of Scatopsciara subciliata. The Salop 3.x.1989, Holmes, Boyce & Reed leg. (PPWM). [= Shropshire] and Herts [= Hertfordshire] records Rotherfield Park, Hampshire, 1 male, 2.x.2002, were Bradysia nigrispina Menzel sp. nov. Chandler leg. (PPCM). Ryton Wood, Bubbenhall, Cov- entry, West Midlands, 4 males, 12.x.2001, J.E. Smith leg. (HRI); 4 males, 25.viii.2002, Menzel & J.E. Smith BRADYSIA SCABRICORNIS TUOMIKOSKI, 1960 leg. (DEI, HRI). Sutton Common, Dorset, 1 male, Literature: Laurence (1994): 116 [in part as Bradysia 24.v.−31.v.1999, George leg. (PPCM). West End Com- triseriata, misidentification]; Menzel (1998): 20. mon, Esher, Surrey, 1 male, 19.x.2001; 1 male, 17.vii.2002, all Chandler leg. (PPCM). Witherslack, New to Irish list. Cumbria, 1 male, 15.vi.1999, Chandler leg. (PPCM). Material examined: Black Cliff, Monmouthshire, 1 Yell, Shetland, 1 male, 3.ix.−19.ix.1990, Laurence leg. male, 9.vii.1985, Chandler leg. (PPCM). Blackditch (PBLN). Wood, Wicklow, 1 male, vi.1989, Withers leg. (PPWM). Ecological notes: very common in broad-leaved and Bolt Wood, Stockerston, Leicestershire, 1 male, carr woodland; amongst fallen pine needles; heath- 24.viii.2003, Menzel leg. (DEI). Coed Tycanol, Pem-

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 62 F. MENZEL ET AL. brokeshire, 1 male, 10.v.−20.v.1999; 1 male, 10.vi.− ford Wood, Moreton-in-Marsh, Warwickshire, 1 male, 27.vi.1999, all Levey & Pavett leg. (PPCM). Corfe 11.x.2001, J.E. Smith leg. [paratype of Bradysia Common, Dorset, 1 male, 29.vi.1998, Chandler leg. smithae] (HRI). Wytham Woods, Wytham, Oxford- (PPCM). Cors Erddreiniog, Isle of Anglesey, 1 male, shire, 3 males, 21.viii.2002, Menzel leg. [holotype and 27.vii.1988; 1 male, 6.x.1988, all Holmes, Boyce & paratypes of Bradysia smithae] (DEI). Reed leg. (PPWM). Epping Forest, Essex, 1 male, ix.- Ecological notes: found in broad-leaved woodland; x.1999, Dagley leg. (PPCM). Falls of Tarnash, Moray, 1 calcareous grassland; school grounds with old pop- male, 9.vii.1936, Coe leg. (BMNH). Gearagh, Cork, 1 lars; formal garden with wooded fringes and lake. v– male, 10.x.1990, de Courcy Williams leg. (DEI). Gors viii, x. Llwyn, Neath/Port Talbot, 1 male, 28.vi.1989, Holmes, Boyce & Reed leg. (PPWM). Greywell Fen, Hampshire, 1 male, 5.viii.2003, Chandler leg. (PPCM). Gwenfro, BRADYSIA SPLENDIDA MOHRIG & KRIVOSHEINA, 1989 Isle of Anglesey, 3 males, 27.vii.1988, Holmes, Boyce & New to British list. Reed leg. (PPWM). Hartslock, Oxfordshire, 1 male, 14.v.2000, Raper leg. (PPCM). Knebworth, Hertford- Material examined: The Loke, Norwich, Norfolk, 1 shire, 1 male, 18.iii.1928, Edwards leg. (BMNH). Llyn male, 7.ix.1987, Laurence leg. (PBLN). Ystumllyn, Gwynedd, 1 male, 26.vii.1988, Holmes, Ecological notes: found in garden. ix. Boyce & Reed leg. (PPWM). Reedham, Norfolk, 1 male, 12.viii.−26.viii.1988, Foster & Procter leg. (PBLN). Rhôs Llawr Cwrt, Ceredigion, 1 male, 29.vii.1987, BRADYSIA STRENUA (WINNERTZ, 1867) Holmes, Boyce & Reed leg. (PPWM). Symond’s Yat, Synonyms: annulata absoloni (Bezzi, 1911); ardua Herefordshire, 1 male, ix.1936, Edwards leg. (BMNH). (Grzegorzek, 1884). Tresillian, Cornwall, 1 male, 7.vii.1983, Chandler leg. (PPWM). Walberswick, Suffolk, 1 male, 1.vi.− Literature: Edwards (1925a): 540 [as Sciara (?) vari- 15.vi.1988, Foster & Procter leg. (PBLN). Windsor For- ans, misidentification]; ? Edwards (1938): 201 [as Sci- est, Berkshire, 1 male, 5.viii.1995, Brock leg. (PPCM). ara (Neosciara) varians, misidentification]; Freeman Woodwalton Fen, Cambridgeshire, 1 male, 24.v.1940, (1983b): 33 [as Bradysia (?) brunnipes, misidentifica- Edwards leg. (BMNH). tion]; Laurence (1994): 118; Laurence (1996): 87 [all as Bradysia brunnipes, misidentification]; Menzel Ecological notes: common in ancient oak and beech (1998): 20. forest, alder carr and other broad-leaved woodland; wetlands including wet meadow with peat, Juncus New to Irish list. and Glyceria, fens, ombrotrophic mire, reed and sedge Material examined: Annahala bog, Gearagh, Cork, 1 beds; grassland and scrub with stream; calcareous male, 13.x.1990, de Courcy Williams leg. (NMID). grassland. Reared from mole nest. iii, v–x. Alconbury, Cambridgeshire, 1 male, 3.v.1976, J.H. Cole leg. (PJCB). Biggleswade, Bedfordshire, 1 male, 6.vii.1927, Edwards leg. (BMNH). Bollin Valley, BRADYSIA SMITHAE MENZEL & HELLER, 2004 Cheshire, 2 males, 16.v.1928 (reared), Britten leg. Literature: Chandler (2001): 200 [as Bradysia norma- (BMNH). Buckingham Palace Garden, Greater Lon- lis, misidentification]; Menzel & Heller (2005): 339. don, 2 males, 20.xi.1995; 1 male, 18.vii.−3.viii.1997, all D.J. Carter & C.W. Plant leg. (PPCM). Coombe Abbey New to British list. Country Park, Coventry, Warwickshire, 1 male, Material examined: Buckingham Palace Garden, 28.viii.2003, Menzel leg. (DEI). Freshfield, Mersey- Greater London, 1 male, 11.vii.−1.viii.1996; 2 males, side, 1 male, 1.vii.1932 (reared), Britten leg. (BMNH). 26.v.−19.vi.1997 [paratypes of Bradysia smithae; Glasshouse Crops Research Station, Littlehampton, recorded as Bradysia normalis by Chandler (2001)]; 1 West Sussex, 1 male, 30.iv.1965; 1 male, 3.v.1965, all male, 18.vii.−13.viii.1997, all D.J. Carter & C.W. Plant Gurney leg. (HRI). Greywell Fen, Hampshire, 1 male, leg. (NMS, PPCM). Lackham Park, Wiltshire, 1 male, 20.vi.2002, Chandler leg. (PPCM). Hogley, Oxford- 24.x.2002, Chandler leg. [paratype of Bradysia shire, 1 male, 14.ii.1909; 1 male, 20.ii.1909 (all smithae] (PPCM). Milham Ford School, Oxford, reared), all Hamm leg. (BMNH). Kirton, Lincolnshire, Oxfordshire, 2 males, 3.v.−11.vi.2002, Webb leg. 4 males, xii.1924 (reared), Miles leg. (BMNH). Leeds, [paratypes of Bradysia smithae] (PPCM). Monkfield West Yorkshire, 2 males, xii.1928, Taylor leg. (BMNH). Wood, Cambridgeshire, 1 male, x.1998, C.W. Plant leg. Letchworth, Hertfordshire, 1 male, v.1917; 1 male, [paratype of Bradysia smithae] (HRI). Sheep Leas, vi.1917, all Edwards leg. (BMNH, ZFMK). Lifton West Horsley, Surrey, 1 male, 16.vi.1991, Chandler Park, Devon, 4 males, 29.x.1935 (reared), Bradshaw leg. [paratype of Bradysia smithae] (PPCM). Which- leg. (BMNH). Marston, Oxfordshire, 1 male, 3.iv.1933

(reared), Basden leg. (NMS). Oxwich, Swansea/ BRADYSIA SUBRUFESCENS MOHRIG & Abertawe, 1 male, 12.vii.1989, Holmes, Boyce & Reed KRIVOSHEINA, 1989 leg. (PPWM). Sale, Greater Manchester, 3 males, New to British list. 23.ii.1928, Buller leg. (BMNH). Taplow, Buckingham- shire, 3 males, xi.1913 (reared), no collector detail Material examined: Salisbury Plain Training Area, (BMNH). Theale Lake, Berkshire, 1 male, 9.vii.2003, Warminster, Wiltshire, 1 male, 30.v.2003, Ismay & Chandler leg. (PPCM). The Loke, Norwich, Norfolk, 1 Schulten leg. (DEI). The Loke, Norwich, Norfolk, 2 male, 10.vi.1987; 1 male, 15.ix.1987; 1 male, v.1992; 1 males, 23.iv.1987, Laurence leg. (PBLN). − male, 23.v. 25.v.1993, all Laurence leg. (PBLN). Ecological notes: found on unimproved calcareous Ecological notes: found in broad-leaved woodland; grassland; garden. iv–v. alder carr woodland and sedge beds by river; partly wooded lake margin; ditch along meadow hedge; cal- BRADYSIA SUBSCABRICORNIS MOHRIG & careous valley; Phragmites; bog; grounds of horticul- MENZEL, 1990 tural research station; gardens. Reared from decaying narcissus bulbs, potatoes, mole nest, angelica root, old Literature: Menzel (1998): 20. ragwort stems. ii, iv–xii. Material examined: Cors Bodeilio, Isle of Anglesey, 1 male, 28.vi.1988, Holmes, Boyce & Reed leg. (PPWM). Salisbury Plain Training Area, Warminster, Wiltshire, BRADYSIA STRIGATA (STAEGER, 1840) 1 male, 30.v.2003, Ismay & Schulten leg. (DEI). Tre Synonym: robusta (Lengersdorf, 1926) [preocc.]. Wilmot, Isle of Anglesey, 1 male, 5.x.1988, Holmes, Boyce & Reed leg. (PPWM). Literature: Freeman (1983b): 35; Menzel (1998): 20. Ecological notes: found on unimproved calcareous Material examined: Cors Llyn Farch a Llyn Fanod, grassland, basin mire; Cladium bed. v–vi, x. Ceredigion, 2 males, 17.ix.1986; 1 male, 24.ix.1986, all Holmes, Boyce & Reed leg. (PPWM). Cothill, Oxford- shire, 1 male, 5.v.1934, Edwards leg. (BMNH). Grey- BRADYSIA TENUICAUDA MOHRIG & MENZEL, 1990 well Fen, Hampshire, 1 male, 2.viii.1998, Chandler New to British list. leg. (PPCM). Hogley, Oxfordshire, 1 male, 10.v.1919, Hamm leg. (BMNH). Leckford Estate, Hampshire, 1 Material examined: Castlemartin Corse, Pem- male, 30.iv.1977; 1 male, 12.iv.1989, all Chandler leg. brokeshire, 1 male, 22.vii.1987, Holmes, Boyce & (PPCM). Rhôs Llawr Cwrt, Ceredigion, 1 male, Reed leg. (PPWM). 29.ix.1986, Holmes, Boyce & Reed leg. (PPWM). Sut- Ecological notes: found in meadow with Juncus and ton Park, Birmingham, West Midlands, 1 male, Iris. vii. 6.viii.1989, J.H. Cole leg. (PJCB). Thompson Common, Norfolk, 1 male, 16.viii.1989, Chandler leg. (NMS). Woodbastwick, Suffolk, 1 male, viii.1932, Edwards leg. BRADYSIA TILICOLA (LOEW, 1850) (BMNH). No location details, 1 male, 1989, no. 89/021, Synonyms: alma (Winnertz, 1871); amoena (Winnertz, ? Disney leg. (PPWM). 1867); coprophila (Lintner, 1895); domestica Frey Ecological notes: found in broad-leaved woodland (1948); incomta (Winnertz, 1867); nanella (Frey, especially carr; heathland; wetlands including water 1936); selecta (Winnertz, 1871); setigera (Winnertz, meadows, wet meadows with Juncus, sedge beds, 1867); signata (Winnertz, 1867); triseriata (Winnertz, basin mire and grassland with pingo pools; parkland. 1867); turbida (Winnertz, 1867); vana (Winnertz, iv–v, viii–ix. 1871); vividula (Winnertz, 1867); volucris (Winnertz, 1867); wendalinae (van Bruggen, 1954). Literature: ? Walker (1848): 108 [as Sciara sylvatica BRADYSIA SUBALPINA FREY, 1948 (recte silvatica)]; Basden (1933): 44 [as Sciara mod- Literature: Freeman (1983b): 38 and 64, fig. 148 [as esta, misidentification]; Freeman (1983b): 36; Lau- Bradysia aprica, misidentification]. rence (1994): 116 [in part as Bradysia triseriata, misidentification]; Menzel (1998): 20; Chandler New to British list. (2001): 200 [all as Bradysia amoena]; Menzel & Heller Material examined: Meall Ghaordie, Stirling, 1 male, (2005): 351 [amoena (Winnertz) as synonym to tilicola 17.vi.1932, Edwards leg. (BMNH). (Loew)]. Ecological notes: found on open montane habitats and Material examined: Belfairs, Essex, 4 males, 23.x.− at 750–1000 m a.s.l. vi. 12.xii.1989 (reared), Withers leg. (PPWM). Berwick-

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 64 F. MENZEL ET AL. upon-Tweed, Northumberland, 1 male, 30.ix.1998; 1 6.v.1992; 1 male, v.1992; 1 male, 29.ix.−3.x.1992; 6 male, vi.1999; 1 male, viii.1999; 2 males, ix.1999; 2 males, 2.iii.1993; 1 male, 14.iii.1993, all Laurence males, x.1999; 1 male, 27.xii.1999, all Laurence leg. leg. (PBLN). Thornham Magna, Suffolk, 2 males, (PBLN). Bishop’s Stortford, Hertfordshire, 4 males, 4 30.x.1988 (reared), Evans leg. (PPWM). Thornton females, 1927 (reared), no collector detail (BMNH). Heath, Greater London, 1 male, 26.ii.1927 (reared), Brampton, Cambridgeshire, 1 male, 13.v.1985; 1 male, Stokes leg. (BMNH). Trawsfynydd, Gwynedd, 1 male, 22.v.1985; 3 males, 10.iii.1994; 2 males, 3.v.−4.v.1997 5.viii.1988, Holmes, Boyce & Reed leg. (PPWM). Tulse (reared), all J.H. Cole leg. (PJCB). Burnham, Bucking- Hill, Greater London, 1 male, 3 females, vii.1934 hamshire, 8 males, 4 females, 11.vi.−12.ix.1934 (reared), Nixon leg. (BMNH). Wellesbourne, Warwick- (reared), Basden leg. (NMS). Byﬂeet, Surrey, 1 male, shire, 1 male, 24.ii.−2.iii.2000; 1 male, 24.v.− iv.1934, no collector detail (BMNH). Cadair Idris, 27.vi.2001; 1 male, 27.vi.−5.vii.2001; 1 male, 13.ix.− Gwynedd, 1 male, 19.vi.1993, Laurence leg. (PBLN). 4.x.2001, all J.E. Smith leg. (HRI). West Harling Com- Cambridge, Cambridgeshire, 1 male, x.1931, Thomas mon, Norfolk, 1 male, 1.iv.1989, Withers leg. (PPWM). leg. (BMNH); 1 male, 21.ix.1940, Edwards leg. Wheatfen Broad, Norfolk, 1 male, 8.vii.−15.vii.1986, (BMNH). Coppets Wood, Greater London, 1 male, Withers leg. (PPWM). No locality details, Berkshire, 1 22.xii.1990; 2 males, 17.vii.1991; 6 males, 25.vii.1991, male, 3 females, 1.v.1925, Fox-Wilson leg. (BMNH). all K.G.V. Smith leg. (PBLN). Cors Lawnt, Powys, 1 Ecological notes: found in damp broad-leaved and male, 11.x.1988, Holmes, Boyce & Reed leg. (PPWM). alder carr woodland; pine plantations; moorland; wet- Dickleburgh, Norfolk, 1 male, 22.i.−29.i.1989; 1 male, lands including fen, bog, sedge bed and valley mire; 2.iv.1991, all Withers leg. (PPWM). Hickling Broad, gardens; grounds of horticultural research station; Norfolk, 1 male, 16.iv.1989 (reared), Evans leg. school grounds with old poplars; open montane habi- (PPWM). How Hill, Norfolk, 1 male, 13.viii.1989, tats; in house; in greenhouse; house-plant compost, in Withers leg. (PPWM). Letchworth, Hertfordshire, 1 old bird nest. Reared from fungi Amanita muscaria, male, x.1911; 3 males, v.1918, Edwards leg. (BMNH). Gymnopilus junonius, Inonotus cuticularis, Lepiota London, Greater London, 2 males, 1 female, x.1982; 1 cristata, under dead poplar bark, larva under pine male, 1 female, ii.1983, all no collector detail (BMNH). bark, tulip bulbs, decaying onions, formicarium, and Market Weston Fen, Suffolk, 1 male, 6.ix.1988 soil from rabbit burrow. Larvae found destroying (reared), Evans leg. (PPWM). Matley Bog, New Forest, young seedlings of ling (Calluna vulgaris) and bulbs of Hampshire, 1 male, 15.xi.1953, Colyer leg. (BMNH). lily (Lilium aurantiacum); in gerbil cage. i–xii. Melksham, Wiltshire, 5 males, 1 female, 21.iii.− 25.vii.2002, Chandler leg. (PPCM). Merton, Greater Taxonomic note: Loew found Sciara tilicola in large London, 1 male, x.1911 (reared), C.B. Williams leg. numbers. From the apparently large type series, only (BMNH). Milham Ford School, Oxford, Oxfordshire, 1 2 males with the original labelling ‘tilicola Lw.’ could male, 18.v.−22.v.2001, Webb leg. (PPCM). Nailsea, still be found in the Loew collection. The lectotype des- Avon, 1 male, v.1944, no collector detail (BMNH). ignation in Menzel & Heller (2005) conforms with the North Wood, Meikleour, Perth & Kinross, 1 male, interpretation of Lengersdorf (1928–30: 52), who con- ii.1996 (reared), MacGowan leg. (NMS). Oxford, sidered that tilicola Loew is identical with amoena Oxfordshire, 1 male, 10.iv.1926 (reared), Hamm leg. Winnertz but listed the species under the incorrect (BMNH). Oxford University Parks, Oxford, Oxford- name ‘modesta Staeger, 1840’. Further information shire, 1 male, 3.iv.1930, Hamm leg. (BMNH). Pett’s about the type material of tilicola (Loew) and about Wood, Greater London, 1 male, 24.vi.1983, Freeman the synonymy of amoena (Winnertz) can be found in leg. (BMNH). Putney, Greater London, 31 males, Menzel & Heller (2005). ix.1917, Cant leg. (BMNH). Rothamsted, Harpenden, Hertfordshire, 3 males, 2 females, i.1933, M.A. Watson leg. (BMNH). Scilly Isles, Cornwall, 1 male, ii.1902, BRADYSIA TRISPINIFERA MOHRIG & Jenkinson leg. (BMNH). Shefﬁeld, South Yorkshire, 4 KRIVOSHEINA, 1979 males, 2 females, ix.1922, L.W. Cole leg. (BMNH). New to British list. Stanford, Norfolk, 1 male, 10.iv.−24.iv.1988, Collier leg. (PPWM). Temple, Berkshire, 1 male, 23.iv.1932; 7 Material examined: Bucklebury Common, Berkshire, males, 3 females, 1934 (all reared), all Basden leg. 1 male, 20.viii.1993, Chandler leg. (PPCM). Coed (BMNH, NMS). The Loke, Norwich, Norfolk, 1 male, Tycanol, Pembrokeshire, 2 males, 10.vi.−29.vi.1999, 20.iv.1987; 1 male, 23.iv.1987; 5 males, 26.iv.1987; Levey & Pavett leg. (PPCM). Collinpark Wood, Glouc- 3 males, 27.iv.1987; 2 males, 8.v.1987; 2 males, estershire, 1 male, 3.ix.2002, Gibbs leg. (PDGB). 5.vii.1987; 2 males, 15.ix.1987; 1 male, 24.x.− Lough Slevin, Westmeath, 1 male, 16.vi.1985, Chan- 25.x.1987; 1 male, 20.iv.1988; 1 male, 23.vi.1988; 1 dler leg. (PPWM). Nutcombe Down, Surrey, 1 male, male, i.-iii.1990; 1 male, 28.iv.−4.v.1990; 1 male, 2.v.− 26.vii.2002, Chandler leg. (PPCM). Redlake, Cornwall,

2 males, 12.vii.2001, Chandler leg. (PPCM). Ryton Fen, Cambridgeshire, 1 male, 26.viii.1980, J.H. Cole Wood, Bubbenhall, Coventry, West Midlands, 2 males, leg. (PJCB). Kelham Bridge, Ravenstone, Leicester- 25.viii.2002, Menzel & J.E. Smith leg. (DEI, HRI). shire, 1 male, 24.viii.2003, J.E. Smith leg. (HRI). Lack- Wheatfen Broad, Norfolk, 1 male, 25.vii.1989, Withers ham Park, Wiltshire, 1 male, 20.ix.2002, Chandler leg. leg. (PPWM). Wychwood, Oxfordshire, 1 male, v.1937, (PPCM). Letchworth, Hertfordshire, 2 males, v.1917; 1 Edwards leg. (BMNH). male, vii.1917; 1 male, x.1917; 3 males, xi.1917, all Edwards leg. (BMNH, ZFMK). Luton Airport, Bed- Ecological notes: widespread in broad-leaved wood- fordshire, 1 male, 20.iv.−6.v.1999, C.W. Plant leg. land (some acid); also found in mixed woodland. vi–ix. (PPCM). Mills Marsh, Norfolk, 1 male, 11.viii.1988, Foster & Procter leg. (PBLN). Moccas Park, Hereford- BRADYSIA TRIVITTATA (STAEGER, 1840) shire, 1 male, 11.vi.2002, Godfrey leg. (PPCM). Monkfield Wood, Cambridgeshire, 1 male, x.1998, Synonyms: basalis (Winnertz, 1867); decipiens (Win- C.W. Plant leg. (PPCM). North Ronaldsay, Orkney, 1 nertz, 1867); devittata Tuomikoski, 1959; lignorum male, 8.ix.1989; 1 male, 10.ix.1989, all Laurence leg. (Kieffer, 1919); spectrum (Winnertz, 1867); versicol- (PBLN). Orford, Suffolk, 1 male, 19.viii.1908, King leg. orea (Lengersdorf, 1940). (BMNH). Oxford University Museum, Oxford, Oxford- Literature: Freeman (1983b): 38 [as Bradysia triseri- shire, 1 male, 10.ix.1918, Hamm leg. (BMNH). Rad- ata, misidentification]; Laurence (1994): 116 [in part well, Hertfordshire, 2 males, 21.x.1917, Edwards as Bradysia triseriata, misidentification]; Laurence leg. (BMNH). Rhôsgoch, Powys, 1 male, 10.x.1989, (1997b): 228 [as Bradysia triseriata (group)]; Menzel Holmes, Boyce & Reed leg. (PPWM). Salisbury Plain (1998): 20. Training Area, Warminster, Wiltshire, 1 male, 29.vi.2003; 5 males, 3.vii.2003; 3 males, 4.vii.2003, all New to Irish list. Ismay & Schulten leg. (DEI, UMO); 1 male, 1.vii.− Material examined: Alconbury, Cambridgeshire, 1 5.vii.2003, Woodcock leg. (UMO). Snailbeach, Shrop- male, 3.v.1976, J.H. Cole leg. (PJCB). Badger Wood, shire, 1 male, 27.vii.−28.vii.1920, Edwards leg. Cambridgeshire, 1 male, x.1998, C.W. Plant leg. (BMNH). Sned Wood, Lower Lye, Herefordshire, 1 (PPCM). Beesands, Devon, 1 male, 8.vi.1920, Edwards male, 1 female, 14.vii.2003, J.E. Smith leg. (HRI). Stal- leg. (BMNH). Bradfield Wood, Suffolk, 1 male, 11.vii.− lode Wash, Suffolk, 1 male, 24.vi.−8.vii.1988, Foster & 16.vii.1990, Laurence leg. (PBLN). Buckingham Pal- Procter leg. (PBLN). Temple, Berkshire, 16 males, 4 ace Garden, Greater London, 1 male, 1996–97, D.J. females 1.iv.−1.v.1934; 13 males, 1 female, 8.iv.− Carter & C.W. Plant leg. (PPCM). Burnham Beeches, 15.iv.1933 (reared), all Basden leg. (BMNH, NMS). Buckinghamshire, 1 male, 10.v.−23.v.199; 1 male, The Loke, Norwich, Norfolk, 2 males, 26.iv.1987; 11 20.iv.1998, all Ismay leg. (PPCM). Catfield, Norfolk, 1 males, 8.v.1987; 3 males, 9.v.1987; 2 males, 8.vi.1987; male, no date, Foster & Procter leg. (PBLN). Chigwell 1 male, 18.vi.1987; 5 males, 5.vii.1987; 1 male, Row Wood, Chigwell Row, Essex, 1 female, 18.vi.2002; 10.viii.1987; 1 male, 7.ix.1987; 7 males, 15.ix.1987; 2 2 males, 28.vi.2002; 1 male, 1 female, 15.ix.2002; 2 males, 10.vi.−24.vi.1988; 1 male, 13.vi.1988; 5 males, males, 8.x.2002; 2 males, 5 females, 16.x.2002; 1 male, 15.viii.−16.viii.1988; 1 male, 16.viii.1988; 1 male, 1 female, 22.x.2002, all Ismay & Schulten leg. (DEI, 18.ix.−20.ix.1988; 3 males, 15.ix.−28.ix.1889; 3 males, UMO). Clovelly, Devon, 1 male, 8.v.1936, Edwards leg. 2.v.−6.v.1992; 1 male, 27.vii.1992; 2 males, 29.ix.− (BMNH). Cock Marsh, Berkshire, 1 male, 25.viii.1999, 3.x.1992; 1 male, 4.xi.−11.xi.1992; 1 male, 30.vii.− Chandler leg. (PPCM). Colwyn Brook, Powys, 1 male, 2.viii.1993; 3 males, 27.viii.−1.ix.1993, all Laurence 18.viii.1989; 2 males, 9.x.1989, all Holmes, Boyce & leg. (PBLN). The Oaks, Cambridgeshire, 2 males, Reed leg. (PPWM). Dinton Pastures Country Park, ix.1998, C.W. Plant leg. (PPCM). Thompson Common, Berkshire, 1 male, 21.vi.1993; 1 male, 3.xi.2001, all Norfolk, 2 males, 13.ix.−27.ix.1988, Foster & Procter Chandler leg. (PPCM). Epping Forest, Essex, 14 leg. (PBLN). Trefeiddan Moor, Pembrokeshire, 1 male, males, 1998–2000, Dagley leg. (PPCM). Farnborough 21.vii.1987, Holmes, Boyce & Reed leg. (PPWM). Park, Warwickshire, 1 female, 10.x.2001, Chandler Tresta, Shetland, 1 male, 5.vi.−6.vi.1998, Laurence leg. (HRI). Ffrwd Fen, Carmarthen, 2 males, 3.x.1989, leg. (PBLN). Unst, Shetland, 1 male, 19.viii.− Holmes, Boyce & Reed leg. (PPWM). Hartslock, 21.viii.1996, Laurence leg. (PBLN). Vicarage Bog, Oxfordshire, 1 male, 6.v.2000, Raper leg. (PDGB). Wrexham, 1 male, 12.x.1988, Holmes, Boyce & Reed Hemingford, Cambridgeshire, 11 males, 9 females, leg. (PPWM). Vidlin, Shetland, 1 male, 18.vi.− 1.x.1938, Edwards leg. (BMNH). Hickling, Norfolk, 1 21.vi.1993, Laurence leg. (PBLN). Wellesbourne, male, v.1988, 1 male, 1988, all Foster & Procter leg. Warwickshire, 5 males, 1999; 3 males, 6.iv.−13.iv.2000; (PBLN). Highgate Wood, Greater London, 1 male, 7 males, 5 females, 17.viii.−31.viii.2000; 2 males, 1998–99, Hackett leg. (PPCM). Hitchin, Hertford- 28.iii.−26.iv.2001; 1 male, 24.v.−27.vi.2001; 1 male, shire, 1 male, ix.1917, Edwards leg. (BMNH). Holme 26.vi.2001; 2 males, 27.vi.−5.vii.2001; 2 males, 13.ix.−

4.x.2001, all J.E. Smith leg. (HRI). Willes Meadow, 1 male, 25.iv.−9.v.1996, D.J. Carter & C.W. Plant leg. Leamington Spa, Warwickshire, 4 males, 1 female, (PPCM). Burnham Beeches, Buckinghamshire, 1 27.viii.2002, Menzel & J.E. Smith leg. (DEI, HRI). male, 26.ix.−9.x.1996, Ismay leg. (PPCM). Cock Wolford Wood, Moreton-in-Marsh, Warwickshire, 1 Marsh, Berkshire, 2 males, 25.viii.1999, Chandler leg. male, 11.x.2001, J.E. Smith leg. (HRI). Woodwalton (PPCM). Den of Alyth, Angus, 1 male, 30.v.1976, Fen, Cambridgeshire, 5 males, 24.v.1940, Edwards leg. Chandler leg. (PPCM). Dinton Pastures Country Park, (BMNH). Wytham Woods, Wytham, Oxfordshire, 4 Berkshire, 1 male, 20.ix.1992; 1 male, 1.v.1993; 1 males, 21.viii.2002, J.E. Smith leg. (HRI). male, 15.v.1993; 1 male, 9.vii.1996; 1 male, 29.x.1996, all Chandler leg. (PPCM). Eday, Orkney, 1 male, 1.vi.− Ecological notes: very common in all types of wood- 6.vi.1988; 3 males, 4.vi.1988, all Laurence leg. land; ancient woodland with oak, hazel and bramble; (PBLN). Epping Forest, Essex, 2 males, 18.ix.1998; 2 heathland; wetlands including fens, bogs and water males, iv.1999, all Dagley leg. (PPCM). Fanore, Clare, meadows; parkland; old hedges; unimproved grass- 1 male, 24.v.1985, J.P. & M.A. O’Connor leg. (NMID). land; near shore; gardens. Reared from nests of song Glen of Drumloch, Moray, 1 male, 16.viii.1937, Coe leg. thrush and moorhen. iv–xi. (BMNH). Hartslock, Oxfordshire, 1 male, 6.v.2000, Raper leg. (PDGB). Hickling Broad, Norfolk, 1 male, viii.1932, Edwards leg. (BMNH). Holme Fen, Cam- BRADYSIA URTICAE MOHRIG & MENZEL, 1992 bridgeshire, 1 male, 27.iv.1987, J.H. Cole leg. (PJCB). Literature: Menzel (1998): 20. Holywood, Down, 1 male, 18.vi.1993, Jordan leg. Material examined: Pumpkin Hill, Burnham Beeches, (UMB). Kelham Bridge, Ravenstone, Leicestershire, Buckinghamshire, 1 male, 22.vii.−5.viii.1996, Ismay 2 females, 24.viii.2003, Menzel leg. (DEI). Kennack leg. (PPCM). Snettisham Beach, Norfolk, 2 males, Cove, Cornwall, 1 male, 4.ix.1983, Chandler leg. 8.v.−13.v.1983, Evans leg. (PPWM). Willes Meadow, (PPCM). Knocksink Wood, Wicklow, 1 male, Leamington Spa, Warwickshire, 2 males, 27.viii.2002, 28.ix.1999, Chandler leg. (PPCM). Lea Meadows, Menzel leg. (DEI). Newtown Linford, Leicestershire, 1 male, 24.viii.2003, Menzel leg. (DEI). Letchworth, Hertfordshire, 1 male, Ecological notes: found in ancient beech woodland; v.1917, Edwards leg. (ZFMK). Loch Fleet, Highland meadow flooded in winter; underground pitfall trap on Region, 1 male, 26.v.2002, Chandler leg. (PPCM). sea shore. v, vii–viii. Lockhaugh Bank Wood, Derwent Valley, Tyne & Wear, 1 male, 24.vii.−13.viii.1990, Jessop leg. (PBLN). Lolly Moor, Norfolk, 2 males, 23.v.1979, Irwin leg. (PPWM). BRADYSIA VAGANS (WINNERTZ, 1868) Luton Airport, Bedfordshire, 1 male, 20.iv.−6.v.1999, Synonyms: angustipennis Frey, 1948 [preocc.]; cal- C.W. Plant leg. (PPCM). Newbridge Fen, Kildare, 1 licera Frey (1948); richardi Gerbachevskaja, 1986. male, 8.ix.−19.ix.2000, Withers leg. (PPWM). North Ronaldsay, Orkney, 1 male, 9.ix.−10.ix.1989; 2 males, Literature: Laurence (1994): 118; Laurence (1997a): 85 ix.1989, all Laurence leg. (PBLN). Occold, Suffolk, 1 [all as Bradysia confinis, misidentification]; Laurence male, 10.v.1990, Withers leg. (PPWM). Oxford Univer- (1997b): 228 [as Bradysia confinis (group) and in part sity Museum, Oxford, Oxfordshire, 1 male, 6.v.1927, as Bradysia pullula, misidentification]; Menzel Hamm leg. (BMNH). Pearman’s Copse, Berkshire, 1 (1998): 20; Chandler et al. (2002): 122. male, 28.viii.1995, Chandler leg. (PPCM). Pett’s Material examined: Alconbury, Cambridgeshire, 1 Wood, Greater London, 1 male, 6.x.1981, Freeman leg. male, 3.v.1976; 1 male, 4.v.1989, all J.H. Cole leg. (BMNH). Raveley Wood, Cambridgeshire, 1 male, (PJCB). Badger Wood, Cambridgeshire, 2 males, 28.ix.1975, J.H. Cole leg. (PJCB). Reedham, Norfolk, 1 1.v.−31.v.1998, C.W. Plant leg. (PPCM). Ban-y-Gor male, 26.viii.−27.viii.1988, Foster & Procter leg. Wood, Gloucestershire, 1 male, 12.ix.2002, Gibbs leg. (PBLN). Salisbury Plain Training Area, Warminster, (PDGB). Bradfield Wood, Suffolk, 8 males, 9.x.− Wiltshire, 1 male, v.2003; 1 male, 27.v.2003; 1 male, 16.x.1990; 2 males, 23.v.−30.v.1991; 1 male, v.1991; 3 31.v.2003, all Ismay & Schulten leg. (DEI, UMO). males, 16.viii.−22.viii.1991, all Laurence leg. (PBLN). Sandy, Bedfordshire, 1 male, 15.v.1919, Edwards leg. Brancaster, Norfolk, 1 male, 28.x.1988, Withers leg. (BMNH). Stronsay, Orkney, 1 male, 25.v.1988, Lau- (PPWM). Brandyquoi, Kirkwall, Orkney, 10 males, v.– rence leg. (PBLN). Sutton Wash, Cambridgeshire, 1 vi.2001, Laurence leg. (PBLN). Blackgang, Isle of male, 18.v.1989, J.H. Cole leg. (PJCB). The Loke, Nor- Wight, 1 male, 25.iv.1975, Jarzembowski leg. (BMNH). wich, Norfolk, 3 males, 23.iv.1987; 2 males, 26.iv.1987; Blair Atholl, Perth & Kinross, 1 male, 28.v.1973, 1 male, 27.iv.1987; 11 males, 8.v.1987; 26 males, Chandler leg. (PPCM). Brampton, Ouse flood meadow, 9.v.1987; 2 males, 18.vi.1987; 1 male, 10.viii.1987; 1 Cambridgeshire, 1 male, 8.v.1977, J.H. Cole leg. male, 15.viii.−16.viii.1988; 3 males, 7.ix.1987; 1 male, (PJCB). Buckingham Palace Garden, Greater London, 8.ix.1987; 6 males, 15.ix.1987; 1 male, 24.x.−25.x.1987;

5 males, 5.v.1988; 7 males, 13.v.1988; 1 male, BRADYSIOPSIS TUOMIKOSKI, 1960 − − 13.vi.1988; 1 male, 18.ix. 20.ix.1988; 9 males, 5.vi. BRADYSIOPSIS VITTATA (MEIGEN, 1830) 13.vi.1989; 3 males, 21.viii.−24.viii.1989; 4 males, Synonyms: leucotricha (Tuomikoski, 1960); quinque- 15.ix.−28.ix.1989; 1 male, 29.ix.−3.x.1989, all Lau- lineata var. alpujarrensis (Strobl, 1900); quinquelin- rence leg. (PBLN). The Oaks, Cambridgeshire, 1 male, eata var. nigripes (Strobl, 1898). ix.1998, C.W. Plant leg. (PPCM). Thundry Meadow, Surrey, 1 male, 15.ix.2001, Chandler leg. (PPCM). Literature: Freeman (1983a): 168; Freeman (1983b): Tomintoul, Moray, 1 male, 8.vii.−11.vii.1937, Coe leg. 30 [all as Lycoriella (Bradysiopsis) leucotricha]; Free- (BMNH). Trefeiddan Moor, Pembrokeshire, 1 male, man (1987): 197; Skidmore (1991): 149; Laurence 21.vii.1997, Holmes, Boyce & Reed leg. (PPWM). (1997a): 85 [all as Lycoriella leucotricha]; Menzel Tresta, Shetland, 1 male, 5.vi.−6.vi.1998, Laurence (1998): 20. leg. (PBLN). Udale Bay, Highland Region, 2 males, Material examined: Alconbury, Cambridgeshire, 1 29.v.2002, Chandler leg. (PPCM). Weem Castle Woods, male, 9.vii.1978, J.H. Cole leg. (PJCB). Aviemore, Perth & Kinross, 1 male, 28.viii.1995, Chandler leg. Highland Region, 1 male, v.1934, Edwards leg. (PPCM). Wellesbourne, Warwickshire, 1 male, 1999; (BMNH). Bradﬁeld Wood, Suffolk, 1 male, 11.vii.− 1 male, 24.ii.−2.iii.2000; 8 males, 6.iv.−13.iv.2000; 16.vii.1990; 1 male, 9.x.−16.x.1990, all Laurence 4 males, 17.viii.−31.viii.2000; 1 female, 28.iii.− leg. (PBLN). King’s Walden, Hertfordshire, 1 male, 26.iv.2001, all J.E. Smith leg. (HRI). Whitacre Heath, 24.iv.1964, Colyer leg. (BMNH). Knott Wood, Rotham- Warwickshire, 1 male, 1 female, 13.x.2001, J.E. Smith sted, Harpenden, Hertfordshire, 3 females, 20.iv.1950; leg. (HRI). Willes Meadow, Leamington Spa, Warwick- 2 males, 29.v.1950 (reared), all Laurence leg. (PBLN). shire, 1 male, 1 female, 27.viii.2002, Menzel leg. (DEI). Nethy Bridge, Highland Region, 1 male, 16.vii.1936, Wistow Wood, Warboys, Cambridgeshire, 1 male, Coe leg. (BMNH). Salisbury Plain Training Area, 4.v.1975, J.H. Cole leg. (PJCB). Yell, Shetland, 1 male, Warminster, Wiltshire, 1 male, 27.v.2003; 3 males, 3.ix.−19.ix.1990, Laurence leg. (PBLN). No location 4.vii.2003, all Ismay & Schulten leg. (DEI, UMO). details, 1 male, no date, Withers leg. (PPWM). Spinningdale Bog, Highland Region, 1 male, Ecological notes: very common in broad-leaved, mixed 17.vi.1976, Chandler leg. (PPCM). Stanford, Norfolk, 1 and carr woodland; heathland and scrub; wetlands male, 27.iii.−10.iv.1988, Collier leg. (PPWM). Upwood including water meadows, Sphagnum bog and sedge Meadow, Cambridgeshire, 1 male, 29.v.1977, J.H. Cole beds, saltmarsh; unimproved calcareous grassland; leg. (PJCB). No locality details, Wales, ‘Clwyd’, 1 male, old hedges; old sand and gravel workings; gardens; 1987, M. Jones leg. (BMNH). near shore; coastal landslip. iii–x. Ecological notes: found in broad-leaved woodland; unimproved calcareous grassland; on ive broomrape; BRADYSIA VERNALIS (ZETTERSTEDT, 1851) hedges; river bank; ditch along meadow hedge; bog New to British and Irish list. with stream. Reared from old cow dung. iii–vii, x. Material examined: Lough Slevin, Westmeath, 1 male, 16.vi.1985, Chandler leg. (PPWM). Reedham, Norfolk, CAMPTOCHAETA HIPPA & VILKAMAA, 1994 2 males, 12.viii.−26.viii.1988, Foster & Procter CAMPTOCHAETA CAMPTOCHAETA (TUOMIKOSKI, 1960) leg. (PBLN). Strumpshaw, Norfolk, 1 male, 13.vi.− 11.vii.1988, Foster & Procter leg. (PBLN). New to British list. Ecological notes: found in mixed woodland; damp Material examined: Dubh Ghleann, Glen Quoich, meadow; sedge bed. vii–viii. Aberdeenshire, 1 male, 21.iv.2000; 4 males, 19.vi.2000, all Godfrey leg. (HRI, PPCM). Upper Quoich, Aber- deenshire, 1 male, 21.vi.2000, Godfrey leg. (PPCM). BRADYSIA ZONATA RUDZINSKI, 1993 Ecological notes: found in ancient pine woodland. iv, New to British list. vi. Material examined: Brynberian Moor, Pembrokeshire, 1 male, 22.vi.1987, Holmes, Boyce & Reed leg. (DEI). Rhôsili Down, Swansea/Abertawe, 1 male, 3.x.1989, CAMPTOCHAETA COEI (FREEMAN, 1983) Holmes, Boyce & Reed leg. (PPWM). Wheatfen Broad, Literature: Freeman (1983a): 164 [in part]; Freeman Norfolk, 1 male, 8.vi.−15.vi.1986; 1 male, 29.v.1990, all (1983b): 29 [all as Corynoptera]; Menzel (1998): 20; Withers leg. (PPWM). Menzel & Mohrig (2000): 196. Ecological notes: found on damp heath; Sphagnum Material examined: California Country Park, Berk- and Myrica ﬂush. v–vi, x. shire, 1 male, 2.vi.1996, Chandler leg. (PPCM). Culbin

Sands, Highland Region, 3 males, 1 female, 5.vii.− Material examined: Burnham Beeches, Buckingham- 7.vii.1936, Coe leg. [including holotype and paratype shire, 1 male, 23.v.−6.vi.1995; 7 males, 8.vii.− of Corynoptera coei] (BMNH). Harpenden, Hertford- 26.xi.1996, all Ismay leg. (HRI, PPCM). Clovelly, shire, 1 male, 1.viii.1950, Laurence leg. (BMNH). Loch Devon, 2 males, 8.v.1936, Edwards leg. [paratypes Park, Moray, 1 male, 3.vii.1936, Coe leg. (BMNH). Tor- of Corynoptera abblanda] (BMNH). Coed y Rhygen, boll Wood, Highland Region, 1 male, 24.x.2000, Chan- Gwynedd, 1 male, 14.vii.−8.ix.1999, Levey & Pavett dler leg. (PPCM). leg. (PPCM). Cotterdale, North Yorkshire, 1 male, 4.v.1976, J.H. Cole leg. (PJCB). Crowborough, East Ecological notes: found in broad-leaved and alder carr Sussex, 1 male, 12.v.1904, Jenkinson leg. [paratype woodland; dunes and conifer plantations. vi–viii, x. of Corynoptera abblanda] (BMNH). Dendles Wood, Devon, 1 male, 4.v.1976, J.H. Cole leg. (PJCB). Epping CAMPTOCHAETA CONSIMILIS (HOLMGREN, 1869) Forest, Essex, 7 males, 12.iv.-xi.1999; 1 male, New to British list. 19.ix.1999; 1 male, ix.1999, all Dagley leg. (NMS, PPCM). Fishpool Valley, Herefordshire, 1 male, Material examined: Cwm Dewi, Pembrokeshire, 1 12.x.1984, Chandler leg. (PPWM). Gearagh, Cork, 1 male, 12.x.1987, Holmes, Boyce & Reed leg. (PPWM). male, 7.iv.1990; 1 male, 10.x.1990, all de Courcy Wil- Dowrog Common, Pembrokeshire, 1 male, 29.ix.1987, liams leg. (DEI, NMID). Hartslock, Oxfordshire, 1 Holmes, Boyce & Reed leg. (DEI). Llyn Fawnog, Den- male, 6.v.2000, Raper leg. (PDGB). J.F. Kennedy Park, bighshire, 1 male, 27.x.1988, Holmes, Boyce & Reed Wexford, 1 male, 29.iii.1989, J.P. & M.A. O’Connor leg. leg. (PPWM). The Ritec, Pembrokeshire, 1 male, (NMID). Kitely Wood, Malham, North Yorkshire, 24.vi.1987; 1 male, 8.vii.1987; 2 males, 22.vii.1987; 2 1 male, 11.ix.−18.ix.1981, Laurence leg. (PBLN). males, 30.ix.1987, all Holmes, Boyce & Reed leg. (DEI, Lancaut Nature Reserve, Gloucestershire, 1 male, PPWM). Trefeiddan Moor, Pembrokeshire, 1 male, 5.ix.2002, Gibbs leg. (PDGB). Lion Wood, Redditch, 21.vii.1987, Holmes, Boyce & Reed leg. (PPWM). Worcestershire, 2 males, 10.iv.2003, J.E. Smith leg. Ecological notes: found on wetlands including fens (HRI). Lisdoonvarna, Clare, 1 male, 21.iv.1982, J.P. with Carex and Phragmites beds. vi–vii, ix–x. O’Connor leg. (NMID). Longborough, Gloucestershire, 5 males, 8.iv.2003, J.E. Smith leg. (HRI). Offley Holes, Hertfordshire, 2 males, iv.1936, Edwards leg. CAMPTOCHAETA STAMMERI (LENGERSDORF, 1940) [paratypes of Corynoptera abblanda] (BMNH). Sels- Literature: Laurence & James (1996): 73; Laurence & don, Greater London, 1 male, 18.iv.1938, Coe leg. James (1997): 166 [all as Corynoptera]; Menzel (1998): (BMNH). Stonybrow Wood, Rotherfield Park, Hamp- 20. shire, 1 male, 3.vii.2002, Chandler leg. (PPCM). Sut- ton Park, Birmingham, West Midlands, 10 males, 1 Material examined: Ben Macdhui, Aberdeenshire, 1 female, 7.iv.2003, J.E. Smith leg. (HRI). Symond’s Yat, male, 18.vii.1982; 1 male, 6.viii.1983, all no collector Herefordshire, 1 male, ix.1936, Edwards leg. [holotype detail (PPWM). Cadair Idris, Gwynedd, 1 male, of Corynoptera abblanda] (BMNH). Wicken Fen, Cam- 3.vii.1963; 1 male, 17.vii.1993; 5 males, 4.ix.1993; 1 bridgeshire, 1 male, 12.v.1990, Withers leg. (PPWM). male, no date, all Laurence leg. (PBLN). Wood Crates, New Forest, Hampshire, 1 male, Ecological notes: found on high and open montane viii.1936, Edwards leg. (BMNH). habitats. vii–ix. Ecological notes: common in broad-leaved and mixed woodland; riverbank with alders; calcareous grass- CAMPTOCHAETA VIVAX (FREY, 1948) land; heath and parkland; wetlands including fen New to British list. meadows and ditches. On fallen beech. iii–xi. Material examined: The Loke, Norwich, Norfolk, 1 male, 26.iv.1987, Laurence leg. (PBLN). CLAUSTROPYGA BREVICHAETA (MOHRIG & ANTONOVA, 1978) Ecological notes: found in garden. iv. Literature: Freeman (1987): 200; Menzel (1998): 20 [all as Corynoptera acanthostyla, misidentification]; CLAUSTROPYGA HIPPA, VILKAMAA & MOHRIG, 2003 Hippa et al. (2003): 488. CLAUSTROPYGA ABBLANDA (FREEMAN, 1983) Material examined: Goyt Valley, Derbyshire, 1 male, Literature: Freeman (1983a): 163; Freeman (1983b): 13.iii.1931, Britten leg. (BMNH). Longdendale, 29; Menzel (1998): 20; Chandler et al. (2002): 123 [all Cheshire, 2 males, 24.ix.1932, Britten leg. (BMNH). as Corynoptera]; Hippa, Vilkamaa & Mohrig (2003): Mid Quoich, Aberdeenshire, 1 male, 21.iv.2000, God- 486. frey leg. (PPCM).

Ecological notes: found in broad-leaved and ancient Beeches, Buckinghamshire, 4 males, 1.vii.−8.vii.1996; pine woodland. iii–iv, ix. 1 male, 5.viii.−31.viii.1996; 1 male, 9.x.−26.xi.1996, all Ismay leg. (HRI, PPCM). Duxmoor Dingle, Shropshire, 1 male, 15.x.1987, Chandler leg. (PPCM). Goathland, CLAUSTROPYGA HETEROCLAUSA (RUDZINSKI, 1991) North Yorkshire, 1 male, 5.ix.1937, Edwards leg. Literature: Menzel (1998): 21 [as Corynoptera]. (BMNH). Hay Wood, Baddesley Clinton, Solihull, West Midlands, 1 male, 25.viii.2002, J.E. Smith leg. (HRI). Material examined: Hartslock, Oxfordshire, 1 male, Headley Common, Hampshire, 1 male, 23.x.1972, 6.v.2000, Raper leg. (PDGB). Malham Tarn, North Cranston leg. (BMNH). Highgate Wood, Greater Lon- Yorkshire, 1 male, 11.ix.−18.ix.1981; 1 male, 2.vi.− don, 1 male, 1998–99, Hackett leg. (PPCM). Hunt- 3.vi.1982, all Disney leg. (PPWM). The Loke, Norwich, house Woods, Worcestershire, 1 male, 13.x.1984, Norfolk, 1 male, 8.v.1987, Laurence leg. (PBLN). Chandler leg. (PPWM). Lion Wood, Redditch, Worces- Ecological notes: found in calcareous grassland adja- tershire, 1 male, 10.iv.2003, J.E. Smith leg. (HRI). cent to hanger woodland; bog and lakeshore; garden. Loch Chon, Stirling, 1 male, 22.vi.1999, Chandler leg. v–vi, ix. (PPCM). Low Pasture, Hawnby, North Yorkshire, 1 male, 12.x.1990, Chandler leg. (PPCM). Rothiemur- chus, by River Druie, Highland Region, 1 male, CORYNOPTERA WINNERTZ, 1867 18.viii.1986, Chandler leg. (PPCM). Slade of Saggart, CORYNOPTERA BICUSPIDATA (LENGERSDORF, 1926) Dublin, 1 male, 26.ix.1999, Chandler leg. (PPCM). Synonym: gymnops Tuomikoski, 1960. Sutton Park, Birmingham, West Midlands, 4 males, 7.iv.2003, J.E. Smith leg. (HRI). New to British list. Ecological notes: found in broad-leaved, mixed and Material examined: Chigwell Row Wood, Chigwell coniferous woodland; heathland and parkland; wet- Row, Essex, 1 male, 15.ix.2002; 2 males, 23.ix.2002, all lands including lakeshore marsh. iv, vi–xi. Ismay & Schulten leg. (DEI, UMO). Selsdon, Greater London, 1 male, 4.vii.1934; 1 male, 6.vii.1934; 1 male, 18.vii.1934; 2 males, 19.vii.1934; 2 males, 26.vii.1934; CORYNOPTERA CAVIPES MOHRIG, 1993 1 male, 23.vii.1935; 1 male, 31.vii.1934, all Coe leg. New to British list. (BMNH) [= Surrey record of Corynoptera perpusilla in Freeman (1983b)]. Material examined: Hartslock, Oxfordshire, 1 male, 14.v.2000; 1 male, 19.v.2000, all Raper leg. (PDGB). Ecological notes: found on heathland (heather, Jun- Letchworth, Hertfordshire, 1 male, v.1917; 1 male, cus, Deschampsia) with ancient and secondary wood- ix.1917; 1 male, x.1917; all Edwards leg. (BMNH). Mil- land. With ants Lasius niger in bungalow. vii, ix. ham Ford School, Oxford, Oxfordshire, 3 males, 3.v.− 5.vi.2002, Webb leg. (PPCM). Pant-y-Sais, Neath/Port CORYNOPTERA BISTRISPINA Talbot, 1 male, 13.vii.1989, Holmes, Boyce & Reed leg. (BUKOWSKI & LENGERSDORF, 1936) (PPWM). Salisbury Plain Training Area, Warminster, Wiltshire, 1 male, 27.v.2003, Ismay & Schulten leg. Synonym: defecta (Frey, 1948). (UMO). The Loke, Norwich, Norfolk, 1 male, 21.v.− New to British list. 23.v.1993, Laurence leg. (PBLN). Wytham Woods, Wytham, Oxfordshire, 1 male, 21.viii.2002, J.E. Smith Material examined: Hartslock, Oxfordshire, 2 males, leg. (HRI). 6.v.2000; 2 males, 14.v.2000, all Raper leg. (HRI, PDGB, PPCM). Ecological notes: found in broad-leaved woodland; wetlands including Phragmites and Molinia bog; Ecological notes: found on calcareous grassland adja- unimproved calcareous grassland; school grounds cent to hanger woodland. v. with old poplars. v–x.

CORYNOPTERA BLANDA (WINNERTZ, 1867) CORYNOPTERA COMPRESSA (WALKER, 1848) Synonyms: curviligula (Lengersdorf, 1952); villica Synonym: pallida (Walker, 1856). (Winnertz, 1867). Literature: Walker (1848): 110; Walker (1856): 56 [all Literature: Freeman (1987): 199; Menzel (1998): 20; as Sciara compressa]; Walker (1856): 56; Edwards Chandler et al. (2002): 123. (1925a): 540 [all as Sciara pallida]; Freeman (1983b): Material examined: Becky Woods, Manaton, Devon, 1 29; Menzel (1998): 20; Menzel & Mohrig (2000): 246; male, 23.vi.2002, J.E. Smith leg. (HRI). Burnham Withers (2002): 114.

Material examined: Baldock, Hertfordshire, 1 male, Material examined: Ban-y-Gor Wood, Gloucestershire, v.1918, Edwards leg. (BMNH). Burnham Beeches, 1 male, 12.ix.2002, Gibbs leg. (PDGB). Buckingham Buckinghamshire, 1 male, 25.v.−1.vi.1999, Ismay leg. Palace Garden, Greater London, 1 male, 26.v.− (PPCM). Killaun Bog, Offaly, 1 male, ix.2000, Withers 19.vi.1997, D.J. Carter & C.W. Plant leg. (PPCM). Cors leg. (PPWM). Letchworth, Hertfordshire, 1 male, Erddreiniog, Isle of Anglesey, 1 male, 14.vi.1988, vi.1917; 2 males, vi.–vii.1936, all Edwards leg. Holmes, Boyce & Reed leg. (PPWM). Dent, Cumbria, (BMNH, ZFMK). Nutcombe Down, Surrey, 1 male, 1 male, 23.vii.1935, Edwards leg. [paratype of 26.vii.2002, Chandler leg. (PPCM). Savernake Forest, Corynoptera curvispinosa] (BMNH). Dickleburgh, Wiltshire, 1 male, 29.ix.1971 (reared), Chandler leg. Norfolk, 1 male, 21.iv.1989, Withers leg. (PPWM). (PPCM). No locality details, ? England, 1 female, 1856, Dinton Pastures Country Park, Berkshire, 1 male, Walker leg. [lectotype of Sciara pallida] (BMNH). No 3.vii.1994, Chandler leg. (PPCM). Epping Forest, locality details, ? England, 1 female, no date, Barnston Essex, 8 males, iv.-ix.1999, Dagley leg. (PPCM). leg. [lectotype of Sciara compressa] (BMNH). Killaun Bog, Offaly, 1 male, ix.2000, Withers leg. (PPWM). Letchworth, Hertfordshire, 1 male, 1922, Ecological notes: found in ancient beech and other Edwards leg. [paratype of Corynoptera curvispinosa] broad-leaved woodland; raised bog. Reared from the (BMNH). Newbridge Fen, Kildare, 1 male, 8.viii.− fungus Hypoxylon fragiforme on beech bark. vi–vii, ix. 19.viii.1998, Good leg. (PPWM). Santon Downham, Suffolk, 1 male, 15.viii.1970, Hutson et al. leg. CORYNOPTERA CONCINNA (WINNERTZ, 1867) [paratype of Corynoptera curvispinosa] (BMNH). Synonym: semiconcinna Mohrig & Krivosheina, 1985. The Oaks, Cambridgeshire, 2 males, ix.1998, C.W. Plant leg. (PPCM). Trawsfynydd, Gwynedd, 1 male, New to British list. 14.x.1988, Holmes, Boyce & Reed leg. (PPWM). Material examined: Berwick-upon-Tweed, Northum- Wicken Fen, Cambridgeshire, 2 males, 12.v.1990, berland, 1 male, vi.1999, Laurence leg. (PBLN). Glass- Withers leg. (PPWM). Wychwood, Oxfordshire, 1 male, house Crops Research Station, Littlehampton, West v.1937, Edwards leg. [holotype of Corynoptera curvis- Sussex, 1 male, 28.i.1958; 1 male, 9.iii.1960; 3 males, 1 pinosa] (BMNH). Wytham Woods, Wytham, Oxford- female, no date, all Hussey leg. (HRI). Letchworth, shire, 2 males, 21.viii.2002, J.E. Smith leg. (HRI). Hertfordshire, 1 male, 25.vi.1934, Edwards leg. Ecological notes: found in broad-leaved woodland; (BMNH). Wellesbourne, Warwickshire, 1 male, wooded bank of river; wetlands including fens, water 17.viii.−31.viii.2000, J.E. Smith leg. (HRI). meadows, raised bog, reed beds and Cladium beds Ecological notes: found in grounds of horticultural with tufa; old hedges; gardens. iv–x. research stations; garden. i, iii, vi, viii.

CORYNOPTERA DENTATA CORYNOPTERA CUNICULATA (LENGERSDORF, 1942) (BUKOWSKI & LENGERSDORF, 1936) Synonym: caldariorum Tuomikoski, 1960. New to British list. New to British list. Material examined: Burnham Beeches, Buckingham- Material examined: Burnham Beeches, Buckingham- shire, 3 males, 14.vi.−5.viii.1996, Ismay leg. (PPCM). shire, 9 males, 28.v.−26.xi.1996; 1 male, 22.vii.− Epping Thicks, Epping Forest, Essex, 1 male, vi.1999, 5.viii.1996, all Ismay leg. (PPCM). Chigwell Row Dagley leg. (PPCM). Lackham Park, Wiltshire, 1 male, Wood, Chigwell Row, Essex, 1 male, 11.vi.2002; 2 2.vii.2002, Chandler leg. (PPCM). Rotherﬁeld Park, females, 18.vi.2002; 1 male, 1 female, 8.x.2002, all Hampshire, 1 male, 3.vii.2002, Chandler leg. (PPCM). Ismay & Schulten leg. (DEI). Monk’s Soham, Suffolk, Ecological notes: found in ancient beech and other 1 male, 5.vi.1915, Morley leg. (BMNH). Sedlescombe, broad-leaved woodland. vi–viii. East Sussex, 1 male, 21.viii.1994, Roper leg. (PBLN). Ecological notes: found in ancient broad-leaved woodland; ancient woodland with oak, hazel and bramble; CORYNOPTERA FLAVICAUDA (ZETTERSTEDT, 1855) heathland (heather) with oak and holly; on fungal Literature: Edwards (1925a): 538 [as Sciara]; Edwards growth on old willow. v–xi. (1938): 201 [as Sciara (Neosciara)]; Freeman (1983b): 28; Menzel (1998): 21; Chandler et al. (2002): 123. CORYNOPTERA CURVISPINOSA FREEMAN, 1983 Material examined: Annahala bog, Gearagh, Cork, Literature: Freeman (1983a): 164; Freeman (1983b): 2 males, 22.iii.1990; 1 male, 7.iv.1990; 6 males, 29; Menzel (1998): 21; Chandler (2001): 201; Withers 1.vi.1990, all de Courcy Williams leg. (DEI, NMID). (2002): 114. Alconbury, Cambridgeshire, 3 males, 4.v.1989, J.H.

Cole leg. (PJCB). Aldermoors, Berkshire, 1 male, Williams leg. (DEI); 1 male, 9.vii.1990, Withers leg. 14.v.1994; 1 male, 11.vi.1994, all Chandler leg. (PPWM). Glen Coiltie, Highland Region, 1 male, (PPCM). Ashridge Estate, Hertfordshire, 2 males, 23.vi.1999; 1 male, 28.v.2002, all Chandler leg. 20.v.1999, Chandler leg. (NMS). Aston Rowant, (PPCM). Golden Park, Devon, 1 male, 16.vi.1989, Oxfordshire, 2 males, 1 female, 24.vi.1972, Chandler Chandler leg. (NMS). Hambleden Great Wood, leg. (PPCM). Badger Wood, Cambridgeshire, 1 male, Buckinghamshire, 1 male, 15.v.1997, Chandler leg. 1.v.−31.v.1998, C.W. Plant leg. (NMS). Bagley (PPCM). Hartslock, Oxfordshire, 2 males, 10.v.− Wood, Oxfordshire, 8 males, 20.v.−21.v.1933; 1 male, 14.v.2000; 1 male, 14.v.2000, all Raper leg. (HRI, 8.v.1936; 1 male, 18.v.1936, all Edwards leg. (BMNH). PDGB). Hawnby, River Rye, North Yorkshire, 1 male, Ballykeeffe Wood, Kilkenny, 1 male, 25.vi.1987, Chan- 19.vii.1996, Perry leg. (NMS). Hayley Wood, Cam- dler leg. (PPCM). Bayfordbury Estate, Hertford, bridgeshire, 1 male, 31.v.1975, J.H. Cole leg. (PJCB). Hertfordshire, 1 male, 22.iv.1975, J.H. Cole leg. Hobb’s Quarry, Gloucestershire, 1 male, 15.v.2003, (PJCB). Bedford Purlieus, Cambridgeshire, 2 males, Gibbs leg. (PDGB). Ivy Thorn Hill, Somerset, 1 male, 23.vi.1975, J.H. Cole leg. (PJCB). Bishop’s Stortford, 4.v.2001; 1 male, 20.vi.2001, all Gibbs leg. (PDGB). Hertfordshire, 1 male, iv.1994, C.W. Plant leg. Knocksink Wood, Wicklow, 1 male, 17.vi.1985, Chan- (PPCM). Bix Bottom, Oxfordshire, 2 males, 9.v.1998, dler leg. (PPWM). Lancaut Nature Reserve, Glouces- Chandler leg. (NMS). Bradﬁeld Wood, Suffolk, 1 male, tershire, 1 male, 28.v.2003, Gibbs leg. (PDGB). 23.v.−30.v.1991, Laurence leg. (PBLN). Brampton Leckford Estate, Hampshire, 1 male, 18.vii.1972, Wood, Cambridgeshire, 1 male, 23.v.1967; 1 male, Chandler leg. (PPCM). Letchworth, Hertfordshire, 1 25.v.1987; 2 males, 13.v.1989, all J.H. Cole leg. (PJCB). male, v.1917; 6 males, vi.1917; 1 male, v.1918, all Brandyquoi, Kirkwall, Orkney, 12 males, v.-vi.2001, Edwards leg. (BMNH). Loxley Wood, Somerset, 1 Laurence leg. (PBLN). Buckingham Thick Copse, male, 5.vii.1985, Chandler leg. (PPCM). Loynton Northamptonshire, 1 male, 9.vi.1992, Warne leg. Moss, Staffordshire, 1 male, 23.v.1994, Chandler leg. (PPCM). California Country Park, Berkshire, 1 male, (PPCM). Monk Wood, Avon, 2 males, 24.v.2000, Gibbs 2.vi.1996, Chandler leg. (PPCM).Castlecomer, Kilk- leg. (PDGB). Monkﬁeld Wood, Cambridgeshire, 2 enny, 2 males, 25.vi.1987, Chandler leg. (NMS). Char- males, 1.v.−31.v.1998, C.W. Plant leg. (PPCM). New lbury, Oxfordshire, 1 male, 4.vii.1936, Edwards Malden, Greater London, 2 males, 18.vi.1932, Coe leg. leg. (BMNH). Charleville Wood, Offaly, 9 males, (BMNH). Oakland Wood, Cambridgeshire, 1 male, 15.vi.1985, Chandler leg. (PPWM). Chedworth, Glouc- 1.v.−31.v.1998, C.W. Plant leg. (NMS). Old Buckenham estershire, 1 male, 5.vi.1936, Edwards leg. (BMNH). Fen, Norfolk, 2 males, 9.vi.1990, Chandler leg. (NMS). Cliveden, Buckinghamshire, 1 male, 3.vii.1984, Purn Hill, Avon, 1 male, 24.v.2001, Gibbs leg. (PDGB). Chandler leg. (PPCM). Coombe, Cornwall, 1 male, Radwell, Hertfordshire, 1 male, v.1918, Edwards leg. 16.vi.1989, Chandler leg. (NMS). Cop Mere, Stafford- (ZFMK). Reelig Glen, Moniack, Highland Region, 1 shire, 1 male, 23.v.1994, Chandler leg. (PPCM). male, 23.vii.1999, Chandler leg. (NMS). Rewell Cothill, Oxfordshire, 1 male, 20.v.1936, Edwards leg. Wood, West Sussex, 1 male, 10.vi.1979, Chandler leg. (BMNH); 1 male, 14.vi.1986, Chandler leg. (PPCM). (PPCM). Rosevear, Cornwall, 1 male, 6.vii.1983, Chan- Crawfordsburn Country Park, Down, 1 male, dler leg. (NMS). Rousdon, Devon, 1 male, 8.vi.1937, 2.vii.1987, Chandler leg. (NMS). Crickhowell, Powys, Edwards leg. (BMNH). Runnymede, Surrey, 1 male, 1 male, v.1937, Edwards leg. (BMNH). Dickleburgh, 8.vii.1973, Chandler leg. (PPCM). Salisbury Plain Norfolk, 3 males, 4.vi.−18.vi.1989, Withers leg. Training Area, Warminster, Wiltshire, 1 male, (PPWM). Dingwall, Highland Region, 1 male, 29.v.2003, Ismay & Schulten leg. (UMO). Selborne 18.vi.1976, Chandler leg. (PPCM). Dinton Pastures Hanger, Hampshire, 1 male, 12.vi.1979, Chandler leg. Country Park, Berkshire, 4 males, 25.iv.−13.vi.1993; 2 (PPCM). Selsdon, Greater London, 2 males, 6.vi.1938, males, 2.v.−12.vi.1994; 1 male, 6.viii.1994, all Chan- Coe leg. (BMNH). Sharpenhoe, Bedfordshire, 1 male, dler leg. (PPCM). Duncombe Park, North Yorkshire, 1 16.vi.1997, Chandler leg. (PPCM). Sheep Leas, West male, 16.vii.1996, Chandler leg. (PPCM). East Grin- Horsley, Surrey, 2 males, 1 female, 21.v.1966; 2 males, stead, West Sussex, 1 male, no date, Stubbs leg. 18.v.1991; 2 males, 16.vi.1991; all Chandler leg. (NMS, (PPCM). East Trewent Wood, Pembrokeshire, 3 males, PPCM). Shortwood, Gloucestershire, 1 male, 12.vii.1986, Chandler leg. (PPCM). Epping Forest, 12.vi.2003, Gibbs leg. (PDGB). Sowley Pond, New For- Essex, 1 male, 25.v.1999, Chandler leg. (PPCM). Farn- est, Hampshire, 1 male, 22.v.1936, Edwards leg. ingham Wood, Kent, 2 males, 18.v.1974, Stubbs (BMNH). The Loke, Norwich, Norfolk, 1 male, 5.vi.− leg. (PPCM). Forge Valley, North Yorkshire, 1 male, 13.vi.1989, Laurence leg. (PBLN). The Moors, Bishop’s 18.vii.1996, Chandler leg. (PPCM). Fowl Mere, Nor- Waltham, Hampshire, 1 male, 13.x.1989, M.N. Pugh folk, 2 males, vi.1939, Edwards leg. (BMNH). Gait leg. (PPCM). Tickenham Hill, Avon, 1 male, 30.v.2001, Barrows, Cumbria, 1 male, 25.v.1994, Chandler leg. Gibbs leg. (PDGB). Trefeiddan Moor, Pembrokeshire, 1 (PPCM). Gearagh, Cork, 1 male, 7.vi.1990, de Courcy male, 21.vii.1987, Holmes, Boyce & Reed leg. (PPWM).

Ufton Bridge, Warwickshire, 2 males, 9.vi.1991, Chan- Chandler leg. [paratypes of Corynoptera flavosignata] dler leg. (NMS, PPCM). Virginia Woods, Cavan. 1 (HRI, PPCM). Theale, Berkshire, 10 males, 7.vii.2003, male, 21.v.1989, J.P. O’Connor leg. (NMID). Weem Chandler leg. [paratypes of Corynoptera flavosignata] Castle Woods, Perth & Kinross, 1 male, 23.vi.1990, (HRI, PPCM). Chandler leg. (NMS). West Walk, Hampshire, 1 male, Ecological notes: found in broad-leaved and carr wood- 6.vi.1998, Chandler leg. (NMS). Wicken Fen, Cam- land; young mixed woodland; wooded river bank; graz- bridgeshire, 1 male, 12.v.1990, Withers leg. (PPWM). ing marshes; water meadows; gravel pits; old hedges. Winmarleigh Moss, Lancashire, 1 male, 14.vi.1999, v–ix. Chandler leg. (NMS). Wistow Wood, Warboys, Cam- bridgeshire, 2 males, 4.v.1975, J.H. Cole leg. (PJCB). Wychwood, Oxfordshire, 1 male, v.1937, Edwards leg. (BMNH). CORYNOPTERA FORCIPATA (WINNERTZ, 1867) Synonyms: echinocordyla Laurence, 1994; involuta Ecological notes: very common in broad-leaved, mixed (Frey, 1948). and carr woodland; partly wooded heathland; wetlands including fen, water meadows and basin mire Literature: Freeman (1983b): 28; Laurence (1994): with Menyanthes bog; gravel pits; ditch along meadow 109 and 118 [as Corynoptera echinocordyla and hedge; old hedges; disused railway cutting; garden; Corynoptera forcipata]; Menzel (1998): 21; Menzel & common on unimproved calcareous grassland. iv–viii, Mohrig (2000): 247; Chandler (2001): 201. x. Material examined: Ashwell Grove, Gloucestershire, 1 male, 13.x.1979, Chandler leg. (PPCM). Badger Wood, − CORYNOPTERA FLAVOSIGNATA MENZEL & HELLER Cambridgeshire, 3 males, 1.v. 31.v.1998, C.W. Plant − SP. NOV. leg. (PPCM). Bagley Wood, Oxfordshire, 1 male, 20.v. 21.v.1933, Edwards leg. (BMNH). Bagpiper’s Tump, New to British list. West Wood, Herefordshire, 2 males, 10.x.1998, Chan- Material examined: Badger Wood, Cambridgeshire, dler leg. (NMS). Baldock, Hertfordshire, 8 males, 1 male, vi.1998, C.W. Plant leg. [paratype of ix.-x.1917, Edwards leg. (BMNH, ZFMK). Beaulieu Corynoptera flavosignata] (PPCM). Bradfield Wood, River, Hampshire, 1 male, 22.v.1936, Edwards leg. Suffolk, 2 males, 6.vii.−11.vii.1990, Laurence leg. (BMNH). Belladrum Burn, Highland Region, 1 male, [paratypes of Corynoptera flavosignata] (PBLN). Cas- 19.vii.1991, Chandler leg. (NMS). Blackgang, Isle of tle Eden Dene, Durham, 1 male, 26.vi.1992, Chandler Wight, 1 male, 25.iv.1975, Jarzembowski leg. leg. [paratype of Corynoptera flavosignata] (PPCM). (BMNH). Bradfield Wood, Suffolk, 1 male, 11.vii.− Charlbury, Oxfordshire, 1 male, 4.vii.1936, Edwards 16.vii.1990, Laurence leg. [paratype of Corynoptera leg. (BMNH). Coombe Abbey Country Park, Coventry, echinocordyla] (PBLN). Brancaster, Norfolk, 3 males, Warwickshire, 1 female, 28.viii.2003, Menzel leg. 2.ix.1988, Laurence leg. (PBLN); 32 males, 16.ix.− [paratype of Corynoptera flavosignata] (DEI). Lack- 28.x.1988; 7 males, 28.x.1988, all Foster & Procter leg. ham Wood, Wiltshire, 1 male, 2.v.2002, Chandler [not 40 males; holotype and paratypes of Corynoptera leg. [paratype of Corynoptera flavosignata] (PPCM). echinocordyla] (DEI, PBLN). Brandon Marsh, Ryton- Letchworth, Hertfordshire, 1 male, v.1918, Edwards on-Dunsmore, West Midlands, 4 females, 12.x.2001, leg. (BMNH). Little Wittenham Wood, Oxfordshire, J.E. Smith leg. (HRI); 15 males, 1 female, 22.viii.2002, 1 male, 3.vii.1985, Chandler leg. [paratype of Menzel & J.E. Smith leg. (DEI, HRI). Bucklebury Corynoptera flavosignata] (PPCM). Monkfield Wood, Common, Berkshire, 1 male, 30.viii.1983, Chandler Cambridgeshire, 1 male, vi.1998, C.W. Plant leg. leg. (PPCM). Burnham Beeches, Buckinghamshire, 4 [paratype of Corynoptera flavosignata] (PPCM). Mor- males, 10.v.−27.v.1995; 1 male, 13.ix.−26.ix.1996, all timer’s Meadow, Dinton Pastures Country Park, Berk- Ismay leg. (HRI, PPCM); 1 male, 20.iv.1998, Chandler shire, 2 males, 26.vi.1993, Chandler leg. [paratypes leg. (PPCM). Castlecomer, Kilkenny, 1 male, of Corynoptera flavosignata] (PPCM). Newbold 25.vi.1987, Chandler leg. (PPCM). Castlefreke, Cork, Comyn Park, Leamington Spa, Warwickshire, 1 male, 1 male, 12.v.1989, Withers leg. (PPWM). Charleville 15.vii.2001, J.E. Smith leg. [paratype of Corynoptera Wood, Offaly, 2 males, 27.v.1984, Chandler leg. flavosignata] (HRI). Pawlett Levels, Somerset, 1 male, (PPCM). Chigwell Row Wood, Chigwell Row, Essex, 2 13.ix.1999, Gibbs leg. [paratype of Corynoptera fla- females, 20.v.2002; 2 males, 23.v.−27.v.2002; 1 male, 3 vosignata] (PDGB). Sandford Copse, Dinton Pastures females, 27.v.2002; 1 male, 28.vi.2002; 3 males, 1 Country Park, Berkshire, 2 males, 3.vii.1994, Chan- female, 15.ix.2002; 2 males, 2 females, 23.ix.2002; 1 dler leg. [paratypes of Corynoptera flavosignata] male, 1 female, 26.ix.2002; 2 males, 8.x.2002; 9 males, (PPCM). Searles Farm, Reading, Berkshire, 4 males, 2 4 females, 16.x.2002, all Ismay & Schulten leg. (DEI, females, 26.vi.2003; 2 males, 1 female, 22.vii.2003, all UMO). Coombe Abbey Country Park, Coventry, War-

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 BRITISH SCIARID FAUNA 73 wickshire, 29 males, 4 females, 28.viii.2003, Menzel & Laurence leg. [paratypes of Corynoptera echino- J.E. Smith leg. (DEI, HRI). Cothill, Oxfordshire, 15 cordyla] (DEI, PBLN). Thompson Common, Norfolk, males, 26.v.1936, Edwards leg. (BMNH). Crickhowell, 1 male, 13.ix.−27.ix.1988, Foster & Procter leg. Powys, 5 males, 1.x.1936; 3 males, v.1937, all Edwards [paratypes of Corynoptera echinocordyla] (PBLN). leg. (BMNH). Daneway, Gloucestershire, 1 male, Tickenham Hill, Avon, 1 male, 30.v.2001, Gibbs leg. 12.x.1979, Chandler leg. (PPCM). Dickleburgh, Nor- (PDGB). Tocil Wood, Warwick University Campus, folk, 1 male, 4.vi.−18.vi.1989, Withers leg. (PPWM). Coventry, Warwickshire, 8 males, 28.viii.2003, Menzel Dinton Pastures Country Park, Berkshire, 1 male, & J.E. Smith leg. (DEI, HRI). Tuddenham Fen, 7.x.1999, Chandler leg. (PPCM). Epping Forest, Suffolk, 1 male, 20.x.1990, Chandler leg. (NMS). Essex, 2 males, 24.v.−25.v.1999, Chandler leg. (NMS); Weston, Hertfordshire, 1 male, 4.x.1931, Edwards leg. 3 males, v.1999, Dagley leg. (PPCM). Falls of Tarnash, (BMNH). Weston Big Wood, Avon, 1 male, 1.ix.2001, Moray, 1 male, 1.vii.1936; 3 males, 9.vii.1936, all Coe Gibbs leg. (PDGB). Weston Common, Hampshire, 1 leg. (BMNH). Forge Valley, North Yorkshire, 1 male, male, 10.v.1984, Chandler leg. (NMS). Weston Green 13.x.1990, Chandler leg. (NMS). Gearagh, Cork, 2 Fen, Oxfordshire, 2 males, 1.vi.1991, Chandler leg. males, 11.v.1990; 2 males, 1 female, 10.x.1990, all de (NMS, PPCM). Wheatfen Broad, Norfolk, 1 male, Courcy Williams leg. (DEI, NMID). Glencar Waterfall, 1.ix.1986; 1 male, 19.v.1990, all Withers leg. (PPWM). Sligo, 1 male, 15.v.1970, Chandler leg. (NMS). Hafod Windsor Forest, Berkshire, 1 male, 8.vi.1979; 3 Garregog, Gwynedd, 1 male, 26.vii.1988, Holmes, males, 6.ix.1986; 1 male, 15.v.1988; 2 males, Boyce & Reed leg. (PPWM). Haugh Wood, Hereford- 22.v.1988; 2 males, 15.v.1994; 1 male, 23.vii.1999, all shire, 1 male, 12.x.1998, Chandler leg. (NMS). Head- Chandler leg. (NMS, PPCM, PPWM). Wistow Wood, ington Wick, Oxfordshire, 1 male, 13.vi.1910, Hamm Warboys, Cambridgeshire, 1 male, 4.v.1975, J.H. leg. (UMO). Hickling, Norfolk, 1 male, v.1989, Foster Cole leg. (PJCB). Witherslack, Cumbria, 1 male, & Procter leg. (PBLN). Holme Fen, Cambridgeshire, 1 13.vi.1977, Chandler leg. (NMS). Witley Common, male, 27.v.1987, J.H. Cole leg. (PJCB). Illtyd Pools, Surrey, 1 male, 16.v.1992, Chandler leg. (PPCM). Powys, 1 male, 11.x.1989, Chandler leg. (PPWM). Jef- Woodwalton Fen, Cambridgeshire, 1 male, 24.v.1940, fry Bog, North Yorkshire, 1 male, 14.vii.1997, Chan- Edwards leg. (BMNH); 2 males, 4.vi.1977, J.H. dler leg. (PPCM). Kennack Cove, Cornwall, 2 males, Cole leg. (PJCB). Wychwood, Oxfordshire, 5 males, 4.ix.1983, Chandler leg. (NMS). Killaun Bog, Offaly, 6 ix.-x.1917, Edwards leg. (BMNH). Wytham Woods, males, ix.2000, Withers leg. (PPWM). Lady’s Wood, Wytham, Oxfordshire, 78 males, 4 females, Cambridgeshire, 1 male, 3.v.1989, J.H. Cole leg. 21.viii.2002, Menzel & J.E. Smith leg. (DEI, HRI). No (PJCB). Lancaut Nature Reserve, Gloucestershire, 1 locality details, England, 1 male, no date, Laurence male, 5.ix.2002, Gibbs leg. (PDGB). Loch Park, Moray, leg. (PBLN). 2 males, 3.vii.1936; 3 males, 12.viii.−14.viii.1937, all Ecological notes: very common in broad-leaved, mixed Coe leg. (BMNH). Lopham Fen, Norfolk, 3 males, and carr woodland; wet and dry heathland; wetlands 20.v.1990, Withers leg. (PPWM). Malham Tarn, North including water meadows, fens, bogs, basin mires, Yorkshire, 1 male, 27.viii.−30.viii.1981; 1 male, 11.ix.− sedge beds and Cladium beds with tufa; former gravel 18.ix.1981; 2 males, 26.vi.−28.vi.1982, all Disney leg. workings; grassland with pingo pools; old hedges; gar- (BMNH, PPWM). Monk Fryston, North Yorkshire, dens; around rotten fallen tree trunk; coastal landslip. 1 male, 7.vi.1988, Chandler leg. (PPCM). Monk’s iv–x. Wood, Cambridgeshire, 1 male, 18.vi.1978; 1 male, 12.vi.1982, all J.H. Cole leg. (PJCB). Muckross House, Kerry, 1 male, 5.v.1981, Chandler leg. (PPCM). Mulgrave Wood, North Yorkshire, 1 male, 23.viii.− CORYNOPTERA FRITZI MOHRIG & RULIK, 2001 3.ix.1937, Edwards leg. (BMNH). Murrough, Wicklow, New to British list. 4 males, 8.v.1989; 2 males, 23.v.1989, all Withers leg. Material examined: Bolt Wood, Stockerston, Leicester- (PPWM). New Close Wood, Brandon, Warwickshire, shire, 1 male, 1 female, 24.viii.2003, J.E. Smith leg. 2 males, 28.viii.2003, Menzel leg. (DEI). Newbridge (HRI). Coombe Abbey Country Park, Coventry, War- Fen, Kildare, 5 males, 8.viii.−19.viii.1998, Good leg. wickshire, 2 males, 2 females, 28.viii.2003, Menzel & (PPWM). Randwick, Gloucestershire, 1 male, J.E. Smith leg. (DEI, HRI). Kelham Bridge, Raven- 11.x.1979, McLean leg. (NMS). Ross Island, Killarney, stone, Leicestershire, 2 males, 24.viii.2003, Menzel Kerry, 1 male, 7.v.1981, Chandler leg. (PPCM). Ryton leg. (DEI). Ryton Wood, Bubbenhall, Coventry, West Wood, Bubbenhall, Coventry, West Midlands, 13 Midlands, 6 males, 25.viii.2002, Menzel & J.E. Smith males, 1 female, 25.viii.2002, Menzel & J.E. Smith leg. leg. (DEI, HRI). (DEI, HRI). The Flits, Herefordshire, 1 male, 11.x.1998, Chandler leg. (PPCM). The Loke, Norwich, Ecological notes: found in broad-leaved woodland; wet Norfolk, 2 males, 18.vi.1987; 2 males, 15.ix.1987, all meadow with reed beds around pools. viii.

CORYNOPTERA FURCATA (HIPPA & VILKAMAA, 1994) 14.vii.2003, J.E. Smith leg. (HRI). The Coombe, Buck- Literature: Freeman (1983b): 29 [as Corynoptera par- inghamshire, 1 male, 17.vi.1997, Chandler leg. vula, misidentification]; Menzel (1998): 21. (PPCM). Windermere, Cumbria, 1 male, vi.1924, Edwards leg. (BMNH). Windsor Forest, Berkshire, 1 Material examined: Becky Woods, Manaton, Devon, 1 male, 15.v.1994, Chandler leg. (PPCM). male, 23.vi.2002, J.E. Smith leg. (HRI). Charlbury, Oxfordshire, 1 male, 4.vii.1936, Edwards leg. Ecological notes: very common in broad-leaved and (BMNH). Lakenheath, Suffolk, 1 male, 30.viii.− carr woodland; mixed woodland and conifer plantation 13.ix.1988, Foster & Procter leg. (PBLN). Letchworth, with broad-leaved fringe; heathland (heather) with Hertfordshire, 1 male, vi.1917, Edwards leg. [paratype oak and holly; wetlands including water meadows and of Camptochaeta furcata] (BMNH). Tarradale House, sedge beds; calcareous grassland. v–vii. Highland Region, 1 male, vs. 2002, Sumner leg. (PPCM). Wytham Woods, Wytham, Oxfordshire, 1 male, 21.viii.2002, Menzel leg. (DEI). CORYNOPTERA GLOBIFORMIS (FREY, 1945) Ecological notes: found in broad-leaved woodland Synonyms: globula Freeman, 1983; paraglobula Rudz- including wet woodland on granite with alder and inski, 1988. birch; damp meadow. v–ix. Literature: Freeman (1983a): 165; Freeman (1983b): 28; Laurence (1997a): 85 [all as Corynoptera globula]; CORYNOPTERA FURCIFERA MOHRIG & MAMAEV, 1987 Menzel (1998): 21; Chandler (2001): 201; Withers (2002): 114. Synonym: vitella Rudzinski & Drissner, 1992. Material examined: Badger Wood, Cambridgeshire, New to British list. 1 male, 1.v.−31.v.1998, C.W. Plant leg. (PPCM). Bal- Material examined: Badger Wood, Cambridgeshire, 1 moral Forest, Aberdeenshire, 2 males, 19.vii.− male, vii.1998, C.W. Plant leg. (PPCM). Becky Woods, 30.vii.1938, Coe leg. (BMNH). Berwick-upon-Tweed, Manaton, Devon, 1 male, 23.vi.2002, J.E. Smith leg. Northumberland, 1 male, viii.1999, Laurence leg. (HRI). Boundless Copse, Surrey, 1 male, 26.vii.2002, (PBLN). Birks of Aberfeldy, Perth & Kinross, 1 male, Chandler leg. (PPCM). Brancaster, Norfolk, 2 males, 30.viii.1987, Chandler leg. (PPCM). Bix Bottom, 17.vi.−1.vii.1988; 1 male, 1.vii.−15.vii.1988, all Foster Oxfordshire, 1 male, 23.ix.2000, Raper leg. (PDGB). & Procter leg. (PBLN). Burnham Beeches, Bucking- Bradfield Wood, Suffolk, 6 males, 9.x.−16.x.1990, Lau- hamshire, 1 male, 26.vi.1993, Chandler leg. (PPCM); 4 rence leg. (PBLN). Buckingham Palace Garden, males, 2 females, 28.v.−5.viii.1996; 1 male, 1.vi.− Greater London, 1 male, 20.x.−20.xi.1995; 2 males, 8.vi.1999; 3 males, no date, all Ismay leg. (HRI, 5.ix.−21.x.1996, all D.J. Carter & C.W. Plant leg. PPCM). Canonteign Falls, Devon, 5 males, 23.vi.2002, (PPCM). Burnham Beeches, Buckinghamshire, 1 J.E. Smith leg. (HRI). Castell Cogan, Carmarthen, 1 male, 10.v.−23.v.1995; 1 male, 15.ix.−4.x.1995; 14 male, 10.vii.1986, Chandler leg. (PPCM). Chigwell males, 5.viii.−26.xi.1996, all Ismay leg. (PPCM). Row Wood, Chigwell Row, Essex, 1 male, 11.vi.2002, Cadair Idris, Gwynedd, 1 male, 16.viii.1993; 1 male, Ismay & Schulten leg. (DEI). Den of Alyth, Angus, 1 4.ix.1993, all Laurence leg. (PBLN). Chigwell Row male, 13.vii.1991, Chandler leg. (PPCM). Devichoys Wood, Chigwell Row, Essex, 3 males, 16.x.2002, Ismay Wood, Cornwall, 1 male, no date, Chandler leg. & Schulten leg. (DEI, UMO). Clapham, North York- (PPCM). Epping Forest, Essex, 4 males, vi.1999, Dag- shire, 1 male, viii.1981, Disney leg. (PPWM). Cock ley leg. (PPCM). Glan y Wern, Gwynedd, 1 male, Marsh, Berkshire, 1 male, 25.viii.1999, Chandler leg. 13.vii.1976, J.H. Cole leg. (PJCB). Glencoe, Highland (PPCM). Convil Braes, Moray, 1 male, no date, Coe leg. Region, 2 males, 13.vi.1976, Chandler leg. (PPCM). (BMNH). Cors Erddreiniog, Isle of Anglesey, 1 male, Hickling, Norfolk, 1 male, v.1988, Foster & Procter leg. 6.x.1988, Holmes, Boyce & Reed leg. (PPWM). Crow- (PBLN). Lackham Park, Wiltshire, 1 male, 2.vii.2002, borough, East Sussex, 1 male, 13.ix.1916, Jenkinson Chandler leg. (PPCM). Lancaut Nature Reserve, leg. [paratype of Corynoptera globula] (BMNH). Eday, Gloucestershire, 1 male, 28.v.2003, Gibbs leg. (PDGB). Orkney, 1 male, 1.vi.−6.vi.1988, Laurence leg. (PBLN). Leckford Estate, Hampshire, 1 male, 25.vii.1998, Farnborough Park, Warwickshire, 1 male, 10.x.2001, Chandler leg. (PPCM). Rotherfield Park, Hampshire, 1 Chandler leg. (HRI). Glen of Drumloch, Moray, 3 male, 3.vii.2002, Chandler leg. (PPCM). Scadbury males, 16.viii.1937, Coe leg. (BMNH). Goathland, Park, Greater London, 1 male, 18.vi.1983, Chandler North Yorkshire, 2 males, 5.ix.1937, Edwards leg. leg. (PPCM). Sedlescombe, East Sussex, 1 male, [paratypes of Corynoptera globula] (BMNH). Hay 29.vi.2002, Roper leg. (PPRS). Sheep Leas, West Hors- Wood, Baddesley Clinton, Solihull, West Midlands, 4 ley, Surrey, 1 male, 29.vii.1991, Chandler leg. (PPCM). males, 25.viii.2002, Menzel leg. (DEI). Highgate Wood, Sned Wood, Lower Lye, Herefordshire, 5 males, Greater London, 2 males, 1998–99, Hackett leg.

(PPCM). Hole of Horcum, North Yorkshire, 4 males, Country Park, Coventry, Warwickshire, 1 male, 31.viii.1937, Edwards leg. [including holotype and 28.viii.2003, J.E. Smith leg. (HRI). Malham Tarn, paratype of Corynoptera globula] (BMNH). Horsell North Yorkshire, 1 male, 24.vi.1981, Disney leg. Common, Surrey, 2 males, 6.ix.1999, Chandler (PPWM). Sned Wood, Lower Lye, Herefordshire, 1 leg. (PPCM). Kergord, Shetland, 3 males, 26.viii.− male, 1 female, 14.vii.2003, J.E. Smith leg. (HRI). 28.viii.1996, Laurence leg. (PBLN). Killaun Bog, Wytham Woods, Wytham, Oxfordshire, 1 male, Offaly, 2 males, ix.2000, Withers leg. (PPWM). Letch- 21.viii.2002, Menzel leg. (DEI). worth, Hertfordshire, 1 male, vi.1917, Edwards leg. Ecological notes: found in broad-leaved woodland; (BMNH). Loch an Eilein, Highland Region, 1 male, conifer plantation with broad-leaved fringe, bog and 18.viii.1986, Chandler leg. (NMS). Loch Park, Moray, 5 lakeshore; river side; open montane habitats. vi–viii. males, 12.viii.−14.viii.1937, Coe leg. (BMNH). Logie, Moray, 1 male, 28.viii.−26.ix.1909, Jenkinson leg. [paratype of Corynoptera globula] (BMNH). Long CORYNOPTERA HEMIACANTHA MOHRIG & Spinney, Moreton-in-Marsh, Warwickshire, 3 females, MAMAEV, 1992 11.x.2001, J.E. Smith leg. (HRI). Malham Tarn, North Yorkshire, 1 male, 18.viii.−25.viii.1981; 2 males, New to British list. − 11.ix. 14.ix.1981, all Disney leg. (PPWM). Oakley Material examined: Aviemore, Highland Region, 1 Wood, Bishops Tachbrook, Warwickshire, 1 male, male, 15.vii.1982, Disney leg. (PBLN). Calcot, Berk- 14.x.2001, Chandler leg. (HRI). Odiham Common, shire, 1 male, 6.viii.2003, Chandler leg. (PPCM). Char- Hampshire, 1 male, 7.vii.1990, Chandler leg. (PPCM). lbury, Oxfordshire, 1 male, 4.vii.1936, Edwards leg. Pentwd Marshes, Ceredigion, 1 male, 6.x.1987, (BMNH). Hartslock, Oxfordshire, 1 male, 14.v.2000, Holmes, Boyce & Reed leg. (PPWM). Rhôs Cilcennin, Raper leg. (PDGB). Letchworth, Hertfordshire, 1 Ceredigion, 1 male, 24.ix.1986, Holmes, Boyce & Reed male, vi.1917, Edwards leg. (BMNH). Milham Ford leg. (PPWM). Rhôs Rhydd, Ceredigion, 1 male, School, Oxford, Oxfordshire, 1 male, 20.v.−4.vi.2001; 1 11.ix.1986, Holmes, Boyce & Reed leg. (PPWM). Rhôs- male, 4.vi.−8.vi.2001, all Webb leg. (PPCM). Salisbury goch, Powys, 1 male, 10.x.1989, Holmes, Boyce & Reed Plain Training Area, Warminster, Wiltshire, 1 male, leg. (PPWM). Ryton Wood, Bubbenhall, Coventry, 29.v.2003; 1 male, 30.v.2003; 9 males, 31.v.2003; 2 West Midlands, 1 male, 12.x.2001, J.E. Smith leg. males, 4.vii.2003, all Ismay & Schulten leg. (DEI, (HRI). Skokholm, Pembrokeshire, 1 male, 24.viii.1934, UMO). Sned Wood, Lower Lye, Herefordshire, 1 male, Edwards leg. [paratype of Corynoptera globula] 14.vii.2003, J.E. Smith leg. (HRI). (BMNH). The Loke, Norwich, Norfolk, 1 male, 15.ix.1987; 2 males, 18.ix.−20.ix.1988; 1 male, 15.ix.− Ecological notes: found in conifer plantation with 28.ix.1989; 1 male, 2.v.−6.v.1992; 1 male, 26.vi.− broad-leaved fringe; calcareous grassland adjacent to 30.vi.1992; 1 male, 29.ix.−3.x.1992; 3 males, 27.viii.− hanger woodland; unimproved calcareous grassland; 1.ix.1997, all Laurence leg. (PBLN). Thompson Com- school grounds with old poplars; river side; ﬂoodplain mon, Norfolk, 1 male, 13.ix.−27.ix.1988, Foster & meadow. v–viii. Procter leg. (PBLN). Tomintoul, Moray, 1 male, 8.viii.− 11.viii.1937, Coe leg. (BMNH). Whichford Wood, More- ton-in-Marsh, Warwickshire, 1 male, 11.x.2001, J.E. CORYNOPTERA HYPOPYGIALIS (LENGERSDORF, 1926) Smith leg. (HRI). Wolford Wood, Moreton-in-Marsh, Synonyms: pachycerca (Frey, 1948); piniphila (Leng- Warwickshire, 1 male, 11.x.2001, J.E. Smith leg. ersdorf, 1940). (HRI). Woodwalton Fen, Cambridgeshire, 1 male, Literature: Menzel (1998): 21. vi.1939, Edwards leg. (BMNH). Material examined: Bix Bottom, Oxfordshire, 1 male, Ecological notes: very common in broad-leaved, mixed 23.ix.2000, Raper leg. (PDGB). Cadair Idris, and carr woodland; also found in ancient pine woodland Gwynedd, 3 males, 19.vi.1993; 4 males, 3.vii.1993; 8 and conifer plantations; heathland; wetlands including males, 17.vii.1993; 11 males, 1.viii.1993; 5 males, fens, basin mires, bogs, water meadows and wooded 16.viii.1993, all Laurence leg. (PBLN). Cors Erd- lake margins; calcareous grassland; gardens; island dreiniog, Isle of Anglesey, 1 male, 6.x.1988, Holmes, with seabird colonies; open montane habitats. v–xi. Boyce & Reed leg. (PPWM). Hartslock, Oxfordshire, 1 male, 6.v.2000, Raper leg. (PDGB). Llanllwch, Car- CORYNOPTERA GROTHAE MOHRIG & MENZEL, 1990 marthen, 1 male, 11.vii.1989, Holmes, Boyce & Reed leg. (PPWM). Loch Callater, Aberdeenshire, 1 male, Literature: Menzel (1998): 21. 22.vii.1938, Coe leg. (BMNH). Luton Airport, Bed- Material examined: Cadair Idris, Gwynedd, 1 male, fordshire, 1 male, 20.iv.−6.v.1999, C.W. Plant leg. 19.vi.1993, Laurence leg. (PBLN). Coombe Abbey (PPCM). Malham Tarn, North Yorkshire, 1 male, no

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 76 F. MENZEL ET AL. date, Disney leg. (PPWM). Natural History Museum, woodland; bog; garden. Reared from dead wood and South Kensington, Greater London, 1 male, rotten log. iv–x. 13.vi.1924, Edwards leg. (BMNH). Park End, North Yorkshire, 1 male, 22.vi.−25.vi.1981, Disney leg. (PPWM). Waun Isaf, Pembrokeshire, 1 male, CORYNOPTERA IRMGARDIS (LENGERSDORF, 1930) 20.vii.1987, Holmes, Boyce & Reed leg. (PPWM). Synonym: subtilissima (Frey, 1948). Wytham Woods, Wytham, Oxfordshire, 1 male, Literature: Freeman (1983b): 37 [as Bradysia subtilis, 21.viii.2002, Menzel leg. (DEI). misidentification]; Freeman (1987): 199; O’Connor & Ecological notes: found on calcareous grassland; damp Ashe (1991): 29 [as Bradysia subtilis, misidentifica- heathland; wetlands including fens with Cladium and tion]; Laurence (1994): 118 [Corynoptera irmgardis Phragmites beds and bogs; open montane habitats. iv– and in part as Bradysia subtilis, misidentification]; x. Laurence (1997b): 228 [as Bradysia subtilis, misidentification]; Menzel (1998): 21 [as Corynoptera irmgardis and in part as Corynoptera subtilis]; Withers CORYNOPTERA INEXSPECTATA TUOMIKOSKI, 1960 (2002): 114. New to British list. Material examined: Blaen y Wergloedd, Conwy, 1 Material examined: Salisbury Plain Training Area, male, 13.x.1988, Holmes, Boyce & Reed leg. (PPWM). Warminster, Wiltshire, 1 male, 30.v.2003, Ismay & Bohernabreena, Dublin, 1 male, 14.vi.1978, Ashe leg. Schulten leg. (DEI). Sedlescombe, East Sussex, 1 (NMID). Bolt Wood, Stockerston, Leicestershire, 2 male, xi.1996, Roper leg. (PPRS). Thompson Common, males, 4 females, 24.viii.2003, Menzel & J.E. Smith Norfolk, 1 male, 25.v.1986, Withers leg. (PPWM). leg. (DEI, HRI). Braedownie, Angus, 1 male, 24.vii.1937, Coe leg. (BMNH). Brancaster, Norfolk, 3 Ecological notes: found in unimproved calcareous males, 17.vi.−1.vii.1988; 9 males, 1.vii.−15.vii.1988; 7 grassland; grassland with pingo pools; grassland at males, 2.ix.−16.ix.1988; 11 males, 28.x.1988, all Foster woodland fringe. v, xi. & Procter leg. (PBLN); 10 males, 2.ix.−16.ix.1988; 4 males, 28.x.1988, all Laurence leg. (PBLN). Brandon CORYNOPTERA INUNDATA FRITZ, 1982 Marsh, Ryton-on-Dunsmore, West Midlands, 1 female, 12.x.2001, J.E. Smith leg. (HRI); 12 males, 5 females, Synonym: minutula (Bukowski & Lengersdorf, 1936) 22.viii.2002, Menzel & J.E. Smith leg. (DEI, HRI). [preocc.]. Bucklebury Common, Berkshire, 1 male, 30.viii.1993, New to British and Irish list. Chandler leg. (PPCM). Charleville Wood, Offaly, 1 male, 15.vi.1985, Chandler leg. (PPWM). Clowes Material examined: Annahala bog, Gearagh, Cork, 1 Wood, Warwickshire, 1 male, 19.iv.1987, Withers leg. male, 24.ix.1990, de Courcy Williams leg. (NMID). (PPWM). Coed Tycanol, Pembrokeshire, 4 males, Archers Wood, Cambridgeshire, 1 male, 18.iv.1977 29.vi.−7.ix.1999, Levey & Pavett leg. (HRI). Colwyn (reared); 1 male, 22.iv.1987, all J.H. Cole leg. (PJCB). Brook, Powys, 2 males, 18.vii.1989, Holmes, Boyce & Cambridge, Cambridgeshire, 1 male, 29.v.1905, Jen- Reed leg. (PPWM). Coombe Abbey Country Park, Cov- kinson leg. (BMNH); 3 males, 2 females, no date, Kei- entry, Warwickshire, 2 males, 28.viii.2003, Menzel leg. lin leg. (BMNH). Cardington, Bedfordshire, 1 male, (DEI). Cors Erddreiniog, Isle of Anglesey, 2 males, 8.viii.1919, Edwards leg. (BMNH). Charlbury, Oxford- 6.x.1988, Holmes, Boyce & Reed leg. (PPWM). Cors shire, 1 male, 4.vii.1936, Edwards leg. (BMNH). Chig- Farlais, Carmarthen, 2 males, 13.vii.1989; 1 male, well Row Wood, Chigwell Row, Essex, 1 male, 3 29.vii.1989, all Holmes, Boyce & Reed leg. (PPWM). females, 8.x.2002, Ismay & Schulten leg. (DEI). Gear- Cors Geirch, Gwynedd, 4 males, 15.vii.1988; 3 males, agh, Cork, 1 female, 4.iv.1990; 1 male, 9.v.1990; 4 7.x.1988, all Holmes, Boyce & Reed leg. (PPWM). Cors males, 10.vi.1990, all de Courcy Williams leg. (DEI, Goch, Isle of Anglesey, 1 male, 2.vii.1989, Holmes, NMID). Milham Ford School, Oxford, Oxfordshire, 4 Boyce & Reed leg. (PPWM). Cors Graianog, Gwynedd, males, 2 females, 6.vii.2001 (reared); 1 male, 25.iv.− 1 male, 28.vii.1988; 1 male, 13.x.1988, all Holmes, 9.v.2002, all Webb leg. (PPCM). The Coombe, Bucking- Boyce & Reed leg. (PPWM). Cors Gyfelog, Gwynedd, 1 hamshire, 1 male, 17.vi.1997, Chandler leg. (PPCM). male, 19.x.1988, Holmes, Boyce & Reed leg. (PPWM). The Loke, Norwich, Norfolk, 1 male, 8.v.1987, Lau- Cors Penally, Pembrokeshire, 1 male, 22.vii.1987; 1 rence leg. (PBLN). Wemmergill Hall, Durham, 1 male, male, 30.ix.1987, all Holmes, Boyce & Reed leg. 25.vi.1983, Chandler leg. (PPCM). (PPWM). Dyffryn Gwaun, Pembrokeshire, 1 male, Ecological notes: found in broad-leaved woodland; 28.ix.1987, Holmes, Boyce & Reed leg. (PPWM). East school grounds with old poplars; heathland (heather, Trewent Wood, Pembrokeshire, 1 male, 12.vii.1986, Juncus, Deschampsia) with ancient and secondary Chandler leg. (PPCM). Gearagh, Cork, 2 males, 1

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 BRITISH SCIARID FAUNA 77 female, 9.iii.1990; 3 males, 26.iv.1990; 2 males, CORYNOPTERA LEVIS TUOMIKOSKI, 1960 22.v.1990; 1 male, 26.ix.1990, all de Courcy Williams New to British list. leg. (DEI, NMID). Hafod Garregog, Gwynedd, 1 male, 26.vii.1988, Holmes, Boyce & Reed leg. (PPWM). Hay Material examined: Moccas Park, Herefordshire, 1 Wood, Baddesley Clinton, Solihull, West Midlands, 1 male, 11.vi.2002, Godfrey leg. (PPCM). male, 25.viii.2002, Menzel leg. (DEI). Hickling, Nor- Ecological notes: found in a site comprising broad- folk, 4 males, x.1988, Foster & Procter leg. (PBLN). leaved woodland and parkland. vi. How Hill, Norfolk, 1 male, 24.vi.1990, Withers leg. (PPWM). Ickenthwaite, Cumbria, 1 male, 18.vi.1999, Chandler leg. (PPCM). Langstone, Monmouthshire, 1 CORYNOPTERA LUTEOFUSCA male, 21.vii.1988, Holmes, Boyce & Reed leg. (PPWM). (BUKOWSKI & LENGERSDORF, 1936) Lea Meadows, Newtown Linford, Leicestershire, 1 New to British list. male, 24.viii.2003, Menzel leg. (DEI). Leebotwood, Shropshire, 1 female, 14.vii.2003, J.E. Smith leg. Material examined: Becky Woods, Manaton, Devon, 4 (HRI). Llangloffan Fen, Pembrokeshire, 1 male, males, 23.vi.2002, J.E. Smith leg. (HRI). Bolt Wood, 27.vi.1987, Holmes, Boyce & Reed leg. (PPWM). Llyn Stockerston, Leicestershire, 1 male, 24.viii.2003, J.E. Gwynllyn, Powys, 1 male, 9.x.1989, Holmes, Boyce & Smith leg. (HRI). Burnham Beeches, Buckingham- Reed leg. (PPWM). Loch Park, Moray, 1 male, 12.viii.− shire, 1 male, 23.v.−7.vi.1995, Ismay leg. (HRI). Cata- 14.viii.1937, Coe leg. (BMNH). Low Pasture, Hawnby, col, Arran, North Ayrshire, 1 male, 29.v.−30.v.1919, North Yorkshire, 1 male, 12.x.1990, Chandler leg. Edwards leg. (BMNH). Lochanhully, Highland Region, (PPCM). Magor Marsh, Monmouthshire, 1 male, 1 male, 17.vii.1991, Chandler leg. (PPCM). 28.ix.1988, Holmes, Boyce & Reed leg. (PPWM). Mid Ecological notes: found in broad-leaved woodland; Yell, Shetland, 1 male, 25.v.1987, Laurence leg. open glen with stream. v–viii. (PBLN). Narborough Bog, Whetstone, Leicestershire, 2 males, 24.viii.2003, J.E. Smith leg. (HRI). New For- est, Hampshire, 1 male, vii.1905, Sharp leg. (BMNH). CORYNOPTERA MACRICULA MOHRIG & Oxwich, Swansea/Abertawe, 1 male, 4.x.1989, Holmes, KRIVOSHEINA, 1986 Boyce & Reed leg. (PPWM). Perry West Wood, Cam- bridgeshire, 2 males, 17.iv.1975, J.H. Cole leg. (PJCB). New to British list. Pinner, Greater London, 1 male, 14.ix.1912, Edwards Material examined: Buckingham Palace Garden, leg. (BMNH). Ross Island, Killarney, Kerry, 2 males, Greater London, 2 males, 11.iv.−9.v.1996, D.J. Carter 11.vii.1990, Withers leg. (PPWM). Ryton Wood, & C.W. Plant leg. (PPCM). Burnham Beeches, Buck- Bubbenhall, Coventry, West Midlands, 27 males, 7 inghamshire, 4 males, 28.v.−14.vi.1996; 2 males, females, 25.viii.2002, Menzel & J.E. Smith leg. (DEI, 14.vi.−1.vii.1996; 1 male, 1.vii.−8.vii.1996; 2 males, HRI). St. David’s Airﬁeld, Pembrokeshire, 1 male, 8.vii.−22.vii.1996; 12 males, 13.ix.−26.ix.1996; 2 12.x.1987, Holmes, Boyce & Reed leg. (PPWM). Sedle- males, 26.ix.−9.x.1996; 6 males, 9.x.−26.xi.1996; 1 scombe, East Sussex, 1 male, 21.ix.1996; 1 male, male, 14.x.−28.xi.1999, all Ismay leg. (PPCM). 19.v.1999, all Roper leg. (PPRS). Sned Wood, Lower Lye, Herefordshire, 2 males, 1 female, 14.vii.2003, J.E. Ecological notes: found in ancient beech and other Smith leg. (HRI). Waun Isaf, Pembrokeshire, 1 male, broad-leaved woodland; formal garden with wooded 28.ix.1987, Holmes, Boyce & Reed leg. (PPWM). West fringes and lake. iv–vii, ix–xi. Harling Common, Norfolk, 1 male, 17.iv.1987, Withers leg. (PPWM). Wolford Wood, Moreton-in-Marsh, War- wickshire, 1 male, 3 females, 11.x.2001, J.E. Smith CORYNOPTERA MELANOCHAETA MOHRIG & leg. (HRI). Woodwalton Fen, Cambridgeshire, 1 male, MENZEL, 1992 24.v.1940, Edwards leg. (BMNH). Wytham Woods, New to British list. Wytham, Oxfordshire, 3 males, 21.viii.2002, Menzel Material examined: Burnham Beeches, Buckingham- leg. (DEI). Yell, Shetland, 1 male, 3.ix.−10.ix.1990, shire, 1 male, 22.vii.−5.viii.1996, Ismay leg. (PPCM). Laurence leg. (PBLN). Monument Field, Epping Forest, Essex, 1 male, vii.1998, Dagley leg. (PPCM). Thompson Common, Ecological notes: very common in broad-leaved and Norfolk, 2 males, 13.ix.−29.ix.1988, Foster & Procter carr woodland; also found in mixed woodland and leg. (PBLN). conifer plantation; damp and wet heathland; wetlands including water meadows, fens, reed beds, basin mires Ecological notes: found in ancient beech and other and bogs; grazed grassland; former gravel workings; broad-leaved woodland; grassland with pingo pools. vegetation beside reservoir. iii–x. vii–ix.

CORYNOPTERA MEMBRANIGERA (KIEFFER, 1903) leg. (PPCM, PPWM). Hafod Garregog, Gwynedd, 1 − Synonym: trispina Tuomikoski (1960). female, 30.vi. 14.ix.1999, Levey & Pavett leg. (PPCM). Llanon, Swansea/Abertawe, 1 male, 3.x.1989, Holmes, Literature: Menzel (1998): 21; Withers (2002): 114. Boyce & Reed leg. (PPWM). Llyn Tyn y Mynydd, Material examined: Becky Woods, Manaton, Devon, 2 Gwynedd, 1 male, 22.vi.1988; 1 male, 20.viii.1988, all males, 1 female, 23.vi.2002, J.E. Smith leg. (HRI). Holmes, Boyce & Reed leg. (PPWM). Malham Tarn, − Burnham Beeches, Buckinghamshire, 1 male, 22.vii.− North Yorkshire, 1 male, 22.vi. 25.vi.1981, Disney 5.viii.1996, Ismay leg. (PPCM). Canonteign Falls, leg. (PPWM). Rhôs Cilcennin, Ceredigion, 1 male, Devon, 1 male, 23.vi.2002, J.E. Smith leg. (HRI). Coed 24.ix.1986; 1 male, 10.x.1986, all Holmes, Boyce & Tycanol, Pembrokeshire, 1 male, 18.vi.−29.vi.1999, Reed leg. (PPWM). Waun Isaf, Pembrokeshire, 1 male, Levey & Pavett leg. (PPCM). Craigmore Wood, 6.vii.1987, Holmes, Boyce & Reed leg. (PPWM). With- Aberfoyle, Stirling, 1 male, 21.vi.1999, Chandler erslack, Cumbria, 2 males, 20 females, vi.1929, leg. (PPCM). Crowborough, East Sussex, 1 male, Edwards leg. (BMNH). Ynys Eidiol, Ceredigion, 1 14.viii.1906, Jenkinson leg. (BMNH). Gearagh, Cork, female, 19.vi.1987, Holmes, Boyce & Reed leg. 1 male, 9.vii.1990, Withers leg. (PPWM). Loch Park, (PPCM). No locality details, England, 1 female, no Moray, 1 male, 3.vii.1936, Coe leg. (BMNH). Sedle- date, Walker leg. [holotype of Sciara brevipennis] scombe, East Sussex, 1 male, 1.vii.1996, Roper leg. (BMNH). (PPRS). Selsdon, Greater London, 1 male, 5.vii.1934, Ecological notes: found in broad-leaved woodland; val- Coe leg. [paratype of Corynoptera curvispinosa] ley fen and wet heathland; damp heath; Molinia bog; (BMNH). Sned Wood, Lower Lye, Herefordshire, 2 Phragmites bed; ungrazed basin mire; ungrazed males, 1 female, 14.vii.2003, J.E. Smith leg. (HRI). heather; lakeshore; open montane habitats. v–x. Ecological notes: found in beech and other broad- leaved woodland; conifer plantation with broad-leaved CORYNOPTERA OBSCURIPILA TUOMIKOSKI, 1960 fringe; grassland at woodland fringe; river side. vi– New to British list. viii. Material examined: Lakenheath, Suffolk, 1 male, 30.viii.−13.ix.1988, Foster & Procter leg. (PBLN). CORYNOPTERA MINIMA (MEIGEN, 1818) Ecological notes: found in damp meadow. viii–ix. Synonyms: brachyptera (Lengersdorf, 1941); brevipennis (Walker, 1848). CORYNOPTERA PARVULA (WINNERTZ, 1867) Literature: Walker (1848): ? 108 and 110; Walker Synonym: uncinata (Hippa & Vilkamaa, 1994). (1856): ? 54 and 56 [all as ? Sciara minima and Sciara brevipennis]; Edwards (1925a): 543 [as Peyerimhofﬁa Literature: Hippa & Vilkamaa (1994): 75 [as Camp- (?) brevipennis]; Freeman (1983b): 29; Laurence & tochaeta uncinata]; Menzel (1998): 21; Menzel & James (1997): 166 [all as Corynoptera brevipennis]; Mohrig (2000): 242; Chandler (2001): 201; Withers Menzel (1998): 21. (2002): 114. Material examined: Ben Chalum, Stirling, 1 female, Material examined: Berwick-upon-Tweed, Northum- 9.vi.−10.vi.1932, Edwards leg. (BMNH). Cadair Idris, berland, 1 male, 24.vi.−27.vi.1996, Laurence leg. Gwynedd, 1 male, 1 female, 3.vii.1963; 1 male, 9 (PBLN). Bix Bottom, Oxfordshire, 1 male, 23.ix.2000, females, 19.vi.1993; 2 males, 2 females, 3.vii.1993; 1 Raper leg. (PDGB). Buckingham Palace Garden, male, 17.vii.1993; 1 female, 16.viii.1993; 5 males, Greater London, 1 male, 18.vii.−13.viii.1997, D.J. 3.vii.1997, all Laurence leg. (PBLN). Cashlie, Glen Carter & C.W. Plant leg. (PPWM). Castlefreke, Cork, 1 Lyon, Perth & Kinross, 5 males, 2 females, 15.vi.1932, male, 12.v.1989, Withers leg. (PPWM). Epping Forest, Edwards leg. (BMNH). Cleddon Bog, Monmouthshire, Essex, 12 males, v.-viii.1999, Dagley leg. (PPCM). 6 females, 24.v.1988; 1 male, 10.vi.1988, all Holmes, Letchworth, Hertfordshire, 1 male, 26.vi.1926, Boyce & Reed leg. (PPWM). Glen Lochay, Stirling, 1 Edwards leg. (BMNH); 1 male, 2.vii.1934, Edwards male, 3.vi.−18.vi.1932, Edwards leg. (BMNH). Glen leg. [paratype of Camptochaeta uncinata] (BMNH). Shiel, Highland Region, 1 female, 1.vii.1934, Richards Salisbury Plain Training Area, Warminster, Wiltshire, leg. (BMNH). Coed Tycanol, Pembrokeshire, 1 female, 1 male, 27.v.2003; 1 male, 3.vii.2003, all Ismay & 27.vi.−7.ix.1999, Levey & Pavett leg. (PPCM). Cors Schulten leg. (DEI, UMO). The Loke, Norwich, Nor- Barfog, Gwynedd, 1 male, 13.vi.1988; 1 male, 1 female, folk, 1 male, 15.v.−18.v.1992, Laurence leg. (PBLN). 25.vii.1988, all Holmes, Boyce & Reed leg. (PPCM, Woodwalton Fen, Cambridgeshire, 1 male, 24.v.1940, PPWM). Cors Caranod, Ceredigion, 3 males, 27.viii.− Edwards leg. [paratype of Camptochaeta uncinata] 2.ix.1985; 2 males, 1.x.1986, all Holmes, Boyce & Reed (BMNH).

Ecological notes: found in broad-leaved and carr wood- Ecological notes: found in broad-leaved and carr woodland; fen; unimproved calcareous grassland; gardens. land; calcareous grassland adjacent to hanger wood- v–ix. land; damp heathland; wetlands including water meadows, sedge bed and bog; former gravel workings; grassland at woodland fringe; grounds of horticultural CORYNOPTERA PERPUSILLA WINNERTZ, 1867 research station; school grounds with old poplars; garden. iii–x. Synonyms: bicornis (Lengersdorf, 1943); dubitata Tuomikoski, 1960. Literature: Freeman (1983b): 29; Menzel (1998): 21. CORYNOPTERA POSTFORCIPATA RUDZINSKI, 1993 New to British list. New to Irish list. Material examined: Sontley Marsh, Wrexham, 1 male, Material examined: Annahala bog, Gearagh, Cork, 1 3.viii.1988, Holmes, Boyce & Reed leg. (PPWM). male, 25.viii.1990, de Courcy Williams leg. (NMID). Berwick-upon-Tweed, Northumberland, 2 males, Ecological notes: found on fen. viii. v.1999; 1 male, vii.1999, all Laurence leg. (PBLN). Brandon Marsh, Ryton-on-Dunsmore, West Mid- lands, 2 males, 22.viii.2002, Menzel leg. (DEI). Char- CORYNOPTERA POSTGLOBIFORMIS MOHRIG, 1993 lbury, Oxfordshire. 1 male, 4.vii.1936, Edwards New to British list. leg. (BMNH). Colt Park Wood, Marrick, North Material examined: Balmoral Forest, Aberdeenshire, Yorkshire, 1 male, 27.viii.−30.viii.1981, Disney leg. 1 male, 1.viii.−4.viii.1937, Coe leg. (BMNH). Bradﬁeld (PPWM). Crymlyn Bog, Swansea/Abertawe 1 male, Wood, Suffolk, 2 males, 9.x.−16.x.1990, Laurence leg. 13.vii.1988; 1 male, 5.x.1989, all Holmes, Boyce & (PBLN). Burnham Beeches, Buckinghamshire, 1 Reed leg. (PPWM). Cwm Crafnant, Gwynedd, 1 male, 5.viii.−31.viii.1996, Ismay leg. (PPCM). Cadair male, 20.vii.1988; 1 male, 4.viii.1988, all Holmes, Idris, Gwynedd, 1 male, 16.viii.1993, Laurence leg. Boyce & Reed leg. (PPWM). Ditchley, Oxfordshire, 1 (PBLN). Coombe Abbey Country Park, Coventry, War- male, v.1937, Edwards leg. (BMNH). Glasshouse wickshire, 1 female, 28.viii.2003, Menzel leg. (DEI). Crops Research Station, Littlehampton, West Cors Erddreiniog, Isle of Anglesey, 1 male, 6.x.1988, Sussex, 1 male, 17.ix.1956, Hussey leg. (HRI). Holmes, Boyce & Reed leg. (PPWM). Epping Forest, Hartslock, Oxfordshire, 1 male, 6.v.2000, Raper leg. Essex, 1 male, 17.viii.1998, Chandler leg. (PPCM). (PDGB). Hickling, Norfolk, 2 males, v.1988, Foster & Hay Wood, Baddesley Clinton, Solihull, West Mid- Procter leg. (PBLN). Lakenheath, Suffolk, 1 male, lands, 2 males, 25.viii.2002, Menzel leg. (DEI). Lea 30.viii.−13.ix.1988, Foster & Procter leg. (PBLN). Meadows, Newtown Linford, Leicestershire, 1 male, Lea Meadows, Newtown Linford, Leicestershire, 1 24.viii.2003, Menzel leg. (DEI). Loch Park, Moray, 1 male, 24.viii.2003, J.E. Smith leg. (HRI). Leebot- male, 12.viii.−14.viii.1937, Coe leg. (BMNH). Lock- wood, Shropshire, 1 male, 14.vii.2003, J.E. Smith haugh Bank Wood, Derwent Valley, Tyne & Wear, 4 leg. (HRI). Letchworth, Hertfordshire, 1 male, males, 24.vii.−13.viii.1990, Jessop leg. (PBLN). vii.1918; 1 male, 25.vi.1923, all Edwards leg. Moorthwaite, Cumbria, 1 male, 21.viii.1988, no collec- (BMNH). Milham Ford School, Oxford, Oxfordshire, tor detail (PPWM). Rothamsted, Harpenden, Hert- 1 male, 3.v.−5.vi.2002, Webb leg. (PPCM). Parham, fordshire, 1 male, 4.viii.1950, Laurence leg. (PBLN). West Sussex, 1 male, 13.xi.1957, Speyer leg. (HRI). Ryton Wood, Bubbenhall, Coventry, West Midlands, 1 Sedlescombe, East Sussex, 1 male, 6.iv.1999, Roper male, 25.viii.2002, Menzel leg. (DEI). St. Kilda, West- leg. (PPRS). Stanford, Norfolk, 1 male, 27.iii.− ern Isles, 1 male, 6.viii.1931, Lack leg. (BMNH). The 10.iv.1988, Withers leg. (PPWM). The Loke, Nor- Loke, Norwich, Norfolk, 1 male, 27.viii.−1.ix.1993; 1 wich, Norfolk, 1 male, 4.vii.1987; 1 male, 15.vi.− male, 28.viii.−1.ix.1993, all Laurence leg. (PBLN). 19.vi.1988; 1 male, 7.ix.1989, all Laurence leg. Tocil Wood, Warwick University Campus, Coventry, (PBLN). The Oaks, Cambridgeshire, 1 male, ix.1998, Warwickshire, 3 males, 2 females, 28.viii.2003, Menzel C.W. Plant leg. (PPCM). Tocil Wood, Warwick Uni- & J.E. Smith leg. (DEI, HRI). Windsor Forest, Berk- versity Campus, Coventry, Warwickshire, 1 male, shire, 1 male, 5.viii.1995, Brock leg. (PPCM). No local- 28.viii.2003, J.E. Smith leg. (HRI). Wellesbourne, ity details, England, 1 male, 16.viii.1950, Freeman leg. Warwickshire, 1 male, 17.viii.−31.viii.2000; 1 male, (PBLN). 24.v.−27.vi.2001, all J.E. Smith leg. (HRI). Willes Meadow, Leamington Spa, Warwickshire, 1 male, Ecological notes: found in broad-leaved and mixed 27.viii.2002, Menzel leg. (DEI). Wolford Wood, woodland; conifer plantations; wet meadow with Jun- Moreton-in-Marsh, Warwickshire, 1 male, 11.x.2001, cus; grassland; grounds of horticultural research sta- J.E. Smith leg. (HRI). tion; garden; open montane habitats. vii–x.

CORYNOPTERA PRAEFORCIPATA MOHRIG & 5.viii.1989, Withers leg. (PPWM). Dinton Pastures MAMAEV, 1987 Country Park, Berkshire, 1 male, 30.vii.1994; 1 male, New to British list. 9.vi.1996, all Chandler leg. (PPCM). Epping Forest, Essex, 1 male, 18.viii.1998, Chandler leg. (PPCM); 3 Material examined: Letchworth, Hertfordshire, 2 males, vi.-viii.1999, Dagley leg. (PPCM). Ffrwd Fen, males, vi.1917, Edwards leg. (BMNH). Woodwalton Carmarthen, 1 male, 3.x.1989, Holmes, Boyce & Fen, Cambridgeshire, 1 male, 24.v.1940, Edwards leg. Reed leg. (PPWM); 1 male, 13.x.1995, Chandler leg. (BMNH). (PPCM). Gearagh, Cork, 1 male, 11.v.1990, de Courcy Ecological notes: one locality comprises fen and carr. Williams leg. (NMID). Kingsbury Water Park, War- v–vi. wickshire, 1 male, 5 females, 13.x.2001, J.E. Smith leg. (HRI). Knebworth, Hertfordshire, 1 male, viii.1936, Edwards leg. [paratype of Corynoptera recurvispina] (BMNH). Lackham Park, Wiltshire, 1 male, CORYNOPTERA PRAEPARVULA MOHRIG & 20.ix.2002, Chandler leg. (PPCM). Lea Meadows, KRIVOSHEINA, 1983 Newtown Linford, Leicestershire, 19 males, 31 New to British list. females, 24.viii.2003, Menzel & J.E. Smith leg. (DEI, Material examined: Letchworth, Hertfordshire, 1 HRI). Llyn Ystumllyn, Gwynedd, 2 males, 5.x.1988, male, v.1918, Edwards leg. (BMNH). Holmes, Boyce & Reed leg. (PPWM). Lopham Fen, Norfolk, 1 male, 20.v.1990, Withers leg. (PPWM). Mal- Ecological notes: possibly found in house or garden. v. ham Tarn, North Yorkshire, 3 males, 24.vi.1982, Disney leg. (PPWM). Narborough Bog, Whetstone, Leicestershire, 3 males, 12 females, 24.viii.2003, Men- CORYNOPTERA RECURVISPINA FREEMAN, 1987 zel & J.E. Smith leg. (DEI, HRI). Pant-y-Sais, Neath/ Literature: Freeman (1983a): 164 [in part as Port Talbot, 1 male, 5.x.1989, Holmes, Boyce & Reed Corynoptera coei, misidentification]; Freeman (1987): leg. (PPWM). Ryton Wood, Bubbenhall, Coventry, 200; Laurence (1994): 118; Laurence (1997a): 85; West Midlands, 19 males, 4 females, 25.viii.2002, Menzel (1998): 21 [as Corynoptera recurvispina and Menzel & J.E. Smith leg. (DEI, HRI). Sedlescombe, Corynoptera dentiforceps]; Menzel & Mohrig (2000): East Sussex, 1 male, 17.v.1997, Roper leg. (PPRS). The 196, 240 and 243 [as Corynoptera recurvispina, in part Loke, Norwich, Norfolk, 1 male, 29.ix.−3.x.1992, Lau- as Corynoptera dentiforceps and under Camptochaeta rence leg. (PBLN). Tocil Wood, Warwick University coei]. Campus, Coventry, Warwickshire, 1 male, 1 female, 28.viii.2003, Menzel leg. (DEI). Tuddenham Fen, Suf- New to Irish list. folk, 2 males, 20.x.1990, Chandler leg. (PPCM, Material examined: Alvecote Pools, Staffordshire, 1 PPWM). Wayland Wood, Norfolk, 1 male, 13.x.2002, male, 3 females, 13.x.2001, J.E. Smith leg. (HRI). Chandler leg. (PPCM). Wemmergill Hall, Durham, 1 Becky Woods, Manaton, Devon, 1 male, 23.vi.2002, male, 25.vi.1981, Chandler leg. (PPCM). West End J.E. Smith leg. (HRI). Bradfield Wood, Suffolk, 1 male, Common, Esher, Surrey, 1 male, 17.vii.2002, Chandler 6.vii.−11.vii.1990; 1 male, 23.v.−30.v.1991; 1 male, leg. (PPCM). West Harling Common, Norfolk, 2 males, 16.viii.−22.viii.1991, all Laurence leg. (PBLN). 20.x.1990, Withers leg. (PPWM). Whichford Wood, Brampton, Cambridgeshire, 1 male, 23.v.1976, J.H. Moreton-in-Marsh, Warwickshire, 1 male, 11.x.2001, Cole leg. (PJCB). Brancaster, Norfolk, 1 male, 17.vi.− J.E. Smith leg. (HRI). Whitacre Heath, Warwickshire, 11.vii.1988; 1 male, 2.ix.−16.ix.1988; 1 male, 16.ix.− 2 males, 4 females, 13.x.2001, J.E. Smith leg. (HRI). 8.x.1988, all Foster & Procter leg. (PBLN). Brandon Whitnash brook, Sydenham, Leamington Spa, War- Marsh, Ryton-on-Dunsmore, Coventry, West Mid- wickshire, 1 female, 4.ix.2003, J.E. Smith leg. (HRI). lands, 1 male, 11.x.2001; 15 females, 12.x.2001, all Willes Meadow, Leamington Spa, Warwickshire, 19 J.E. Smith leg. (HRI); 1 male, 3 females, 25.viii.2002, males, 6 females, 27.viii.2002, Menzel & J.E. Smith Menzel leg. (DEI). Burnham Beeches, Buckingham- leg. (DEI, HRI). Woodwalton Fen, Cambridgeshire, 12 shire, 1 male, 14.vi.−1.vii.1996; 1 male, 2.vii.− males, 24.v.1940; 4 males, 7.vi.1940, all Edwards leg. 5.viii.1996, all Ismay leg. (PPCM). Catfield, Norfolk, 1 [holotype and paratypes of Corynoptera recurvispina] male, 27.ix.−18.x.1988, Foster & Procter leg. (PBLN). (BMNH). No locality details, England, 1 male, no date, Coombe Abbey Country Park, Coventry, Warwick- Laurence leg. (PBLN). shire, 2 males, 2 females, 28.viii.2003, Menzel & J.E. Smith leg. (DEI, HRI). Cothill, Oxfordshire, 2 males, Ecological notes: very common in broad-leaved and 26.v.1936, Edwards leg. [paratypes of Corynoptera coniferous woodland; common in carr woodland; wet recuvispina; 1 male also paratype of Corynoptera coei] heathland; wetlands including water meadows, fens, (BMNH). Dickleburgh, Norfolk, 2 males, 28.vii.− reed and sedge beds, bogs, river banks and lake

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 BRITISH SCIARID FAUNA 81 shores; grassland at woodland fringe; old sand and haugh Bank Wood, Derwent Valley, Tyne & Wear, 1 gravel workings; parkland; old hedges; gardens. v–x. male, 24.vii.−13.viii.1990, Jessop leg. (PBLN). Milham Ford School, Oxford, Oxfordshire, 1 male, 29.v.− Taxonomic note: The type series of Corynoptera recur- 5.vi.2002, Webb leg. (PPCM). Monkfield Wood, Cam- vispina Freeman comprises 19 males [holotype and 18 bridgeshire, 1 male, viii.1998, C.W. Plant leg. (PPCM). paratypes] (Freeman, 1987). The total type material Narborough Bog, Whetstone, Leicestershire, 1 male, examined was [9 males complete on slides; 10 males 24.viii.2003, Menzel leg. (DEI). Sedlescombe, East pinned – of which 1 male also had a genitalia prepa- Sussex, 1 male, 22.viii.1994, Roper leg. (PBLN). ration (no. 121) and 3 males with separately mounted The Clappers, Sharpenhoe, Bedfordshire, 1 male, genitalia]. A check on the data shows that only 11 16.vi.1997, Chandler leg. (PPCM). The Loke, Norwich, paratypes with the label ‘Hunts., Wood Walton, Norfolk, 1 male, 15.v.−18.v.1992; 1 male, 18.v.− 24.v.1940, B.M.140–147’ and 2 males with ‘Berks., 23.v.1992, all Laurence leg. (PBLN). The Oaks, Cam- Cothill, 26.v.1936, B.M.136–336’ are available. This bridgeshire, 1 male, ix.1998, C.W. Plant leg. (PPCM). incorrect recording of the type data by Freeman is Tocil Wood, Warwick University Campus, Coventry, herein corrected. Warwickshire, 2 males, 28.viii.2003, Menzel leg. All records of Corynoptera dentiforceps (Bukowski & (DEI). Wellesbourne, Warwickshire, 1 male, 17.viii.− Lengersdorf, 1936) from Great Britain to date were 31.viii.2000, J.E. Smith leg. (HRI). West End Common, incorrectly identified [recte Corynoptera recurvispina Esher, Surrey, 1 male, 17.viii.1982, Chandler leg. Freeman, 1987]. The misidentifications are based on (PPCM). Woodwalton Fen, Cambridgeshire, 1 male, Menzel & Mohrig (2000) [in part], on small differences 24.v.1940, Edwards leg. (BMNH). Wytham Woods, in the base of the genitalia as well as the shape of Wytham, Oxfordshire, 1 male, 21.viii.2002, Menzel spines on the inner side of the gonostyles. There is, leg. (DEI). however, variation within species and a large part of the error is attributed to the preparation of the Ecological notes: common in broad-leaved and carr specimens (deformed and/or angled genitalia). After woodland; calcareous grassland adjacent to hanger examination of about 350 specimens from the entire woodland; grassland at woodland fringe; wet meadow Palaearctic region (including the types of Corynoptera with reed bed on peat; fen; sedge bed; school grounds dentiforceps and Corynoptera recurvispina), it was with old poplars; garden. iv–x. established that Corynoptera dentiforceps was, to date, only known from the Ukraine (holotype from the Crimea), Greece, Portugal and Spain. A redescription CORYNOPTERA SAETISTYLA MOHRIG & with figures of Corynoptera dentiforceps with a KRIVOSHEINA, 1985 detailed comparison of species will be produced in Synonym: densiseta Mohrig & Menzel, 1990. another publication. Literature: Laurence (1994): 108 [as Corynoptera densiseta]; Menzel (1998): 21. CORYNOPTERA SACCATA TUOMIKOSKI, 1960 New to Irish list. Synonym: arboris Fritz, 1982. Material examined: Brandon Marsh, Ryton-on-Dun- Literature: Laurence (1994): 118; Laurence (1997a): 85 smore, West Midlands, 1 male, 22.viii.2002, J.E. Smith [all as Corynoptera curvispinosa, misidentification]. leg. (HRI). Cors Erddreiniog, Isle of Anglesey, 3 males, New to British list. 6.x.1988, Holmes, Boyce & Reed leg. (PPWM). Hick- ling, Norfolk, 13 males, v.1989, Foster & Procter leg. Material examined: Berwick-upon-Tweed, Northum- (DEI, PBLN). Killaun Bog, Offaly, 3 males, ix.2000, berland, 1 male, 24.vi.−27.vi.1996, Laurence leg. Withers leg. (PPWM). Lea Meadows, Newtown Lin- (PBLN). Bradfield Wood, Suffolk, 1 male, 11.vii.− ford, Leicestershire, 3 males, 24.viii.2003, Menzel leg. 16.vii.1990; 1 male, 1 female, 23.v.−30.v.1991; 2 males, (DEI). Malham Tarn, North Yorkshire, 3 males, 2.vi.− 16.viii.1991; 1 female, 16.viii.−22.viii.1991, all Lau- 3.vi.1982; 1 male, 21.vi.−24.vi.1982, all Disney leg. rence leg. (PBLN). Brandyquoi, Kirkwall, Orkney, 3 (PBLN, PPWM). Rotherfield Park, Hampshire, 1 male, males, 28.v.−8.vi.2001; 6 males, v.-vi.2001, all Lau- 2.x.2002, Chandler leg. (PPCM). Spartum Fen, rence leg. (PBLN). Burnham Beeches, Buckingham- Oxfordshire, 1 male, 15.x.1999, Chandler leg. (PPCM). shire, 1 male, 5.viii.−31.viii.1996, Ismay leg. (PPCM). Wytham Woods, Wytham, Oxfordshire, 2 males, Charlbury, Oxfordshire, 2 males, 4.vi.1936, Edwards 21.viii.2002, Menzel leg. (DEI). leg. (BMNH). Epping Forest, Essex, 9 males, iv.- x.1999, Dagley leg. (PPCM). Hartslock, Oxfordshire, 1 Ecological notes: found in broad-leaved, mixed and male, 14.v.2000, Raper leg. (PDGB). Hickling, Norfolk, carr woodland; wetlands including water meadows, 1 male, v.1988, Foster & Procter leg. (PBLN). Lock- fens with Cladium and Phragmites bed; reed and

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 82 F. MENZEL ET AL. sedge beds, bogs and lakeshore; former gravel work- Ecological notes: found in broad-leaved woodland; ings. v, viii–x. ancient woodland with oak, hazel and bramble; calcareous grassland adjacent to hanger woodland; school grounds with old poplars; garden. v–vii. CORYNOPTERA SEDULA MOHRIG & KRIVOSHEINA, 1985 Synonym: consumpta (Freeman, 1987). CORYNOPTERA SETOSA FREEMAN, 1983 Literature: Freeman (1987): 202 [as Bradysia con- Literature: Freeman (1983a): 165; Freeman (1983b): sumpta]; Menzel (1998): 21; Chandler (2001): 201. 29; Menzel (1998): 21. Material examined: Bangor, Gwynedd, 3 males, 1985, Material examined: Aylesbeare, Devon, 1 male, Calvert leg. [paratypes of Bradysia consumpta] 25.v.2000, C.W. Plant leg. (PPCM). Clovelly, Devon, 2 (BMNH). Buckingham Palace Garden, Greater Lon- males, 8.v.1936, Edwards leg. [holotype and paratype don, 5 males, 27.iii.−27.xi.1999, D.J. Carter & C.W. of Corynoptera setosa] (BMNH). Cors Caron, Ceredi- Plant leg. (PPCM). Burnham Beeches, Buckingham- gion, 1 male, 27.vii.1987, Holmes, Boyce & Reed leg. shire, 4 males, 14.vi.−9.x.1996, Ismay leg. (PPCM). (PPWM). Cors Goch, Isle of Anglesey, 1 male, Canonteign Falls, Devon, 1 male, 22.vi.2001, J.E. 18.vii.1989, Holmes, Boyce & Reed leg. (PPWM). Cwm Smith leg. (HRI). Cors Penally, Pembrokeshire, 1 Ystwyth, Ceredigion, 1 male, 16.vi.1987, Holmes, male, 30.ix.1987, Holmes, Boyce & Reed leg. (PPWM). Boyce & Reed leg. (PPWM). Savernake Forest, Wilt- Epping Forest, Essex, 5 males, 21.iii.−26.ix.1998, shire, 1 male, 31.v.1990, Porter leg. (PPCM). Dagley leg. (PPCM); 1 male, 28.vi.2000, Chandler leg. (PPCM). Harpenden, Hertfordshire, 5 males, Ecological notes: found in ancient beech and mixed 14.viii.1950, Laurence leg. [holotype and paratypes of woodland; grazed river fen; Cladium bed; watershed Bradysia consumpta] (BMNH). Llangloffan Fen, Pem- mire; raised bog. v–vii. brokeshire, 1 male, 23.vi.1987, Holmes, Boyce & Reed leg. (PPWM). Luton Airport, Bedfordshire, 5 males, 20.iv.−6.v.1999, C.W. Plant leg. (HRI, PPCM). Malham CORYNOPTERA SPHENOPTERA TUOMIKOSKI, 1960 Tarn, North Yorkshire, 2 males, 2.vi.−3.vi.1982, Dis- Literature: Menzel (1998): 21. ney leg. (PBLN). Rothamsted, Harpenden, Hertford- Material examined: Newbold Comyn Park, Leaming- shire, 1 male, 16.viii.1950, Laurence leg. (PBLN). ton Spa, Warwickshire, 1 male, 15.vii.2001, J.E. Smith Ryton Wood, Bubbenhall, Coventry, West Midlands, 2 leg. (HRI). males, 25.viii.2002, Menzel & J.E. Smith leg. (DEI, HRI). St. David’s Airﬁeld, Pembrokeshire, 2 males, Ecological notes: found in a site comprising young 29.ix.1987, Holmes, Boyce & Reed leg. (PPWM). Sal- mixed woodland and damp meadow. vii. isbury Plain Training Area, Warminster, Wiltshire, 2 males, 31.v.2003, Ismay & Schulten leg. (DEI, UMO). Stanford, Norfolk, 1 male, 27.iii.−10.iv.1988, Collier CORYNOPTERA SPINIFERA TUOMIKOSKI, 1960 leg. (PPWM). New to British list. Ecological notes: found in ancient broad-leaved wood- Material examined: Coed Tycanol, Pembrokeshire, 1 land; wet heath with Typha; grazed fen; unimproved male, 29.viii.−7.ix.1999, Levey & Pavett leg. (PPCM). calcareous grassland; grounds of horticultural Ecological notes: found in broad-leaved woodland. research station; formal garden with wooded fringes viii–ix. and lake. iii–xi.

CORYNOPTERA SPOECKERI (LENGERSDORF, 1930) CORYNOPTERA SEMISACCATA MOHRIG & MAMAEV, 1987 Synonym: venerata Rudzinski, 1994. New to British list. Literature: Menzel & Heller (2005): 352 [venerata Rudzinski as synonym to spoeckeri (Lengersdorf)]. Material examined: Badger Wood, Cambridgeshire, 1 male, 1.v.−31.v.1998; 1 male, 1.x.1998, all C.W. New to British list. Plant leg. (PPCM). Hartslock, Oxfordshire, 1 male, Material examined: Chigwell Row Wood, Chigwell 14.v.2000, Raper leg. (PDGB). Milham Ford School, Row, Essex, 1 male, 23.v.−27.v.2002, Ismay & Schulten Oxford, Oxfordshire, 1 male, 11.vi.−18.vi.2002, Webb leg. (DEI). leg. (PPCM). The Loke, Norwich, Norfolk, 1 male, 5.vii.1987; 1 male, 23.v.−25.v.1993; 1 male, 27.viii.− Ecological notes: found in ancient woodland with oak, 1.ix.1993, all Laurence leg. (PBLN). hazel and bramble. v.

Taxonomic note: The identity of Corynoptera spoeckeri isbury Plain Training Area, Warminster, Wiltshire, 1 and Corynoptera venerata was only recently brought male, v.2003; 2 males, 27.v.2003; 2 males, 29.v.2003; 1 to light [comparison of types in Menzel & Heller male, 30.v.2003; 4 males, 31.v.2003, all Ismay & (2005)]. Until now, it was accepted that Corynoptera Schulten leg. (DEI, UMO). The Loke, Norwich, Nor- spoeckeri was only known from caves and mines [areas folk, 1 male, 10.viii.1987; 1 male, 19.vii.1988; 4 males, around entrance to deeper regions]. To date, surface 15.viii.−16.viii.1988; 4 males, 16.viii.1988; 3 males, ﬁnds only exist from England and Germany [Bavaria: 21.viii.−24.viii.1989; 1 male, 18.v.−23.v.1992; 12 males, beech–spruce mixed woodland in Schöngeising]. 27.vii.1992; 6 males, 21.viii.1992; 1 male, 29.ix.− 3.x.1992; 2 males, 22.viii.−1.ix.1993; 3 males, 27.v.1997, all Laurence leg. (PBLN). Theale, Berk- CORYNOPTERA SUBDENTATA MOHRIG, 1985 shire, 2 males, 6.viii.2003, Chandler leg. (PPCM). New to British list. Welwyn, Hertfordshire, 1 male, 21.vii.−24.vii.1934, Edwards leg. (BMNH). Willes Meadow, Leamington Material examined: Lackham Park, Wiltshire, 1 male, Spa, Warwickshire, 1 male, 27.viii.2002, Menzel leg. 20.ix.2002; 1 male, 24.x.2002, all Chandler leg. (DEI). (PPCM). Ecological notes: found in broad-leaved and carr wood- Ecological notes: found in broad-leaved woodland. ix– land; water meadows; by river; unimproved calcareous x. grassland; near shore; garden. v, vii–x.

CORYNOPTERA SUBFURCIFERA MOHRIG & CORYNOPTERA SUBTILIS (LENGERSDORF, 1929) HÖVEMEYER, 1992 Synonyms: longicornis (Bukowski & Lengersdorf, Synonyms: differa Rudzinski, 1994. 1936); signhildae (Frey, 1948). New to British and Irish list. Literature: Menzel (1998): 21. Material examined: Bradfield Wood, Suffolk, 1 male, New to British list. 6.vii.−11.vii.1990, Laurence leg. (PBLN). Ennis, Clare, 1 male, 30.v.1984, J.P. O’Connor leg. (NMID). Falls of Material examined: Aviemore, Highland Region, 1 Tarnash, Moray, 1 male, 1.vii.1936, Coe leg. (BMNH). male, 9.vii.1982, Disney leg. (PBLN). Brancaster, Nor- Sned Wood, Lower Lye, Herefordshire, 1 male, folk, 4 males, 2.ix.−16.ix.1988; 4 males, 25.x.1988; 2 14.vii.2003, J.E. Smith leg. (HRI). The Clappers, males, 28.x.1988, all Foster & Procter leg. (PBLN). Sharpenhoe, Bedfordshire, 1 male, 16.vi.1997, Chan- Cadair Idris, Gwynedd, 1 male, 19.vi.1993; 3 males, dler leg. (PPCM). 3.vii.1993; 11 males, 4.ix.1993, all Laurence leg. (PBLN). Clowes Wood, Warwickshire, 1 male, Ecological notes: found in beech and other broad- 19.iv.1987, Withers leg. (PPWM). Killaun Bog, Offaly, leaved woodland; conifer plantation with broad-leaved 1 male, ix.2000, Withers leg. (PPWM). Letchworth, fringe; river side. v–vii. Hertfordshire, 1 male, viii.1917, Edwards leg. (BMNH). Malham Tarn, North Yorkshire, 1 male, 8.v.− CORYNOPTERA SUBPARVULA TUOMIKOSKI, 1960 11.v.1981, Disney leg. (PPWM). Literature: Freeman (1983b): 29 [as Corynoptera par- Ecological notes: found in broad-leaved woodland; vula, misidentification]; Hippa & Vilkamaa (1994): 77 sedge bed; bog and lakeshore; raised bog; open mon- [as Camptochaeta]; Laurence (1994): 118; Laurence tane habitats. iv–x. (1997b): 228 [all as Corynoptera parvula, misidentifi- Taxonomic note: earlier references in the British liter- cation]; Menzel (1998): 21. ature [as Bradysia or Corynoptera subtilis in Freeman Material examined: Bix Bottom, Oxfordshire, 1 male, (1983b: 77), O’Connor & Ashe (1991: 29), Laurence 6.vii.2002, Chandler leg. (PPCM); 2 males, 23.ix.2000, (1994: 118) and Menzel (1998: 21)] referred to Raper leg. (PDGB). Castlemartin Corse, Pem- Corynoptera irmgardis so Corynoptera subtilis is cor- brokeshire, 5 males, 14.x.1987, Holmes, Boyce & Reed rectly recorded here as new to the British list. leg. (PPWM). Epping Forest, Essex, 2 males, ix.2000, Dagley leg. (PPCM). Lackham Park, Wiltshire, 1 male, 5.viii.2002, Chandler leg. (PPCM). Lakenheath, Suf- CORYNOPTERA TETRACHAETA TUOMIKOSKI, 1960 folk, 4 males, 30.viii.−13.ix.1988, Foster & Procter Literature: Menzel (1998): 21 [as Corynoptera inexpec- leg. (PBLN). Hemingford, Cambridgeshire, 2 males, tata (recte inexspectata), misidentification based on ix.1938, Edwards leg. (BMNH). North Ronaldsay, unpublished material by Laurence]; Leigh & Ware Orkney, 1 male, 3.ix.1989, Laurence leg. (PBLN). Sal- (2003): [as Bradysia albanensis, misidentification].

New to British list. female, 28.vi.2002; 1 male, 23.ix.2002, all Ismay & Schulten leg. (DEI). Coed Tycanol, Pembrokeshire, 12 Material examined: Brandon Marsh, Ryton-on- males, 10.vi.−7.ix.1999, Levey & Pavett leg. (HRI, Dunsmore, West Midlands, 2 males, 22.viii.2002, PPCM). Colt Park Wood, Marrick, North Yorkshire, 1 Menzel leg. (DEI). Bix Bottom, Oxfordshire, 1 male, male, 27.viii.−30.viii.1981, Disney leg. (PBLN). Comin 23.ix.2000, Gibbs leg. (PDGB). Burnham Beeches, Esgair Maen, Ceredigion, 1 male, 23.vii.1987, Holmes, Buckinghamshire, 1 male, 14.vi.−1.vii.1996; 1 male, Boyce & Reed leg. (PPWM). Cowbeck, North York- 22.vii.−5.viii.1996; 1 male, 13.ix.−26.ix.1996, all Ismay shire, 1 male, 18.viii.−25.viii.1981, Disney leg. leg. (PPCM). Cadair Idris, Gwynedd, 3 males, (PBLN). Devichoys Wood, Cornwall, 1 male, no date, 3.vii.1993; 9 males, 17.vii.1993; 7 males, 1.viii.1993; 3 Merrifield leg. (PPCM). Dinton Pastures Country males, 4.ix.1993, all Laurence leg. (DEI, PBLN). Cors Park, Berkshire, 1 male, 3.xi.2001, Chandler leg. Erddreiniog, Isle of Anglesey, 1 male, 27.vii.1988, (PPCM). Epping Forest, Essex, 1 male, iv.1999, Dagley Holmes, Boyce & Reed leg. (PPWM). Epping Forest, leg. (PPCM). Ffrwd Fen, Carmarthen, 1 male, Essex, 1 male, iv.1999; 1 male, v.1999, all Dagley leg. 3.x.1989, Holmes, Boyce & Reed leg. (PPWM). Galms- (PPCM). Highgate Wood, Greater London, 2 males, dale, Isle of Eigg, Highland Region, 1 male, 10.iv.1969, 1998–99, Hackett leg. (PPCM). Lea Meadows, New- Hutson leg. (BMNH). Hafod Garregog, Gwynedd, 1 town Linford, Leicestershire, 4 males, 24.viii.2003, male, 16.vi.−30.vi.1999, Levey & Pavett leg. (PPCM). Menzel & J.E. Smith leg. (DEI, HRI). Natural History Hay Wood, Baddesley Clinton, Solihull, West Mid- Museum, South Kensington, Greater London, 4 males, lands, 137 males, 66 females, 22.viii.2002, Menzel & 9.viii.−23.viii.2002, no collector detail. (BMNH). Ryton J.E. Smith leg. (DEI, HRI). Haytor, Devon, 2 males, 1 Wood, Bubbenhall, Coventry, West Midlands, 1 male, female, 23.vi.2002, J.E. Smith leg. (HRI). Lea Mead- 25.viii.2002, Menzel leg. (DEI). Ufton Bridge, War- ows, Newtown Linford, Leicestershire, 2 males, wickshire, 1 male, 9.vi.1991, Chandler leg. (PPCM). 24.viii.2003, Menzel leg. (DEI). Leebotwood, Shrop- Wytham Woods, Wytham, Oxfordshire, 1 male, 1 shire, 5 females, 14.vii.2003, J.E. Smith leg. (HRI). female, 21.viii.2002, Menzel leg. (DEI). Luibeg, Aberdeenshire, 1 male, 21.x.2000, Chandler Ecological notes: common in broad-leaved and carr leg. (PPCM). Moccas Park, Herefordshire, 1 male, woodland; wetlands including water meadow, fen with 11.vi.2002, Godfrey leg. (PPCM). Oxwich, Swansea/ Cladium and Phragmites bed; former gravel workings; Abertawe, 2 males, 28.vi.1989, Holmes, Boyce & Reed calcareous grassland; wildlife gardens; disused rail- leg. (PPWM). Rhôsili Down, Swansea/Abertawe, 1 way cutting; open montane habitats. iv–ix. male, 12.vii.1989, Holmes, Boyce & Reed leg. (PPWM). Ryton Wood, Bubbenhall, Coventry, West Midlands, 6 males, 12.x.2001, J.E. Smith leg. (HRI); 151 males, 17 CORYNOPTERA TREPIDA (WINNERTZ, 1867) females, 25.viii.2002, Menzel & J.E. Smith leg. (DEI, Synonyms: clinochaeta Tuomikoski, 1960; subflava HRI). Santon Downham, Suffolk, 1 male, 15.viii.1970, (Lengersdorf, 1941). Hutson et al. leg. (BMNH). Sned Wood, Lower Lye, Herefordshire, 1 male, 5 females, 14.vii.2003, J.E. Literature: Laurence (1994): 118 [in part as Bradysia Smith leg. (HRI). Waggoner’s Wells, Hampshire, 1 subtilis, misidentification]; Menzel (1998): 21. male, 8.x.2002, Chandler leg. (PPCM). Wangford, Suf- Material examined: Becky Woods, Manaton, Devon, 2 folk, 2 males, 10.vi.−24.vi.1980, Foster & Procter leg. males, 2 females, 23.vi.2002, J.E. Smith leg. (HRI). (PBLN). Willes Meadow, Leamington Spa, Warwick- Blelham Bog, Cumbria, 1 male, 17.vi.1989, Chandler shire, 1 male, 3 females, 27.viii.2002, Menzel leg. leg. (PPCM). Bolt Wood, Stockerston, Leicestershire, 5 (DEI). Woodwalton Fen, Cambridgeshire, 1 male, males, 4 females, 24.viii.2003, Menzel leg. (DEI). 24.v.1940, Edwards leg. (BMNH). Wytham Woods, Brandon Marsh, Ryton-on-Dunsmore, West Midlands, Wytham, Oxfordshire, 6 females, 21.viii.2002, J.E. 6 males, 13 females, 22.viii.2002, Menzel & J.E. Smith Smith leg. (HRI). leg. (DEI, HRI). Brownsham, Devon, 1 male, Ecological notes: very common in broad-leaved, mixed 18.vii.2001, Chandler leg. (PPCM). Burnham Beeches, and carr woodland; conifer plantation with broad- Buckinghamshire, 7 males, 8.vii.−9.x.1996, Ismay leg. leaved fringe, heathland including Breckland, dry and (PPCM); 1 male, 4.x.1999, Chandler leg. (PPCM). damp heath; moorland on granite; wetlands including Cadair Idris, Gwynedd, 6 males, 19.vi.1993; 9 males, water meadows, Phragmites, stream side and fens; old 3.vii.1993; 1 male, 1.viii.1993; 6 males, 4.ix.1993; 2 hedges; parkland; grassland; open montane habitats. males, 22.x.1993; 1 male, 13.xi.1993, all Laurence leg. iv–xi. (PBLN). Canonteign Falls, Devon, 4 males, 3 females, 22.vi.2002, J.E. Smith leg. (HRI). Catfield, Norfolk, 3 males, 13.xii.−29.ii.1989, Foster & Procter leg. CORYNOPTERA UNCATA MENZEL & SMITH SP. NOV. (PBLN). Chigwell Row Wood, Chigwell Row, Essex, 1 New to British list.

Material examined: Burnham Beeches, Buckingham- Rydd, Ceredigion, 1 male, 8.x.1987, Holmes, Boyce & shire, 1 male, 8.vii.−22.vii.1996; 1 male, 22.vii.− Reed leg. (PPWM). Rhôs-y-gad, Isle of Anglesey, 4 5.viii.1996, all Ismay leg. [paratypes of Corynoptera males, 5.x.1988, Holmes, Boyce & Reed leg. (PPWM). uncata] (PPCM). Fedw Fawr, Isle of Anglesey, 3 males, Strumpshaw, Norfolk, 1 male, 13.vi.−11.vii.1988, Fos- 13.vii.1988, Holmes, Boyce & Reed leg. (PPWM). ter & Procter leg. (PBLN). Welsh Moor, Swansea/Aber- Highgate Woods, Greater London, 1 male, 1998–1999, tawe, 1 male, 12.vii.1989, Holmes, Boyce & Reed leg. Hackett leg. [paratype of Corynoptera uncata] (PPWM). Ynys Uchaf, Carmarthen, 2 males, 3.x.1989, (PPCM). Kelham Bridge, Ravenstone, Leicestershire, Holmes, Boyce & Reed leg. (PPWM). 2 males, 24.viii.2003, Menzel leg. [paratypes of Ecological notes: found in carr woodland; humid and Corynoptera uncata] (DEI). Ryton Wood, Bubbenhall, wet heathland; common in all types of fen and bog, Coventry, West Midlands, 16 males, 25.viii.2002, basin and ombrotrophic mires; Cladium beds with Menzel & J.E. Smith leg. [holotype and paratypes tufa; former gravel workings. iv, vi–x. of Corynoptera uncata] (DEI, HRI). Welsh Moor, Swansea/Abertawe, 1 male, 14.vi.1989, Holmes, Boyce & Reed leg. (PPWM). Y Gors, Powys, 1 male, CORYNOPTERA VERRUCIFERA (LENGERSDORF, 1952) 17.vii.1989, Holmes, Boyce & Reed leg. (PPWM). New to British list. Ecological notes: found in broad-leaved woodland; humid and wet heathland; ﬂoodplain mire; wet Material examined: Epping Forest, Essex, 1 male, meadow with reed beds around pools; pond banks in 15.iii.1999, Dagley leg. (PPCM). Salisbury Plain damp meadow. vii–viii. Training Area, Warminster, Wiltshire, 1 male, 2.vii.2003, Ismay & Schulten leg. (UMO). Ecological notes: found in ancient broad-leaved wood- CORYNOPTERA VAGULA TUOMIKOSKI, 1960 land; unimproved calcareous grassland. iii, vii. Literature: Menzel (1998): 21; Withers (2002): 114. Material examined: Aberithon Turbary, Powys, 1 male, 3.vii.1989, Holmes, Boyce & Reed leg. (PPWM). CORYNOPTERA WALTRAUDIS MOHRIG & MAMAEV, 1987 Annahala bog, Gearagh, Cork, 1 male, 29.iv.1990, de Literature: Menzel (1998): 21. Courcy Williams leg. (DEI). Brandon Marsh, Ryton- Material examined: Burnham Beeches, Buckingham- on-Dunsmore, Coventry, West Midlands, 1 male, shire, 1 male, 10.v.−23.v.1995, Ismay leg. (PPCM). 11.x.2001, Chandler leg. (HRI). Comin Esgair Maen, Cors Goch, Isle of Anglesey, 1 male, 14.vii.1988, Ceredigion, 2 males, 23.vii.1987, Holmes, Boyce & Holmes, Boyce & Reed leg. (PPWM). Llanon, Swansea/ Reed leg. (PPWM). Cors Bodeilio, Isle of Anglesey, 1 Abertawe 1 male, 13.vii.1989, Holmes, Boyce & Reed male, 13.vii.1988, Holmes, Boyce & Reed leg. (PPWM). leg. (PPWM). Sutton Broad, Norfolk, 1 male, 21.viii.− Cors Caranod, Ceredigion, 1 male, 24.ix.1986, Holmes, 4.ix.1989, Foster & Procter leg. (PPCM). Boyce & Reed leg. (PBLN). Cors Clegyrog, Isle of Anglesey, 1 male, 27.vii.1988; 1 male, 6.x.1988, all Ecological notes: found in ancient beech woodland; Holmes, Boyce & Reed leg. (PPWM). Cors Goch, Isle of carr woodland and fen; Molinia bog, ungrazed; Cla- Anglesey, 1 male, 14.vii.1988, Holmes, Boyce & Reed dium bed. v, vii–ix. leg. (PPWM). Cors Graianog, Gwynedd, 3 males, 15.vii.1988; 1 male, 28.vii.1988; 2 males, 7.x.1988, all Holmes, Boyce & Reed leg. (PPWM). Cors Gyfelog, CORYNOPTERA WINNERTZI MOHRIG, 1993 Gwynedd, 2 males, 26.vii.1988, Holmes, Boyce & Reed New to British list. leg. (PPWM). Cwm Crafnant, Gwynedd, 1 male, Material examined: Hartslock, Oxfordshire, 1 male, 20.vii.1988, Holmes, Boyce & Reed leg. (PPWM). Gors 14.v.2000, Raper leg. (PDGB). Llwyn, Neath/Port Talbot, 1 male, 28.vi.1989, Holmes, Boyce & Reed leg. (PPWM). Killaun Bog, Offaly, 2 Ecological notes: found on calcareous grassland adja- males, ix.2000, Withers leg. (PPWM). Llyn Coethlyn, cent to hanger woodland. v. Powys, 2 males, 1.viii.1988, Holmes, Boyce & Reed leg. (PPWM). Llyn Tyn y Mynydd, Gwynedd, 1 male, 18.viii.1988, Holmes, Boyce & Reed leg. (PPWM). Llyn CRATYNA WINNERTZ, 1867 Ystumllyn, Gwynedd, 1 male, 26.vii.1988; 2 males, CRATYNA (CRATYNA) AMBIGUA (LENGERSDORF, 1934) 19.x.1988, all Holmes, Boyce & Reed leg. (PPWM). Synonyms: latiforceps (Bukowski & Lengersdorf, Newbridge Fen, Kildare, 1 male, 27.vi.2002, Withers 1936); lignea (Lengersdorf, 1941); prima (Frey, 1942). leg. (PPWM). Oxwich, Swansea/Abertawe, 1 male, 4.x.1989, Holmes, Boyce & Reed leg. (PPWM). Rhôs New to British list.

Material examined: Bevills Wood, Abbot’s Ripton, CRATYNA (CRATYNA) SCHINERI (WINNERTZ, 1867) Cambridgeshire, 1 male, 8.vi.1980, J.H. Cole leg. Synonyms: arenaria (Beling, 1872); glabricollis (Win- (PJCB). Bix Bottom, Oxfordshire, 1 male, 23.ix.2000, nertz, 1871); intermedia (Winnertz, 1867); stokesi Raper leg. (PDGB). Bradfield Wood, Suffolk, 2 males, (Edwards, 1925a). 23.v.−30.v.1991, Laurence leg. (PBLN). Bridge of Walls, Shetland, 1 male, vii.1985, Laurence leg. Literature: Edwards (1925b): 223 [as Psilosciara (PBLN). Burnham Beeches, Buckinghamshire, 1 stokesi]; Freeman (1983b): 24 [as Plastosciara (Plas- male, 26.ix.−9.x.1996; 1 male, 1.vi.−8.vi.1999, all tosciara)]; Menzel (1998): 21; Coldwell (1999): 25 [as Ismay leg. (PPCM). Coed Tycanol, Pembrokeshire, 2 Plastosciara (Plastosciara)]; Menzel & Mohrig (2000): males, 29.vi.−7.ix.1999, Levey & Pavett leg. (HRI, 276. PPCM). Milham Ford School, Oxford, Oxfordshire, 1 Material examined: Barnsley, South Yorkshire, 1 male, 18.v.2001, Webb leg. (PPCM). Roydon Common, female, v.1924 (reared), Bayford leg. (BMNH). Wim- Norfolk, 1 male, 5.vi.1993, Chandler leg. (PPCM). Sut- bledon, Greater London, 125 males, 50 females, − ton Common, Dorset, 1 male, 17.v. 24.v.1990, George iv.1925 (reared), Stokes leg. [lectotype and paralecto- leg. (PPCM). types of Psilosciara stokesi] (BMNH, ZFMK). Ecological notes: found in ancient beech and other Ecological notes: Reared from rotten wood. iv–v. broad-leaved woodland; mixed woodland on clay; ancient beech woodland; school grounds with old poplars; calcareous grassland. v–vii, ix–x. CRATYNA (CRATYNA) ULIGINOSA (LENGERSDORF, 1929) Literature: Menzel (1998): 21; Withers (2002): 114. Material examined: Alice Holt Forest, Hampshire, 1 CRATYNA (CRATYNA) CONTRACTA MOHRIG & male, 10.v.1984, Chandler leg. (PPCM). Alvecote Pools, RÖSCHMANN, 1996 Staffordshire, 2 males, 13.x.2001, J.E. Smith leg. New to British list. (HRI). Blaen Glaswen, Powys, 1 male, 16.vi.1989, Holmes, Boyce & Reed leg. (PPWM). Boundless Copse, Material examined: Migdale Wood, Highland Region, Surrey, 2 males, 26.vii.2002, Chandler leg. (PPCM). 2 males, 31.v.2002, Chandler leg. (PPCM). Burnham Beeches, Buckinghamshire, 2 males, 28.v.− Ecological notes: found in alder and birch carr by 9.vii.1996, Ismay leg. (PPCM); 1 male, 4.x.1999; 1 stream. v. male, 23.x.2001, all Chandler leg. (PPCM). Bwlch y Dwr, Gwynedd, 1 male, 14.vii.1989, Holmes, Boyce & Reed leg. (PPWM). Cadair Idris, Gwynedd, 2 males, 19.vi.1997; 1 male, 4.ix.1993, all Laurence leg. CRATYNA (CRATYNA) PERNITIDA (EDWARDS, 1915) (PBLN). Catfield, Norfolk, 1 male, 29.ix.1988, Foster Literature: Edwards (1915a): 164; Edwards (1925a): & Procter leg. (PBLN). Cefn Grwhyd, Neath/Port 542; Perry & Stubbs (1978): 71; Freeman (1983a): 169 Talbot, 1 male, 4.x.1989, Holmes, Boyce & Reed [all as Plastosciara]; Freeman (1983b): 24 [as Plastos- leg. (PPWM). Chobham Common, Surrey, 1 male, ciara (Plastosciara)]; Hancock, 1997: 28 [as Plastosci- 16.x.1984, Chandler leg. (PPCM). Collinpark Wood, ara]; Menzel (1998): 21; Menzel & Mohrig (2002): 276; Gloucestershire, 1 male, 3.ix.2002, Gibbs leg. (PPCM). Withers (2002): 114. Cors Caranod, Ceredigion, 1 male, 25.vi.1987; 1 male, 23.vii.1987, all Holmes, Boyce & Reed leg. (PPWM). Material examined: Auchindachy, Moray, 3 males, 7 Cors Farlais, Carmarthen, 1 male, 5.x.1989, Holmes, females, 30.vi.1936, Coe leg. (BMNH). Farnham, Boyce & Reed leg. (PPWM). Cors Goch, Isle of Buckinghamshire, 4 males, 4 females, 24.v.1930 Anglesey, 1 male, 18.vii.1989, Holmes, Boyce & Reed (reared), Mansbridge leg. (BMNH). Gorge of Avon, leg. (PPWM). Cors Graianog, Gwynedd, 1 male, South Lanarkshire, 1 female, 2.vi.1906, King leg. 28.vii.1988; 2 males, 7.x.1988, all Holmes, Boyce & (BMNH). Killaun Bog, Offaly, 3 males, ix.2000, Reed leg. (PPWM). Cors-y-Llyn, Powys, 3 males, Withers leg. (PPWM). Offley, Hertfordshire, 5 10.x.1989, Holmes, Boyce & Reed leg. (PPWM). Denny males, 9 females, v.1936 (reared), Edwards leg. Wood, New Forest, Hampshire, 1 male, 17.iv.1954, (BMNH). Stanmore, Greater London, 8 males, 12 Colyer leg. (BMNH). Epping Forest, Essex, 1 male, females, 3.v.1914 (reared), Blair leg. [lectotype and 27.v.1999, Dagley leg. (PPCM); 1 male, 17.x.2000, paralectotypes of Plastosciara pernitida] (BMNH, Merrifield leg. (PPCM). Hafod Garregog, Gwynedd, 1 ZFMK). male, 13.x.1988, Holmes, Boyce & Reed leg. (PPWM). Ecological notes: found in broad-leaved woodland; Hay Wood, Baddesley Clinton, Solihull, West Mid- mixed woodland by river; raised bog. Reared from rot- lands, 26 males, 25.viii.2002, Menzel & J.E. Smith leg. ten pine stump and beech log. v, vi, ix. (DEI, HRI). Hickling, Norfolk, 2 males, x.1988, Foster

& Procter leg. (PBLN); 2 males, 16.iv.1989, Withers Literature: Laurence (1994): 106 [as Plastosciara (Ter- leg. (PPWM). Killaun, Offaly, 2 males, vi.2000, With- mitosciara) taractica]; Menzel (1998): 21; Menzel & ers leg. (PPWM). Llyn Creiniog, Denbighshire, 1 male, Mohrig (2000): 292. 13.x.1988, Holmes, Boyce & Reed leg. (PPWM). Llyn Material examined: Blaen Cilieni, Powys, 1 male, Pencraig, Gwynedd, 1 male, 27.x.1988, Holmes, Boyce 5.vii.1989, Holmes, Boyce & Reed leg. (PPWM). East & Reed leg. (PPWM). Llynoedd Ieuan, Ceredigion, 1 Harling, Norfolk, 1 male, 14.vi.−28.vi.1988, Foster & male, 18.vi.−20.vii.1988, Holmes, Boyce & Reed leg. Procter leg. [paratype of Plastosciara taractica] (PPWM). Malham Tarn, North Yorkshire, 1 male, (PBLN). Strumpshaw, Norfolk, 8 males, 13.vi.− 24.vi.−28.vi.1982, Disney leg. (PPWM). Migneint, 11.vii.1988, Foster & Procter leg. [paratypes of Conwy, 1 male, 14.x.1988, Holmes, Boyce & Reed leg. Plastosciara taractica] (PBLN). Thompson Common, (PPWM). Nûg, Conwy, 5 males, 12.x.1988; 1 male, Norfolk, 7 males, 13.ix.−27.ix.1988, Foster & Procter 27.x.1988, all Holmes, Boyce & Reed leg. (PPWM). leg. [holotype and paratypes of Plastosciara taractica] Oakley Wood, Bishops Tachbrook, Warwickshire, 1 (DEI, PBLN). male, 14.x.2001, Chandler leg. (HRI). Rhôs Cilcennin, Ceredigion, 1 male, 24.ix.1986, Holmes, Boyce & Reed Ecological notes: found in broad-leaved woodland; leg. (PPWM). Rhôs Rydd, Ceredigion, 1 male, 8.x.1987, heathland with Carex paniculata and Molinia; grass- Holmes, Boyce & Reed leg. (PPWM). Rhôsgoch, Powys, land with pingo pools; damp meadow; glacial pool. vi– 2 males, 17.vii.1989, Holmes, Boyce & Reed leg. vii, ix. (PPWM). Rhôsili Down, Swansea/Abertawe, 1 male, 3.x.1989, Holmes, Boyce & Reed leg. (PPWM). Rother- ﬁeld Park, Hampshire, 2 males, 2.x.2002, Chandler CRATYNA (PEYERIMHOFFIA) CURTIPENNIS leg. (PPCM). Roydon Common, Norfolk, 1 male, (EDWARDS, 1926) 5.vi.1993, Chandler leg. (PPCM). Ryton Wood, Literature: Edwards (1926a): 112; Freeman (1983a): Bubbenhall, Coventry, West Midlands, 1 male, 169; Freeman (1983b): 27 [all as Hyperlasion]; Menzel 12.x.2001, Chandler leg. (HRI); 1 male, 25.viii.2002, (1998): 21; Menzel & Mohrig (2000): 286. Menzel leg. (DEI). Stokeford Heath, Dorset, 1 male, 20.v.2001, Gibbs leg. (PDGB). Trawsfynydd, Gwynedd, Material examined: Basingstoke, Hampshire, 2 lar- 2 males, 14.x.1988, Holmes, Boyce & Reed leg. vae, 20.ii.1936, Speyer leg. (HRI). Clacton-on-Sea, (PPWM). Tuddenham Fen, Suffolk, 1 male, 20.x.1990, Essex, 2 males, 1 female, 5 larvae, 28.xii.1925, Speyer Chandler leg. (PPCM). Vicarage Bog, Wrexham, 1 leg. [lectotype and paralectotypes of Hyperlasion cur- male, 12.x.1988, Holmes, Boyce & Reed leg. (PPWM). tipennis] (BMNH). Rockbeare, Exeter, Devon, 3 pupae, Waggoner’s Wells, Hampshire, 2 males, 8.x.2002, 14.ii.1943; 6 larvae, 15.ii.1943, all Speyer leg. (HRI). Chandler leg. (PPCM). West End Common, Esher, Ecological notes: larvae in large aggregation on soil of Surrey, 1 male, 19.x.2001; 1 male, 16.vii.2002, all tomato house. ii, xii. Chandler leg. (PPCM). Ecological notes: very common in broad-leaved, mixed CRATYNA (PEYERIMHOFFIA) HYBRIDA and carr woodland; conifer plantations; sandy, damp, grazed wet and humid heathland; wetlands including (MOHRIG & MAMAEV, 1974) fens, mires, bogs, Cladium and Phragmites beds and New to British list. Sphagnum kettlehole; open montane habitats. iv–x. Material examined: ‘Horwood’, Basingstoke, Hamp- shire, 1 male, 26.vi.1956, Hussey leg. (HRI).

CRATYNA (DIVERSICRATYNA) PERORNATA Ecological notes: no habitat details. vi. (MOHRIG & RÖSCHMANN, 1993) New to British list. CRATYNA (PEYERIMHOFFIA) PERNICIOSA Material examined: Sned Wood, Lower Lye, Hereford- (EDWARDS, 1922) shire, 2 males, 14.vii.2003, J.E. Smith leg. (HRI). Literature: Edwards (1922): 160; Edwards (1925a): Ecological notes: found in conifer plantation with 543; Madwar (1937): 50; Freeman (1983a): 169 [all as broad-leaved fringe by river. vii. Plastosciara]; Freeman (1983b): 25 [as Plastosciara (Termitosciara)]; Menzel (1998): 21; Menzel & Mohrig (2000): 288. CRATYNA (DIVERSICRATYNA) SPICULOSA Material examined: Charing Cross, Greater London, 4 (RUDZINSKI, 1993) females, ix.1933; 2 females, ii.1934, all no collector Synonym: taractica (Laurence, 1994). detail (BMNH). Glasshouse Crops Research Station,

Littlehampton, West Sussex, 1 male, 17.xii.1956, Coventry, Warwickshire, 1 male, 28.viii.2003, J.E. Hussey leg. (HRI). Hoddesdon, Hertfordshire, 2 Smith leg. (HRI). Cors Penally, Pembrokeshire, 1 females, 15.ix.1922, Speyer leg. (HRI). Natural His- male, 22.vii.1987, Holmes, Boyce & Reed leg. (PPWM). tory Museum, South Kensington, Greater London, 1 Cotterill Clough, Cheshire, 1 female, 6.viii.1939, Brit- female, 2.ix.1926, Edwards leg. (BMNH). Nottingham, ten leg. (BMNH). Coventry, West Midlands, 1 male, Nottinghamshire, 18 females, iii.1934; 1 female, vii.1998, C.W. Plant leg. (PPCM). Dawlish Warren, 18.xi.1935, all H.P. Jones leg. (BMNH). Slough, Berk- Devon, 1 male, 29.vi.1988, J.H. Cole leg. (PJCB). shire, 2 males, 3 larvae, 24.vii.1941, Rochford leg. Delamere, Cheshire, 1 male, 1 female, 16.vii.1939, (HRI); 5 males, 2 females, 2 pupae, 29.vii.1941, Speyer Britten leg. (BMNH). Epping Forest, Essex, 1 male, leg. (HRI). Worthing, West Sussex, 6 males, 19 28.vi.2000, Chandler leg. (PPCM). Falls of Tarnash, females, 25.v.1922 (reared), Speyer leg. [including lec- Moray, 1 male, 1.vii.1936, Coe leg. (BMNH). Frithsden totype and paralectotypes of Plastosciara perniciosa] Beeches, Hertfordshire, 1 male, 30.vii.1997, Chandler (BMNH); 1 male, 1 female, 2.vi.1922, Speyer leg. leg. (PPCM). Gearagh, Cork, 2 males, 22.v.1990; 1 (HRI). Woodwalton Fen, Cambridgeshire, 35 males, male, 12.vi.1990, all de Courcy Williams leg. (DEI, vi.1940, Edwards leg. (BMNH). NMID); 1 male, 9.vii.1990, Withers leg. (PPWM). Glasshouse Crops Research Station, Littlehampton, Ecological notes: found in alder carr and fen; grounds West Sussex, 2 males, 20.viii.1956; 1 male, of horticultural research station; glasshouses; office 30.vii.1957, all Hussey leg. (HRI). Glen Lochay, buildings. Reared from cucumber; birch log with ants Stirling, 1 male, 8.vi.1932, Edwards leg. (BMNH). Myrmica ruginodis. ii–iii, v–vii, ix, xii. Haytor, Devon, 1 male, 23.vi.2002, J.E. Smith leg. (HRI). Hickling, Norfolk, 1 male, v.1988, Foster & Procter leg. (PBLN). Lea Meadows, Newtown Linford, CRATYNA (PEYERIMHOFFIA) VAGABUNDA Leicestershire, 2 males, 24.viii.2003, J.E. Smith leg. (WINNERTZ, 1867) (HRI). Leckford Estate, Hampshire, 1 male, 1.ix.1999, Synonyms: alata (Frey, 1948); brachyptera (Kieffer, Chandler leg. (PPCM). Leebotwood, Shropshire, 2 1903). males, 14.vii.2003, J.E. Smith leg. (HRI). Letchworth, Hertfordshire, 1 male, 26.vi.1932, Edwards leg. Literature: Donisthorpe (1913): 267; Edwards (1925a): (BMNH). Llyn Gwynant, Gwynedd, 1 male, 543; Donisthorpe (1924): 73; Donisthorpe (1927): 139 13.vii.1914, Edwards leg. (BMNH). Loch Garten, [all as Peyerimhoffia brachyptera]; O’Toole (1978): 163 Highland Region, 1 male, 24.vii.1997, Chandler leg. [as Plastosciara brachyptera]; Freeman (1983b): 25 [as (PPCM). Loch Morlich, Highland Region, 1 male, Plastosciara (Peyerimhoffia) brachyptera]; Laurence 24.vi.1933, Coe leg. (BMNH). Loch Pityoulish, High- (1994): 118; Laurence (1996): 87; Laurence & James land Region, 1 male, 3.vii.1933, Coe leg. (BMNH). (1997): 166 [all as Plastosciara brachyptera]; Menzel Lough Slevin, Westmeath, 1 male, 16.vi.1985, Chan- (1998): 21; Withers (2002): 114. dler leg. (PPWM). Lundy Island, Devon, 3 males, Material examined: Annahala bog, Gearagh, Cork, 1 20.vii.−27.vii.1972, Lane leg. (BMNH). Magor Marsh, male, 22.iii.1990, de Courcy Williams leg. (NMID). Monmouthshire, 1 male, 10.vi.1988, Holmes, Boyce & Badger Wood, Cambridgeshire, 1 male, vi.1998, C.W. Reed leg. (PPWM). Milham Ford School, Oxford, Plant leg. (NMS). Beaulieu River, Hampshire, 2 males, Oxfordshire, 2 males, 24.v.−4.vi.2001, Webb leg. 22.v.1936, Edwards leg. (BMNH). Blorenge, Mon- (PPCM). Monkfield Wood, Cambridgeshire, 1 male, mouthshire, 1 male, 23.vi.1988, Holmes, Boyce & Reed 1.v.−31.v.1998, C.W. Plant leg. (NMS). Murrough, leg. (PPWM). Braedownie, Angus, 1 male, 24.vii.1937, Wicklow, 1 male, 20.v.−3.vi.1990, Blackith leg. Coe leg. (BMNH). Brynberian Moor, Pembrokeshire, 1 (PPWM). New Malden, Greater London, 1 male, male, 22.vi.1987, Holmes, Boyce & Reed leg. (PPWM). 25.vi.1931, Coe leg. (BMNH). Newbold Comyn Park, Buckingham Thick Copse, Northamptonshire, 1 male, Leamington Spa, Warwickshire, 2 males, 15.vii.2001, 9.vi.1992, Warne leg. (PPCM). Burnham Beeches, J.E. Smith leg. (HRI). Nûg, Conwy, 1 male, 3.viii.1988, Buckinghamshire, 6 males, 20.vii.−2.viii.1995; 1 male, Holmes, Boyce & Reed leg. (PPWM). Oakland Wood, 1.vii.−8.vii.1996; 2 males, 5.vii.−20.vii.1999; 1 male, Cambridgeshire, 1 male, 1.v.−31.v.1998; 1 male, 1.vi.− 1.x.−28.x.1999, all Ismay leg. (HRI, PPCM). Bushy 30.vi.1998, all C.W. Plant leg. (NMS, PPCM). Rhôs-y- Park, Greater London, 1 male, vi.1940, no collector gad, Isle of Anglesey, 1 male, 27.vii.1988, Holmes, detail (BMNH). Cadair Idris, Gwynedd, 9 males, Boyce & Reed leg. (PPWM). Rhôsgyll Fawr, Gwynedd, 19.vi.1993; 1 male, 3.vii.1993, all Laurence leg. 1 male, 28.vii.1988, Holmes, Boyce & Reed leg. (PBLN). Cambridge, Cambridgeshire, 1 male, (PPWM). Salisbury Plain Training Area, Warminster, 23.v.1905, Jenkinson leg. (BMNH). Chigwell Row Wiltshire, 1 male, 30.v.2003; 4 males, 31.v.2003, all Wood, Chigwell Row, Essex, 1 male, 18.vi.2002, Ismay Ismay & Schulten leg. (DEI, UMO). Sedlescombe, East & Schulten leg. (DEI). Coombe Abbey Country Park, Sussex, 1 male, 27.vi.1996, Roper leg. (PPRS). Sned

Wood, Lower Lye, Herefordshire, 3 males, 14.vii.2003, Coe leg. (BMNH). Loch Tummel, Perth & Kinross, J.E. Smith leg. (HRI). Stackpole Quay, Pembrokeshire, 1 male, 8.vii.1937, Collin leg. (BMNH). Lochan an 1 male, 12.viii.1986, Chandler leg. (PPCM). The Brit- Daimh, Perth & Kinross, 1 male, 26.viii.1995, Chan- ish, Monmouthshire, 1 male, 24.vi.1992, Gibbs leg. dler leg. (PPCM). Mid Quoich, Aberdeenshire, 1 male, (PDGB). The Dingle, Isle of Anglesey, 1 male, 15.vi.2003, Chandler leg. (PPCM). Moineach Mararu- 5.vii.1987, Chandler leg. (NMS). The Loke, Norwich, lin, Skye, Highland Region, 1 male, 10.vii.1991, J.H. Norfolk, 2 males, 4vii.−5.vii.1987; 2 males, Cole leg. (PJCB). Rake Beck, North Yorkshire, 1 male, 15.viii.1987; 2 males, 13.vii.−19.vii.1989; 1 male, 9.vi.1976, J.H. Cole leg. [holotype of Plastosciara 21.viii.−24.viii.1989; 1 male, 15.v.−18.v.1992; 4 males, colei] (BMNH). Rhôs Cilcennin, Ceredigion, 1 male, 27.vii.1992; 1 male, 30.vii.−2.viii.1993, all Laurence 1.x.1986, Holmes, Boyce & Reed leg. (PPWM). Thomp- leg. (PBLN). Ryton Wood, Bubbenhall, Coventry, son Common, Norfolk, 1 male, 1.vi.1985, Chandler leg. West Midlands, 1 male, 25.viii.2002, Menzel leg. (PPCM). Udale Bay, Highland Region, 1 male, (DEI). Wellesbourne, Warwickshire, 1 male, 17.viii.− 29.v.2002, Chandler leg. (PPCM). Woodbastwick, Nor- 31.viii.2000; 1 male, 1 female, 27.vi.−5.vii.2001, all folk, 3 males, viii.1932, Edwards leg. (BMNH). J.E. Smith leg. (HRI). Winfrith, Dorset, 1 male, Ecological notes: found in broad-leaved, mixed and 17.viii.1910, Haines leg. (BMNH). Woodwalton Fen, carr woodland; ancient pine woodland; moorland Cambridgeshire, 1 male, 24.v.1940, Edwards leg. flushes; wetlands including fen, open marsh by pool, (BMNH). grazed Molinia bog with Carex rostrata, acid bog, Ecological notes: very common in broad-leaved and wooded stream gorge and water meadows by river; carr woodland; also found in mixed woodland; conifer grassland with pingo pools; open montane habitats. plantations and ancient pine woodland; heathland; iv–x. moorland; wetlands including fens, mires, bogs, water meadows, sedge beds and riverside habitats; unimproved calcareous grassland; grounds of horticultural CRATYNA (SPATHOBDELLA) EGERTONI research station; school grounds with old poplars; gar- (EDWARDS, 1934) den; fixed dune grassland and scrub; coastal cliffs; Literature: Edwards (1934a): 141 [as Plastosciara]; open montane habitats; mining spoil heaps. Reared Perry & Stubbs (1978): 71 [as Sciara]; Freeman from dung. iii, v–x. (1983a): 170 [as Plastosciara]; Freeman (1983b): 25 [as Plastosciara (Termitosciara)]; Menzel (1998): 21; Menzel & Mohrig (2000): 281. CRATYNA (SPATHOBDELLA) COLEI (FREEMAN, 1990) Literature: Edwards (1925a): 541 [as Sciara long- Material examined: Rostherne, Cheshire, 2 males, 18 ispina, misidentification]; Freeman (1983b): 25 [as females, v.1932 (reared), Britten leg. [lectotype and Plastosciara (Spathobdella) brachialis, misidentifica- paralectotypes of Plastosciara egertoni] (BMNH). tion]; Freeman (1990): 52 [as Plastosciara (Spathob- Ecological notes: reared from under bark of dead oak. della)]; Laurence (1997b): 228; Laurence & James v. (1997): 166 [all as Plastosciara falcifera, misidentification]; Menzel (1998): 21; Menzel & Mohrig (2000): 281; Chandler et al. (2002): 123. CRATYNA (SPATHOBDELLA) FALCATA Material examined: Annahala bog, Gearagh, Cork, 1 (TUOMIKOSKI, 1960) male, 29.iv.1990, de Courcy Williams leg. (DEI). Bwlch New to British list. y Ddeufaen, Conwy, 1 male, 5.vii.1987, J.H. Cole leg. Material examined: Balmoral Forest, Aberdeenshire, (PJCB). Cadair Idris, Gwynedd, 1 male, 4.ix.1993, 2 males, 1.viii.−4.viii.1937, Coe leg. (BMNH). Ben Laurence leg. (PBLN). Corriegills, Arran, North Ayr- Macdhui, Aberdeenshire, 19 males, 6.viii.1937, Coe shire, 1 male, 2.vi.−4.vi.1919, Edwards leg. (BMNH). leg. (BMNH). Feshiebridge, Highland Region, 1 male, Crawcwellt, Gwynedd, 1 male, 18.ix.1968, Vane- 1.vii.1933, Coe leg. (BMNH). Loch Callater, Aberdeen- Wright leg. (BMNH). Eday, Orkney, 2 males, 1.vi.1988, shire, 2 males, 28.vii.−29.vii.1937; 1 male, 22.vii.1938; Laurence leg. (PBLN). Glen Lochay, Stirling, 1 male, 11 males, 27.vii.1938, all Coe leg. (BMNH). Lockhaugh 3.vi.−18.vi.1932, Edwards leg. (BMNH). Kenmare Bank Wood, Derwent Valley, Tyne & Wear, 1 male, Estate, Killarney, Kerry, 1 male, 8.ix.1981, J.P. & M.A. 24.vii.−13.viii.1990, Jessop leg. (PBLN). O’Connor leg. (NMID). Lea Meadows, Newtown Lin- ford, Leicestershire, 46 males, 3 females, 24.viii.2003, Ecological notes: found in broad-leaved woodland; Menzel & J.E. Smith leg. (DEI, HRI). Leckford Estate, conifer plantations; wooded river bank; at flowers of Hampshire, 1 male, 20.v.1971, Stubbs leg. (PPCM). moss campion; high montane habitat; at 1309 m a.s.l. Loch Callater, Aberdeenshire, 10 males, 22.vii.1938, vii–viii.

CRATYNA (SPATHOBDELLA) FALCIFERA damp heathland; moorland; wetlands including fen, (LENGERSDORF, 1933) sedge bed, valley and basin mire, Molinia and raised Literature: Freeman (1983b): 25 [as Plastosciara bog, Cladium beds with tufa, water meadows; grass- (Spathobdella)]; Laurence (1994): 118 [as Plastosci- land with pingo pools. iv–x. ara]; Menzel (1998): 21; Withers (2002): 114.

Material examined: Annahala bog, Gearagh, Cork, 1 CRATYNA (SPATHOBDELLA) KEILINI (EDWARDS, 1915) male, 24.ix.1990, de Courcy Williams leg. (NMID). Literature: Edwards (1915b): 263; Edwards (1925a): − Balmoral Forest, Aberdeenshire, 4 males, 19.vii. 542 [all as Plastosciara]; Perry & Stubbs (1978): 71 [as 30.vii.1938, Coe leg. (BMNH). Beinn Eighe, Highland Sciara]; Freeman (1983a): 169 [as Plastosciara]; Free- Region, 1 male, 10.vi.1984, Chandler leg. (NMS). Cal- man (1983b): 24 [as Plastosciara (Spathobdella)]; ifornia Country Park, Berkshire, 1 male, 1.vi.1997; 4 Menzel (1998): 21; Menzel & Mohrig (2000): 283. males, viii.2000, all Chandler leg. (HRI, PPCM). Clay- bokie, Aberdeenshire, 1 male, 12.vii.1977, Chandler Material examined: Barton Mills, Suffolk, 1 male, 1 leg. (PPCM). Cors Gorsgoch, Ceredigion, 1 male, female, 8.v.1915, Keilin leg. [lectotype and paralecto- 9.ix.1986, Holmes, Boyce & Reed leg. (PPWM). Cors type of Plastosciara keilini] (BMNH). Bradﬁeld Wood, Fochno, Ceredigion, 1 male, 24.vii.1987, Holmes, Suffolk, 1 female, 23.v.−30.v.1991, Laurence leg. Boyce & Reed leg. (PPWM). Cwm Crafnant, Gwynedd, (PBLN). 1 male, 4.viii.1988, Holmes, Boyce & Reed leg. (NMS). Ecological notes: found in broad-leaved woodland. Dartmoor, Bush Down, Devon, 1 male, 28.viii.1989, Reared from rotten wood. v. J.H. Cole leg. (PJCB). Glen Einich, Highland Region, 1 male, 30.vi.1933, Coe leg. (BMNH). Hickling, Norfolk, 1 male, 15.viii.−29.viii.1980; 1 male, 18.v.1989, all Fos- CRATYNA (SPATHOBDELLA) NOBILIS (WINNERTZ, 1867) ter & Procter leg. (NMS, PBLN); 1 male, 27.v.1991, Synonyms: brachialis (Winnertz, 1867); cunctans Withers leg. (PPWM). How Hill, Norfolk, 1 male, (Winnertz, 1871). 27.v.1989, Withers leg. (PPWM). Leckford Estate, Hampshire, 2 males, 12.iv.1981, Chandler leg. (NMS, Literature: ? Walker (1848): 109 [as Sciara nemoralis, PPCM). Loch Garten, Highland Region, 1 male, misidentiﬁcation]; Edwards & Collin (1932): 264 [as 4.vii.1933, Coe leg. (BMNH). Loch Loy, Highland Sciara]; Freeman (1983b): 25; O’Connor & Ashe Region, 1 male, 14.vi.1984, Chandler leg. (PPCM). (1991): 29 [all as Plastosciara (Spathobdella)]; Lau- Loch Shieldaig, Highland Region, 1 male, 10.vi.1984, rence (1997b): 228; Laurence & James (1997): 166 [all Chandler leg. (PPCM). Market Weston Fen, Suffolk, 1 as Plastosciara]; Menzel (1998): 21. male, 12.v.1991, Withers leg. (PPWM). Mills Marsh, Material examined: Annahala bog, Gearagh, Cork, 4 Norfolk, 1 male, 11.viii.1988, Foster & Procter leg. males, 1.vi.1990; 1 male, 24.ix.1990, all de Courcy Wil- − (PBLN). Newbridge Fen, Kildare, 2 males, 8.viii. liams leg. (DEI, NMID). Beacon Bog, Carmarthen, 1 19.viii.1998, Good leg. (PPWM). Puttles Bridge, New male, 11.vii.1989, Holmes, Boyce & Reed leg. (PPWM). Forest, Hampshire, 3 males, 8.vii.1990, Chandler Becky Falls, Manaton, Devon, 1 male, 28.v.1978, J.H. leg. (PPCM). Rhôs Cilcennin, Ceredigion, 3 males, Cole leg. (PJCB). Beinn a′ Chreachain, Perth & Kin- 4.ix.1986; 2 males, 10.x.1986, all Holmes, Boyce & ross, 1 male, 14.vi.1932, Edwards leg. (BMNH). Beinn Reed leg. (PPWM). Rhôs Llawr-Cwrt, Ceredigion, 1 Eighe, Highland Region, 1 male, 10.vi.1984, Chandler male, vi.1987, J.H. Cole leg. (PJCB). Rhôs Rhydd, leg. (PPCM). Ben Macdhui, Aberdeenshire, 2 males, Ceredigion, 2 males, 25.ix.1986, Holmes, Boyce & 6.viii.1983, no collector detail (PPWM). Bix Bottom, Reed leg. (PPWM). Spinningdale Bog, Highland Oxfordshire, 1 male, 23.ix.2000, Raper leg. (PDGB). Region, 1 male, 17.vi.1976, Chandler leg. (NMS). Blaen Cilieni, Powys, 1 male, 21.vi.1989, Holmes, Stokeford Heath, Dorset, 1 male, 28.viii.2001, Gibbs Boyce & Reed leg. (PPWM). Boundless Copse, Surrey, leg. (PDGB). Sutton Park, Birmingham, West Mid- 1 male, 26.vii.2002, Chandler leg. (PPCM). Brandy- lands, 1 male, 6.viii.1989, J.H. Cole leg. (PJCB). The quoi, Kirkwall, Orkney, 1 male, 28.v.−8.vi.2001, Lau- Quill, Wicklow, 1 male, 7.v.1989, Withers leg. (PPWM). rence leg. (PBLN). Brechfa Pool, Powys, 1 male, Thompson Common, Norfolk, 1 male, 1.vi.1985, Chan- 26.vi.1976, Chandler leg. (PPCM). Burley, New Forest, dler leg. (NMS). Waun Isaf, Pembrokeshire, 1 male, Hampshire, 1 male, 30.vii.1953, Colyer leg. (BMNH). 22.vi.1987, Holmes, Boyce & Reed leg. (PPWM). Wood Cadair Idris, Gwynedd, 1 male, 3.vii.1993; 1 male, Crates, New Forest, Hampshire, 1 male, 8.vii.1990, 1.viii.1993; 2 males, 4.ix.1993; 1 male, 22.x.1993, all Chandler leg. (PPCM). Woodbastwick, Norfolk, 3 Laurence leg. (PBLN). California Country Park, Berk- males, viii.1932, Edwards leg. (BMNH). shire, 1 male, 29.vi.2002, Chandler leg. (PPCM). Chig- Ecological notes: common in broad-leaved, mixed, pine well Row Wood, Chigwell Row, Essex, 2 males, and carr woodland; conifer plantations; sandy and 15.ix.2002; 1 male, 23.ix.2002; 1 male, 16.x.2002, all

Ismay & Schulten leg. (DEI, UMO). Corriegills, 8.vii.1990, Chandler leg. (NMS). Reydon Wood, Suf- Arran, North Ayrshire, 1 male, 24.vi.1919, Edwards folk, 1 male, 11.v.1986, Withers leg. (PPWM). Rhôs leg. (BMNH). Cors Caranod, Ceredigion, 2 males, Rhydd, Ceredigion, 1 male, 11.ix.1986, Holmes, Boyce 24.ix.1986; 2 males, 11.vi.1987, all Holmes, Boyce & & Reed leg. (PPWM). Rhôsgyll Fawr, Gwynedd, 1 Reed leg. (PPWM). Cors Geirch, Gwynedd, 1 male, male, 28.vii.1988; 2 males, 7.x.1988, all Holmes, Boyce 16.vi.1988; 1 male, 15.vii.1988, all Holmes, Boyce & & Reed leg. (PPWM). Ryton Wood, Bubbenhall, Cov- Reed leg. (PPWM). Cors Gyfelog, Gwynedd, 1 male, entry, West Midlands, 6 males, 1 female, 25.viii.2002, 30.vi.1988, Holmes, Boyce & Reed leg. (PPWM). Cors Menzel & J.E. Smith leg. (DEI, HRI). Salem, Ceredi- Llyn Farch a Llyn Fanod, Ceredigion, 1 male, gion, 1 male, 1.viii.1987, Holmes, Boyce & Reed 2.ix.1986; 1 male, 17.x.1986, all Holmes, Boyce & Reed leg. (PPWM). Savernake Forest, Wiltshire, 1 male, leg. (PPWM). Crymlyn Bog, Swansea/Abertawe, 1 31.v.1990, Porter leg. (NMS). Stokeford Heath, Dorset, male, 5.x.1989, Holmes, Boyce & Reed leg. (PPWM). 1 male, 20.v.2001, Gibbs leg. (PDGB). Tomintoul, Cwm Ystwyth, Ceredigion, 1 male, 27.viii.1986; 3 Moray, 1 male, 8.viii.−11.viii.1937, Coe leg. (BMNH). males, 19.ix.1986; 1 male, 2.x.1986; 1 male, 16.vi.1987, Trawsfynydd, Gwynedd, 1 male, 5.viii.1988, Holmes, all Holmes, Boyce & Reed leg. (PPWM). Dingwall, Boyce & Reed leg. (PPWM). Turville Heath, Bucking- Highland Region, 1 male, 17.viii.1909, King leg. hamshire, 4 males, 20.vii.2002, Chandler leg. (PPCM). (BMNH). Dyffryn Gwaun, Pembrokeshire, 1 male, Upton Broad, Norfolk, 1 male, 12.v.−19.v.1988, With- 22.vi.1987; 1 male, 20.vii.1987, all Holmes, Boyce & ers leg. (PPWM). Waterperry Wood, Oxfordshire, 1 Reed leg. (PPWM). Eday, Orkney, 1 male, 1.vi.1988, male, 5.v.1940; 12 males, v.1940, all Edwards leg. Laurence leg. (PBLN). Eyeworth Wood, New Forest, (BMNH). Welton Wood, Lincolnshire, 1 male, 1.v.1988, Hampshire, 1 male, 2.vi.1985, Chandler leg. (PPCM). Hoff leg. (NMS). West End Common, Esher, Surrey, 5 Figyn Blaen Breﬁ, Ceredigion, 1 male, 4.ix.1986; 1 males, 16.vii.2002, Chandler leg. (PPCM). Wistmans male, 18.ix.1986, all Holmes, Boyce & Reed leg. Wood, Devon, 1 male, 28.v.1978, J.H. Cole leg. (PJCB). (PPCM, PPWM). Fleet Pond, Hampshire, 1 male, Witherslack, Cumbria, 1 male, vi.1922, Edwards leg. 15.vii.1990, Chandler leg. (PPCM). Gearagh, Cork, 1 (BMNH). Woodwalton Fen, Cambridgeshire, 10 males, male, 31.iii.1990; 1 male, 2.iv.1990; 1 male, 11.v.1990; 24.v.1940, Edwards leg. (BMNH). 1 male, 12.vi.1990, all de Courcy Williams leg. (DEI, Ecological notes: very common in broad-leaved and NMID). Glen Einich, Highland Region, 1 male, mixed woodland; conifer plantations; ancient pine 30.vi.1933, Coe leg. (BMNH). Greywell Fen, Hamp- woodland; stunted oak woodland on moorland; carr shire, 1 male, 15.vii.1990, Chandler leg. (NMS). woodland; common on dry and damp heathland; com- Gwaun Garthenor, Ceredigion, 2 males, 2.x.1987, mon on wetlands including all types of fen, mires, Holmes, Boyce & Reed leg. (PPWM). Hickling, Nor- bogs, ditches, sedge beds and water meadows; sea cliff; folk, 1 male, 16.iv.1989, Withers leg. (PPWM). Ipley grazed meadow; calcareous grassland; open and high Manor, New Forest, Hampshire, 1 male, 13.vii.1990, montane habitats. iii–x. Chandler leg. (PPCM). Killaun Bog, Offaly, 1 male, ix.2000, Withers leg. (PPWM). Lea Meadows, New- town Linford, Leicestershire, 3 males, 24.viii.2003, CRATYNA (SPATHOBDELLA) PERPLEXA Menzel & J.E. Smith leg. (DEI, HRI). Llyn Llennyrch, (WINNERTZ, 1867) Gwynedd, 1 male, 16.vii.1976, Chandler leg. (PPCM). Llyn Pencraig, Gwynedd, 2 males, 20.vii.1988, Synonyms: brevicornis (Tuomikoski, 1957); dispar Holmes, Boyce & Reed leg. (PPWM). Llyn Ystumllyn, (Beling, 1885) [preocc.]; gregaria (Beling, 1872); pilosa Gwynedd, 1 male, 26.vii.1988, Holmes, Boyce & Reed (Rübsaamen, 1894) [preocc.]; socialis (Winnertz, leg. (PPWM). Loch Garten, Highland Region, 1 male, 1871). v.1981, Owen leg. (BMNH). Loch Park, Moray, 1 male, New to British list. 12.viii.−14.viii.1937, Coe leg. (BMNH). Lough Slevin, Westmeath, 1 male, 16.vi.1985, Chandler leg. Material examined: Burnham Beeches, Buckingham- (PPWM). Lundy Island, Devon, 1 male, 20.vii.− shire, 1 male, 20.iv.1998, Chandler leg. (PPCM). 27.vii.1972, Lane leg. (BMNH). Meall nan Tarmachan, Ecological notes: found in ancient beech woodland. iv. Perth & Kinross, 1 male, 2.vi.1932, Edwards leg. (BMNH). Mongan Bog, Offaly, 1 male, 14.vi.1983, Ashe leg. (NMID). New Close Wood, Brandon, War- CRATYNA (SPATHOBDELLA) PHILI MENZEL, 2002 wickshire, 2 males, 28.viii.2003, Menzel & J.E. Smith New to Irish list. leg. (DEI, HRI). Pont Croesor, Gwynedd, 1 male, 2.vi.1987, Chandler leg. (PPCM). Porthcawl, Brid- Material examined: Annahala bog, Gearagh, Cork, 1 gend, 1 male, 17.vi.1906, Yerbury leg. (BMNH). Put- male, 13.x.1990, de Courcy Williams leg. (NMID). tles Bridge, New Forest, Hampshire, 1 male, Gearagh, Cork, 1 male, 7.iv.1990, de Courcy Williams

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 92 F. MENZEL ET AL. leg. (DEI). Victoria Bridge, River Liffey, Kildare, 1 Essex, 2 males, 1.iv.1991, Withers leg. (PPWM). Ben male, 30.ix.2000, Chandler leg. (PPCM). Chalum, Stirling, 1 male, 1 female, 9.vi.−10.vi.1932, Edwards leg. (BMNH). Bersham, Wrexham, 1 female, Ecological notes: found in damp broad-leaved wood- 15.iii.1953, Colyer leg. (BMNH). Black Wood, Ran- land; riverbank; bog. iv, ix–x. noch, Perth & Kinross, 2 males, 27.viii.1995, Chandler leg. (PPCM). Blean Wood, Kent, 6 females, 19.v.1973, CRATYNA (SPATHOBDELLA) TUBERCULATA Parmenter leg. (BMNH). Blaen y Wergloedd, Conwy, 2 (TUOMIKOSKI, 1960) males, 4.viii.1988, Holmes, Boyce & Reed leg. (PPWM). Bolt Wood, Stockerston, Leicestershire, 1 New to British list. female, 24.viii.2003, J.E. Smith leg. (HRI). Box Hill, Material examined: Ben Macdhui, Aberdeenshire, 2 Surrey, 1 male, 11.vii.1991, Chandler leg. (PPCM). males, 6.viii.1937, Coe leg. (BMNH). Bradford, West Yorkshire, 1 male, 31.v.1949, Fidler leg. (BMNH). Braelangwell, Highland Region, 1 male, Ecological notes: found in high montane habitat; at 1.ix.1984, Chandler leg. (NMS). Brampton Wood, ﬂowers of moss campion; at 1309 m a.s.l. viii. Cambridgeshire, 1 male, 25.v.1987, J.H. Cole leg. (PJCB). Brancaster, Norfolk, 3 males, 28.x.1988, Fos- ter & Procter leg. (PBLN). Brandon Marsh, Ryton- CTENOSCIARA TUOMIKOSKI, 1960 on-Dunsmore, Coventry, West Midlands, 4 females, CTENOSCIARA HYALIPENNIS (MEIGEN, 1804) 12.x.2001, J.E. Smith leg. (HRI); 1 female, Synonyms: annulata (Meigen, 1818); autumnalis 22.viii.2002, Menzel leg. (DEI). Brendon, Devon, 2 (Winnertz, 1867); electa (Grzegorzek, 1884); eximia males, viii.1934, Edwards leg. (BMNH). Brodick, (Winnertz, 1867); insularis (Frey, 1936); rufa (Grze- Arran, North Ayrshire, 1 female, 22.v.−25.v.1919, gorzek, 1884); sordidella (Zetterstedt, 1851). Edwards leg. (BMNH). Bromley, Greater London, 1 male, 10.v.1966; 1 male, iv.1967; 1 male, 12.x.1974, all Literature: Edwards (1925a): 537; Edwards (1926b): Chandler leg. (NMS, PPCM). Buckingham Palace Gar- 142; ? Basden (1933): 44; Edwards (1934b): 34 [all as den, Greater London, 1 male, 14.ii.−27.iii.1997, D.J. Sciara autumnalis]; Chandler (1978): 209; Freeman Carter & C.W. Plant leg. (PPCM). Burnham Beeches, (1983b): 32; Skidmore et al. (1987): 119; Laurence Buckinghamshire, 1 male, 2 females, 4.xi.1990; 2 (1994): 178; Laurence (1996): 86; Laurence (1997a): males, 20.iv.−21.iv.1998; 2 males, 19.x.1998, all Chan- 85; Laurence (1997b): 228; Laurence & James (1997): dler leg. (NMS, PPCM); 1 male, 7.vi.−21.vi.1995; 4 166; Menzel (1998): 21; Chandler (2001): 201. males, 21.vi.−5.vii.1995; 1 male, 15.ix.−4.x.1995; 1 Material examined: Aber Valley, Gwynedd, 1 male, male, 14.vi.−1.vii.1996; 1 male, 8.vii.−22.vii.1996, all 7.vii.1987, Chandler leg. (NMS). Affpuddle Heath, Ismay leg. (NMS, PPCM). Bushy Park, Teddington, Dorset, 1 male, 16.x.1993, Chandler leg. (PPCM). Greater London, 1 male, 14.iv.1964, Harris leg. Alvecote Pools, Staffordshire, 1 male, 17 females, (BMNH). Cadair Idris, Gwynedd, 6 males, 19.vi.1993; 13.x.2001, J.E. Smith leg. (HRI). Annahala bog, Gear- 1 male, 3.vii.1993; 3 males, 1.viii.1993; 10 males, agh, Cork, 1 male, 23.vi.1990; 2 males, 24.ix.1990, de 4.ix.1993, all Laurence leg. (PBLN). Cairngorm, Courcy Williams leg. (DEI, NMID). Archers Wood, Highland Region, 1 male, 16.vi.1984, Chandler leg. Cambridgeshire, 1 male, 29.iv.1987 (reared), J.H. Cole (PPWM). California Country Park, Berkshire, 1 male, leg. (PJCB). Arne, Dorset, 1 male, 17.v.−3.vi.1997, C.W. 6.viii.1997, Chandler leg. (PPCM). Canonteign Falls, Plant leg. (NMS). Arthog Bog, Gwynedd, 1 male, Devon, 2 females, 23.vi.2002, J.E. Smith leg. (HRI). 17.x.1988, Holmes, Boyce & Reed leg. (PPWM). Catacol, Arran, North Ayrshire, 1 female, 29.v.− Ashridge Estate, Hertfordshire, 1 female, 22.iv.1972; 1 30.v.1919, Edwards leg. (BMNH). Cefn Grwhyd, male, 8.x.1999, all Chandler leg. (NMS). Avoca Wood, Neath/Port Talbot, 1 male, 12.vii.1989, Holmes, Boyce Wicklow, 1 male, 10.xi.1986, Chandler leg. (NMS). & Reed leg. (PPWM). Chelsham, Surrey, 4 males, 2 Aysgarth Force, North Yorkshire, 1 male, 5.x.1985, females, vi.1940, Coe leg. (BMNH). Chigwell Row Withers leg. (PPWM). Baldock, Hertfordshire, 1 male, Wood, Chigwell Row, Essex, 1 female, 20.v.2002; 1 ix.-x.1917; 1 female, v.1918, all Edwards leg. (BMNH). female, 23.v.−27.v.2002; 9 females, 11.vi.2002; 8 Barra, Western Isles, 1 male, 1 female, 17.viii.− females, 18.vi.2002; 9 females, 28.vi.2002; 4 males, 6 28.viii.1936, A.R. Waterston leg. (NMS). Bayford, females, 15.ix.2002; 1 male, 5 females, 23.ix.2002; 4 Hertfordshire, 1 male, 15.x.1931, Edwards leg. males, 3.x.2002; 4 males, 7 females, 8.x.2002; 5 males, (BMNH). Beckenham Place Park, Greater London, 1 17 females, 16.x.2002; 4 males, 1 female, 22.x.2002, all male, 4.ix.1966, Vane-Wright leg. (BMNH). Beinn Ismay & Schulten leg. (DEI, UMO). Chobham Com- Heasgarnich, Perth & Kinross, 2 females, 11.vi.1932, mon, Surrey, 1 male, vi.1940, Coe leg. (BMNH); 1 Edwards leg. (BMNH). Beinn nan Eachan, Stirling, 2 female, 8.ix.1974, Chandler leg. (NMS). Clovelly, males, 31.v.1932, Edwards leg. (BMNH). Belfairs, Devon, 1 male, 8.v.1936, Edwards leg. (BMNH).

Clowes Wood, Warwickshire, 3 males, 19.iv.1987, 1 male, 9.vi.1990; 1 female, 18.ix.1990; 1 male, Withers leg. (PPWM). Coire Raibeirt, Cairngorm, 10.x.1990, all de Courcy Williams leg. (DEI, NMID). Highland Region, 1 male, 16.vi.1984, Chandler leg. Glen Croe, Argyll & Bute, 1 male, 25.iv.1973, Dear (NMS). Collinpark Wood, Gloucestershire, 1 male, leg. (BMNH). Glen of Drumloch, Moray, 2 males, 3.ix.2002, Gibbs leg. (PDGB). Comin Esgair Maen, 16.viii.1937, Coe leg. (BMNH). Glen of the Downs, Ceredigion, 1 male, 23.vii.1987; 1 male, 15.x.1987, Wicklow, 1 male, 25.vi.1975, Chandler leg. (NMS). all Holmes, Boyce & Reed leg. (PPWM). Consall Godlingstone Heath, Dorset, 1 male, 1.vii.1998, Chan- Wood, Staffordshire, 1 male, 12.x.1997, Chandler leg. dler leg. (PPCM). Grantown-on-Spey, Highland (PPCM). Convil Braes, Moray, 8 males, 11.viii.1937, Region, 1 male, 17.vii.1991, Chandler leg. (NMS). Coe leg. (BMNH). Corriegills, Arran, North Ayrshire, 1 Grassholm, Pembrokeshire, 2 males, 25.viii.1934, female, 2.vi.−4.vi.1919, Edwards leg. (BMNH). Cors Edwards leg. (BMNH). Gwenfro, Isle of Anglesey, 1 Caron, Ceredigion, 1 male, 27.vii.1987; 2 males, male, 27.vii.1988, Holmes, Boyce & Reed leg. (PPWM). 5.x.1987; 1 male, 27.x.1987, all Holmes, Boyce & Reed Haugh Wood, Herefordshire, 1 male, 10.x.1998, Chan- leg. (PPWM). Cors Erddreiniog, Isle of Anglesey, 2 dler leg. (PPCM). Hay Wood, Baddesley Clinton, Soli- males, 6.x.1988, Holmes, Boyce & Reed leg. (PPWM). hull, West Midlands, 32 males, 25.viii.2002, Menzel & Cors Gorsgoch, Ceredigion, 2 males, 6.x.1987, Holmes, J.E. Smith leg. (DEI, HRI). Hickling, Norfolk, 1 male, Boyce & Reed leg. (PPWM). Cors Llyn Farch a Llyn 1988, Foster & Procter leg. (PBLN). Headley Common, Fanod, Ceredigion, 1 male, 17.ix.1986; 1 male, Hampshire, 1 male, 23.x.1972, Cranston leg. (BMNH). 1.x.1986; 1 male, 15.x.1987, all Holmes, Boyce & Reed Hermon Copper Bog, Gwynedd, 1 male, 25.vii.1988, leg. (PPWM). Cors Penally, Pembrokeshire, 1 male, Holmes, Boyce & Reed leg. (PPWM). Hitchin, Hert- 30.ix.1987; 1 male, 14.x.1987, all Holmes, Boyce & fordshire, 1 female, x.1917, Edwards leg. (BMNH). Reed leg. (PPWM). Coventry, West Midlands, 1 male, Hole of Horcum, North Yorkshire, 1 male, 31.viii.1937, vii.1998, C.W. Plant leg. (NMS). Crickhowell, Powys, 1 Edwards leg. (BMNH). Holme Fen, Cambridgeshire, 2 male, 1.x.1936, Edwards leg. (BMNH). Culzean Cas- males, 27.iv.1987, J.H. Cole leg. (PJCB). Hosey Com- tle, South Ayrshire, 1 male, 16.vii.1995, Chandler leg. mon, Westerham, Kent, 2 males, 3 females, 25.ix.1966, (PPCM). Cwm Bychan, Gwynedd, 1 male, 12.vii.1976, Chandler leg. (NMS, PPCM). Kergord, Shetland, 1 Chandler leg. (PPCM). Darenth Wood, Kent, 1 female, male, 16.viii.−18.viii.1996, Laurence leg. (PBLN). Kil- 17.ix.1966, Chandler leg. (NMS). Delamere, Cheshire, laun Bog, Offaly, 1 male, ix.2000, Withers leg. 1 female, 23.viii.1953; 1 male, 7.vii.1957, all Colyer (PPWM). Kingsbury Water Park, Warwickshire, 6 leg. (BMNH). Derrybawn, Wicklow, 1 male, 13.xi.1984, males, 3 females, 13.x.2001, J.E. Smith leg. (HRI). Kis- Chandler leg. (PPCM). Derrycunihy Cascade, Kerry, 1 meldon Meadow, Devon, 1 male, 15.x.1988, Chandler female, 16.x.1973, Chandler leg. (PPCM). Devil’s Glen, leg. (PPCM). Knebworth, Hertfordshire, 1 male, Wicklow, 1 male, 4.x.1980, Chandler leg. (PPCM). 24.iii.1923, Edwards leg. (BMNH). Knole Park, Kent, Devil’s Punchbowl, Surrey, 2 males, 23.vi.1990; 1 2 females, 6.ix.1966; 1 female, 1.x.1967; 2 females, male, 1.x.1995, all Chandler leg. (NMS). Dowrog Com- 8.x.1973, all Chandler leg. (NMS, PPCM). Lairig mon, Pembrokeshire, 1 male, 29.ix.1987; 1 male, Ghru, Highland Region, 1 male, 22.vii.1933, Coe leg. 13.x.1987, all Holmes, Boyce & Reed leg. (PPWM). (BMNH). Lake Windermere, Cumbria, 1 male, 1 Dromore Forest, Kerry, 1 male, 15.x.1973, Chandler female, 9.vii.1927, Edwards leg. (BMNH). Lea Mead- leg. (NMS). Drumguish, Highland Region, 1 male, ows, Newtown Linford, Leicestershire, 19 females, 17.viii.1986, Chandler leg. (PPCM). Duddleswell, East 24.viii.2003, Menzel & J.E. Smith leg. (DEI, HRI). Sussex, 1 female, 9.x.1975, Chandler leg. (NMS). Dyf- Leckford Estate, Hampshire, 1 male, 15.iii.1992; 1 fryn Gwaun, Pembrokeshire, 1 male, 20.vii.1987, male, 30.x.1998, all Chandler leg. (NMS). Leebotwood, Holmes, Boyce & Reed leg. (PPWM). Ebbor Gorge, Shropshire, 3 females, 14.vii.2003, J.E. Smith leg. Somerset, 1 male, 16.x.1986, Stubbs leg. (PPCM). (HRI). Leigh-on-Sea, Essex, 1 male, 28.iv.1986, With- Edford Wood, Somerset, 2 males, 18.x.1986, Chandler ers leg. (PPWM). Lion Wood, Redditch, Worcester- leg. (NMS). Epping Forest, Essex, 1 male, 17.viii.1998, shire, 3 males, 5 females, 10.iv.2003, J.E. Smith leg. Chandler leg. (NMS); 1 male, 25.viii.1998; 3 males, (HRI). Llyn Coethlyn, Powys, 1 male, 1.viii.1988, 28.i.−19.iv.1999, all Dagley leg. (PPCM). Ewden Valley, Holmes, Boyce & Reed leg. (PPWM). Llyn Tyn y Shefﬁeld, South Yorkshire, 1 female, 21.ix.1971, Chan- Mynydd, Gwynedd, 1 male, 18.viii.1988, Holmes, dler leg. (PPCM). Figyn Blaen Breﬁ, Ceredigion, 1 Boyce & Reed leg. (PPWM). Loch Callater, Aberdeen- male, 2.x.1986, Holmes, Boyce & Reed leg. (PPWM). shire, 2 males, 1 female, 22.vii.−29.vii.1937, Coe Freshwater East, Pembrokeshire, 2 males, leg. (BMNH). Loch Loy, Highland Region, 1 male, 11.vii.1986, Chandler leg. (NMS, PPCM). Frithsden 24.vii.1997, Chandler leg. (PPCM). Lochan na Lairige, Beeches, Hertfordshire, 2 males, 16.x.1997; 1 male, Perth & Kinross, 1 male, 12.vi.−13.vi.1932, Edwards 15.x.1998, all Chandler leg. (NMS, PPCM). Gearagh, leg. (BMNH). Lockhaugh Bank Wood, Derwent Valley, Cork, 1 male, 2 females, 9.iii.1990; 1 male, 26.iv.1990; Tyne & Wear, 6 males, 24.vii.−13.viii.1990, Jessop leg.

(PBLN). Long Spinney, Moreton-in-Marsh, Warwick- (BMNH). Slepe Heath, Dorset, 1 female, 19.ix.1975, shire, 1 female, 11.x.2001, J.E. Smith leg. (HRI). Chandler leg. (NMS). Sliddery, Arran, North Ayrshire, Lopham Fen, Norfolk, 1 male, 11.x.1990, Withers leg. 1 male, viii.1925, J. Waterston leg. (BMNH). Sned (PPWM). Lundy Island, Devon, 1 male, 20.vii.− Wood, Lower Lye, Herefordshire, 9 males, 8 females, 27.vii.1972, Lane leg. (BMNH). Meall a′ Churain, 14.vii.2003, J.E. Smith leg. (HRI). Spinningdale, High- Stirling, 1 male, 6.vi.1937, Edwards leg. (BMNH). land Region, 1 male, 14.v.1994, Entwistle leg. (PPCM). Meall Ghaordie, Stirling, 5 males, 17.vi.1932, St. David’s Airfield, Pembrokeshire, 1 male, 12.x.1987, Edwards leg. (BMNH). Mereworth Wood, Kent, 2 Holmes, Boyce & Reed leg. (PPWM). Stanford, Nor- males, 1 female, 29.viii.1966, Chandler leg. (PPCM). folk, 4 males, 27.iii.−10.iv.1988, Collier leg. (PPWM). Migdale Wood, Highland Region, 1 male, 18.x.1995, Sutton Park, Birmingham, West Midlands, 19 males, Chandler leg. (PPCM). Monkfield Wood, Cam- 17 females, 7.iv.2003, J.E. Smith leg. (HRI). The Din- bridgeshire, 1 male, 3.vii.2002, C.W. Plant leg. gle, Isle of Anglesey, 1 male, 5.vii.1987, Chandler (PPCM). Narborough Bog, Whetstone, Leicestershire, leg. (NMS). The Loke, Norwich, Norfolk, 2 males, 1 female, 24.viii.2003, J.E. Smith leg. (HRI). New For- 26.iv.1987; 1 male, 24.x.−25.x.1987; 1 male, est, Hampshire, 4 males, 1937, no collector detail 19.vii.1988; 1 male, 18.ix.−20.ix.1988; 1 male, no date, (BMNH). Newbold Comyn Park, Leamington Spa, all Laurence leg. (PBLN). Thetford Forest, Norfolk, 1 Warwickshire, 3 females, 15.vii.2001, J.E. Smith leg. female, 10.iv.1973, Chambers leg. (BMNH). Thunder- (HRI). Nûg, Conwy, 1 male, 12.x.1988, Holmes, Boyce sley Glen, Essex, 2 males, 30.iv.1986, Withers leg. & Reed leg. (PPWM). Nutcombe Down, Surrey, 2 (PPWM). Trefeiddan Moor, Pembrokeshire, 1 male, males, 26.vii.2002, Chandler leg. (PPCM). Oaken 29.ix.1987, Holmes, Boyce & Reed leg. (PPWM). Tub- Wood, Barming, Kent, 2 males, 20.ix.1966, Chandler ney, Oxfordshire, 1 male, 20.iv.1908, Hamm leg. leg. (PPCM). Oakley Wood, Bishops Tachbrook, War- (UMO). Upton Broad, Norfolk, 1 male, 15.x.1983, wickshire, 2 males, 1 female, 14.x.2001, Chandler & Withers leg. (PPWM). Vicarage Bog, Wrexham, 4 J.E. Smith leg. (HRI). Orlestone Wood, Kent, 1 female, males, 12.x.1988, Holmes, Boyce & Reed leg. (PPWM). 11.ix.1966, Chandler leg. (PPCM). Oxshott Heath, Victoria Falls, Highland Region, 1 male, 19.ix.1989, Surrey, 1 female, 15.x.1966, Chandler leg. (PPCM). Chandler leg. (NMS). Walberswick, Suffolk, 2 males, 2 Pant-y-Sais, Neath/Port Talbot, 1 male, 5.x.1989, females, 7.iii.1973, Chambers leg. (BMNH). Waun Holmes, Boyce & Reed leg. (PPWM). Pen-y-Ghent, Isaf, Pembrokeshire, 1 male, 28.ix.1987, Holmes, North Yorkshire, 1 male, v.1933, Edwards leg. Boyce & Reed leg. (PPWM). West Harling Common, (BMNH). Pine Wood, Berkshire, 1 male, 22.vi.2001, Norfolk, 1 male, 1.iv.1989; 2 males, 23.iv.1991, all Gibbs leg. (PDGB). Pont Aber, Sawdde Gorge, Car- Withers leg. (PPWM). Western Cleddau, Pem- marthen, 1 male, 12.x.1995, Chandler leg. (PPCM). brokeshire, 1 male, 7.vii.1987, Holmes, Boyce & Reed Powler’s Piece, Devon, 1 male, 13.x.1988, Chandler leg. (PPWM). Westhide, Herefordshire, 1 male, leg. (NMS). Priddy, Somerset, 1 male, 16.x.1986, 9.x.1998, Chandler leg. (NMS). Weston, Hertfordshire, Chandler leg. (NMS). Rhôs Llawr Cwrt, Ceredigion, 1 1 male, 4.x.1931, Edwards leg. (BMNH). Whichford male, 23.x.1987, Holmes, Boyce & Reed leg. (PPWM). Wood, Moreton-in-Marsh, Warwickshire, 2 males, 2 Rhôs Rhydd, Ceredigion, 1 male, 2.vii.1987; 3 males, females, 11.x.2001, J.E. Smith leg. (HRI). Willes 18.x.1987, all Holmes, Boyce & Reed leg. (PPWM). Meadow, Leamington Spa, Warwickshire, 1 female, Richmond Park, Greater London, 2 males, 1 female, 27.viii.2002, Menzel leg. (DEI). Windbury, Devon, 1 6.vi.1992, Chandler leg. (NMS). Rothamsted Wood, male, 18.v.1987, Chandler leg. (NMS). Windsor Forest, Harpenden, Hertfordshire, 1 male, vii.1951, Laurence Berkshire, 1 male, 24.vi.1950, Colyer leg. (BMNH); 1 leg. (PBLN). Rotherfield Park, Hampshire, 1 male, female, 28.iii.1971; 1 male, 13.vi.1987, all Chandler 3.vii.2002, Chandler leg. (PPCM). Rough Hill Wood, leg. (NMS, PPCM). Wisley, Surrey, 1 male, 1.iii.1923; 1 Studley, Warwickshire, 2 males, 1 female, 10.iv.2003, male, 5.ix.1923, all Dudgeon leg. (BMNH). Wokefield J.E. Smith leg. (HRI). Ryton Wood, Bubbenhall, Cov- Common, Berkshire, 1 male, 1.x.1972, Chandler leg. entry, West Midlands, 22 males, 26 females, 12.x.2001, (NMS). Wolford Wood, Moreton-in-Marsh, Warwick- Chandler & J.E. Smith leg. (HRI); 25 males, 4 females, shire, 8 males, 4 females, 11.x.2001, Chandler & J.E. 25.viii.2002, Menzel & J.E. Smith leg. (DEI, HRI). Smith leg. (HRI). Woodwalton Fen, Cambridgeshire, 1 Savernake Forest, Wiltshire, 1 male, 21.iv.−13.vi.1990, male, 24.v.1940, Edwards leg. (BMNH). Wyre Forest, Porter leg. (PPCM). Selsdon, Greater London, 1 male, Worcestershire, 1 male, 18.vii.1971; 1 male, 14.x.1984, 4 females, 25.iv.1937; 1 male, 1 female, 9.v.1937, all all Chandler leg. (NMS). Wytham Woods, Wytham, Coe leg. (BMNH). Shefford, Bedfordshire, 2 males, 1 Oxfordshire, 2 females, 21.viii.2002, Menzel & J.E. female, x.1917; 1 female, 17.xi.1917, all Edwards leg. Smith leg. (DEI, HRI). Yell, Shetland, 1 male, 3.ix.− (BMNH). Silchester Common, Hampshire, 1 female, 19.ix.1990, Laurence leg. (PBLN). Ynys Eidiol, Cere- 1.x.1972, Chandler leg. (PPCM). Silwood Park, Berk- digion, 5 males, 9.x.1987; 1 male, 28.x.1987, all shire, 2 males, 2 females, 25.ix.1957, Richards leg. Holmes, Boyce & Reed leg. (PPWM).

Ecological notes: very common in broad-leaved, mixed, 12.x.2001, J.E. Smith leg. (HRI). Burnham Beeches, pine and carr woodland; conifer plantations; dry and Buckinghamshire, 1 male, 31.viii.−13.ix.1996, Ismay wet heathland; wetlands including fens, mires, bogs, leg. (PPCM). Cadair Idris, Gwynedd, 1 male, wet meadows, sedge bed and Sphagnum kettlehole; 13.xi.1993, Laurence leg. (PBLN). Farnham Royal, parkland; grounds of horticultural research station; Buckinghamshire, 1 male, 20.v.1931 (reared), Basden gardens; grassland and scrub; marginal vegetation by leg. (BMNH). Freshfield, Merseyside, 3 males, 22.vi.− upland lake; thrift flush; sea cliff; salt marshes and 23.vi.1932 (reared), Warwick leg. (BMNH). Kingsbury reed beds; rocky island with seabird colonies; very Water Park, Warwickshire, 1 male, 13.x.2001, J.E. common in open montane habitats. Reared from decay- Smith leg. (HRI). Lockhaugh Bank Wood, Derwent ing vegetable matter; orchids and rotten log. ii–xi. Valley, Tyne & Wear, 1 male, 24.vii.−13.viii.1990, Jes- sop leg. (PBLN). Rothamsted, Harpenden, Hertford- shire, 1 male, 3.viii.1950, Laurence leg. (PBLN). The EPIDAPUS HALIDAY, 1851 Loke, Norwich, Norfolk, 1 male, 21.iv.1988, Laurence EPIDAPUS (EPIDAPUS) ALNICOLA (TUOMIKOSKI, 1957) leg. (PBLN). Wellesbourne, Warwickshire, 1 male, New to British list. 13.ix.−4.x.2001, J.E. Smith leg. (HRI). Material examined: Burnham Beeches, Buckingham- Ecological notes: found in broad-leaved and carr wood- shire, 1 male, 1.vi.−8.vi.1999, Ismay leg. (PPCM). land; wooded lake margin with reed beds; fen with Epping Thicks, Epping Forest, Essex, 1 male, Carex paniculata, Carex riparia, Cladium, Equisetum, viii.1999, Dagley leg. (PPCM). Old Slade, Bucking- Phragmites, Schoenus nigricans and Sphagnum; hamshire, 5 males, 21.iv.−12.v.1972 (reared), Chandler grazed Molinia and Myrica; blanket bog; former leg. (PPCM). Plas Islyn Bog, Conwy, 1 male, gravel workings; grazed grassland; parkland; gardens; 5.viii.1988, Holmes, Boyce & Reed leg. (PPWM). Ryton grounds of horticultural research station; open mon- Wood, Bubbenhall, Coventry, West Midlands, 1 male, tane habitats. Reared from nest of blue tit and dun- 25.viii.2002, J.E. Smith leg. (HRI). Temple, Berkshire, nock. iv–xi. 1 male, 3.iv.1934 (reared), Basden leg. (BMNH). West End Common, Esher, Surrey, 1 male, 17.vii.2002, Chandler leg. (PPCM). Woodwalton Fen, Cam- EPIDAPUS (EPIDAPUS) GRACILIS (WALKER, 1848) bridgeshire, 21 males, 7.vi.1940, Edwards leg. Synonyms: aptera (Kieffer, 1903); edwardsi Freeman, (BMNH). 1983; gracilis (Winnertz, 1853) [preocc.]; longicornis Ecological notes: found in broad-leaved and carr (Lengersdorf, 1941); pulicina (Frey, 1952). woodland; fen; rotten birch log with ants Myrmica Literature: Walker (1848): 109; Walker (1856): 55 [all ruginodis. Reared from rotten elm log and rotting as Sciara]; Freeman (1983a): 170; Freeman (1983b): wood. iv–viii. 26 [all in part as Epidapus edwardsi]; Laurence (1996): 85; Laurence (1997a): 87; Laurence & James EPIDAPUS (EPIDAPUS) ATOMARIUS (DE GEER, 1778) (1997): 166 [all in part as Epidapus atomarius, misidentification]; Menzel (1998): 21; Menzel & Mohrig Synonyms: acutehomerata (Venturi, 1964); brevipalpis (2000): 319; Withers (2002): 114. (Kieffer, 1903); degeerii (Bigot, 1854); melina (Schmitz, 1915); monocalcarata (Vimmer, 1928); Material examined: Balmoral Forest, Aberdeenshire, moravica (Czizek, 1917); pumila (Winnertz, 1853); 1 male, 1.viii.−4.viii.1937, Coe leg. (BMNH). Bradfield soudeki (Vimmer, 1926); venaticus Haliday in Walker, Wood, Suffolk, 1 female, x.1990; 6 males, v.1991; 2 1856. males, 3 females, 16.viii.−22.viii.1991, all Laurence leg. (PBLN). Belch-y-dwr, Gwynedd, 1 male, Literature: Haliday in Walker (1856): 56 [as Epidapus 14.vii.1989, Holmes, Boyce & Reed leg. (PPWM). Chig- venaticus]; Freeman (1983b): 26; Laurence (1996): 87 well Row Wood, Chigwell Row, Essex, 1 male, [in part]; Laurence (1997a): 85 [in part]; Laurence & 16.x.2002, Ismay & Schulten leg. (DEI). Cors Geirch, James (1997): 166 [in part]; Menzel (1998): 21; Menzel Gwynedd, 1 male, 7.x.1988, Holmes, Boyce & Reed leg. & Mohrig (2000): 315. (PPWM). Cwm Crafnant, Gwynedd, 1 male, 13.x.1988, Material examined: Becky Woods, Manaton, Devon, 1 Holmes, Boyce & Reed leg. (PPWM). Dickleburgh, male, 23.vi.2002, J.E. Smith leg. (HRI). Berwick-upon- Norfolk, 1 male, 7.v.1988, Withers leg. (PPWM). Tweed, Northumberland, 1 male, iv.1999, Laurence Ditchley, Oxfordshire, 1 male, v.1937, Edwards leg. leg. (PBLN). Bradfield Wood, Suffolk, 1 male, v.1991; 1 (BMNH). Gors Goch, Powys, 1 male, 18.vii.1989, male, 23.v.−30.v.1991; 1 male, 16.viii.−22.viii.1991, all Holmes, Boyce & Reed leg. (PPWM). Killaun Bog, Laurence leg. (PBLN). Brandon Marsh, Ryton-on- Offaly, 1 male, ix.2000, Withers leg. (PPWM). Llanon, Dunsmore, Coventry, West Midlands, 1 male, Swansea/Abertawe, 1 male, 13.vii.1989, Holmes,

Boyce & Reed leg. (PPWM). Luton Airport, Bedford- Ecological notes: found in broad-leaved woodland; carr shire, 1 male, 20.iv.−6.v.1999, C.W. Plant leg. (PPCM). woodland and fen; blanket bog remnant; bog and lake- New Close Wood, Brandon, Warwickshire, 1 male, shore; garden. Reared from beech detritus; the fungus 28.viii.2003, J.E. Smith leg. (HRI). Stanford, Norfolk, Hypholoma velutinum. iv–ix. 2 males, 27.iii.−10.iv.1988, Collier leg. (PPWM). Todd’s Green, Hertfordshire, 1 male, 4.v.1938, no collector detail (BMNH). No locality details, England, 1 male, EPIDAPUS (EPIDAPUS) SCHILLEI (BÖRNER, 1903) no date, Walker leg. [lectotype of Sciara gracilis] Synonyms: absoloni Czizek, 1915; titan Frey (1948). (BMNH). Literature: Edwards (1925a): 544 [as Epidapus atom- Ecological notes: found in broad-leaved woodland; arius, misidentification]; Menzel (1998): 21. conifer plantations; heathland (heather, Juncus, Des- Material examined: Balmoral Forest, Aberdeenshire, champsia) with ancient and secondary woodland; 1 male, 1.viii.−4.viii.1937, Coe leg. (BMNH). Bix Bot- blanket bog; Molinia bog; raised bog; ungrazed tom, Oxfordshire, 1 male, 23.ix.2000, Gibbs leg. heather; Schoenus nigricans, Sphagnum and Phrag- (PDGB). Blaen y Wergloedd, Conwy, 1 male, 13.x.1988, mites; grazed Molinia and Myrica; calcareous grass- Holmes, Boyce & Reed leg. (PPWM). Bradfield land. iii–v, vii–x. Wood, Suffolk, 1 male, 23.v.−30.v.1991, Laurence leg. (PBLN). Burnham, Buckinghamshire, 1 male, EPIDAPUS (EPIDAPUS) MICROTHORAX (BÖRNER, 1903) 15.ix.1934, Basden leg. (BMNH). Burnham Beeches, Buckinghamshire, 1 male, 14.vi.−17.vi.1996; 5 males, Synonyms: gracilicornis (Lengersdorf, 1926); intermit- 28.v.−26.xi.1996, all Ismay leg. (PPCM). Cadair Idris, tens Tuomikoski, 1959; obenbergeri (Vimmer, 1926); Gwynedd, 1 male, 13.xi.1993, Laurence leg. (PBLN). subdetrita (Kratochvil, 1936); subdetrita var. bispina Coed Tycanol, Pembrokeshire, 1 male, 29.vi.1994, (Kratochvil, 1936); subdetrita var. monospina (Kra- Levey & Pavett leg. (PPCM). Cors Erddreiniog, Isle of tochvil, 1936); subdetrita var. trispina (Kratochvil, Anglesey, 1 male, 27.vii.1988, Holmes, Boyce & Reed 1936). leg. (PPWM). Cors Geirch, Gwynedd, 1 male, 7.x.1988, Literature: Edwards (1925a): 544; Perry & Stubbs Holmes, Boyce & Reed leg. (PPWM). Cors Penally, (1978): 71 [all as Epidapus gracilis, misidentification]; Pembrokeshire, 1 male, 14.x.1987, Holmes, Boyce & Freeman (1983b): 26 [as Epidapus edwardsi, misiden- Reed leg. (PPWM). Dart Head, Devon, 1 male, tification]; Laurence (1997a): 85 [as Epidapus 1.vi.1920, Edwards leg. (BMNH). Epping Forest, edwardsi, misidentification]. Essex, 1 male, v.1999, Dagley leg. (PPCM). Farnham Royal, Buckinghamshire, 1 male, 18.v.1931 (reared), New to British list. Basden leg. (BMNH). Glasshouse Crops Research Sta- Material examined: Bradfield Wood, Suffolk, 1 male, tion, Littlehampton, West Sussex, 2 males, 23.x.1956, 9.iv.−16.iv.1990; 1 male, 23.v.−30.v.1991, all Laurence Hussey leg. (HRI). Glen Coiltie, Highland, 1 male, leg. (PBLN). Brandyquoi, Kirkwall, Orkney, 1 male, v.- 28.v.2002, Chandler leg. (PPCM). Glen Lochay, Perth vi.1999; 1 male, v.-vi.2001, all Laurence leg. (PBLN). & Kinross, 1 male, 3.vi.−18.vi.1932, Edwards leg. Cothill, Oxfordshire, 1 male, 1.v.1940, Edwards leg. (BMNH). Hay Wood, Baddesley Clinton, Solihull, West (BMNH). Crickhowell, Powys, 1 male, v.1937, Midlands, 20 males, 25.viii.2002, Menzel & J.E. Smith Edwards leg. (BMNH). Gors Maen-llwyd, Denbighs- leg. (DEI, HRI). Kingsbury Water Park, Warwickshire, hire, 1 female, 25.vi.1988, Holmes, Boyce & Reed leg. 1 male, 13.x.2001, J.E. Smith leg. (HRI). Lea Mead- (PPCM). Granish, Highland Region, 1 female, ows, Newtown Linford, Leicestershire, 1 male, 17.vi.1968, Aubrook leg. (BMNH). Letchworth, Hert- 24.viii.2003, Menzel leg. (DEI). Lion Wood, Redditch, fordshire, 1 male, v.1917; 1 male, vii.1917; 1 male, Worcestershire, 3 males, 10.iv.2003, J.E. Smith leg. ix.1936, all Edwards leg. (BMNH). Malham Tarn, (HRI). Llanbrynmair Moor, Powys, 1 male, 1.viii.1988, North Yorkshire, 1 male, 3.vi.−5.vi.1982, Disney leg. Holmes, Boyce & Reed leg. (PPWM). New Close Wood, (BMNH). New Malden, Greater London, 1 male, Brandon, Warwickshire, 4 males, 28.viii.2003, Menzel 31.v.1932, Coe leg. (BMNH). Sand, Shetland, 1 male, leg. (DEI). Ryton Wood, Bubbenhall, Coventry, West 11.vi.−12.vi.1998, Laurence leg. (PBLN). Sidmouth, Midlands, 2 males, 12.x.2001, J.E. Smith leg. (HRI); 5 Devon, 1 male, 17.viii.−21.viii.1926, Edwards leg. males, 25.viii.2002, Menzel & J.E. Smith leg. (DEI, (BMNH). Tewin, Hertfordshire, 1 male, ix.1922 HRI). Sned Wood, Lower Lye, Herefordshire, 3 males, (reared), Edwards leg. (BMNH). The Loke, Norwich, 14.vii.2003, J.E. Smith leg. (HRI). Stonebyres, Walker Norfolk, 1 male, 18.v.−23.v.1992; 1 male, 21.v.− Farm, South Lanarkshire, 1 male, 23.v.1977, no col- 23.v.1993, all Laurence leg. (PBLN). Walker Farm, lector detail (BMNH). Tocil Wood, Warwick Uni- Stonebyres, South Lanarkshire, 1 male, 23.v.1977, no versity Campus, Coventry, Warwickshire, 5 males, collector detail (BMNH). 28.viii.2003, Menzel & J.E. Smith leg. (DEI, HRI).

Warren Wood, Honingham, Norfolk, 1 male, 1.xi.1980, 25.vi.1987, Holmes, Boyce & Reed leg. (PPWM). Crym- Evans leg. (BMNH). Western Cleddau, Pem- lyn Bog, Swansea/Abertawe, 1 male, 13.vii.1988, brokeshire, 1 male, 23.vi.1987, Holmes, Boyce & Reed Holmes, Boyce & Reed leg. (DEI). leg. (PPWM). Whiteﬁeld, Greater Manchester, 1 male, Ecological notes: found on damp heath; Carex panicu- 23.vi.1932 (reared), Warwick leg. (BMNH). Wood lata and Molinia heath; valley fen; Molinia bog. v–vii. Crates, New Forest, Hampshire, 1 male, viii.1936, Edwards leg. (BMNH). Wytham Woods, Wytham, Taxonomic note: Until now, only the type of this spe- Oxfordshire, 2 males, 21.viii.2002, Menzel leg. (DEI). cies from North Korea was known (Mohrig, Menzel & Kozánek, 1992). In the British specimens the palpi Ecological notes: very common in broad-leaved wood- were sometimes only 2-segmented and in contradic- land; also found in mixed woodland; conifer plantation to the original description there were often only 3 tions; wooded lake margin; reed beds; grounds of strong spines of similar length on the apex of the gono- horticultural research station; calcareous grassland; styles [one dorsal spine almost on the outer angle; parkland, wet meadow; open montane habitats. with inward/downward-pointing spines arranged in 2 Reared from the fungus Stereum sanguinolentum and pairs subapically below]. nests of robin and dunnock. iv–xi.

HYPERLASION SCHMITZ, 1918 EPIDAPUS (EPIDAPUS) SUBGRACILIS MENZEL & MOHRIG SP. NOV. HYPERLASION WASMANNI SCHMITZ, 1918 New to British list. Literature: Laurence (1994): 108; Laurence (1997a): 85; Menzel (1998): 21 [as Cratyna (Cratyna)]. Material examined: Epping Forest, Monument Field, Essex, 1 male, 29.iii.1999, Dagley leg. [paratype of Material examined: Bradfield Wood, Suffolk, 1 male, 7 Epidapus subgracilis] (PPCM). females, 6.vii.−11.vii.1990; 1 female, vii.1990; 7 females, 16.viii.−22.viii.1991, all Laurence leg. (DEI, Ecological notes: found in grassland adjacent to PBLN). Burnham Beeches, Buckinghamshire, 9 ancient broad-leaved woodland. iii. males, 7 females, 20.vii.−16.viii.1995; 21 males, 2.viii.−26.viii.1995, all Ismay leg. (HRI, PPCM). Chig- well Row Wood, Chigwell Row, Essex, 22 males, 20 EPIDAPUS (PSEUDOAPTANOGYNA) ABSCONDITUS females, 15.ix.2002; 27 males, 15 females, 23.ix.2002; (VIMMER, 1926) 1 male, 3.x.2002; 1 male, 22.x.2002, all Ismay & Synonym: flabellata (Lengersdorf, 1942). Schulten leg. (DEI, UMO). Doncaster, South York- New to British list. shire, 53 males, 61 females, 30.vii.1993, Skidmore leg. (DEI). Epping Forest, Essex, 1 male, vii.1999, Dagley Material examined: Coed Tycanol, Pembrokeshire, 1 leg. (PPCM). Hatfield Moor, Doncaster, South York- male, 1999, Levey & Pavett leg. (PPCM). Epping For- shire, 28 males, 20.viii.1996, Skidmore leg. (DEI). est, Essex, 2 males, viii.1998; 1 male, vi.-vii.1999; 1 Hickling, Norfolk, 1 female, 15.viii.−29.viii.1990, Fos- male, ix.2000, all Dagley leg. (PPCM). Tocil Wood, ter & Procter leg. (PPCM). Linwood, New Forest, Warwick University Campus, Coventry, Warwickshire, Hampshire, 1 male, viii.1936, Edwards leg. (BMNH). 3 males, 28.viii.2003, Menzel leg. (DEI). Whitacre Lockhaugh Bank Wood, Derwent Valley, Tyne & Wear, Heath, Warwickshire, 3 males, 13.x.2001, J.E. Smith 25 females, 24.vii.−13.viii.1990, Laurence leg. (PBLN); leg. (HRI). 1 male, 14.vii.−13.viii.1990; 3 females, 24.vii.− Ecological notes: found in broad-leaved woodland; wil- 13.viii.1990, all Jessop leg. (PBLN). Newbold Comyn low carr with birch and reed beds; old sand and gravel Park, Leamington Spa, Warwickshire, 2 females, workings; wetland and grassland. vi–x. 29.vii.2001, J.E. Smith leg. (HRI). Nûg, Conwy, 1 male, 3.viii.1988, Holmes, Boyce & Reed leg. (PPWM). Ecological notes: common in broad-leaved and carr EPIDAPUS (PSEUDOAPTANOGYNA) ECHINATUM woodland; also found in young mixed woodland; MOHRIG & KOZÁNEK, 1992 heathland (heather, Juncus, Deschampsia); sedge bed; New to British and European list. raised bog; damp meadow. vii-x. Material examined: Blaen Cilieni, Powys, 1 male, Taxonomic note: Only ‘1919’ is printed on the cover of 5.vii.1989, Holmes, Boyce & Reed leg. (PPWM). the issue 1–2 of volume 61 of ‘Tijdschrift voor Ento- Cleddon Bog, Monmouthshire, 1 male, 24.v.1988; 1 mologie’, in which is the article by Hermann Schmitz male, 10.vi.1988, all Holmes, Boyce & Reed leg. ‘Neue Beiträge zur Kenntnis der Sciariden mit reduz- (DEI, PPWM). Cors Caranod, Ceredigion, 1 male, ierten Maxillarpalpen’, which is incorrect. On the 2nd

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 98 F. MENZEL ET AL. page of the cover there is the following entry that was LEPTOSCIARELLA (LEPTOSCIARELLA) CERIFERA not recorded to date: ‘Aﬂevering 1 en 2 (blz. 1–142) uit- MOHRIG & MENZEL, 1997 = gegeven 15 Juli 1918’ [ ‘Entries 1 and 2 (pages 1–142) Literature: Chandler et al. (2002): 123. issued on 15 July 1918’]. After that the original descriptions of Hyperlasion Schmitz und Hyperlasion Material examined: Afon Tywi, Carmarthen, 1 male, wasmanni Schmitz are correctly dated as 1918. 17.x.1995, McLean leg. (PPCM). Ban-y-Gor Wood, Gloucestershire, 1 male, 12.ix.2002, Gibbs leg. (PDGB). Crickley Hills Country Park, Gloucester- LEPTOSCIARELLA TUOMIKOSKI, 1960 shire, 1 male, 12.x.1979, Chandler leg. (PPCM). LEPTOSCIARELLA (HIRTIPENNIA) HIRTIPENNIS Dinefwr Park, Carmarthen, 1 male, 12.x.1995, Chan- (ZETTERSTEDT, 1838) dler leg. (PPCM). Dromore Forest, Kerry, 1 male, Synonyms: absurda (Winnertz, 1867); hirtipennis var. 15.x.1973, Chandler leg. (PPCM). Edge Hill, Warwick- minor (Frey, 1948); jugicola (Strobl, 1898); parcepilosa shire, 1 male, 14.x.2001, Chandler leg. (HRI). Fishpool var. opacicollis (Strobl, 1902). Valley, Herefordshire, 1 male, 12.x.1989, Chandler leg. (PPWM). Folly Farm, Avon, 1 male, 12.ix.2000, Gibbs Literature: Edwards (1925a): 533 [as Trichosia leg. (PDGB). Gearagh, Cork, 1 male, 2 females, absurda]; Edwards (1926b): 142 [as Trichosia]; Free- 26.ix.1990, de Courcy Williams leg. (DEI, NMID). man (1983b): 20 [as Trichosia (Leptosciarella) Hobb’s Quarry, Gloucestershire, 1 male, 4.ix.2002; 1 absurda]; King (1991): 43 [as Trichosia]; Mohrig & male, 15.v.2003, all Gibbs leg. (PDGB). Lancaut Menzel (1997): 46; Menzel (1998): 21. Nature Reserve, Gloucestershire, 1 male, 5.ix.2002, Material examined: Craigellachie, Aviemore, High- Gibbs leg. (PDGB). Longborough, Gloucestershire, 2 land Region, 1 male, 30.v.2002, Chandler leg. (PPCM). males, 8.iv.2003, J.E. Smith leg. (HRI). Monk Wood, Glen Glass, Highland Region, 1 male, 15.vi.1984, Avon, 1 male, 22.ix.2000, Gibbs leg. (PDGB). Ridley Vane-Wright leg. (PPCM). Hungerford, Berkshire, 1 Wood, Hampshire, 1 male, 15.iv.1949, Colyer leg. female, 14.v.1927, Hurst leg. (BMNH). Llanmynach (BMNH). Rough Hill Wood, Studley, Warwickshire, 1 Hills, Shropshire, 1 male, 23.v.1994, J.H. Cole leg. male, 10.iv.2003, J.E. Smith leg. (HRI). Scadbury (PJCB). New Forest, Hampshire, 1 female, Park, Greater London, 1 male, 22.x.1983, Chandler 23.viii.1902; 1 female, 23.v.1907, all Adams leg. leg. (PPWM). Stroat, Gloucestershire, 1 male, (BMNH). 13.x.1979, Stubbs leg. (NMS). Symond’s Yat, Hereford- shire, 2 males, ix.1936, Edwards leg. (BMNH). Ecological notes: found in birch and other broad- leaved woodland; grassland and scrub in old limestone Ecological notes: widespread in broad-leaved wood- quarry. v–vi, viii. land (incl. calcareous and neutral); parkland. iv–v, ix–x.

LEPTOSCIARELLA (HIRTIPENNIA) PARCEPILOSA (STROBL, 1900) LEPTOSCIARELLA (LEPTOSCIARELLA) DIMERA (TUOMIKOSKI, 1960) Synonym: quadristrigata (Strobl, 1909). New to British list. New to British list. Material examined: Epping Forest, Essex, 3 males, Material examined: Kelham Bridge, Ravenstone, 16.viii.−18.ix.1998; 4 males, v.–vii.1999; 2 males, Leicestershire, 1 female, 24.viii.2003, Menzel leg. vi.2000, all Dagley leg. (PPCM). Newbold Comyn (DEI). Park, Leamington Spa, Warwickshire, 4 males, Ecological notes: found in wet meadow with reed beds 29.vii.2001, J.E. Smith leg. (HRI). Windsor Forest, around pools. viii. Berkshire, 1 male, 6.ix.1986, Chandler leg. (PPWM). Ecological notes: found in ancient broad-leaved wood- LEPTOSCIARELLA (LEPTOSCIARELLA) BREVIPALPA land; young mixed woodland and damp meadow. v–ix. (MOHRIG & MENZEL, 1992)

New to British list. LEPTOSCIARELLA (LEPTOSCIARELLA) FUSCIPALPA Material examined: Milham Ford School, Oxford, (MOHRIG & MAMAEV, 1979) Oxfordshire, 1 male, 18.vi.−25.vi.2002, Webb leg. New to British and Irish list. (PPCM). Material examined: Badger Wood, Cambridgeshire, 1 Ecological notes: found in school grounds with old pop- male, vi.1998, C.W. Plant leg. (PPCM). Bagley Wood, lars. vi. Oxfordshire, 1 male, 20.v.−21.v.1933, Edwards leg.

(BMNH). Blair Atholl, Perth & Kinross, 1 male, Cambridgeshire, 2 males, 1.vi.−30.vi.1998, C.W. Plant 28.v.1973, Chandler leg. (NMS). Blean Wood, Kent, 1 leg. (PPCM). Oakley Inclosure, New Forest, Hamp- male, 30.iv.1965, Parmenter leg. (BMNH); 2 males, shire, 1 male, 17.iv.1953, Colyer leg. (BMNH). Reelig 19.v.1973, no collector detail (BMNH). Bromley, Glen, Moniack, Highland Region, 1 male, 20.v.1994, Greater London, 1 male, 15.v.1971, Chandler leg. Chandler leg. (PPCM). Rosemarkie, Highland Region, (PPCM). Buckingham Thick Copse, Northampton- 1 male, 29.v.2002, Merrifield leg. (PPCM). Sannox, shire, 1 male, 9.vi.1992, Warne leg. (PPCM). Burnham Arran, North Ayrshire, 1 male, 26.v.−28.v.1919, Beeches. Buckinghamshire, 2 males, 6.v.1914, Edwards leg. (BMNH). Savernake Forest, Wiltshire, 4 Edwards leg. (BMNH); 3 males, 21.iv.1998, Chandler males, 3.v.1993, Chandler leg. (PPCM). Selsdon, leg. (PPCM). Cambridge, Cambridgeshire, 1 male, Greater London, 1 male, 16.v.1932, Coe leg. (BMNH). 12.v.1913, Edwards leg. (BMNH). Chigwell Row Wood, Sheep Leas, West Horsley, Surrey, 1 male, 18.v.1991, Chigwell Row, Essex, 1 female, 11.vi.2002; 1 female, Chandler leg. (PPCM). Shortwood, Gloucestershire, 18.vi.2002; 1 female, 28.vi.2002; 1 male, 26.ix.2002, all 1 male, 12.vi.2002, Gibbs leg. (PDGB). Sidmouth, Ismay & Schulten leg. (DEI). Clothall, Hertfordshire, Devon, 1 male, 10.v.1936; 1 male, 1.vi.1938, all 1 male, 4.v.1918, Edwards leg. (BMNH). Clovelly, Edwards leg. (BMNH). Skinner’s Pastures, Cumbria, 2 Devon, 7 males, 8.v.1936, Edwards leg. (BMNH). males, 17.vi.1999, Chandler leg. (PPCM). Spreacombe Coombe, Cornwall, 1 male, 7.vii.1986, Chandler Chapel Wood, Devon, 1 male, 6.vii.1986, Chandler leg. leg. (NMS). Corrimony, Highland Region, 1 male, (PPCM). Staffhurst Wood, Surrey, 2 males, 18.v.− 28.v.2002, Chandler leg. (PPCM). Craigellachie, Avi- 19.v.1991, Chandler leg. (PPCM). Stonybrow Wood, emore, Highland Region, 1 male, 30.v.2002, Chandler Rotherfield Park, Hampshire, 1 male, 3.vii.2002, leg. (PPCM). Crickhowell, Powys, 1 male, v.1937, Chandler leg. (PPCM). Tarradale House, Highland Edwards leg. (BMNH). Cuckoo Wood, Downe, Greater Region, 1 male, v.2002, Sumner leg. (PPCM). Tynron, London, 1 male, 23.vi.1979, Chandler leg. (NMS). Dumfries & Galloway, 1 male, 26.v.1945, Duncan leg. Dinton Pastures Country Park, Berkshire, 1 male, (NMS). Verwood, Dorset, 1 male, 15.iv.1953, Colyer 25.iv.1993; 1 male, 20.ix.1998, all Chandler leg. leg. (BMNH). Windsor Forest, Berkshire, 1 male, (PPCM). Ditchley, Oxfordshire, 2 males, v.1937, 8.vi.1979; 1 male, 21.v.1988; 1 male, 15.vi.1991, all Edwards leg. (BMNH). Ebbor Gorge, Somerset, 1 Chandler leg. (NMS, PPCM). Wytham Woods, male, 3.vii.1985, Chandler leg. (PPCM). Epping For- Wytham, Oxfordshire, 1 male, 1 female, 21.viii.2002, est, Essex, 7 males, 1 female, 19.iv.−25.iv.1999, Dagley Menzel leg. (DEI). leg. (HRI, PPCM); 1 male, 24.v.1999, Chandler leg. Ecological notes: very common in broad-leaved, mixed (PPCM). Eyeworth Wood, New Forest, Hampshire, 1 and carr woodland; ancient woodland with oak, hazel male, 6.vi.1987, Chandler leg. (PPCM). Falls of Tar- and bramble; pine woodland; heathland (heather) nash, Moray, 1 male, 1.vii.1936, Coe leg. (BMNH). with oak and holly; sedge bed; lakeshore marsh; Gearagh, Cork, 1 male, 22.iv.1990, de Courcy Williams wooded river bank; water meadows and gravel pits, leg. (NMID). Glen Coiltie, Highland Region, 1 male, old hedges; calcareous grassland; garden. iv–ix. 28.v.2002, Chandler leg. (PPCM). Glen Lochay, Stirling, 3 males, 3.vi.−8.vi.1932, Edwards leg. (BMNH). Gunnerside, North Yorkshire, 1 male, LEPTOSCIARELLA (LEPTOSCIARELLA) MELANOMA 27.vi.1981, Chandler leg. (PPCM). Harrow, Greater (MOHRIG & MENZEL, 1990) London, 1 male, 24.v.1914, Edwards leg. (BMNH). Literature: Mohrig & Menzel (1997): 66; Menzel Hartslock, Oxfordshire, 1 male, 6.v.2000; 2 males, (1998): 21; Chandler et al. (2002): 123. 14.v.2000, all Raper leg. (PDGB). Hickling, Norfolk, 1 male, 8.vi.−22.vi.1989, Foster & Procter leg. (PPCM). Material examined: Bagley Wood, Oxfordshire, 1 male, Hod Hill, Dorset, 1 male, 22.v.1968, Parmenter leg. 18.v.1936, Edwards leg. (BMNH). Ballyvourney, Cork, (BMNH). Holker Moss, Cumbria, 1 male, 11.vii.− 1 male, 10.vii.1990, Withers leg. (PPWM). Burnham 13.vii.1923, Edwards leg. (BMNH). Ingleton, North Beeches, Buckinghamshire, 4 males, 18.v.−29.v.1999; 8 Yorkshire, 1 male, 20.vi.1924, Edwards leg. (BMNH). males, 25.v.−1.vi.1999, all Ismay leg. (HRI, PPCM); Letchworth, Hertfordshire, 3 males, vi.1917; 1 male, 1 male, 1.vi.−8.vi.1999, Chandler leg. (HRI). Culbin iv.1918, all Edwards leg. (BMNH). Linn of Dee, Aber- Sands, Highland Region, 1 male, 5.viii.−7.viii.1936, deenshire, 1 male, 31.v.1998, Chandler leg. (PPCM). Coe leg. (BMNH). Epping Forest, Essex, 1 male, Loch Chon, Stirling, 1 male, 22.vi.1999, Chandler leg. vi.2000, Dagley leg. (PPCM). Falls of Tarnash, Moray, (NMS). Loch Meiklie, Highland Region, 1 male, 1 male, 1.vii.1936, Coe leg. (BMNH). Glen of Drum- 13.vi.1984, Chandler leg. (PPCM). Lodge Wood, Berk- loch, Moray, 1 male, 10.vii.1936, Coe leg. (BMNH). shire, 1 male, 2.v.1994, Chandler leg. (PPCM). Glen of Knocknarea, Sligo, 1 male, 1.x.1977, Chandler Monkfield Wood, Cambridgeshire, 1 male, 1.v.− leg. (PPCM). Lea Meadows, Newtown Linford, Leices- 31.v.1998, C.W. Plant leg. (NMS). Oakland Wood, tershire, 1 male, 24.viii.2003, Menzel leg. (DEI). Loch

Park, Moray, 2 males, 3.vii.1936, Coe leg. (BMNH). langwell, Highland Region, 3 males, 29.v.2002, Raindale, North Yorkshire, 1 male, 19.vii.1996, God- Chandler leg. (PPCM). Brodick, Arran, North Ayr- frey leg. (PPCM). Scadbury Park, Greater London, 1 shire, 1 male, 22.v.−25.v.1919, Edwards leg. [paratype male, 18.vi.1983, Chandler leg. (PPCM). The Ritec, of Sciara subspinulosa] (BMNH). Charleville Wood, Pembrokeshire, 1 male, 22.vii.1987, Holmes, Boyce & Offaly, 3 males, 15.vi.1985, Chandler leg. (PPWM). Reed leg. (PPWM). Wheatfen Broad, Norfolk, 1 male, Cilkenny Dingle, Powys, 1 male, 22.v.1979, Chandler 19.v.1990, Withers leg. (PPWM). Woodwalton Fen, leg. (PPCM). Devil’s Glen, Wicklow, 1 male, 10.vi.1985, Cambridgeshire, 1 male, 24.v.1940, Edwards leg. Chandler leg. (PPCM). Feshiebridge, Highland (BMNH). Region, 1 male, 29.v.1975, Chandler leg. (PPCM). Gearagh, Cork, 1 male, 9.vii.1990, Withers leg. Ecological notes: common in broad-leaved woodland; (PPWM). Hickling Broad, Norfolk, 1 male, 3.viii.1988, mixed woodland; dunes and conifer plantations; Withers leg. (PPWM). Lancaut Nature Reserve, Glouc- wooded gorge; fen and alder carr; Phragmites; wet estershire, 1 male, 28.v.2003, Gibbs leg. (PDGB). meadow. v–viii, x. Migdale Wood, Highland Region, 1 male, 31.v.2002, Chandler leg. (PPCM). Morrone Birkwood, Aberdeen- LEPTOSCIARELLA (LEPTOSCIARELLA) NIGROSETOSA shire, 1 male, 31.v.1998; 1 male, 14.vi.2003, all Chan- (FREEMAN, 1990) RESTIT. dler leg. (NMS, PPCM). Phoenix Park, Dublin, 2 Synonym: truncatula Mohrig & Menzel, 1997 syn. nov. males, 24.viii.1994 (reared), Dyte leg. (PPCM). Rhôsili Down, Swansea/Abertawe, 1 male, 28.vi.1989, Literature: Freeman (1990): 54; Laurence (1997a): 85 Holmes, Boyce & Reed leg. (PPWM). Sannox, Arran, [all as Corynoptera]; Mohrig & Menzel (1997): 80; North Ayrshire, 1 male, 26.v.−28.v.1919, Edwards leg. Menzel (1998): 22 [all as Leptosciarella truncata, mis- [holotype of Sciara subspinulosa] (BMNH). Tarradale identification and incorrectly synonymized from Lep- House, Highland Region, 1 male, v.2002, Sumner leg. tosciarella nigrosetosa to Leptosciarella truncata]. (PPCM). Tummel Forest, Perth & Kinross, 1 male, Material examined: Bradfield Wood, Suffolk, 3 males, 28.v.1973, Chandler leg. (NMS). − 23.v. 30.v.1991, Laurence leg. (PBLN). Monk’s Wood, Ecological notes: widespread in broad-leaved (incl. cal- Cambridgeshire, 1 male, 18.vi.1978, J.H. Cole leg. careous) and carr woodland; also found in mixed wood- [holotype of Corynoptera nigrosetosa] (BMNH). land; conifer plantations; pine woodland; damp heath; Ecological notes: found in broad-leaved woodland; sedge bed; wooded river bank; parkland with old trees. ancient woodland on clay. v–vi. Reared from debris in rot hole in beech. v–viii. Taxonomic note: Mohrig & Menzel (1997) incorrectly Taxonomic note: Chandler et al. (2002) wrote that synonymized Corynoptera nigrosetosa Freeman with pilosa (Staeger) and subspinulosa (Edwards) were dif- Leptosciarella truncata (Tuomikoski). Corynoptera ferent species and that the synonymy by Mohrig & nigrosetosa Freeman is identical to Leptosciarella Menzel (1997) is not correct. The authors based their truncatula Mohrig & Menzel. The result of the com- diagnosis on two characteristics: (a) the macrotrichia parison of the types for this Leptosciarella-species is on the wing veins x and y; (b) the presence of a number that ‘nigrosetosa (Freeman, 1990)’ is the correct name of awl-like spines on the inner side on the gonostyles and ‘truncatula Mohrig & Menzel, 1997’ will be a syn- below the tooth. Chandler et al. found fairly constant onym of nigrosetosa (Freeman). characteristics during their examination of a relatively small number of examples from the British Isles. A check of both characteristics in 26 males from LEPTOSCIARELLA (LEPTOSCIARELLA) PILOSA Germany, England, Austria and the Czech or Slovak (STAEGER, 1840) republics has shown that there is strong variation in Synonyms: elegans var. defecta (Strobl, 1910); sub- both characteristics and that even on one example spinulosa (Edwards, 1925a). alone there can be distinct differences. Examination of the Types of pilosa (Staeger) and subspinulosa Literature: Edwards (1925a): 536 [as Sciara subspinu- (Edwards) as well as the DEI material found that the losa]; Freeman (1983b): 20 [as Trichosia (Leptoscia- x–y macrotrichia varied from ‘complete’ to ‘totally rella) subspinulosa]; Chandler et al. (1996): 258 [as absent’ with every variation between. In addition, the Trichosia (Leptosciarella) verburyi [recte yerburyi], subapical spines below the apical tooth ranged from misidentification]; Mohrig & Menzel (1997): 72; Men- the presence of 4–5 pale spines together with several zel (1998): 21; Chandler et al. (2002): 123 [as Leptos- stronger bristles, to a denser group of about 10–15 ciarella pilosa and Leptosciarella subspinulosa]. dark spines. In preparations where the genitalia lie at Material examined: Balnaguard Burn, Perth & Kin- an angle, the spines on the gonostyles appear to be ross, 1 male, 1.vi.1998, Chandler leg. (NMS). Brae- almost totally absent. However, the spines are present

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 BRITISH SCIARID FAUNA 101 but lie on the side. They can be seen under high mag- (NMS). Clifton, Cumbria, 1 male, 2.vi.1973, Chandler nification. The correlation between the presence of leg. (PPCM). Cogley Wood, Somerset, 1 male, macrotrichia on x and y with the presence of only a few 19.x.1986, Stubbs leg. (NMS). Coldborough Park, subapical spines on the gonostyles and, likewise, the Herefordshire, 1 male, 31.v.1972, Chandler leg. absence of macrotrichia on y with the presence of (PPCM). Coom Wood, Cork, 1 male, 10.vii.1990, With- strong spines on the inner side of the gonostyles as ers leg. (PPWM). Cop Mere, Staffordshire, 1 male, stated by Chandler et al. (2002) is not always the case. 23.v.1994, Chandler leg. (PPCM). Coppets Wood, For identification of the species, it is established that Greater London, 1 male, 21.vi.1991, K.G.V. Smith the gonostyles (Mohrig & Menzel, 1997: 74, fig. 21a) leg. (PBLN). Corfheath Firs, Isle of Wight, 1 male, are distinctive. On the basis of the variability 2.vii.1980, Chandler leg. (PPCM). Cothill, Oxford- described, subspinulosa (Edwards) is here confirmed shire, 1 male, 3.vi.1990, Chandler leg. (NMS). Craw- as a synonym of pilosa (Staeger) and the error in the fordsburn Country Park, Down, 1 male, 3.vii.1987, key in Mohrig & Menzel (1997) is corrected. Chandler leg. (NMS). Crickhowell, Powys, 1 male, v.1937, Edwards leg. (BMNH). Dinton Pastures Country Park, Berkshire, 1 male, 28.v.1995; 1 male, LEPTOSCIARELLA (LEPTOSCIARELLA) REJECTA 6.v.1997, all Chandler leg. (PPCM). Ditchley, Oxford- (WINNERTZ, 1867) shire, 2 males, v.1937, Edwards leg. (BMNH). Eye- Synonyms: echinata (Lengersdorf, 1926); elongata worth Wood, New Forest, Hampshire, 1 male, (Winnertz, 1867) [preocc.]; hispida (Winnertz, 1871). 2.vi.1985, Chandler leg. (NMS). Epping Forest, Essex, 1 male, 19.iv.1999; 3 males, 3.v.−9.v.1999, all Dagley Literature: Edwards (1925a): 536 [as Sciara pilosa, leg. (HRI, PPCM); 2 males, 25.v.−26.v.1999, Chandler misidentification]; Perry & Stubbs (1978): 71 [as Tri- leg. (NMS). Falls of Tarnash, Moray, 3 males, chosia pilosa, misidentification]; Freeman (1983b): 21 1.vii.1936, Coe leg. (BMNH). Farnborough Park, War- [as Trichosia (Leptosciarella) pilosa, misidentifica- wickshire, 1 female, 10.x.2001, Chandler leg. (HRI). tion]; Skidmore (1991): 149; Chandler et al. (1996): Felbrigg Wood, Norfolk, 1 male, 10.vi.1993, Chandler 258 [all as Trichosia pilosa, misidentification]; Mohrig leg. (PPCM). Ffrwd Fen, Carmarthen, 1 male, & Menzel (1997): 63; Menzel (1998): 21; Chandler 13.x.1995, Chandler leg. (PPCM). Foxhole, Highland (2001): 201; Chandler et al. (2002): 123. Region, 1 male, 28.v.2002, Chandler leg. (PPCM). Material examined: Abbot’s Wood, Hampshire, 1 male, Frilford Heath, Oxfordshire, 1 male, 25.v.−15.vi.1991, 25.v.1992, Chandler leg. (NMS). Annahala bog, Gear- Porter leg. (NMS). Gearagh, Cork, 2 males, 7.vi.1990, agh, Cork, 4 males, 1.vi.1990, de Courcy Williams leg. de Courcy Williams leg. (DEI, NMID). Glen Lochay, (DEI, NMID). Ashridge Estate, Hertfordshire, 1 male, Stirling, 1 male, 3.vi.−18.vi.1922, Edwards leg. 20.v.1999; 1 male, 8.vii.1999; 1 male, 8.x.1999, all (BMNH). Glendalough, Wicklow, 1 male, 8.vii.1969, Chandler leg. (NMS). Aviemore, Highland Region, 1 Chandler leg. (PPCM). Goring-on-Thames, Oxford- male, 24.vi.−29.vi.1933, Coe leg. (BMNH). Badger shire, 1 male, 7.vi.1962, J.H. Cole leg. (PJCB). Grant- Wood, Cambridgeshire, 1 male, 1.v.−31.v.1998, C.W. own-on-Spey, Highland Region, 1 male, 15.vi.1984, Plant leg. (NMS). Bedford Purlieus, Cambridgeshire, 1 Chandler leg. (PPWM). Great Ellerside, Cumbria, 1 male, 14.v.1988, J.H. Cole leg. (PJCB). Bix Bottom, male, 15.vi.1999, Chandler leg. (NMS). Greywell Fen, Oxfordshire, 1 male, 9.v.1998, Chandler leg. (NMS); 1 Hampshire, 1 male, 3.vii.1992, Chandler leg. (NMS). male, 23.ix.2000, Raper leg. (PDGB). Blair Atholl, Hartslock, Oxfordshire, 1 male, 14.v.2000, Raper leg. Perth & Kinross, 1 male, 28.v.1973, Chandler leg. (PDGB). High Dam, Cumbria, 1 male, 15.vi.1999, (PPCM). Brampton Wood, Cambridgeshire, 1 male, Chandler leg. (PPCM). Hobb’s Quarry, Gloucester- 9.v.1987, J.H. Cole leg. (PJCB). Brandyquoi, Kirkwall, shire, 1 male, 15.v.2003, Gibbs leg. (PDGB). Holme Orkney, 4 males, 1 female, v.-vi.2001, Laurence leg. Fen, Cambridgeshire, 2 males, 27.iv.1987, J.H. Cole (PBLN). Bromley, Greater London, 1 male, 15.v.1971, leg. (PJCB). Ilfracombe, Devon, 1 male, 6.vii.1986, Chandler leg. (PPCM). Burnham Beeches, Bucking- Chandler leg. (NMS). Killarney, Kerry, 2 males, 25.v.− hamshire, 1 male, 15.v.1972, Chandler leg. (PPCM). 27.v.1929, Edwards leg. (BMNH). Knebworth, Hert- Caerhays Castle, Cornwall, 1 male, 7.vii.1983, Chan- fordshire, 1 male, vi.1922, Edwards leg. (BMNH). dler leg. (PPCM). California Country Park, Berkshire, Leckford Estate, Hampshire, 1 male, 9.v.1975; 1 male, 1 male, 15.vi.1996, Chandler leg. (PPCM). Castlerosse, 14.vi.1987, all Chandler leg. (NMS, PPCM). Letch- Kerry, 1 male, 2.v.1981, Chandler leg. (PPCM). Cata- worth, Hertfordshire, 1 male, v.1917; 1 male, vi.1917; col, Arran, North Ayrshire, 1 male, 29.v.−30.v.1919, 1 male, v.1918, all Edwards leg. (BMNH). Littlehemp- Edwards leg. (ZFMK); 1 male, ix.1929, J. Waterston ston, Devon, 1 male, 3.vi.1974, Chandler leg. (NMS). leg. (BMNH). Charleville Wood, Offaly, 1 male, Maen Porth, Cornwall, 1 male, 5.x.1983, Chandler leg. 15.vi.1985, Chandler leg. (PPWM). Chippenham Fen, (NMS). Malham Tarn, North Yorkshire, 1 male, 3.vi.− Cambridgeshire, 1 male, 2.vii.1988, Chandler leg. 5.vi.1982; 1 male, 27.vi.−29.vi.1982, all Disney leg.

(PPWM). Monk Wood, Avon, 1 male, 24.v.2000, Gibbs Berkshire, 1 male, 8.vi.1979; 1 male, 30.v.1987; 1 leg. (PDGB). Monkﬁeld Wood, Cambridgeshire, 1 male, 13.vi.1987; 2 males, 15.vii.1991; 1 male, male, 1.v.−31.v.1998, C.W. Plant leg. (PPCM). Monk’s 15.v.1994; 1 male, 21.v.1998; 1 male, 15.vii.1999, all Wood, Muckross, Kerry, 3 males, 4.v.−7.v.1981, Chan- Chandler leg. (NMS, PPCM). Witherslack, Cumbria, 1 dler leg. (PPCM). Morrone Birkwood, Aberdeenshire, 1 male, 15.vi.1999, Chandler leg. (NMS). Woodwalton male, 31.v.1998, Chandler leg. (PPCM). Murrough, Fen, Cambridgeshire, 3 males, 24.v.1940, Edwards leg. Wicklow, 1 male, vi.1989, Blackith leg. (PPWM). New (BMNH). Wychwood, Oxfordshire, 3 males, v.1937, Forest, Hampshire, 4 males, 17.x.1937, Edwards Edwards leg. (BMNH). leg. (BMNH). New Malden, Greater London, 1 male, Ecological notes: very common in all types of wood- 15.vi.1932, Coe leg. (BMNH). Newmarket, Cam- land; wetlands including fen, sedge beds, reed beds, bridgeshire, 1 male, 5.ix.1894, Verrall leg. (UMO); 1 bogs and water meadows; gravel pits; old hedges; neu- male, 18.v.1908, Collin leg. (UMO). Oakland Wood, tral and calcareous grassland; gardens; on window. ii– Cambridgeshire, 2 males, 1.v.−30.vi.1998, C.W. Plant x. leg. (PPCM). Old Slade Wood, Buckinghamshire, 1 male, 12.v.1972 (reared), Chandler leg. (PPCM). Pass of Killiecrankie, Perth & Kinross, 1 male, 31.viii.1984, LEPTOSCIARELLA (LEPTOSCIARELLA) SCUTELLATA Chandler leg. (PPCM). Pearman’s Copse, Berkshire, 2 (STAEGER, 1840) males, 8.v.1995, Chandler leg. (PPCM). Penzance, Synonyms: bilineata (Staeger, 1840); elegans (Win- Cornwall, 1 male, 14.v.1940, Collin leg. (UMO). nertz, 1867); inhonesta (Winnertz, 1867); interrupta Rhôs Pwllygawnen, Carmarthen, 1 male, 11.vii.1989, (Strobl, 1895); obscuripennis (Winnertz, 1867). Holmes, Boyce & Reed leg. (PPWM). Riverstick, Cork, 1 male, 11.vii.−15.vii.1994, Withers leg. (PPWM). Literature: Mohrig & Menzel (1997): 59; Menzel Rotherﬁeld Park, Hampshire, 4 males, 2.x.2002, (1998): 22; Chandler et al. (2002): 123. Chandler leg. (PPCM). Rydal Water, Cumbria, 1 male, New to Irish list. 1.vi.1968, Hutson leg. (BMNH). Sannox, Arran, North Ayrshire, 1 male, 26.v.−28.v.1919, Edwards Material examined: Alice Holt Forest, Hampshire, 1 leg. (BMNH). Savernake Forest, Wiltshire, 1 male, male, 25.v.1991, Chandler leg. (PPCM). Annahala bog, 31.v.1990, Porter leg. (PPCM). Scarning Fen, Norfolk, Gearagh, Cork, 1 male, 10.viii.1990, de Courcy Will- 1 male, 8.vi.−22.vi.1988, Foster & Procter leg. iams leg. (NMID). Ashridge Estate, Hertfordshire, 3 (PPWM). Sheep Leas, West Horsley, Surrey, 1 male, males, 8.x.1999, Chandler leg. (PPCM). Ballielogue, 18.v.1991, Chandler leg. (NMS). Shortwood, Glouces- Kilkenny, 1 male, 25.vi.1987, Chandler leg. (PPCM). tershire, 2 males, 12.vi.2002, Gibbs leg. (PDGB). Becky Woods, Manaton, Devon, 1 male, 1 female, Spreacombe Chapel Wood, Devon, 1 male, 6.vii.1986, 23.vi.2002, J.E. Smith leg. (HRI). Berry, Devon, 1 Chandler leg. (PPCM). Staffhurst Wood, Surrey, 1 male, 27.vii.1953, Colyer leg. (BMNH). Birks of Aber- male, 18.v.1991, Chandler leg. (NMS). Strathrory, feldy, Perth & Kinross, 1 male, 30.viii.1987, Chandler Highland Region, 1 male, 15.vi.1976, Chandler leg. leg. (NMS). Bishop Wood, North Yorkshire, 1 male, (PPCM). Stroat, Gloucestershire, 1 male, 13.x.1979, 20.vii.1997, Chandler leg. (PPCM). Box Hill, Surrey, 1 Stubbs leg. (PPCM). The Clappers, Sharpenhoe, Bed- male, 11.vii.1992, Chandler leg. (NMS). Brockenhurst, fordshire, 1 male, 16.vi.1997, Chandler leg. (PPCM). Hampshire, 1 male, 10.ix.1977, Chandler leg. (NMS). The Loke, Norwich, Norfolk, 3 males, 26.iv.1987; 1 Bucklebury Common, Berkshire, 1 male, 27.v.1990; 1 male, 5.v.1988; 2 males, 5.vi.−13.vi.1989; 1 male, male, 6.vi.1993, all Chandler leg. (NMS, PPCM). 28.iv.−4.v.1990; 2 males, 15.v.−18.v.1992, all Laurence Burnham Beeches, Buckinghamshire, 1 male, 28.iv.− leg. (PBLN). The Mound, Highland Region, 1 male, 10.v.1995; 1 male, viii.-ix.1996, all Ismay leg. (NMS). 27.vii.1936, Collin leg. (UMO). Thornham Magna, Suf- Cadnam, New Forest, Hampshire, 1 male, 17.vi.1990, folk, 1 male, 25.iv.1991, Withers leg. (PPWM). Thur- Chandler leg. (PPCM). California Country Park, Berk- garton, Norfolk, 1 male, 29.v.1991, Carr leg. (BMNH). shire, 1 male, 6.vii.1997; 1 male, 22.v.1999, all Chan- Vidlin, Shetland, 2 males, 16.vi.−21.vi.1993, Laurence dler leg. (PPCM). Charleville Wood, Offaly, 1 male, leg. (PBLN). Wellesbourne, Warwickshire, 1 male, 27.v.1984, Chandler leg. (PPCM). Charterhouse Alder 24.ii.−2.iii.2000, J.E. Smith leg. (HRI). Westbury Holt, Surrey, 1 male, 16.vii.1989, Chandler leg. (NMS). Wood, Hertfordshire, 1 male, 14.viii.1918, Edwards Cliveden, Buckinghamshire, 1 male, 3.vii.1984, Chan- leg. (BMNH). Weston Common, Hampshire, 1 male, dler leg. (NMS). Devil’s Punchbowl, Surrey, 1 male, 10.v.1984, Chandler leg. (NMS). Weston Green Fen, 23.vi.1990, Chandler leg. (NMS). Dingwall, Highland Oxfordshire, 1 male, 1.vi.1991, Chandler leg. (PPCM). Region, 1 male, 8.viii.1909, King leg. (BMNH). Epping Wheatfen Broad, Norfolk, 1 male, 19.v.1990, Withers Thicks, Epping Forest, Essex, 4 males, 9.v.−16.v.1999, leg. (PPWM). Wicken Fen, Cambridgeshire, 2 males, Dagley leg. (HRI). Fingle Bridge, Devon, 1 male, 12.v.1990, Withers leg. (PPWM). Windsor Forest, 8.vii.1992, Chandler leg. (PPCM). Frithsden Beeches,

Hertfordshire, 3 males, 6.vi.−30.vii.1997, Chandler 17.vii.1987, C.W. Plant leg. (NMS); 1 male, 7.viii.1988, leg. (PPCM). Fungarth, Perth & Kinross, 1 male, Chandler leg. (PPCM). Wytham Woods, Wytham, 30.v.1998, Chandler leg. (PPCM). Gearagh, Cork, 1 Oxfordshire, 1 male, 21.viii.2002, Menzel leg. (DEI). male, 31.vi.1990; 1 male, 18.ix.1990, all de Courcy Ecological notes: very common in all types of wood- Williams leg. (DEI, NMID). Glasdrum, Argyll & Bute, land; wetlands including fen, sedge bed, lakeshore 1 male, 7.vi.1974, Chandler leg. (NMS). Glen marsh, Juncus and Molinia bog. v–x. Coiltie, Highland Region, 1 male, 24.vii.1997; 1 male, 23.vi.1999, all Chandler leg. (PPCM). Grandtully, Perth & Kinross, 1 male, 30.v.1998, Chandler leg. (NMS). Grantown-on-Spey, Highland Region, 1 male, LEPTOSCIARELLA (LEPTOSCIARELLA) SUBPILOSA 15.vi.1984, Chandler leg. (PPWM). Grondra Great (EDWARDS, 1925) Wood, Monmouthshire, 1 male, 9.vii.1986, Chandler Synonym: subelegans (Mohrig & Mamaev, 1985). leg. (NMS). Haugh Wood, Herefordshire, 1 male, Literature: Edwards (1925a): 536; Edwards (1926b): 13.vi.1997, M.N. Pugh leg. (PPCM). Hay Wood, Bad- 142 [all as Sciara]; Freeman (1983b): 21 [as Trichosia desley Clinton, Solihull, West Midlands, 1 male, (Leptosciarella)]; Laurence 1994: 118; Laurence 25.viii.2002, Menzel leg. (DEI). Hickling Broad, Nor- (1996): 87; Laurence (1997a): 85 [all as Trichosia folk, 1 male, 3.viii.1988, Withers leg. (PPWM). Knole pilosa, misidentification]; Mohrig & Menzel (1997): 61; Park, Kent, 1 male, 8.x.1972, Chandler leg. (NMS). Menzel (1998): 22; Menzel & Mohrig (2000): 366. Loch Chon, Stirling, 1 male, 22.vi.1999, Chandler leg. (NMS). Loch Tummel, Perth & Kinross, 1 male, Material examined: Ale Water, Ancrum, Scottish Bor- 30.vi.1937, Collin leg. (BMNH). Lockhaugh Bank, ders, 1 male, 11.vii.1988, J.H. Cole leg. (PJCB). Derwent Valley, Tyne & Wear, 1 male, 7.vi.− Archers Wood, Cambridgeshire, 1 male, 27.v.1990, 23.vii.1990, Jessop leg. (PBLN). Monk Wood, Avon, 1 J.H. Cole leg. (PJCB). Aunt Mary’s Bottom, Dorset, 2 male, 24.v.2000, Gibbs leg. (PDGB). Newcastle, Down, males, 28.vi.1998, Chandler leg. (PPCM). Aviemore, 1 male, 34.viii.1917, King leg. (BMNH). Oakland bank of Spey, Highland Region, 1 male, 9.vi.1984, Wood, Cambridgeshire, 1 male, 1.vi.−30.vi.1998, C.W. Chandler leg. (PPCM). Beltra Lough, Sligo, 1 male, Plant leg. (NMS). Odiham Common, Hampshire, 1 17.vi.1985, Chandler leg. (PPCM). Bradfield Wood, male, 7.vii.1990, Chandler leg. (NMS). Rhôs Pwllyga- Suffolk, 1 male, 6.vii.−11.vii.1990, Laurence leg. wnen, Carmarthen, 1 male, 11.vii.1989, Holmes, (PBLN). Brigsteer Park, Cumbria, 1 male, 14.vi.1999, Boyce & Reed leg. (PPWM). Rosemarkie, Highland Chandler leg. (PPCM). Buckingham Thick Copse, Region, 2 males, 23.vii.1997, Chandler leg. (PPCM). Northamptonshire, 1 male, 16.v.1992, Warne leg. Rotherfield Park, Hampshire, 1 male, 2.x.2002, Chan- (PPCM). Burnham Beeches, Buckinghamshire, 1 dler leg. (PPCM). Savernake Forest, Wiltshire, 1 male, male, 27.vi.1993, Chandler leg. (PPCM); 1 male, 4.vii.−25.vii.1990, Porter leg. (NMS). Scadbury Park, 5.viii.−31.viii.1996, Ismay leg. (PPCM). Chigwell Row Greater London, 1 male, 18.vi.1983; 1 male, Wood, Chigwell Row, Essex, 4 males, 2 females, 23.v.− 21.vii.1985, all Chandler leg. (NMS). Sned Wood, 27.v.2002; 1 male, 11.vi.2002, all Ismay & Schulten Lower Lye, Herefordshire, 2 males, 14.vii.2003, J.E. leg. (DEI, UMO). Culbin Forest, Highland Region, 2 Smith leg. (HRI). The Lythes, Selborne, Hampshire, 1 males, 27.v.2002, Chandler leg. (PPCM). Dingwall, male, 28.v.1988, Chandler leg. (PPCM). Ton Dingle, Highland Region, 1 male, 10.viii.1909, King leg. Powys, 1 male, 15.x.1987, Stubbs leg. (PPCM). Urqu- (BMNH). Dinton Pastures Country Park, Berkshire, 1 hart Bay, Drumnadrochit, Highland Region, 1 male, male, 6.viii.1994, Chandler leg. (PPCM). Ffrith, Flint- 13.vi.1984, Chandler leg. (PPCM). Wamphray Glen, shire, 1 male, 7.vi.−9.vi.1919, Edwards leg. [paratype Dumfries & Galloway, 1 male, 1.vi.1973, Chandler of Sciara subpilosa] (BMNH). Gearagh, Cork, 1 male, leg. (NMS). Warren’s Court Wood, Cork, 1 male, 12.vi.1990, de Courcy Williams leg. (NMID). Grange, 11.vii.1994, Withers leg. (PPWM). Way Beck, Cum- Lancashire, 1 male, vii.1923, Edwards leg. [holotype of bria, 1 male, 14.vi.1999, Chandler leg. (PPCM). West Sciara subpilosa] (BMNH). Greywell Fen, Hampshire, End Common, Esher, Surrey, 1 male, 16.vii.2002, 1 male, 2.viii.1998, Chandler leg. (PPCM). Hartslock, Chandler leg. (PPCM). Whitecross Green Wood, Oxfordshire, 1 male, 14.v.2000, Raper leg. (PDGB). Oxfordshire, 1 male, 7.x.1999, Chandler leg. (PPCM). Holker Moss, Cumbria, 1 male, 11.vii.−13.vii.1923, Windsor Forest, Berkshire, 1 male, 27.v.1991; 1 male, Edwards leg. [paratype of Sciara subpilosa] (BMNH). 15.vi.1999, all Chandler leg. (NMS). Wood Crates, Ickenthwaite, Cumbria, 1 male, 18.vi.1999, Chandler New Forest, Hampshire, 1 male, 8.vii.1990, Chandler leg. (PPCM). Lackham Park, Wiltshire, 1 male, leg. (PPCM). Woodwalton Fen, Cambridgeshire, 1 7.vii.2002, Chandler leg. (PPCM). Loch Chon, Stirling, male, 24.v.1940, Edwards leg. (BMNH). Wychwood 1 male, 29.vi.1999, Chandler leg. (PPCM). Lockhaugh Forest, Oxfordshire, 1 male, 12.ix.1998, Chandler Bank, Derwent Valley, Tyne & Wear, 1 male, 7.vi.− leg. (NMS). Wyre Forest, Worcestershire, 1 male, 23.vi.1990; 1 male, 24.vii.−13.viii.1990, all Jessop leg.

(PBLN). Long Wood, Nettleton, Lincolnshire, 1 male, male, 2.vii.1977, J.H. Cole leg. (PJCB). Aldermoors, v.1997, Godfrey leg. (PPCM). Mark Ash Wood, New Berkshire, 2 males, 14.v.1994, Chandler leg. (PPCM). Forest, Hampshire, 1 male, 4.vi.1989, Chandler leg. Arcan Mains, Highland Region, 1 male, 28.v.1994, (PPCM). Monk’s Wood, Cambridgeshire, 1 male, Chandler leg. (PPCM). Ashridge Estate, Hertford- 12.vi.1972; 1 male, 10.vii.1972, all J.H. Cole leg. shire, 4 males, 20.v.1999, Chandler leg. (NMS). (PJCB). Nairn, Highland Region, 1 male, 26.v.1905, Austwick, North Yorkshire, 2 males, v.1934, Edwards Yerbury leg. (BMNH). Pine Wood, Berkshire, 1 male, 1 leg. (BMNH). Aviemore, Highland Region, 2 males, female, 22.vi.2001, Gibbs leg. (PDGB). Purn Hill, v.1934, Edwards leg. (BMNH). Begwary Brook, Bed- Avon, 1 male, 21.viii.2001, Gibbs leg. (PDGB). Ryton fordshire, 1 male, 20.v.1999, J.H. Cole leg. (PJCB). Wood, Bubbenhall, Coventry, West Midlands, 1 male, Bettws, Conwy, 2 females, 16.vi.1887, Verrall leg. 25.viii.2002, J.E. Smith leg. (HRI). Spreacombe (UMO). Bevills Wood, Abbot’s Ripton, Cam- Chapel Wood, Devon, 1 male, 6.vii.1986, Chandler leg. bridgeshire, 1 male, 31.v.1976, J.H. Cole leg. (PJCB). (PPCM). Stoke Gabriel, Devon, 1 male, 7.vi.1920, Bonhill, West Dunbartonshire, 1 female, 22.vi.1907, Edwards leg. [paratype of Sciara subpilosa] (BMNH). Malloch leg. (NMS). Bovey Valley, Devon, 1 male, Tempsford, Bedfordshire, 1 male, 20.ix.1922, Edwards 31.v.1978, Chandler leg. (NMS). Brandon Marsh, leg. (BMNH). Thompson Common, Norfolk, 1 male, Ryton-on-Dunsmore, West Midlands, 1 male, 1.vi.1985; 1 male, 9.vi.1990, all Chandler leg. (PPCM). 22.viii.2002, Menzel leg. (DEI). Buckingham Palace Valency Valley, Cornwall, 1 male, 29.v.2000, J.H. Cole Garden, 1 male, 27.iii.−24.iv.1997, D.J. Carter & C.W. leg. (PJCB). West End Common, Esher, Surrey, 1 male, Plant leg. (PPCM); 3 males, 13.viii.1997, Chandler leg. 16.vii.2002, Chandler leg. (PPCM). Wicken Fen, Cam- (PPCM). Buckingham Thick Copse, Northampton- bridgeshire, 1 male, 12.v.1990, Withers leg. (PPWM). shire, 1 male, 1.vi.1993; 1 male, 9.vi.1992, all Warne leg. (NMS, PPCM). Bulwell Hall Park, Nottingham- Ecological notes: common in all types of woodland; shire, 1 male, 27.v.1919, Carr leg. (BMNH). Burnham ancient woodland with oak, hazel and bramble; heath- Beeches, Buckinghamshire, 2 females, 2.v.1929 land; wetlands including fen, sedge beds, water mead- (reared), Richards leg. (BMNH); 1 male, 1 female, ows, river banks and lake shores; gravel pits; old 13.v.1990, Chandler leg. (PPCM); 1 male, 10.v.− hedges; grassland with pingo pools; calcareous grass- 23.v.1998, Ismay leg. (PPCM). California Country land. v–viii. Park, Berkshire, 2 males, 22.v.1999, Chandler leg. (PPCM). Carie, Rannoch, Perth & Kinross, 2 males, 29.v.1998, Chandler leg. (PPCM). Carton, Kildare, 1 LEPTOSCIARELLA (LEPTOSCIARELLA) SUBVIATICA male, 29.iv.1987, J.P. O’Connor leg. (NMID). Castle MOHRIG & MENZEL, 1997 Neroche, Somerset, 1 male, 12.v.1936, Edwards leg. New to British list. (BMNH). Cawdor Wood, Highland Region, 1 male, Material examined: Sutton, Norfolk, 2 males, 9.vi.− 12.vi.1984, Stubbs leg. (NMS). Charleville Wood, 23.vi.1988, Foster & Procter leg. (DEI). The Loke, Offaly, 1 male, 27.v.1984, Chandler leg. (PPCM). Chig- Norwich, Norfolk, 1 male, 26.iv.1987, Laurence leg. well Row Wood, Chigwell Row, Essex, 1 female, − (PBLN). 20.v.2002; 1 male, 1 female, 23.v. 27.v.2002; 1 female, 11.vi.2002; 3 males, 1 female, 18.vi.2002; 1 female, Ecological notes: found in carr woodland and fen; gar- 15.ix.2002; 2 females, 26.ix.2002; 2 females, 8.x.2002, den. iv, vi. all Ismay & Schulten leg. (DEI, UMO). Clovelly, Devon, 2 males, 2 females, 8.v.1936, Edwards leg. (BMNH). Cod Wood, Devon, 2 males, 29.v.1978, Chan- LEPTOSCIARELLA (LEPTOSCIARELLA) TROCHANTERATA dler leg. (NMS). Coire Raibeirt, Cairngorm, Highland (ZETTERSTEDT, 1851) Region, 1 female, 16.vi.1984, Chandler leg. (NMS). Synonyms: coarctata (Winnertz, 1867); hirsutissima Colsterworth, Lincolnshire, 1 male, 3.vi.1987, Chan- (Strobl, 1895); prisca (Winnertz, 1867); saltuum (Win- dler leg. (PPCM). Colwich Park, Staffordshire, 2 nertz, 1868); splendens (Winnertz, 1867) [Sciara]. males, 8.vi.1889, Verrall leg. (BMNH, UMO). Coombe, Cornwall, 1 male, 19.v.1987, Chandler leg. (PPCM). Literature: Edwards (1925a): 537; Basden (1933): 44 Cop Mere, Staffordshire, 1 male, 23.v.1994, Chandler [all as Sciara hispida, misidentiﬁcation]; Perry & leg. (PPCM). Crianlarich, Stirling, 1 male, 1 female, Stubbs (1978): 71 [as Trichosia coarctata]; Freeman 12.vi.1996, Chandler leg. (NMS, PPCM). Culbin (1983b): 20; O’Connor & Ashe (1991): 29 [all as Tricho- Sands, Highland Region, 1 male, 27.v.2002, Chandler sia (Leptosciarella) coarctata]; Mohrig & Menzel leg. (PPCM). Derrycunihy Cascade, Kerry, 1 female, (1997): 55; Menzel (1998): 22; Chandler (2001): 201. 16.x.1975, Chandler leg. (PPCM). Dinton Pastures Material examined: Abbot’s Wood, Hampshire, 1 male, Country Park, Berkshire, 1 male, 15.vi.1993; 1 male, 25.v.1992, Chandler leg. (PPCM). Abernyte, Angus, 1 4.vi.1994; 1 male, 6.viii.1994; 1 male, 6.v.1997, all

Chandler leg. (PPCM). Dorchester, Dorset, 2 males, male, 1.v.−31.v.1998, C.W. Plant leg. (PPCM). Pen- 11.vi.1888, Verrall leg. (UMO). Dovedale, Derbyshire, tyrch, Cardiff, 1 male, 27.v.1968, Stubbs leg. (PPCM). 1 male, 11.vi.1988, Verrall leg. (BMNH). Dromahaire, Radwell, Hertfordshire, 2 males, v.1918, Edwards Leitrim, 1 male, 18.v.1970, Chandler leg. (PPCM). leg. (BMNH). Raveley Wood, Cambridgeshire, 1 male, Dromore Forest, Kerry, 3 males, 15.x.1973, Chandler 26.v.1975; 2 males, 21.v.1978, all J.H. Cole leg. (PJCB). leg. (NMS, PPCM). Drumnadrochit, Highland Region, Rotherfield Park, Hampshire, 1 male, 3.vi.2002, Chan- 1 male, 13.vi.1976, Chandler leg. (NMS). Epping dler leg. (PPCM). Royston, Hertfordshire, 3 males, 3 Forest, Essex, 2 males, 16.viii.−26.ix.1998; 3 males, females, iv.1918, Edwards leg. (BMNH). Rudge Wood, 10.v.−16.v.1999, all Dagley leg. (HRI, PPCM); 1 male, Bovey Valley, Devon, 1 male, 31.v.1978, Chandler leg. 25.v.1999, Chandler leg. (PPCM). Farnham Royal, (PPCM). Sarsgrove Wood, Oxfordshire, 1 male, Buckinghamshire, 8 males, 1 female, 12.v.−2.vi.1931; 12.vi.1977, Chandler leg. (NMS). Savernake Forest, 5 males, 4.vi.1931; 1 female, 10.v.1934; 1 male, 2 Wiltshire, 3 males, 2.v.−22.vi.1991, Porter leg. females, 11.v.1934 (all reared), all Basden leg. (PPCM); 1 male, 3.v.1993, Chandler leg. (PPCM). (BMNH, NMS). Fowl Mere, Norfolk, 1 male, vi.1939, Scratch Wood, Greater London, 1 male, 1 female, Edwards leg. (BMNH). Frilford Heath, Oxfordshire, 1 7.vii.−14.vii.1967 (reared), Bradford leg. (PPCM). Sea male, 25.iv.−25.v.1991, Porter leg. (PPCM). Fungarth, Mere, Norfolk, 2 males, 1 female, 29.v.1975, K.G.V. Perth & Kinross, 1 male, 30.v.1998, Chandler leg. Smith leg. (BMNH). Shotover, Oxfordshire, 1 female, (NMS). Glen Lochay, Stirling, 1 male, 3.vi.−18.vi.1932, 22.v.1915, Hamm leg. (UMO). Shrawley Wood, Worces- Edwards leg. (BMNH). Glendalough, Wicklow, 1 male, tershire, 1 male, 3.vi.−10.vi.1995, Cooter leg. (NMS). 1 female, 10.vi.1985, Chandler leg. (NMS, PPCM). Spartum Fen, Oxfordshire, 1 male, 1.vi.1991, Chan- Gorge of Avon, South Lanarkshire, 1 male, 30.v.1903, dler leg. (PPCM). Stodmarsh, Kent, 1 female, King leg. (BMNH). Grantown-on-Spey, Highland 1.vi.1977, Chandler leg. (PPCM). Temple, Berkshire, 1 Region, 1 male, 15.vi.1984, Chandler leg. (PPWM). male, 4.v.1934; 4 males, 9.v.−21.v.1934 (all reared), all Headington Hill, Oxfordshire, 1 male, 8.v.1915, Hamm Basden leg. (NMS). The Loke, Norwich, Norfolk, 1 leg. (UMO). Ivybridge, Devon, 1 male, 18.v.1890, male, 26.iv.1987; 1 male, 1 female, 8.v.1987; 3 males, Verrall leg. (UMO). Jonny Wood, Cumbria, 1 male, 10.viii.1987; 1 male, 5.vi.−13.vi.1989; 1 male, 21.viii.− 11.ix.1991, Gibbs leg. (PDGB). Knebworth, Hertford- 24.viii.1989, all Laurence leg. (PBLN). Thompson shire, 1 male, vi.1922; 1 male, 22.v.1925, all Edwards Common, Norfolk, 1 male, 1.vi.1985, Chandler leg. leg. (BMNH). Knocksink Wood, Wicklow, 1 male, (NMS). Tilberthwaite Ghyll, Cumbria, 1 male, 17.vi.1985, Chandler leg. (PPWM). Lancelot Clark 26.v.1938, Edwards leg. (BMNH). Udale Bay, High- Storth Nature Reserve, Cumbria, 1 male, 27.viii.1985, land Region, 1 male, 29.v.2002, Chandler leg. (PPCM). J.H. Cole leg. (PJCB). Leckford Estate, Hampshire, 2 Ufton Bridge, Warwickshire, 1 male, 9.vi.1991, Chan- males, 3.vi.−4.vi.1972, Chandler leg. (NMS). Letch- dler leg. (NMS). Wentbridge, West Yorkshire, 2 males, worth, Hertfordshire, 1 male, v.1917; 1 male, vi.1917; 20.vi.1987, Chandler leg. (NMS). Weston, Hertford- 1 male, v.1918; 1 male, vi.1924, all Edwards leg. shire, 3 males, 3 females, iv.-v.1924 (reared), Edwards (BMNH). Little Paxton Pits, Cambridgeshire, 1 male, leg. (BMNH). Windsor Great Park, Berkshire, 1 male, 19.v.2002, J.H. Cole leg. (PJCB). Loch Achnacloich, 14.vi.1972, Chandler leg. (NMS). Windsor Forest, Highland Region, 1 male, 15.vi.1976, Chandler leg. Berkshire, 1 male, 21.v.1988, Chandler leg. (NMS). (NMS). Loch Meiklie, Highland Region, 1 male, Woodwalton Fen, Cambridgeshire, 1 male, 15.v.1919; 2 13.vi.1984, Chandler leg. (PPCM). Lockhaugh Bank, males, 24.v.1940, all Edwards leg. (BMNH). Wych- Derwent Valley, Tyne & Wear, 1 male, 21.vi.− wood, Oxfordshire, 1 male, 11.v.−26.v.1989, Porter leg. 3.vii.1990, Jessop leg. (PBLN). Lough Ballynafid, (PPCM). Westmeath, 1 male, 16.vi.1985, Chandler leg. Ecological notes: very common in all types of wood- (PPWM). Mark Ash Wood, New Forest, Hampshire, 1 land; ancient woodland with oak, hazel and bramble; male, 2.vi.1984, Stubbs leg. (NMS). Matlock, Derby- heathland; wetlands including fen, water meadows shire, 2 males, 17.vi.1888, Verrall leg. (BMNH, UMO). and wooded river bank; former gravel workings; Maulden Wood, Bedfordshire, 1 male, 5.vi.1980, J.H. dunes; old hedges; parkland with rough grassland and Cole leg. (PJCB). Migdale Wood, Highland Region, 1 scrub; grassland with pingo pools; disused railway cut- male, 31.v.2002, Chandler leg. (PPCM). Monkfield ting; gardens; open montane habitat. Reared from rot- Wood, Cambridgeshire, 1 male, 1.v.−31.v.1998; 2 ting pine stump, from nests of blackbird and song males, viii.1998, all C.W. Plant leg. (NMS, PPCM). thrush (form with dark coxae only). iii–x. Monk’s Wood, Cambridgeshire, 1 male, 17.vi.1972, J.H. Cole leg. (PJCB). New Forest, Hampshire, 1 male, 18.v.1968, Parmenter leg. (BMNH). Newmarket, Cam- LEPTOSCIARELLA (LEPTOSCIARELLA) VIATICA bridgeshire, 1 female, 2.vi.1888; 1 male, 18.v.1894, all (WINNERTZ, 1867) Verrall leg. (UMO). Oakland Wood, Cambridgeshire, 1 Synonym: proxima (Winnertz, 1867).

Literature: Freeman (1983b): 20; O’Connor & Ashe Kent, 1 male, iv.1935, Nurse leg. (BMNH). Welches (1991): 29 [all as Trichosia (Leptosciarella)]; Mohrig & Dam, Cambridgeshire, 1 male, 13.iii.1999, J.H. Cole Menzel (1997): 75; Menzel (1998): 22. leg. (PJCB). Windsor Forest, Berkshire, 6 males, 19.xi.1989, Owen leg. (NMS). Withyham, East Sussex, Material examined: Aversley Wood, Cambridgeshire, 2 1 male, 21.iii.1946, Grensted leg. (UMO). males, 2.iv.1988, J.H. Cole leg. (PJCB). Blean Wood, Ecological notes: very common in broad-leaved and Kent, 1 male, 9.iv.1965, Parmenter leg. (BMNH). carr woodland; heathland; water meadows and gravel Bramfield, Hertfordshire, 1 male, 18.iv.1964, Colyer pits; old hedges; within Ouse Washes bird hide; leg. (BMNH). Brampton Wood, Cambridgeshire, 1 wooded river bank; calcareous grassland; parkland; male, 6.iii.1966, J.H. Cole leg. (PJCB). Brasenose, gardens. Reared from rotten wood. iii–vii, ix, xi. Oxfordshire, 1 male, 2 females, 17.iii.1925 (reared), Richards leg. (BMNH). Burnham Beeches, Bucking- hamshire, 1 male, 15.iv.1915, Edwards leg. (BMNH); 1 LEPTOSCIARELLA (LEPTOSCIARELLA) YERBURYI male, 20.iv.1998, Chandler leg. (PPCM). Caerlaverock, (FREEMAN, 1983) Dumfries & Galloway, 1 male, 16.vii.1980, Duncan leg. Literature: Freeman (1983a): 163; Freeman (1983b): (NMS). Cefn-y-Bedd, Flintshire, 1 male, 15.iii.1953, 20 [all as Trichosia (Leptosciarella)]; Mohrig & Menzel Colyer leg. (BMNH). Chigwell Row Wood, Chigwell (1997): 73; Menzel (1998): 22; Menzel & Mohrig Row, Essex, 1 male, 18.vi.2002, Ismay & Schulten leg. (2000): 369. (DEI). Crowborough, East Sussex, 2 males, 2 females, 8.iv.−16.iv.1904; 1 male, 31.iii.−14.iv.1906; 1 male, Material examined: Annahala bog, Gearagh, Cork, 2 12.iv.−27.iv.1916, all Jenkinson leg. (BMNH). Dickle- males, 22.iii.1990, de Courcy Williams leg. (DEI, burgh, Norfolk, 1 male, 19.iii.−14.iv.1989, Withers leg. NMID). Ballindalloch, Highland Region, 1 male, (PPWM). Dinton Pastures Country Park, Berkshire, 24.vi.1905, Yerbury leg. [holotype of Trichosia yer- 1 male, 27.iii.1994, Chandler leg. (PPCM). Dunstall buryi] (BMNH). Charleville Wood, Offaly, 1 male, Wood, Shoreham, Kent, 1 male, 18.ix.1964, Chandler 15.vi.1985, Chandler leg. (PPWM). Coverham, North leg. (PPCM). Epping Forest, Essex, 6 males, 4 females, Yorkshire, 1 male, 23.vi.1924, Edwards leg. [paratype 1.iii.−29.iii.1999; 2 males, 1 female, iii.2000, all Dagley of Trichosia yerburyi] (BMNH). Culbin Forest, High- leg. (NMS, PPCM). Eynsford, Kent, 1 male, 7.iv.1964, land Region, 2 males, 27.v.2002, Chandler leg. Chandler leg. (PPCM). Falls of Tarnash, Moray, 1 (PPCM). Ditchley, Oxfordshire, 1 male, v.1937, male, 9.vii.1936, Coe leg. (BMNH). Felden, Hertford- Edwards leg. [paratype of Trichosia yerburyi] shire, 1 male, 21.iii.1897; 1 female, 25.iii.1897, all (BMNH). Grantown-on-Spey, Highland Region, 1 A. Piffard leg. (BMNH). Godmanchester, Cam- male, 15.vi.1984, Chandler leg. (PPWM). Holne, bridgeshire, 1 male, 16.iii.1999, J.H. Cole leg. (PJCB). Devon, 1 male, 3.vi.1974, Chandler leg. (PPCM). Kin- Grwyne Fechan, Powys, 1 male, 18.iv.1927, Richards rara, Highland Region, 1 male, 19.vi.1976, Chandler leg. (BMNH). Hartslock Wood, Oxfordshire, 2 males, leg. (NMS). Leigh Wood, Avon, 1 male, 30.v.1950, 15.iv.1963, J.H. Cole leg. (PJCB). Hickling Broad, Nor- d’Assis-Fonseca leg. (PPCM). Loch Loy, Highland folk, 1 male, 16.iv.1989, Withers leg. (PPWM). Iving- Region, 1 male, 14.vi.1984, Chandler leg. (PPCM). hoe, Buckinghamshire, 2 males, 7.iv.1938, Edwards Nairn, Highland Region, 1 male, 12.vi.1905, Yerbury leg. (BMNH). Killoughrim, Wexford, 1 male, 4.iv.1988, leg. [paratype of Trichosia yerburyi] (BMNH). Pass of J.P. O’Connor leg. (NMID). Lopham Fen, Norfolk, 1 Killiecrankie, Perth & Kinross, 1 male, 31.viii.1989, male, 22.xi.1990, Withers leg. (PPWM). New Forest, Chandler leg. (PPCM). Shortwood, Gloucestershire, 1 Hampshire, 1 female, 31.iv.1897; 1 male, 20.iv.1909; 1 male, 12.vi.2002, Gibbs leg. (PDGB). Swindale Beck, male, 22.iv.1909, all Adams leg. (BMNH); 6 males, 1 Cumbria, 1 male, 20.vi.1981, McLean leg. (PPCM). female, 27.iii.1948, Colyer leg. (BMNH). Offley Holes, Tarradale House, Highland Region, 1 male, v.2002, Hertfordshire, 1 female, iv.1936, Edwards leg. Sumner leg. (PPCM); 1 male, 1.vi.2002, Chandler leg. (BMNH). Oxford, Oxfordshire, 1 male, 24.iii.1953, (PPCM). No locality details, Hampshire, 1 male, Grensted leg. (UMO). Rhinefields Inclosure, New 12.v.1951, Colyer leg. (PBLN). Forest, Hampshire, 1 male, 20.iv.1973, Cranston Ecological notes: common in broad-leaved woodland; leg. (BMNH). Savernake Forest, Wiltshire, 1 male, also found in mixed and pine woodland; conifer plan- 3.v.1993, Chandler leg. (PPCM). Sidmouth, Devon, 1 tations; wetlands including saltmarsh and bog; rotten female, 19.iv.1925, Edwards leg. (BMNH). Stoke Court stump. iii, v–vi, viii. Park, Buckinghamshire, 1 female, 31.iii.1968, Chan- dler leg. (PPCM). The Loke, Norwich, Norfolk, 1 male, LYCORIELLA FREY, 1942 20.iv.1987; 1 male, 8.v.1987; 1 male, 5.vi.−13.vi.1989, all Laurence leg. (PBLN). Tintern, Wexford, 1 male, LYCORIELLA ( COELOSTYLINA) FREYI TUOMIKOSKI, 1960 3.iv.1988, J.P. O’Connor leg. (NMID). Tunbridge Wells, New to British list.

Material examined: Spey bank, Aviemore, Highland, 1 sedge and reed beds, raised mire, lake and river mar- male, 9.vi.1984, Chandler leg. (PPWM). gins and wet meadows; old sand and gravel workings; grassland with pingo pools. v–xi. Ecological notes: found in broad-leaved woodland on river bank. vi. LYCORIELLA (HEMINEURINA) INFLATA (WINNERTZ, 1867) LYCORIELLA (HEMINEURINA) CONSPICUA Synonyms: difficilis (Grzegorzek, 1884); interdicta (WINNERTZ, 1867) (Grzegorzek, 1884); nitens (Winnertz, 1867); sub- Literature: ? Walker (1848): 105; ? Walker (1856): 53; venosa Mohrig & Krivosheina, 1983. Edwards (1925a): 539 [all as Sciara hyalipennis, Literature: Edwards (1925a): 539 [as Sciara]; Free- misidentification]; Edwards (1938): 201 [as Sciara man (1983b): 30; O’Connor & Ashe (1991): 29; Menzel (Neosciara) hyalipennis, misidentification]; Freeman (1998): 22 [all as Lycoriella venosa, misidentification]; (1983b): 30; Laurence (1994): 118; Menzel (1998): 22; Menzel (1998): 22 [as Lycoriella inflata]. Chandler et al. (2002): 124. Material examined: Aviemore, Highland Region, 1 Material examined: Austwick, North Yorkshire, 1 male, 9.vi.1984, Chandler leg. (PPWM). Bagley, male, vi.1930, Cheetham leg. (BMNH); 1 male, 26.v.− Oxfordshire, 2 males, 9.vi.1917, Hamm leg. (BMNH, 27.v.1934, Edwards leg. (BMNH). Aviemore, Highland UMO). Bagpiper’s Tump, West Wood, Herefordshire, 1 Region, 1 male, v.1934, Edwards leg. (BMNH). Brae- male, 10.x.1998, Chandler leg. (NMS). Bix Bottom, downie, Angus, 1 male, 24.vii.1937, Coe leg. (BMNH). Oxfordshire, 2 males, 6.vii.2002, Chandler leg. Cairngorm, Highland Region, 2 males, 15.vi.1984, (PPCM). Brandon Marsh, Ryton-on-Dunsmore, West Chandler leg. (PPWM). Cardington, Bedfordshire, 2 fe- Midlands, 5 males, 6 females, 22.viii.2002, Menzel & males, 6.viii.1914, Edwards leg. (BMNH). Fallowfield, J.E. Smith leg. (DEI, HRI). Burnham Beeches, Buck- Greater Manchester, 1 male, 1 female, 26.ix.1920, Brit- inghamshire, 1 male, 5.vii.−20.vii.1995, Ismay leg. ten leg. (BMNH). Falls of Tarnash, Moray, 1 female, (HRI). Castle Eden Dene, Durham, 1 male, 12.ix.1989, 9.vii.1936, Coe leg. (BMNH). Gearagh, Cork, 1 male, J.H. Cole leg. (PJCB). Collinpark Wood, Gloucester- 30.vi.1990; 1 male, 10.x.1990, de Courcy Williams leg. shire, 1 male, 3.ix.2002, Gibbs leg. (PDGB). Coombe (DEI, NMID). Grafham Water, Savage’s Creek, Cam- Abbey Country Park, Coventry, Warwickshire, 1 bridgeshire, 1 male, 21.viii.1976, J.H. Cole leg. (PJCB). female, 28.viii.2003, J.E. Smith leg. (HRI). Crawfords- Gwenfro, Isle of Anglesey, 1 male, 14.vi.1988, Holmes, burn Country Park, Down, 1 male, 2.vii.1987, Chan- Boyce & Reed leg. (PPWM). Harpenden, Hertfordshire, dler leg. (PPCM). Epping Forest, Essex, 1 male, 3 females, 1934, C.B. Williams leg. (BMNH). Hickling, 28.vi.2000, Chandler leg. (PPCM); 1 male, 29.vi.2000, Norfolk, 6 males, x.1988, Foster & Procter leg. (PBLN). Henshaw leg. (PPCM). Glyn Wood, Monmouthshire, 1 Kennack Cove, Cornwall, 1 male, 4.ix.1983, Chandler male, 9.vii.1986, Chandler leg. (NMS). Goring Heath, leg. (PPCM). Loch Park, Moray, 1 male, 12.viii.− Oxfordshire, 1 male, 20.iii.1964, Burtt leg. (BMNH). 14.viii.1937, Coe leg. (BMNH). Nuremore Lake, Car- Greywell Fen, Hampshire, 3 males, 1 female, rickmacross, Monaghan, 1 male, 19.vi.1979, J.H. Cole 29.viii.2002, Chandler leg. (HRI). Hobb’s Quarry, leg. (PJCB). Rye Water, Kildare, 1 male, 9.viii.1981, Gloucestershire, 1 male, 4.ix.2002, Gibbs leg. (PDGB). J.P. & M.A. O’Connor leg. (NMID). Searles Farm, Ivy Thorn Hill, Somerset, 1 male, 20.vi.2001, Gibbs Reading, Berkshire, 1 male, 22.vii.2003, Chandler leg. leg. (PDGB). Jeffry Bog, North Yorkshire, 1 male, (PPCM). Shefford, Bedfordshire, 1 male, 1.vii.1915, 14.vi.1997, Chandler leg. (PPCM). Lackham Park, Edwards leg. (BMNH). Stallode Wash, Suffolk, 14 Wiltshire, 1 male, 20.ix.2002, Chandler leg. (PPCM). males, 30.viii.−13.ix.1988, Foster & Procter leg. Lea Meadows, Newtown Linford, Leicestershire, 3 (PBLN). Straffan, Kildare, 1 male, 14.xi.1981, J.P. males, 2 females, 24.viii.2003, Menzel & J.E. Smith O’Connor leg. (NMID). Thompson Common, Norfolk, 1 leg. (DEI, HRI). Littlecote Park, Wiltshire, 1 male, male, 13.x.2002, Chandler leg. (PPCM). Wellesbourne, 25.viii.1973, Chandler leg. (NMS). Longborough, Warwickshire, 1 male, 17.viii.−31.viii.2000, J.E. Smith Gloucestershire, 1 male, 8.iv.2003, J.E. Smith leg. leg. (HRI). Weston, Hertfordshire, 1 male, 4.x.1931, (HRI). Narborough Bog, Whetstone, Leicestershire, 1 Edwards leg. (BMNH). Wheatfen Broad, Norfolk, 1 female, 24.viii.2003, Menzel leg. (DEI). Newtown St. male, 19.v.1990, Withers leg. (PPWM). Whitacre Boswell’s, Scottish Borders, 1 male, 15.vii.1986, Chan- Heath, Warwickshire, 3 males, 13.x.2001, J.E. Smith dler leg. (NMS). Odiham Common, Hampshire, 1 leg. (HRI). Woodwalton Fen, Cambridgeshire, 1 male, male, 7.vii.1990, Chandler leg. (PPCM). Passage East, 7.vi.1940, Edwards leg. (BMNH). Waterford, 1 male, 13.vii.1989, J.P. & M.A. O’Connor Ecological notes: found in broad-leaved and carr wood- leg. (NMID). Sned Wood, Lower Lye, Herefordshire, 2 land; ancient pine woodland; wetlands including fen, males, 14.vii.2003, J.E. Smith leg. (HRI). Stradey

Wood, Carmarthen, 3 males, 13.x.1995, McLean & fens, water meadows and reed beds; near gravel pit; Chandler leg. (PPCM). The Dingle, Isle of Anglesey, 1 old hedges; parkland. v–x. male, 5.vii.1987, Chandler leg. (PPCM). Tomintoul, Moray, 1 male, 8.viii.−11.viii.1937, Coe leg. (BMNH). West End Common, Esher, Surrey, 1 male, 16.vii.2002, LYCORIELLA (LYCORIELLA) CASTANESCENS Chandler leg. (PPCM). West Harling Common, Nor- (LENGERSDORF, 1940) folk, 1 male, 1.iv.1989, Withers leg. (PPWM). Windsor Synonyms: agarici Loudon, 1978; difficilis (Frey, 1948) Forest, Berkshire, 1 male, 6.ix.1986, Chandler leg. [preocc.]; fucorum (Frey, 1948); paucisetulosa (Frey, (NMS). Wyre Forest, Worcestershire, 1 male, 1948); rufotincta Tuomikoski, 1959. 6.viii.1988, Chandler leg. (PPCM). Wytham Woods, Wytham, Oxfordshire, 1 male, 1 female, 21.viii.2002, Literature: Symes (1921): 142, 188 and 234 [as Sciara Menzel leg. (DEI). praecox, misidentification]; Edwards (1925a): 540 [in part as Sciara praecox, misidentification]; Basden Ecological notes: common in broad-leaved and carr (1933): 44; Madwar (1937): 43 [all as Sciara auripila, woodland; mixed woodland and conifer plantations; misidentification]; Binns (1973): 119; Binns (1975): 1; water meadows; reed and sedge beds; river bank; Chandler (1978): 209; Skidmore (1978): 76; Binns former gravel workings; grassland (incl. calcareous). (1980): 143; Binns (1981): 417; Freeman (1983b): 31; iii–iv, vi–x. White (1986): 11; Freeman (1987): 201; White & Jar- rett (1990): 373; Pethybridge (1991): 453; Laurence LYCORIELLA (HEMINEURINA) MODESTA (1994): 118; Laurence (1997a): 85; Laurence (1996): (STAEGER, 1840) 85; Laurence (1997b): 228; Laurence & James (1997): 166; White (1997): 29 [all as Lycoriella auripila, mis- Synonyms: arctica (Holmgren, 1869); ecalcarata identification]; Menzel (1998): 22; White et al. (2000): (Holmgren, 1869); frigida (Holmgren, 1869) [preocc.]; 208; White & Smith (2000): 165; Chandler (2001): 201. fumatella (Lundbeck, 1898); groenlandica (Holmgren, 1872); holmgreni (Rübsaamen, 1894). New to Irish list. Literature: Freeman (1983a): 168; Freeman (1983b): Material examined: Alconbury, Cambridgeshire, 1 30; Menzel (1998): 22; Chandler et al. (2002): 124. male, 3.v.1976, J.H. Cole leg. (PJCB). Austwick, North Yorkshire, 1 male, v.1933, Edwards leg. (BMNH). Avon Material examined: Blaise Wood, Avon, 1 female, Valley Wood, Devon, 1 male, 3.i.1978 (reared), Hurrell 4.vii.1985, Ackland leg. (NMS). Castlecomer, Kilkenny, leg. (BMNH). Badger Wood, Cambridgeshire, 1 male, 2 males, 25.vi.1987, Chandler leg. (PPCM). Charlbury, vi.1998, C.W. Plant leg. (PPCM). Beaconsfield, Oxfordshire, 1 male, 4.vii.1936, Edwards leg. (BMNH). Buckinghamshire, 5 males, 24.iii.−2.vi.1934 (reared), Collinpark Wood, Gloucestershire, 1 male, 3.ix.2002, Basden leg. (NMS). Berwick-upon-Tweed, Northum- Gibbs leg. (PDGB). Dinton Pastures Country Park, berland, 3 males, vi.1999; 1 male, vii.1999; 1 male, Berkshire, 1 male, 30.vii.1994, Chandler leg. (PPCM). viii.1999, all Laurence leg. (PBLN). Black Park, Buck- Dunster, Somerset, 1 male, 8.vii.1986, Chandler leg. inghamshire, 2 males, 20.vi.−26.vi.1934; 1 male, (NMS). Earith Gravel Pits, Cambridgeshire, 1 male, 26.vi.1934 (all reared), all Basden leg. (BMNH, NMS). 26.vi.1976, J.H. Cole leg. (PJCB). Glen of Drumloch, Bockmer End, Buckinghamshire, 1 male, 10.iii.1932 Moray, 4 males, 16.viii.1937, Coe leg. (BMNH). How (reared), Basden leg. (BMNH). Boveney, Buckingham- Hill, Norfolk, 3 males, 24.vi.1990, Withers leg. shire, 1 male, 23.iii.1933 (reared), Basden leg. (NMS). (PPWM). Kingsbury Water Park, Warwickshire, 3 Braemar, Aberdeenshire, 1 male, 19.vii.−30.vii.1938, males, 1 female, 13.x.2001, J.E. Smith leg. (HRI). Nes- Coe leg. (BMNH). Bradfield Wood, Suffolk, 1 male, bitt Dene, Durham, 1 male, 23.vii.1990, Chandler leg. 6.vii.−11.vii.1990; 1 male, 23.v.−30.v.1991; 1 male, (PPCM). Scargill, Durham, 1 male, 27.vii.1985, Chan- v.1991; 1 male, 16.viii.−22.viii.1991, all Laurence leg. dler leg. (PPCM). Sidmouth, Devon, 1 male, 10.v.1936, (PBLN). Brampton Wood, Cambridgeshire, 1 male, Edwards leg. (BMNH). Skenfrith, Monmouthshire, 1 20.xi.1995; 2 males, 23.xi.1995 (all reared), all J.H. male, 12.vi.1997, Chandler leg. (PPCM). Spey Bay, Cole leg. (PJCB). Brancaster, Norfolk, 1 male, Highland Region, 1 male, 2.vii.1936, Coe leg. (BMNH). 2.ix.1988, Foster & Procter leg. (PBLN). Buckingham Tregate Bridge, Monmouthshire, 2 males, 12.vi.1997, Palace Garden, Greater London, 1 male, 27.iii.− Chandler leg. (PPCM). Wicken Fen, Cambridgeshire, 1 24.iv.1997, D.J. Carter & C.W. Plant leg. (PPCM). male, 8.vii.1989, Chandler leg. (NMS). Wytham Wood, Bucklebury Common, Berkshire, 9 males, v.1997 Oxfordshire, 1 male, 8.vii.1962, K.G.V. Smith leg. (reared), Storey leg. (PPCM). Burnham, Buckingham- (BMNH). shire, 1 male, 14.vii.1934; 1 male, 7.viii.1934; 1 male, Ecological notes: found in broad-leaved and carr wood- 12.ix.1934 (all reared), all Basden leg. (NMS). Byfleet, land (incl. acid and limestone); wetlands including Surrey, 1 male, iv.1939, Edwards leg. (BMNH). Cat-

field, Norfolk, 1 male, 26.viii.1988, Foster & Procter males, xi.1917; 1 male, ii.1918; 1 male, 27.ix.1923, all leg. (PBLN). Cheltenham, Gloucestershire, 23 males, Edwards leg. (BMNH). Littlehampton, West Sussex, 4 12.v.1995, White leg. (DEI). Cheshunt, Hertfordshire, males, 1986, White leg. (HRI). Liverpool, Merseyside, 1 male, 26.xii.1928, Speyer leg. (BMNH). Chesswood 1 male, 15.xi.1901, Newstead leg. (BMNH). Llyn Nurseries, Pulborough, West Sussex, 33 males, 1 Creiniog, Denbighshire, 1 male, 13.x.1988, Holmes, female, 24.v.1995, White leg. (DEI). Coed Dolgarrog, Boyce & Reed leg. (PPWM). London, Greater London, Conwy, 1 male, 28.vi.1988, Holmes, Boyce & Reed 2 males, 1921, no collector detail (BMNH). Lye Hill, leg. (PPWM). Coed Tycanol, Pembrokeshire, 1 male, Oxfordshire, 1 male, 7.iii.1912, Hamm leg. (BMNH). 29.vi.−7.vii.1999, Levey & Pavett leg. (PPCM). Cors Magor Marsh, Monmouthshire, 1 male, 28.ix.1987; 1 Erddreiniog, Isle of Anglesey, 1 male, 27.vii.1988, male, 7.vii.1988; 1 male, 28.ix.1988, all Holmes, Boyce Holmes, Boyce & Reed leg. (PPWM). Cotswold Mush- & Reed leg. (PPWM). Maidenhead, Berkshire, 11 room Farm, Cheltenham, Gloucestershire, 2 males, males, 25.v.1995, White leg. (DEI); 3 males, 1996, J.E. 12.v.1995, White leg. (HRI). Dickleburgh, Norfolk, 2 Smith leg. (HRI). Market Weston Fen, Suffolk, 1 male, males, 23.iii.1986; 1 male, 27.xii.1988–2.i.1989, all 6.ix.1988 (reared), Withers leg. (PPWM). Marston, Withers leg. (PPWM). Dinton Pastures Country Park, Oxfordshire, 3 males, 21.iii.−27.iii.1933 (reared), Bas- Berkshire, 1 male, 7.xi.1994, Chandler leg. (PPCM). den leg. (NMS). Medmenham, Buckinghamshire, 1 Dorchester, Dorset, 5 males, xi.1921, Bottomley leg. male, 17.iii.1934, Basden leg. (BMNH). Milham Ford (BMNH). Epping Forest, Essex, 1 male, v.1999, Dagley School, Oxford, Oxfordshire, 1 male, 6.vi.−11.vi.2002, leg. (PPCM). Felixstowe, Suffolk, 19 males, 15.vi.1995, Webb leg. (PPCM). Monkfield Wood, Cambridgeshire, White leg. (DEI). Gatewen Marsh, Denbighshire, 3 1 male, x.1998, C.W. Plant leg. (PPCM). Nazeing, males, 3.viii.1988; 1 male, 21.vi.1988, all Holmes, Essex, 5 males, 31.vii.1975; 8 males, 3.viii.1975, all Boyce & Reed leg. (PPWM). Glasshouse Crops Stephens leg. (HRI). Newbury, Berkshire, 2 males, Research Station, Littlehampton, West Sussex, 1 31.v.1935 (reared), Brown leg. (BMNH). North Ronald- male, 13.viii.1956; 4 males, 20.viii.1956; 1 male, say, Orkney, 9 males, 9.ix.−10.ix.1989, Laurence leg. 27.viii.1956; 2 males, 5.ix.1956; 1 male, 10.ix.1956; 2 (PBLN). Oxford, Oxfordshire, 2 males, 5.ii.1917, males, 17.ix.1956; 5 males, 1 female, 18.ix.1956; 1 Hamm leg. (BMNH). Penarth, Vale of Glamorgan, 1 male, 24.ix.1956; 3 males, 3.x.1956; 1 male, 8.x.1956; male, v.1934 (reared), Hallett leg. (BMNH). Pulbor- 1 male, 15.x.1956; 1 male, 22.x.1956; 17 males, ough, West Sussex, 1 male, 1996, J.E. Smith leg. 23.x.1956; 1 male, 9.xi.1956; 1 male, 12.xi.1956; 1 (HRI). Reedham, Norfolk, 4 males, 27.vi.−11.vii.1988; male, 19.xi.1956; 2 males, 23.xi.1956; 1 male, 14 males, 11.vii.−12.viii.1988; 4 males, 13.viii.− 26.xi.1956; 1 male, 17.xii.1956; 5 males, 15.x.1958; 1 27.viii.1988, all Foster & Procter leg. (PBLN). Rye male, 1 female, no date, all Hussey leg. (HRI); 1 male, Harbour, East Sussex, 1 male, 27.iii.2002 (reared), 7.iv.1961; 1 male, 26.iii.1962; 1 male, 27.i.1965; 1 Roper leg. (PPRS). Ryton Wood, Bubbenhall, Coventry, male, 26.iv.1965; 5 males, 4.v.1965, all Gurney leg. West Midlands, 1 male, 12.x.2001, J.E. Smith leg. (HRI); 2 males, vii.1963, Warner leg. (HRI); 4 males, (HRI). St. David’s Airfield, Pembrokeshire, 1 male, ii.1975; 6 males, 26.viii.1975; 10 males, viii.1975; 19 29.ix.1987, Holmes, Boyce & Reed leg. (PPWM). St. males, 11.ix.1975; 1 male, 24.ix.1975; 7 males, 2 Ives, Cornwall, 12 males, 18.v.1995, White leg. (DEI); 2 females, 1975; 6 males, 4 females, 1977; 4 males, 2 males, 1996, J.E. Smith leg. (HRI). Salisbury Plain females, no date, all Binns leg. (HRI). Grange- Training Area, Warminster, Wiltshire, 1 male, over-Sands, Cumbria, 1 male, 20.iii.1996, Birkett 4.vii.2003, Ismay & Schulten leg. (UMO). Snetterton, leg. (PPCM). Great Missenden, Buckinghamshire, 2 Norfolk, 1 male, 11.x.1987, Withers leg. (PPWM). Stal- males, 1996, J.E. Smith leg. (HRI). Greywell Fen, lode Wash, Suffolk, 1 male, 24.vi.−8.vii.1988, Foster Hampshire, 1 male, 29.viii.2002, Chandler leg. & Procter leg. (PBLN). Stanford, Norfolk, 2 males, (PPCM). Hailsham, Kent, 8 males, 25.v.1995, White 27.iii.−10.iv.1988, Collier leg. (PPWM). Stronsay, leg. (DEI); 2 males, 1996, J.E. Smith leg. (HRI). Hick- Orkney, 4 males, 21.v.1988, Laurence leg. (PBLN). ling, Norfolk, 1 male, v.1988; 1 male, vii.1988; 12 Strumpshaw, Norfolk, 2 males, 13.vi.−11.vii.1988, Fos- males, x.1988, all Foster & Procter leg. (PBLN). Hill ter & Procter leg. (PBLN). Temple, Berkshire, 1 male, End, Oxfordshire, 1 male, 13.x.1999, Chandler leg. 27.iii.1923; 4 males, 13.iii.−24.iii.1932; 2 males, (PPCM). Inglecroft Nursery, Littlehampton, West 20.iii.1932; 1 male, 29.iv.1933; 2 males, 11.v.− Sussex, 1 male, 9.ii.1958, Gurney leg. (HRI). Ipswich, 12.v.1933; 1 male, 10.iii.1934 (all reared), all Basden Suffolk, 3 males, 1996, J.E. Smith leg. (HRI). leg. (BMNH, NMS). The Loke, Norwich, Norfolk, Kelham Bridge, Ravenstone, Leicestershire, 2 males, 5 males, 23.iv.1987; 2 males, 26.iv.1987; 1 male, 24.viii.2003, Menzel leg. (DEI). Kingsbury Water 9.v.1987; 3 males, 4.vii.1987; 1 male, 5.vii.1987; 1 Park, Warwickshire, 1 female, 13.x.2001, J.E. Smith male, 10.viii.1987; 1 male, 15.viii.1987; 1 male, leg. (HRI). Land’s End, Cornwall, 2 males, 22.i.1998, 8.ix.1987; 2 males, 15.viii.−16.viii.1988; 1 male, 2.v.− Roper leg. (PPRS). Letchworth, Hertfordshire, 4 6.v.1992; 5 males, v.1992; 1 male, 26.vi.−30.vi.1998, all

Laurence leg. (PBLN). The Oaks, Cambridgeshire, 2 sfield, Buckinghamshire, 1 male, 15.iii.1933, Basden males, ix.1998, C.W. Plant leg. (PPCM). Thompson leg. (BMNH). Berwick-upon-Tweed, Northumberland, Common, Thompson, Norfolk, 1 male, 10.vi.− 2 males, vii.1999; 1 male, ix.1999; 1 male, x.1999; 1 24.vi.1988, Foster & Procter leg. (PBLN). Tomintoul, male, 4.xi.1999, all Laurence leg. (PBLN). Boat of Moray, 1 male, 8.viii.−11.viii.1937, Coe leg. (BMNH). Garten, Highland Region, 1 male, v.1934, Edwards Trefeiddan Moor, Pembrokeshire, 1 male, 21.vii.1987; leg. (BMNH). Buckingham Palace Garden, Greater 3 males, 29.ix.1987,all Holmes, Boyce & Reed leg. London, 1 male, 21.x.−28.xi.1996, D.J. Carter & (PPWM). Vicarage Bog, Wrexham, 1 male, 12.x.1988, C.W. Plant leg. (PPCM). Burnham, Buckinghamshire, Holmes, Boyce & Reed leg. (PPWM). Wellesbourne, 6 males, 7.viii.−29.xii.1934 (reared); 1 male, Warwickshire, 2 males, 24.ii.−2.iii.2000; 2 males, 25.viii.1934, all Basden leg. (BMNH, NMS). Burnham 17.viii.−31.viii.2000; 3 males, 24.v.−27.vi.2001; 1 male, Beeches, Buckinghamshire, 1 male, 27.viii.1934 27.vi.−5.vii.2001, all J.E. Smith leg. (HRI). Wicken (reared), Basden leg. (NMS); 1 male, 9.x.−26.x.1996, Fen, Cambridgeshire, 1 male, 12.v.1990, Withers leg. Ismay leg. (PPCM). Chigwell Row Wood, Chigwell (PPWM). Willes Meadow, Leamington Spa, Warwick- Row, Essex, 1 male, 23.v.−27.v.2002; 1 male, shire, 1 male, 27.viii.2002, Menzel leg. (DEI). Wotton- 11.vi.2002; 1 male, 18.vi.2002; 1 male, 15.ix.2002, all under-Edge, Gloucestershire, 1 male, 20.iv.1965, Gur- Ismay & Schulten leg. (DEI, UMO). Chingford, ney leg. (HRI); 1 male, no date, Hussey leg. (HRI). Yeo- Greater London, 6 males, v.1916, Nicholson leg. vil, Somerset, 10 males, 17.v.1995, White leg. (DEI). (BMNH). Cobham, Kent, 1 male, 1 female, 27.ii.1967 No locality details, England, 1 male, 1 female, 1 pupa, (reared), Hutson leg. (BMNH). Collinpark Wood, 2 larvae, no date, Speyer leg. (HRI). Gloucestershire, 1 male, 2.ix.2002, Gibbs leg. (PPCM). East Burnham, Buckinghamshire, 9 males, 14.iii.− Ecological notes: common in broad-leaved and carr 23.v.1933 (reared); 10 males, 15.x.−23.xi.1933; 1 male, woodland; also found in mixed woodland; wet and 17.iii.1934; 1 male, 24.iii.1934; 1 male, 23.v.1934, all humid heathland; wetlands including fen; reed Basden leg. (NMS). Epping Forest, Essex, 5 males, and sedge beds, Sphagnum kettlehole, raised mire 22.iii.-viii.1999, Dagley leg. (PPCM). Farnham Com- and water meadows; gravel pits; old hedges; grass- mon, Buckinghamshire, 2 males, 24.vi.1933; 11 land with pingo pools; unimproved calcareous grass- males, 1.x.−28.x.1933; 3 males, 14.x.1933; 1 male, land; parkland; gardens; grounds of horticultural 24.iv.1934 (all reared), all Basden leg. (BMNH, NMS). research station; school grounds with old poplars; Farnham Royal, Buckinghamshire, 21 males, 3.iv.− decaying vegetation by race track; vegetable detritus 26.vi.1934; 1 male, 18.v.1934; 10 males, 25.vi.− on cliff top with salty spray; shingle beach; green- 27.vi.1934 (all reared), all Basden leg. (BMNH, NMS). houses; mushroom farms. Reared from nests of dor- Glasshouse Crops Research Station, Littlehampton, mouse, mole, moorhen, coot, linnet and blackbird; West Sussex, 1 male, 10.ix.1956; 1 male, rotten fungus-covered wood; daffodil bulbs; haystack; 26.xi.1956; 1 male, 17.xii.1956; 1 male, 30.vii.1957, all old wasp nest; rotting onions; stable manure; mush- Hussey leg. (HRI). Grange-over-Sands, Cumbria, 1 room compost; the fungi Laetiporus sulphureus and male, 20.iii.1996, Birkett leg. (PPCM). Hook Farm, Lepiota cristata; soil from rabbit burrow; crowns of Bromley, Greater London, 1 male, 9.i.1965 (reared), sea kale. i–xii. Chandler leg. (PPCM). Kinsealy Research Centre, Dublin, 3 males, vii.-viii.1986 (reared), Dunne leg. LYCORIELLA (LYCORIELLA) CELLARIS (NMID). Lackham Park, Wiltshire, 1 male, 24.x.2002, (LENGERSDORF, 1934) Chandler leg. (PPCM). Loch Rannoch, Perth & Kin- ross, 1 male, 21.x.2000, Chandler leg. (PPCM). Lon- Synonyms: rufula Tuomikoski, 1959; stramentorum don, Greater London, 2 males, 1921, no collector (Frey, 1948). detail (BMNH). Medmenham, Buckinghamshire, 1 Literature: Basden (1933): 44 [as Sciara agraria, mis- male, 16.i.1932 (reared), Basden leg. (NMS). Oxford identification]; O’Toole (1978): 163 [as Plastosciara University Museum, Oxford, Oxfordshire, 1 male, 1 vanderwieli, misidentification]; Freeman (1983b): female, 19.x.1920, Hamm leg. (BMNH). Pett’s Wood, 31 [in part as Lycoriella agraria, misidentification]; Greater London, 1 male, 22.i.1982; 1 male, no date, Dunne & O’Connor (1989): 65 [as Lycoriella solani, all Freeman leg. (BMNH). Pinkney’s Green, Berk- misidentification]; Laurence (1994): 118 [in part as shire, 1 male, 21.iv.1934 (reared), Basden leg. Lycoriella mali]; Menzel (1998): 22; Chandler (2001): (BMNH). Plymouth, Devon, 1 male, 8.ii.1985, J.H. 201 [as Lycoriella ingenua, misidentification]. Cole leg. (PJCB). Radwell, Hertfordshire, 1 male, 21.x.1917, Edwards leg. (BMNH). Seer Green, Buck- New to Irish list. inghamshire, 13 males, 4.i.−22.ii.1933; (reared), all Material examined: Balmoral Forest, Aberdeenshire, Basden leg. (BMNH, NMS). Skomer Island, Pem- 1 male, 19.vii.−30.vii.1938, Coe leg. (BMNH). Beacon- brokeshire, 1 male, x.1995, A.R. Plant leg. (PPRS).

Stonor, Oxfordshire, 1 male, 12.iii.1933 (reared), LYCORIELLA (LYCORIELLA) INGENUA (DUFOUR, 1839) Basden leg. (NMS). Temple, Berkshire, 8 males, Synonyms: ? agraria (Felt, 1897); bigoti (Laboulbène, − − 1.vi. 18.vi.1931; 1 male, 30.i.1932; 3 males, 3.ii. 1863); brevipila Tuomikoski, 1960; celer (Winnertz, − 15.ii.1932; 1 male, 25.iii.1932; 13 males, 7.vi. 1867); debilis (Winnertz, 1867); decliva (Winnertz, 29.vi.1933; 2 males, 4.ix.1933; 2 males, 4.x.1933; 1 1867); flammulinae (Sasakawa, 1983); flaviventris male, 24.iii.1934; 1 male, 30.iv.1934 (all reared), (Winnertz, 1867); humilis (Winnertz, 1867); mali all Basden leg. (BMNH, NMS). The Loke, (Fitch, 1856); mycorum (Frey, 1948); pauciseta (Felt, Norwich, 2 males, 20.iv.1987; 1 male, 26.iv.1987; 1 1897); ramicola (Kieffer, 1919); segnis (Winnertz, − male, 18.vi.1987; 1 male, 24.x. 25.x.1987; 1 male, 1871); solani (Winnertz, 1871); velox (Winnertz, 1867); − 19.vii.1988; 1 male, 15.viii. 16.viii.1988; 1 male, venusta (Winnertz, 1867). 21.viii.−24.viii.1989; 4 males, 28.iv.−4.v.1990; 1 male, 2.v.−6.v.1992; 1 male, 26.vi.−30.vi.1992; 1 male, Literature: Edwards (1925a): 540 [as Sciara agraria 29.ix.−3.x.1992, all Laurence leg. (DEI, PBLN). and Sciara ? nitidicollis, misidentification]; Edwards Wellesbourne, Warwickshire, 1 male, 24.v.−27.vi.2001, (1934b): 34 [as Sciara (?) fenestralis, misidentifica- J.E. Smith leg. (HRI). Wisley, Surrey, 1 male, no date, tion]; Madwar (1937): 49 [as Sciara fenestralis, misi- Buxton leg. (BMNH). Wye, Kent, 3 males, vii.1932, dentification]; Chandler (1978): 209 [as Lycoriella Austin leg. (BMNH). agraria and Lycoriella solani]; Binns (1981): 417 [as Lycoriella solani and Lycoriella mali]; Freeman Ecological notes: common in broad-leaved woodland; (1983b): 31 [as Lycoriella agraria and Lycoriella ancient woodland with oak, hazel and bramble; conifer solani]; Freeman (1987): 201 [as Lycoriella mali]; Lau- plantations; heathland (heather) with oak and holly; rence (1994): 118 [in part as Lycoriella mali]; Menzel grounds of horticultural research station; gardens; (1998): 22; White et al. (2000): 208; Leigh & Ware island with seabird colonies. Reared from the fungi (2003): 124. Hypholoma fasciculare, Sparassis crispa; soil taken Material examined: Addington, Greater London, 1 from rabbit burrow; nests of blackbird, sand martin, male, 3 females, v.-vi.1943 (reared), Coe leg. (BMNH). song thrush, house sparrow, linnet, rook, moorhen, Afon Bran, Powys, 1 male, 1 female, 11.vi.1979 magpie, little owl, grey squirrel, brown rat and hedge- (reared), Chandler leg. (PPCM). Ashford, Kent, 15 hog; wasp nest; tit nestbox; decoy nest; from mush- males, 25.v.1995, White leg. (DEI). Badger Wood, rooms and poinsettia compost. i–xii. Cambridgeshire, 1 male, x.1998, C.W. Plant leg. (PPCM). Baldock, Hertfordshire, 1 male, ix.-x.1917, LYCORIELLA (LYCORIELLA) FELIX (SCHMITZ, 1918) Edwards leg. (BMNH). Beaconsfield, Buckingham- shire, 1 male, 19.iii.1933 (reared), Basden leg. New to British list. (BMNH). Belstead House, Suffolk, 1 male, 1984 Material examined: Austwick, North Yorkshire, 1 (reared), Evans leg. (PPWM). Berwick-upon-Tweed, male, v.1933, Edwards leg. (BMNH). Northumberland, 1 male, ix.1999, Laurence leg. (PBLN). Boston Common, Norfolk, 1 male, 5.vii.1982, Ecological notes: found on a raised mire. v. Evans leg. (PPWM). Brampton, Cambridgeshire, 4 males, 6 females, 3.v.−4.v.1997 (reared), J.H. Cole leg. LYCORIELLA (LYCORIELLA) INCONSPICUA (PJCB). Bristol, Avon, 3 males, 1998, J.E. Smith TUOMIKOSKI, 1960 leg. (HRI). Bromham, Bedfordshire, 1 male, vi.1983 (reared), Rands leg. (BMNH). Bromley, Greater Lon- Literature: Laurence & James (1997): 166 [as Lycori- don, 5 males, 2 females, i.1966; 1 male, 1 female, ella lundstromi, misidentification]. 2.ix.1966; 6 males, 4 females, xii.1971 (reared), all New to British list. Chandler leg. (NMS, PPCM). Brown’s Folly, Avon, 1 male, 24.viii.2000, Gibbs leg. (PDGB). Burnham Material examined: Cadair Idris, Gwynedd, 3 males, Beeches, Buckinghamshire, 1 male, 28.v.−14.vi.1996; 1 19.vi.1993; 2 males, 3.vii.1993; 1 male, 16.viii.1993; 4 male, 9.x.−26.xi.1996, all Ismay leg. (PPCM). Burton, males, 4.ix.1993; 1 male, 22.x.1993; 1 male, 13.xi.1993, Staffordshire, 9 males, 3 females, 15.viii.1995, White all Laurence leg. (PBLN). Clovelly, Devon, 1 male, leg. (DEI). Castletown, Highland Region, 1 male, 8.v.1936, Edwards leg. (BMNH). Hafod Garregog, 3.viii.1972, Chandler leg. (PPCM). Chattenden Wood, Gwynedd, 1 male, 30.vi.−14.vii.1999, Levey & Pavett Kent, 1 male, 4 females, 26.xi.1966 (reared), Chandler leg. (PPCM). Haytor, Devon, 1 male, 23.vi.2002, J.E. leg. (PPCM). Chigwell Row Wood, Chigwell Row, Smith leg. (HRI). Essex, 2 males, 23.v.−27.v.2002; 1 female, 11.vi.2002; 3 Ecological notes: found in broad-leaved woodland; females, 23.ix.2002; 1 female, 16.x.2002, all Ismay & mixed woodland; moorland on granite; open montane Schulten leg. (DEI, UMO). Costessey, Norfolk, 1 male, habitats. v–xi. 31.vi.1984 (reared), Evans leg. (PPWM). Cothill,

Oxfordshire, 1 male, 14.x.1999, Chandler leg. (NMS). leg. (PPCM). Letchworth, Hertfordshire, 1 male, Crofton Heath, Greater London, 1 male, 1 female, ix.1917, Edwards leg. (BMNH). London, Greater Lon- 16.x.1966, Chandler leg. (NMS). Cuckoo Wood, Downe, don, 1 male, no date, Wycherley leg. (PBLN). Lopham Greater London, 5 males, 5 females, 31.i.1967 Fen, Norfolk, 1 male, vii.1978 (reared), Evans leg. (reared), Chandler leg. (PPCM). Cupar, Fife, 12 males, (PPWM). Manchester, Greater Manchester, 6 males, 27.vii.1995, White leg. (DEI). Ditchley, Oxfordshire, 1 10.xi.1931 (reared), Britten leg. (BMNH). Maynooth, male, v.1937, Edwards leg. (BMNH). Epping Forest, Kildare, 2 males, 2 females, 1988, Al-Amidi leg. Essex, 3 males, 2 females, 18.ix.−20.ix.1998 (reared), (BMNH). Milham Ford School, Oxford, Oxfordshire, 7 Chandler leg. (NMS, PPCM); 1 male, 1.iii.1999, Dag- males, 3 females, v.2001, Webb leg. (HRI). Narford ley leg. (PPCM). Evesham, Worcestershire, 8 males, 6 Hall, Norfolk, 1 male, 2 females, x.1984 (reared), females, 8.viii.1995, White leg. (DEI); 2 males, 1996, Evans leg. (PPCM). Natural History Museum, South J.E. Smith leg. (HRI). Fairﬁeld Experimental Horticul- Kensington, Greater London, 1 male, 17.xi.1891 ture Station, Kirkham, Lancashire, 1 male, 1 female, (reared), Austen leg. (BMNH); 3 males, xi.1933 no date, Hussey leg. (HRI). Farnborough Park, (reared), Edwards leg. (BMNH); 7 males, 1 female, Warwickshire, 1 male, 10.x.2001, Chandler leg. (HRI). xi.1938 (reared), Edelsten leg. (BMNH). New Forest, Farnham Common, Buckinghamshire, 3 males, 28.v.− Hampshire, 3 males, 17.x.1937, Edwards leg. 31.v.1934 (reared), Basden leg. (NMS). Felthorpe (BMNH). North Berwick, East Lothian, 10 males, Wood, Norfolk, 1 male, 1.xi.1983; 1 male, 1984; 1 male, 27.vii.1995, White leg. (DEI). Norwich, Norfolk, 1 15.v.1987 (all reared), all Evans leg. (PPWM). Foxley male, 25.ix.1980; 2 males, 5.iv.1988 (all reared), all Wood, Norfolk, 1 male, ii.1972 (reared), Evans leg. Evans leg. (PPWM); 17 males, 3 females, 15.vi.1995, (PPWM). Gerrard’s Cross, Buckinghamshire, 3 males, White leg. (DEI). Old Warden, Bedfordshire, 2 males, xi.1951 (reared), Buxton leg. (BMNH). Milton of viii.1922 (reared), Edwards leg. (BMNH). Oxford, Campsie, Glasgow, City of Glasgow, 6 males, Oxfordshire, 1 male, 1 female, 21.ii.1923 (reared), 26.vii.1995, White leg. (DEI). Glasshouse Crops Hamm leg. (BMNH). Oxford University Museum, Research Station, Littlehampton, West Sussex, 2 Oxford, Oxfordshire, 1 male, 3.xii.1930, Hamm leg. males, 17.ix.1956; 1 male, 24.ix.1956; 2 males, (UMO). Parham, West Sussex, 2 males, no date, Hus- 3.x.1956; 2 males, 23.x.1956; 1 male, 12.xi.1956; 1 sey leg. (HRI). Pett’s Wood, Greater London, 1 male, male, 30.vii.1957; 25 males, 13.viii.1957; 5 males, 21.xi.−22.xi.1974 (reared), Chandler leg. (PPCM); 1 15.x.1958; 2 males, no date, all Hussey leg. (HRI); 2 male, 22.i.1982, Freeman leg. (BMNH). Port Appin, males, 6.i.1964; 1 male, 1 female, 8.i.1964; 1 female, Argyll & Bute, 1 male, iii.1951 (reared), Pelham-Clin- 10.i.1964; 2 males, 26.iv.1965; 3 males, 4.v.1965; 1 ton leg. (NMS). Preston, Lancashire, 20 males, 2 female, 28.v.1965; 1 male, 1 female, 3.vii.1965; 2 females, 8.vi.1995, White leg. (DEI). Quidenham, Nor- males, 1 female, 9.vii.1965; 1 male, 2.iii.1966, all Gur- folk, 1 male, 31.vii.1987 (reared), Evans leg. (PPWM). ney leg. (HRI); 1 male, 24.ix.1975; 5 males, 2 females, Ripley, Surrey, 1 male, 3 females, 1922 (reared), Fox- 1975; 3 males, 1 female, 1977, all Binns leg. (HRI). Wilson leg. (BMNH). Rothamsted, Harpenden, Hert- Great Missenden, Buckinghamshire, 11 males, fordshire, 4 males, i.1933 (reared), M.A. Watson leg. 24.v.1995, White leg. (DEI). Greenlands, Buckingham- (BMNH); 4 males, 25.iii.1950; 14 males, 27.iii.− shire, 18 males, 4.ix.−25.xi.1933; 2 males, 28.x.1933; 1 28.iii.1950; 2 males, 29.iii.1950; 25 males, 14.iv.− male, 10.xi.1933 (all reared), all Basden leg. (BMNH, 22.iv.1950; 24 males, 1950; 4 males, no date, all Lau- NMS). Hartslock, Oxfordshire, 1 male, 6.v.2000, Raper rence leg. (PBLN). Savernake Forest, Wiltshire, 4 leg. (PDGB). Hickling Broad, Norfolk, 1 male, males, 1 female, x.1971; 1 male, x.1972 (all reared), all 16.iv.1989 (reared), Evans leg. (PPWM). Holt Country Chandler leg. (NMS, PPCM). Selby, North Yorkshire, 6 Park, Norfolk, 1 male, 18.vi.1988 (reared), Evans leg. males, 2 females, 8.vi.1995, White leg. (DEI). Selsdon, (PPWM). Honingham Fen, Norfolk, 1 male, 9.vii.1978 Greater London, 2 males, iv.1936; 1 male, 1.v.1936 (all (reared), Evans leg. (PPWM). Hosey Common, Wester- reared), all Coe leg. (BMNH). Send, Surrey, 2 males, 2 ham, Kent, 1 male, viii.1967; 4 males, 3 females, females, no date, Harris leg. (BMNH). Sevenoaks, xii.1968 (all reared), all Chandler leg. (PPCM). Iver Kent, 2 males, i.1933 (reared), Clout leg. (BMNH). Heath, Buckinghamshire, 2 males, 1 female, Slough, Berkshire, 1 male, 17.x.1942, Richards leg. 13.xi.1914 (reared), Edwards leg. (BMNH). Kendal, (BMNH). Snape Maltings, Suffolk, 1 male, 11.vi.1988 Cumbria, 19 males, 1 female, 27.vii.1995, White leg. (reared), Evans leg. (PPWM). St. David’s Airﬁeld, (DEI). Kilgetty, Pembrokeshire, 18 males, 8.vi.1995, Pembrokeshire, 1 male, 29.ix.1987, Holmes, Boyce & White leg. (DEI). Lackham Park, Wiltshire, 1 male, Reed leg. (PPWM). Taplow, Buckinghamshire, 2 24.v.2002, Chandler leg. (PPCM). Langton Hall, North males, xi.1913 (reared), no collector detail (BMNH). Yorkshire, 1 male, 28.vii.1985, Withers leg. (PPWM). The Loke, Norwich, Norfolk, 2 males, 5.vii.1987; 1 Leckford Estate, Hampshire, 2 males, 1 female, 15.iv.− male, 15.ix.1987; 1 male, 5.v.1988; 1 male, 19.vii.1988; 21.iv.1972 (reared); 1 male, 16.iii.1974, all Chandler 2 males, 15.viii.−16.viii.1988; 1 male, 18.ix.−

20.ix.1988; 1 male, 5.vi.−13.vi.1989; 3 males, 29.ix.− LYCORIELLA (LYCORIELLA) LATISTYLA FREEMAN, 1987 3.x.1992, all Laurence leg. (PBLN). Thursby, Cumbria, Literature: Freeman (1987): 201; Menzel (1998): 22; 13 males, 1 female, 25.vii.1995, White leg. (DEI). Menzel & Mohrig (2000): 397. Thompson Common, Norfolk, 1 male, 1 female, xi.1983 (reared), Evans leg. (PPCM). Torc Falls, Killarney, Material examined: Letchworth, Hertfordshire, 1 Kerry, 1 male, 11.vii.1990, Withers leg. (PPWM). male, v.1917, Edwards leg. [holotype of Lycoriella lati- Wakefield, West Yorkshire, 13 males, 8.vi.1995, White styla] (BMNH). leg. (DEI). Warboys Wood, Cambridgeshire, 1 male, Ecological notes: probably found in house or garden. v. 20.vi.1913 (reared), Fryer leg. (BMNH). Warren Wood, Norfolk, 1 male, 1980 (reared), Evans leg. (PPWM). Wellesbourne, Warwickshire, 3 males, xi.1997; 27 LYCORIELLA (LYCORIELLA) LUNDSTROMI (FREY, 1948) males, 12 females, 15.i.1998; 9 males, 21.i.1998; 3 males, 1 female, 6.i.1999; 3 males, 1 female, 20.i.1999; Literature: Freeman (1983a): 168; Freeman (1983b): 3 males, 11.viii.1999; 1 male, 17.ii.−24.ii.2000; 1 male, 31; Menzel (1998): 22 [as Lycoriella lundstroemi (recte 5.vii.−17.viii.2000; 1 female, 17.viii.−31.viii.2000, all lundstromi)]. J.E. Smith leg. (DEI, HRI). Weston-on-the-Green, Material examined: Beaconsfield, Buckinghamshire, Oxfordshire, 1 male, 9.iii.1923 (reared), Hamm 1 male, 15.iii.1933; 7 males, 21.iii.−25.vi.1933 (all leg. (BMNH). Weybourne Dunes, Norfolk, 1 male, reared), all Basden leg. (BMNH, NMS). Bix Bottom, 13.vi.1988 (reared), Evans leg. (PPWM). White Downs, Oxfordshire, 1 male, 24.v.1934 (reared), Basden leg. Surrey, 3 males, 1 female, xii.1968 (reared), Chandler (NMS). Buckfastleigh, Devon, 1 male, 4.vii.− leg. (PPCM). Windsor Forest, Berkshire, 3 males, 5.vii.1968, Hutson leg. (BMNH). Burnham Beeches, − 4.viii. 5.viii.1999 (reared), Chandler leg. (NMS, Buckinghamshire, 1 male, 15.vi.1931 (reared), Basden PPCM). Windsor Great Park, Berkshire, 1 male, leg. (BMNH); 3 males, 22.vii.−26.xi.1996; 1 male, 31.viii.1980, Chandler leg. (PPCM). Withington, 17.ix.−1.x.1999, all Ismay leg. (HRI, PPCM). Collum Greater Manchester, 1 male, 19.x.1920 (reared), J.H. Green, Buckinghamshire, 1 male, 6.xi.1932; 1 male, Watson leg. (BMNH). Yetminster, Dorset, 40 males, 15.xi.1932 (all reared), all Basden leg. (BMNH, NMS). 31 females, 5.xi.1996, White leg. (DEI). No locality Dropmore, Buckinghamshire, 1 male, 10.v.1933 details, England, 1 female, no date, Speyer leg. (HRI); (reared), Basden leg. (NMS). Epping Forest, Wake Val- 1 male, no date, Hussey leg. (HRI). ley, Essex, 1 male, 18.viii.1998, Dagley leg. (PPCM). Farnham Royal, Buckinghamshire, 1 male, 24.v.1934 Ecological notes: very common in broad-leaved and (reared), Basden leg. (NMS). Fawley, Buckingham- carr woodland; also found in mixed woodland; park- shire, 3 males, 1.vi.−19.vi.1934; 1 male, 25.vi.1934 (all land with ancient oaks; ancient woodland with oak, reared), all Basden leg. (BMNH, NMS). Glen of Drum- hazel and bramble; wooded lake margin; wet and dry loch, Moray, 1 male, 16.viii.1937, Coe leg. (BMNH). heathland; wetlands including fen, sedge bed and Greenlands, Buckinghamshire, 1 male, no date water meadows; grassland with pingo pools; grounds (reared), Basden leg. (NMS). Leckford, Hampshire, 1 of horticultural research station; school grounds with male, 17.iv.1982, S.A. Williams leg. (PPCM). Lye Hill, old poplars; gardens; in houses; pest on mushroom Oxfordshire, 1 male, 20.iii.1911; 2 males, 12.iii.− farms, in rubbish. Reared from the fungi Agaricus 19.iii.1916; 1 male, 17.iv.1920 (all reared), all Hamm campestris, Agrocybe praecox, Amanita rubescens, leg. (BMNH). Marston, Oxfordshire, 1 male, 29.iv.1933 Bjerkandera adusta, Calocybe gambosa, Collybia (reared), Basden leg. (BMNH). Mayford, Surrey, 2 dryophila, Grifola gigantea, decayed Ganoderma males, 28.iv.1934; 2 males, 11.v.1934; 20 males, 14.iv.− applanatum, decayed Polyporus on pine trunk, Hebe- 22.v.1934 (all reared), all Basden leg. (BMNH, NMS). loma species, Hirneola auricula-judae, Hypoxylon Medmenham, Buckinghamshire, 11 males, 30.iv.− rubiginosum, Inonotus cuticularis, I. hispidus, 18.v.1934 (reared), Basden leg. (NMS). Moccas Park, I. nodulosus, I. radiatus, Kuehneromyces mutabilis, Herefordshire, 1 male, 11.vi.2002, Godfrey leg. Lactarius species, Lepiota cristata, Peziza vesiculosa, (PPCM). Temple, Berkshire, 2 males, 7.iv.1932; 1 Pleurotus cornucopiae, old P. ostreatus, Polyporus male, 17.iii.1934 (all reared), all Basden leg. (BMNH, squamosus and other Polyporus species, Russula atro- NMS). The Coombe, Buckinghamshire, 1 male, purpurea, R. pectinata, R. vesca, Trametes versicolor, 17.vi.1997, Chandler leg. (PPCM). The Loke, Norwich, rotten onions, oak root gall; nests of rabbit, grey squir- Norfolk, 1 male, 8.v.1987, Laurence leg. (PBLN). War- rel and brown rat; potatoes badly attacked by scab; ren Row, Berkshire, 2 males, 7.iv.1932; 3 males, 7.iv.− propolis and wax in beehive; slug Testacella; flower 22.iv.1932 (all reared), all Basden leg. (BMNH, NMS). heads of hoary ragwort; dead poplar bark; decaying bulbs; rotten oak stump; narcissus bulbs; rhubarb Ecological notes: found in ancient beech and other roots; rotten potatoes. i–xi. broad-leaved woodland; garden; parkland. Reared

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 114 F. MENZEL ET AL. from nests of brown rat, barn owl, linnet, sand martin, Literature: Haliday (1833): 148; Walker (1848): 108; mole, house sparrow, rabbit. iii–x. Walker (1856): 55 [all as Sciara]; Edwards (1925a): 534; Parmenter (1950): 110 [all as Phorodonta]; Lau- rence (1994): 118; Freeman (1983b): 32; Menzel LYCORIELLA (LYCORIELLA) MICRIA MOHRIG & (1998): 22; Coldwell (1999): 25. MENZEL, 1990 New to British list. Material examined: Aber Mawr, Pembrokeshire, 1 male, 13.x.1987, Holmes, Boyce & Reed leg. (PPWM). Material examined: Burnham Beeches, Buckingham- Alvecote Pools, Staffordshire, 1 male, 13.x.2001, J.E. shire, 1 male, 22.vii.−5.viii.1996, Ismay leg. (PPCM). Smith leg. (HRI). Badbury Rings, Dorset, 1 male, Temple, Berkshire, 2 females, 18.vi.1934; 2 females, 1.vii.1966, Parmenter leg. (BMNH). Bagley Wood, 27.vi.1934; 7 males, 13.ix.1934 (all reared), all Basden Oxfordshire, 1 female, 21.v.1961, K.G.V. Smith leg. leg. (NMS). (BMNH). Ban-y-Gor Wood, Gloucestershire, 1 male, Ecological notes: found in ancient beech woodland. 12.ix.2002, Gibbs leg. (PDGB). Barra, Western Isles, 1 Reared from rotten lime wood. iv–ix. female, 17.viii.−28.viii.1936, A.R. Waterston leg. (NMS). Bedgebury Pinetum, Kent, 1 male, 27.x.1968, Chandler leg. (NMS). Betws-y-Coed, Conwy, 1 male, LYCORIELLA (LYCORIELLA) PARVA (HOLMGREN, 1869) 24.ix.1966, Vane-Wright leg. (BMNH). Bix Bottom, Synonyms: curvispina Tuomikoski, 1960; difficilis var. Oxfordshire, 1 male, 23.ix.2000, Raper leg. (PDGB). obscuratipes (Frey, 1948). Blaise Wood, Avon, 1 female, 24.vi.1949, d’Assis-Fon- seca leg. (NMS). Blean Wood, Kent, 2 females, 24.vi.− Literature: Laurence (1996): 87; Laurence & James 29.vi.1964; 6 males, 14.vii.1964; 1 male, 23.vii.1964; 1 (1997): 166 [all as Lycoriella mali, misidentification]. female, 20.iv.1965; 1 female, 17.vi.1965; 2 males, New to British list. 14.ix.1965; 11 females, 19.v.1973, all Parmenter leg. (BMNH). Bolt Wood, Stockerston, Leicestershire, 1 Material examined: Austwick, North Yorkshire, 3 male, 24.viii.2003, J.E. Smith leg. (HRI). Bonhill, West males, v.1933, Edwards leg. (BMNH). Cadair Idris, Dunbartonshire, 1 female, 8.viii.1906; 1 female, Gwynedd, 13 males, 19.vi.1993; 2 males, 3.vii.1993; 1 25.viii.1906; 1 male, 1 female, 27.viii.1907; 1 female, male, 1.viii.1993; 51 males, 4.ix.1993, all Laurence leg. 7.ix.1907; 1 male, 5.vii.1909, all Malloch leg. (NMS). (PBLN). Bookham Common, Surrey, 2 males, 14.x.1945; 6 Ecological notes: found on raised mire; open montane males, 1 female, 14.xi.1948; 2 males, 16.xi.1948; 3 habitats. v–ix. males, 1 female, 12.xii.1948; 1 male, 8.iv.1950; 1 male, 8.x.1950; 1 male, 10.v.1964; 1 male, 12.v.1964, all Par- menter leg. (BMNH). Brackloon Wood, Mayo, 1 male, LYCORIELLA (LYCORIELLA) SUBTERRANEA 4.viii.1911, Grimshaw leg. (NMS). Brancaster, Nor- (MÄRKEL, 1844) folk, 5 males, 1.vii.−15.vii.1988; 4 males, 2.ix.1988, all Synonym: vanderwieli (Schmitz, 1920). Foster & Procter leg. (PBLN). Brandon Marsh, Ryton- on-Dunsmore, West Midlands, 2 males, 11.x.2001, Literature: Freeman (1983b): 31 [as Lycoriella Chandler leg. (HRI); 3 males, 4 females, 22.viii.2002, vanderwieli]. Menzel & J.E. Smith leg. (DEI, HRI). Brithdir-Isaf, Material examined: Burnham Beeches, Buckingham- Gwynedd, 9 males, 13 females, 25.x.1971, Hutson shire, 1 male, 26.ix.−9.x.1996, Ismay leg. (PPCM). leg. (BMNH). Brockenhurst, Hampshire, 1 male, Oxshott, Surrey, 1 male, 1902, Donisthorpe leg. 27.v.1910, Yerbury leg. (UMO); 1 female, 23.v.1969, (BMNH). Tubney, Oxfordshire, 3 males, 20.iv.1908, Hutson leg. (BMNH). Burnham Beeches, Bucking- Hamm leg. (BMNH). hamshire, 1 female, 6.v.1914, Edwards leg. (BMNH). Butley Thicks, Suffolk, 1 male, 5.vi.1910, Collin leg. Ecological notes: found in ancient beech and other (UMO). Canonteign Falls, Devon, 2 males, 22.vi.2002, broad-leaved woodland; heathland and pine woodland; J.E. Smith leg. (HRI). Catfield, Norfolk, 3 males, in ant Formica rufa nests. iv, ix–x. 13.x.−12.xii.1988, Foster & Procter leg. (PBLN). Char- leville Wood, Offaly, 1 male, 15.vi.1985, Chandler leg. (PPWM). Charterhouse, Surrey, 1 male, 26.vi.1967, PHYTOSCIARA FREY, 1942 Stubbs leg. (PPCM). Chippenham Fen, Cam- PHYTOSCIARA (DOLICHOSCIARA) FLAVIPES bridgeshire, 1 male, 14.ix.1943, Collin leg. (UMO); 1 (MEIGEN, 1804) female, 4.xi.1984, J.H. Cole leg. (PJCB). Chipstead, Synonyms: flavipes var. nigrithorax (Strobl, 1898); Surrey, 1 male, 10.v.1965, Parmenter leg. (BMNH). fugax (Grzegorzek, 1884). Cleeve Heronry, Avon, 1 male, 11.ix.2000, Gibbs leg.

(PDGB). Clovelly, Devon, 3 males, 8.v.1936, Edwards (BMNH). Hollybush Hill, Stoke Poges, Buckingham- leg. (BMNH). Coed Dol y Bebin, Gwynedd, 1 male, shire, 1 male, 29.x.1967, Chandler leg. (PPCM). Hors- 12.vii.1976, J.H. Cole leg. (PJCB). Coed y Brenin, ley, Surrey, 1 male, 21.v.1966, Chandler leg. (PPCM). Gwynedd, 1 female, 18.ix.1968, Vane-Wright leg. Hosey Common, Westerham, Kent, 1 female, (BMNH). Collinpark Wood, Gloucestershire, 1 male, 25.ix.1966, Chandler leg. (NMS). Hothfield Bog, Kent, 3.ix.2002, Gibbs leg. (PDGB). Comin Esgair Maen, 1 female, 17.x.1970, Vane-Wright leg. (BMNH). Ceredigion, 1 male, 23.vii.1987, Holmes, Boyce & Howth, Dublin, 1 male, 1 female, 17.vii.1971, Chan- Reed leg. (PPWM). Coneyberry Wood, Goring-on- dler leg. (NMS, PPCM). Ivybridge, Devon, 2 females, Thames, Oxfordshire, 1 female, 17.v.1964, J.H. Cole 18.v.1890, Verrall leg. (BMNH, UMO). Killarney, leg. (PJCB). Cors Caron, Ceredigion, 1 male, Kerry, 1 male, 5.viii.1940, Beirne leg. (BMNH). Kil- 27.x.1987, Holmes, Boyce & Reed leg. (PPWM). Cors laun Bog, Offaly, 3 males, 7.ix.1999, Withers leg. Erddreiniog, Isle of Anglesey, 1 male, 27.vii.1988; 1 (PPWM). Kingsbury Water Park, Warwickshire, 1 male, 20.x.1988, all Holmes, Boyce & Reed leg. male, 13.x.2001, Chandler leg. (HRI). Knebworth, (PPWM). Cors Geirch, Gwynedd, 1 male, 28.vii.1988, Hertfordshire, 1 female, 12.x.1920; 1 male, 10.vi.1923, Holmes, Boyce & Reed leg. (PPWM). Cors Gorsgoch, all Edwards leg. (BMNH). Knole Park, Kent, 1 male, Ceredigion, 1 male, 15.vii.1987, Holmes, Boyce & 12.xi.1967; 1 male, 16.x.1968, all Chandler leg. Reed leg. (PPWM). Cors Graianog, Gwynedd, 1 male, (PPCM). Lancaut Nature Reserve, Gloucestershire, 1 28.vii.1988; 1 male, 7.x.1988, all Holmes, Boyce & male, 1 female, 5.ix.2002, Gibbs leg. (PDGB). Leigh Reed leg. (PPWM). Coulsdon, Greater London, 1 male, Wood, Avon, 1 male, 19.x.2000, Gibbs leg. (PDGB). 23.iii.1957, Parmenter leg. (BMNH). Cuckoo Wood, Lelant, Cornwall, 3 males, 2 females, 22.viii.− Downe, Greater London, 1 male, 25.ix.1966, Chandler 31.viii.1912, Yerbury leg. (BMNH). Lephinmore, leg. (PPCM). Cudham, Greater London, 2 males, Argyll & Bute, 1 male, 1.x.1951, Pelham-Clinton leg. 27.viii.1966, Vane-Wright leg. (BMNH). Dartmouth, (NMS). Limpsfield Common, Surrey, 1 male, Devon, 1 female, 5.vi.−12.vi.1920, Edwards leg. 4.vii.1937; 3 males, 24.v.−28.vi.1964, all Parmenter (BMNH). Dolgellau, Gwynedd, 1 female, 13.vi.1887; 1 leg. (BMNH). Linwood, New Forest, Hampshire, 1 male, 23.vii.1888; 1 male, 24.vii.1888, all Verrall leg. female, 18.iv.1954, Colyer leg. (BMNH). Llay, Wrex- (BMNH, UMO). Dowrog Common, Pembrokeshire, 1 ham, 1 male, 31.vi.1972, Stubbs leg. (BMNH). Llyn male, 29.ix.1987; 1 male, 13.x.1987, all Holmes, Boyce Coethlyn, Powys, 1 male, 11.x.1988, Holmes, Boyce & & Reed leg. (PPWM). Ewden Valley, Sheffield, South Reed leg. (PPWM). Llyn Gwynant, Gwynedd, 1 Yorkshire, 1 male, 21.ix.1971, Chandler leg. (PPCM). female, 13.viii.1914, Edwards leg. (BMNH). Lodore, Farningham Wood, Kent, 1 male, 18.v.1974, Stubbs Cumbria, 1 female, 21.vi.1889, Verrall leg. (UMO). leg. (PPCM). Felden, Hertfordshire, 4 males, 21.xi.− Lord’s Wood, Avon, 1 male, 25.vi.2002, Gibbs leg. 30.xi.1894, A. Piffard leg. (BMNH). Fernworthy Res- (PDGB). Lough Conn, Mayo, 1 male, 20.ix.2000, ervoir, Dartmeet, Devon, 1 male, 17.viii.1986, Banks Chandler leg. (PPCM). Lundy Island, Devon, 2 males, & Vane-Wright leg. (BMNH). Ffrith, Flintshire, 1 2 females, 20.vii.−22.vii.1972, Lane leg. (BMNH); 1 male, 7.vi.−9.vi.1919, Edwards leg. (BMNH). Fisher’s male, 26.vii.1972, Hutson leg. (BMNH). Lyndhurst, Wood, Bromley, Greater London, 1 female, 8.viii.1966, Hampshire, 1 male, 11.viii.1885, Verrall leg. (UMO); 1 Chandler leg. (PPCM). Folly Farm, Avon, 1 male, female, 27.iii.1901, Adams leg. (UMO); 1 male, 12.ix.2000; 1 male, 28.ix.2001, all Gibbs leg. (PDGB). 5.v.1911, Collin leg. (UMO). Matley Bog, New Forest, Frant, East Sussex, 1 male, 16.vi.1887, Verrall leg. Hampshire, 1 female, viii.1936, Edwards leg. (BMNH). Glencoyne Wood, Ullswater, Cumbria, 1 (BMNH). Monk Wood, Avon, 1 male, 24.v.2000; 1 male, male, 1 female, 8.x.1971, Hutson leg. (BMNH). Glen- 22.ix.2000, all Gibbs leg. (PDGB). Monk’s Wood, Cam- dalough, Wicklow, 1 male, 1 female, 13.xi.1984, Chan- bridgeshire, 1 male, 10.vii.1972; 1 male, 25.vii.1972, dler leg. (PPCM). Glendalough State Forest, Wicklow, all J.H. Cole leg. (PJCB). Murrough, Wicklow, 1 male, 2 females, 17.ix.1968, Chandler leg. (PPCM). Glengar- 8.v.1989, Withers leg. (PPWM). Myross Wood, Leap, riff State Forest, Cork, 1 male, 4.vii.1969, Chandler Cork, 1 male, 11.viii.1977, Chainey leg. (BMNH). New leg. (NMS). Godmanchester, Cambridgeshire, 1 male, Forest, Hampshire, 5 females, 10.ix.−4.x.1896; 1 male, 1 female, 29.v.2000, J.H. Cole leg. (PJCB). Hackhurst 25.iii.1897; 1 female, 5.v.1897; 1 female, 25.ix.1897; 1 Down, Surrey, 1 male, 27.viii.1966, Chandler leg. female, 22.iv.1901; 1 female, 7.ix.1907; 1 female, (PPCM). Hay Wood, Baddesley Clinton, Solihull, West 16.vii.1909; 1 female, 27.vii.1909; 3 males, 26.ix.− Midlands, 1 male, 25.viii.2002, J.E. Smith leg. (HRI). 28.ix.1909; 1 female, 17.x.1909; 1 female, 13.iv.1910; Hayley Wood, Cambridgeshire, 1 male, 31.v.1975, all Adams leg. (NMS, BMNH); 1 male, 18.x.1937, J.H. Cole leg. (PJCB). Helston, Cornwall, 1 male, Edwards leg. (BMNH); 1 male, 12.v.1951, Colyer leg. 11.v.1890, Verrall leg. (UMO). Hitch Wood, Hertford- (BMNH); 1 male, 20.v.1966; 1 male, 16.v.1968; 2 shire, 1 male, 9.vii.1964, Colyer leg. (BMNH). Holt males, 18.v.1968; 1 male, 31.v.1968, all Parmenter leg. Forest, Dorset, 1 male, 13.v.1951, no collector detail (BMNH). Newmarket, Cambridgeshire, 1 female,

20.viii.1901, Verrall leg. (UMO). Nuremore Lake, Car- Wells, Kent, 1 female, 12.vi.1886, Verrall leg. rickmacross, Monaghan, 1 female, 19.vi.1979, J.H. (BMNH); 1 male, iv.1925; 1 male, vi.1925, all Nurse Cole leg. (PJCB). Oaken Wood, Barming, Kent, 1 male, leg. (BMNH). Valency Valley, Cornwall, 1 male, 1 20.ix.1966, Chandler leg. (NMS). Old Coulsdon, female, 29.v.2000, J.H. Cole leg. (PJCB). Vaynor Greater London, 1 female, 9.ix.1954, Britten leg. Wood, Gwynedd, 1 male, 2.ix.1966, Vane-Wright (BMNH). One Tree Hill, Sevenoaks, Kent, 1 male, leg. (BMNH). Warren’s Court Wood, Cork, 1 male, 5.v.1968, Chandler leg. (PPCM). Oxford University 11.vii.1994, Withers leg. (PPWM). Whichford Wood, Museum, Oxford, Oxfordshire, 1 male, 1.xii.1920; 1 Moreton-in-Marsh, Warwickshire, 3 males, 13.x.2001, female, 2.xii.1920; 1 female, 3.i.1921; 1 female, J.E. Smith & Chandler leg. (HRI). Whitacre Heath, 27.i.1921; 1 male, 4.x.1922, all Hamm leg. (UMO). Warwickshire, 2 males, 13.x.2001, J.E. Smith & Chan- Oxford University Parks, Oxford, Oxfordshire, 1 dler leg. (HRI). Whitmoor Vale, Hampshire, 1 male, female, 9.vii.1931, Hamm leg. (UMO). Pamber Forest, 12.vii.1990, Chandler leg. (NMS). Wolford Wood, Hampshire, 1 male, 20.vi.1967, Stubbs leg. (PPCM); 1 Moreton-in-Marsh, Warwickshire, 1 male, 11.x.2001, male, 5.v.1971, J.H. Cole leg. (PJCB). Pennington, Chandler leg. (HRI). Woodditton Wood, Cam- Hampshire, 1 male, 12.v.1951, Colyer leg. (BMNH). bridgeshire, 1 female, 12.v.1920; 1 male, 8.v.1929; 1 Pentyrch, Cardiff, 1 female, 27.v.1968, Chandler leg. male, 6.v.1939, all Collin leg. (UMO). Wotton, Surrey, (PPCM). Penzance, Cornwall, 1 female, 14.v.1890, Ver- 1 male, 27.viii.1966, Chandler leg. (NMS). Wyre For- rall leg. (UMO). Peppercombe, Devon, 1 female, est, Worcestershire, 2 females, 14.ix.1922, Edwards 14.vi.1989, J.H. Cole leg. (PJCB). Pond Wood, Chisle- leg. (BMNH); 1 male, 2.viii.1974, Ackland leg. (NMS). hurst, Greater London, 1 female, 7.xi.1966, Chandler Wytham Woods, Wytham, Oxfordshire, 1 male, 1 leg. (NMS). Reedham, Norfolk, 4 males, 27.x.1988, female, 21.viii.2002, J.E. Smith leg. (HRI). No locality Foster & Procter leg. (PBLN). Rhineﬁelds, New For- details [? Holywood, Down], 2 specimens [without est, Hampshire, 1 male, 10.iv.1966, Colyer leg. abdomen], no date, Haliday leg. (NMID). (BMNH). Rhôs Rhydd, Ceredigion, 1 male, 8.x.1987, Ecological notes: very common in broad-leaved, mixed Holmes, Boyce & Reed leg. (PPWM). Rickmansworth, and carr woodland; arboretum of conifers; grazed wet Hertfordshire, 2 males, 29.vii.−31.vii.1919, Wallace heathland; wetlands especially fen and bog; also reed leg. (BMNH). Ridley Wood, Hampshire, 1 female, and sedge beds, valley and basin mire, pond and lake- 9.v.1964, Colyer leg. (BMNH). River East Lyn, Rock- side marshes; former gravel workings; calcareous ford, Devon, 1 male, 16.vi.1989, J.H. Cole leg. (PJCB). grassland; parkland. iii–xii. Ryton Wood, Bubbenhall, Coventry, West Midlands, 1 male, 25.viii.2002, J.E. Smith leg. (HRI). Scilly Is, Cornwall, 1 female, iii.1902, Edwards leg. (BMNH). PHYTOSCIARA (DOLICHOSCIARA) ORNATA Sheviock, Cornwall, 2 males, 1 female, 9.ix.− (WINNERTZ, 1867) 14.ix.1912, Yerbury leg. (BMNH). Shortwood, Glouces- tershire, 1 male, 27.vi.2002, Gibbs leg. (PDGB). Sho- New to Irish list. tover, Oxfordshire, 2 males, 1 female, 5.viii.1918, Material examined: Gearagh, Cork, 1 male, 4.v.1990, Hamm leg. (UMO). Simonshyde, Hertfordshire, 1 de Courcy Williams leg. (DEI). female, 16.iv.1957, Colyer leg. (BMNH). Sontley Marsh, Wrexham, 1 male, 19.vii.1988, Holmes, Boyce Ecological notes: found in damp broad-leaved wood- & Reed leg. (PPWM). Stock Ghyll, Cumbria, 4 males, 2 land. v. females, 5.x.1971, Hutson leg. (BMNH). Stoke Poges, Buckinghamshire, 1 male, 29.x.1967, Chandler leg. PHYTOSCIARA (DOLICHOSCIARA) SAETOSA (NMS). Stonybrow Wood, Rotherﬁeld Park, Hamp- (LENGERSDORF, 1929) shire, 1 male, 2.x.2002, Chandler leg. (PPCM). Strel- ley, Nottinghamshire, 1 male, 21.ix.1922, Edwards New to British list. leg. (BMNH). Symond’s Yat, Herefordshire, 1 male, Material examined: Thompson Common, Thompson, ix.1936, Edwards leg. (BMNH). Three Bridges, West Norfolk, 1 male, 13.ix.−27.ix.1988, Foster & Procter Sussex, 1 female, 11.viii.1888; 2 males, 1.viii.1889; 1 leg. (PBLN). male, 15.vi.1897, all Verrall leg. (UMO). Thundersley Glen, Essex, 1 male, 30.iv.1986, Withers leg. (PPWM). Ecological notes: found in a site comprising broad- Tickenham Hill, Avon, 1 male, 30.v.2001, Gibbs leg. leaved woodland and grassland with pingo pools. ix. (PDGB). Tocil Wood, Warwick University Campus, Coventry, Warwickshire, 1 male, 28.viii.2003, J.E. Smith leg. (HRI). Torc Falls, Killarney, Kerry, 1 male, PHYTOSCIARA (PHYTOSCIARA) HALTERATA 11.vii.1990, Withers leg. (PPWM). Torcross, Devon, 1 (LENGERSDORF, 1926) female, 26.iv.1909, Verrall leg. (UMO). Tunbridge Literature: Freeman (1983b): 31; Menzel (1998): 22.

Material examined: Headington Hill, Oxfordshire, 1 leburgh, Norfolk, 1 male, 6.vii.1987; 1 male, 7.v.1988, male, 31.viii.1916, Hamm leg. (BMNH). Spartum Fen, all Withers leg. (PPWM). Epping Thicks, Epping For- Oxfordshire, 1 male, 13.v.−2.vi.1988, Porter leg. est, Essex, 1 male, v.1999, Dagley leg. (PPCM). Letch- (PPCM). Warburton, Greater Manchester, 1 female, worth. Hertfordshire, 1 male, 2.vii.1937, Edwards leg. 10.vi.1937 (reared); 1 male, 15.vi.1937 (reared), all (BMNH). Rogate, West Sussex, 1 male, 10.vi.1979, Cohen leg. (BMNH). Chandler leg. (PPCM). Wicken Fen, Cambridgeshire, 1 male, 12.v.1990, Withers leg. (PPWM). Wildmoor Ecological notes: found in carr woodland. Reared from Heath, Berkshire, 1 male, 15.x.1999, Chandler leg. larvae feeding exposed on leaves of buttercup. v–vi, (NMS). viii. Ecological notes: found in ancient broad-leaved, mixed and carr woodland; heathland; fen meadows and PHYTOSCIARA (PROSCIARA) PRODUCTA ditches; gardens. v–viii, x. TUOMIKOSKI, 1960 Literature: Ashe & O’Connor (1991): 16; Menzel (1998): 22. PNYXIA JOHANNSEN, 1912 Material examined: Epping Thicks, Epping Forest, PNYXIA SCABIEI (HOPKINS, 1895) Essex, 3 males, 3.v.−9.v.1999; 2 males, iv.1999; 1 male, Synonyms: dispar Schmitz, 1927; ruffoi (Venturi, vii.1999, all Dagley leg. (HRI, PPCM). Killoughrim 1970); subterranea (Schmitz, 1913). Forest, Wexford, 1 male, 4.vi.1987, J.P. O’Connor leg. Literature: Edwards (1925a): 584; ? Donisthorpe (NMID). Lockhaugh Bank Wood, Derwent Valley, Tyne (1927): 139 [as Allostoomma subterranea]; Skidmore & Wear, 1 male, 7.vi.−23.vi.1990, Jessop leg. (PBLN). (1978): 76; Freeman (1983b): 27; Skidmore (1991): 101 The Loke, Norwich, Norfolk, 1 male, 23.vi.1988, Lau- and 149; Menzel (1998): 22. rence leg. (PBLN). Material examined: Cheshunt, Hertfordshire, 3 Ecological notes: found in beech, oak and other broad- females, 31.v.1922; 1 female, 1.viii.1922 (all reared), leaved woodland; garden. iv–vii. all Speyer leg. (HRI). Enﬁeld Wash, Greater London, 2 males, 5 larvae, 12.i.1922, Speyer leg. (HRI). Lifton PHYTOSCIARA (PROSCIARA) PROSCIAROIDES Park, Devon, 5 males, 30.x.1935 (reared), Bradshaw (TUOMIKOSKI, 1960) leg. (BMNH). Natural History Museum, South Kens- ington, Greater London, 1 male, 4.viii.1931, Edwards Literature: Freeman (1983a): 169; Freeman (1983b): leg. (BMNH). No locality details, ? England, 2 males, 37 [all as Bradysia]; Menzel (1998): 22. 3.vii.1887 (reared), Verrall leg. (BMNH); England, 2 Material examined: How Hill, Norfolk, 1 male, males, 2 females, 2 larvae, no date, Speyer leg. (HRI). 13.viii.1989, Withers leg. (PPWM). Sutton Broad, Nor- Ecological notes: found in museum; pot plants. Reared folk, 1 male, 21.viii.−1.ix.1989, Foster & Procter leg. from narcissus bulbs; cucumber; potato and tomato (PPCM). Woodbastwick, Norfolk, 1 male, viii.1932; 1 seedlings; manure. i, v, vii–viii, x. male, viii.1993, all Edwards leg. (BMNH). Ecological notes: found in damp broad-leaved woodland; carr woodland and fen; peat hollows and fen. PSEUDOLYCORIELLA MENZEL & MOHRIG, 1998 viii–ix. PSEUDOLYCORIELLA BRUCKII (WINNERTZ, 1867) Synonym: virgultorum (Winnertz, 1867). PHYTOSCIARA (PROSCIARA) UNGULATA Literature: Freeman (1983b): 30 [as Lycoriella (WINNERTZ, 1867) (Bradysiopsis)]; Menzel, 1998: 22; Chandler (2001): Literature: Freeman (1983b): 32; Ashe & O’Connor 201. (1991): 16; Menzel (1998): 22; Chandler (2001): 201. Material examined: Beech, Devil’s Ditch, Cam- Material examined: Brampton Wood, Cambridgeshire, bridgeshire, 4 males, 15.vi.1936, Collin leg. (UMO). 2 males, 7.vi.1986, J.H. Cole leg. (PJCB). Buckingham Bishop’s Stortford, Hertfordshire, 1 male, vi.1993; 1 Palace Garden, Greater London, 2 males, 18.vii.− male, vi.1994, all C.W. Plant leg. (PPCM). Brasenose, 13.viii.1997, D.J. Carter & C.W. Plant leg. (PPCM, Oxfordshire, 1 male, 12.vi.1915, Hamm leg. (UMO). PPWM). Burnham Beeches, Buckinghamshire, 1 Buckingham Palace Garden, Greater London, 1 male, male, 28.v.−14.vi.1996; 1 male, 17.v.−25.v.1999, all 23.vi.−11.vii.1996; 1 male, 27.vi.−11.viii.1996; 3 males, Ismay leg. (PPCM). California Country Park, Berk- 26.v.−17.vi.1997, all D.J. Carter & C.W. Plant leg. shire, 1 male, 19.v.1998, Chandler leg. (PPCM). Dick- (NMS, PPCM). Buckingham Thick Copse, Northamp-

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 118 F. MENZEL ET AL. tonshire, 1 male, 29.vi.1992, Warne leg. (NMS). Burr- Material examined: Bedford Purlieus, Cam- ington Combe, Somerset, 1 male, 2.vii.1985, J.H. Cole bridgeshire, 1 male, 20.vi.1985; 1 male, 23.vi.1985, all leg. (PJCB). Cambridge, Cambridgeshire, 1 male, J.H. Cole leg. [including holotype of Corynoptera pol- 13.vi.1904, Jenkinson leg. (BMNH). Cothill, Oxford- liciformis] (BMNH, PJCB). Brockenhurst, Hampshire, shire, 1 male, 13.vii.1985, Chandler leg. (NMS). 1 male, 10.vii.1907, King leg. (BMNH). Burnham Epping Forest, Essex, 1 male, vi.1999, Dagley leg. Beeches, Buckinghamshire, 1 male, 8.vi.−18.vi.1999, (PPCM). Letchworth, Hertfordshire, 1 male, v.- Ismay leg. (HRI). Charlbury, Oxfordshire, 6 males, vi.1925; 1 female, 27.vi.1926, all Edwards leg. 4.vii.1936, Edwards leg. (BMNH). Coed Gorswen, (BMNH). Little Paxton Pits, Cambridgeshire, 1 male, Gwynedd, 1 male, 6.vii.1974, Chandler leg. (PPCM). 15.vi.1993, J.H. Cole leg. (PJCB). Milham Ford School, Monk’s Wood, Cambridgeshire, 1 male, 22.vi.1975, Oxford, Oxfordshire, 1 male, 20.vi.2002, Webb leg. J.H. Cole leg. (PJCB). Pingle Wood, Warboys, Cam- (PPCM). Oxford University Museum, Oxford, Oxford- bridgeshire, 1 male, 25.vi.1988, J.H. Cole leg. (PJCB). shire, 1 male, 24.vi.1926, Hamm leg. (BMNH). Sharp- Ecological notes: found in ancient beech and other enhoe, Bedfordshire, 1 male, 11.vii.1997, Chandler broad-leaved woodland. vi–vii. leg. (PPCM). Sheep Leas, West Horsley, Surrey, 1 male, 16.vi.1991, Chandler leg. (PPCM). Therﬁeld Heath, Hertfordshire, 1 male, 29.v.1993, J.H. Cole leg. PSEUDOLYCORIELLA SEMIALATA (EDWARDS, 1913) (PJCB). Wansford Meadow, Cambridgeshire, 1 male, 1 female, 24.vi.1989, J.H. Cole leg. (PJCB). Wicken Literature: Edwards (1913a): 210; Edwards (1925a): Fen, Cambridgeshire, 2 males, 15.vi.1990; 1 male, 538; Madwar (1937): 47; Skidmore (1978): 76 [all as 5.vii.1991, all J.H. Cole leg. (PJCB). Wye Downs, Kent, Sciara]; Freeman (1983a): 169; Freeman (1983b): 23; 1 male, 6.vi.1936, Collin leg. (UMO). Freeman (1987): 197 [all as Heterosciara]; Menzel (1998): 22; Menzel & Mohrig (2000): 478. Ecological notes: found in broad-leaved and carr woodland; fen meadows and ditches; common on calcareous Material examined: Brampton, Cambridgeshire, 8 − grassland; stream in rocky gully; school grounds with males, 9 females, 25.iv. 27.iv.1997 (reared), J.H. Cole old poplars; gardens. v–viii. leg. (PJCB, PPCM). Clapham, West Sussex, 4 males, 8 females, 26.iii.1913 (reared), Collins leg. (BMNH). Knott Wood, Rothamsted, Hertfordshire, 4 males, PSEUDOLYCORIELLA BRUNNEA 22.v.−23.v.1950; 3 males, 23.v.1950 (reared), all Lau- (BUKOWSKI & LENGERSDORF, 1936) rence leg. (BMNH, PBLN). Norwood, Greater London, New to British and Irish list. 12 males, 9 females, 22.iv.1913, Distant leg. [including lectotype and paralectotypes of Sciara semialata] Material examined: Coed Tycanol, Pembrokeshire, 1 (BMNH, ZFMK). male, 10.vi.−29.vi.1999, Levey & Pavett leg. (PPCM). Newbridge Fen, Kildare, 2 males, 8.viii.−19.viii.1998, Ecological notes: found in grounds of horticultural Withers leg. (PPWM). research station and garden. Reared from cow dung and soil on dead plant roots. iii–v. Ecological notes: found in broad-leaved woodland and open fen. vi, viii. PSEUDOLYCORIELLA SUBBRUCKII (MOHRIG & HÖVERMEYER, 1992) PSEUDOLYCORIELLA KOREENSIS (MOHRIG & MENZEL, 1992) Literature: Laurence (1994): 110 [as Lycoriella (Bradysiopsis)]; Menzel (1998): 22. New to British list. New to Irish list. Material examined: Stokeford Heath, Dorset, 1 male, 20.v.2001, Gibbs leg. (PDGB). Material examined: Aberithon Turbary, Powys, 1 male, 19.vi.1989; 1 male, 3.vii.1989; 1 male, Ecological notes: found on sandy heathland. v. 17.vii.1989, all Holmes, Boyce & Reed leg. (DEI). Blaen y Wergloedd, Conwy, 1 male, 1 female, PSEUDOLYCORIELLA PALUDUM (FREY, 1948) 4.viii.1988, Holmes, Boyce & Reed leg. (PPWM). Cat- ﬁeld, Norfolk, 2 males, 21.vi.−5.vii.1988; 5 males, 2 Synonyms: leucocera (Mohrig & Menzel, 1990); pollic- females, 28.viii.−29.ix.1988; 1 male, 29.ix.−12.x.1988, iformis (Freeman, 1990). all Foster & Procter leg. (DEI, PBLN). Colwyn Brook, Literature: Freeman (1990): 54 [as Corynoptera pollic- Powys, 1 male, 20.vi.1989, Holmes, Boyce & Reed leg. iformis]; Menzel (1998): 22; Menzel & Mohrig (2000): (PPWM). Cors Bodeilio, Isle of Anglesey, 1 male, 474. 26.vii.1988, Holmes, Boyce & Reed leg. (PPCM). Cors

Erddreiniog, Isle of Anglesey, 9 males, 29.vi.1988; 1 [as Sciara (Scatopsciara) vivida]; Buxton (1954): 167 male, 14.vii.1988; 5 males, 27.vii.1988; 1 male, [as Scaptosciara vivida (recte Scatopsciara)]; Chan- 6.x.1988, all Holmes, Boyce & Reed leg. (PPWM). dler (1978): 209; O’Toole (1978): 163; Skidmore (1978): Cors Graianog, Gwynedd, 1 male, 28.vii.1988, 76; Freeman (1983b): 39; Skidmore (1991): 149; Lau- Holmes, Boyce & Reed leg. (PPWM). Crymlyn Bog, rence (1994): 118; Laurence (1996): 86; Laurence Swansea/Abertawe, 2 males, 13.vii.1989, Holmes, (1997a): 85; Laurence (1997b): 228 [all as Scatopsciara Boyce & Reed leg. (PPWM). Cwm Crafnant, vivida]; Menzel (1998): 22; Chandler (2001): 201; Gwynedd, 1 male, 6.viii.1988, Holmes, Boyce & Reed Chandler et al. (2002): 124. leg. (PPWM). Gatewen Marsh, Denbighshire, 2 males, 3.viii.1988, Holmes, Boyce & Reed leg. (DEI, PPWM). Material examined: Ach na Bourin, Highland Region. Killaun Bog, Offaly, ix.2000, Withers leg. (PPWM). 1 male, 1.vii.1972, Chandler leg. (PPCM). Acton, Llyn Pencraig, Gwynedd, 1 male, 1 female, Greater London, 3 males, no date [19th century] 4.viii.1988, Holmes, Boyce & Reed leg. (DEI). Llyn (reared), Waterhouse leg. (BMNH). Alconbury, Ystumllyn, Gwynedd, 2 males, 16.vi.1988; 1 male, Cambridgeshire, 1 male, 29.iv.1976; 2 males, 9.v.1978, 28.vi.1988; 1 male, 26.vii.1988; 1 male, 5.x.1988; all all J.H. Cole leg. (PJCB). Annahala bog, Gearagh, Holmes, Boyce & Reed leg. (PPWM). Middle Harling, Cork, 4 males, 1.vi.1990, de Courcy Williams leg. Norfolk, 1 male, 1.viii.−15.viii.1988, Foster & Procter (DEI, NMID). Aviemore, River Spey, Highland Region, leg. (PBLN). Mills Marsh, Norfolk, 20 males, 27.vi.− 1 male, 9.vi.1984, Chandler leg. (PPWM). Babraham, 10.vii.1988; 16 males, 27.vi.−11.vii.1988; 10 males, 2 Cambridgeshire, 1 male, 25.i.1918, Edwards leg. females, 11.viii.1988, all Foster & Procter leg. (BMNH). Badger Wood, Cambridgeshire, 1 male, (PBLN). Moorthwaite, Cumbria, 3 males, 21.viii.1988, x.1998, C.W. Plant leg. (PPCM). Beaconsfield, Buck- no collector detail (PPWM). Nûg, Conwy, 1 male, inghamshire, 11 males, 18 females, 14.iii.−17.vii.1934 3.viii.1988; 1 male, 27.x.1988, all Holmes, Boyce & (reared), Basden leg. (NMS). Becky Woods, Manaton, Reed leg. (PPWM). Oxwich, Swansea/Abertawe, 2 Devon, 3 females, 23.vi.2002, J.E. Smith leg. (HRI). males, 12.vii.1989, Holmes, Boyce & Reed leg. Berwick-upon-Tweed, Northumberland, 1 male, (PPWM). Pant-y-Sais, Neath/Port Talbot, 1 male, iv.1999; 2 males, viii.1999; 1 male, 4.xi.1999, all Lau- 13.vii.1989, Holmes, Boyce & Reed leg. (PPWM). rence leg. (PBLN). Black Park, Buckinghamshire, 1 Rhôsgoch, Powys, 1 male, 10.x.1989, Holmes, Boyce & male, 3 females, 19.vi.−23.vi.1934 (reared), Basden Reed leg. (PPWM). St. David’s Airfield, Pem- leg. (NMS). Bockmer End, Buckinghamshire, 1 male, brokeshire, 1 male, 24.ix.1987, Holmes, Boyce & Reed 4.iii.1932 (reared), Basden leg. (NMS). Boveney, Buck- leg. (PPWM). Sea Mere, Norfolk, 1 male, 6.vii.− inghamshire, 2 males, 16.iv.1981, Chandler leg. 2.ix.1988, Foster & Procter leg. (PBLN). Thompson (PPCM). Bradfield Wood, Suffolk, 1 male, 6.vii.− Common, Norfolk, 1 male, 7.vi.1986, Withers leg. 11.vii.1990; 4 males, 16.viii.−22.viii.1991; 3 males, (PPWM); 4 males, 13.ix.−27.ix.1988, Foster & Procter viii.1991, all Laurence leg. (PBLN). Braedownie, leg. (PBLN). Trawsfynydd, Gwynedd, 1 male, Angus, 1 male, 24.vii.1937, Coe leg. (BMNH). Bramp- 28.x.1988, Holmes, Boyce & Reed leg. (PPWM). Wal- ton, Cambridgeshire, 1 male, 19.ii.1966, J.H. Cole leg. berswick, Suffolk, 2 males, 13.ix.−27.ix.1988, Foster & (PJCB). Brendon, Devon, 1 male, viii.1935, Edwards Procter leg. (PBLN). leg. (BMNH). Buckingham Palace Garden, Greater London, 1 male, 14.i.−14.ii.1997; 1 male, 28.vi.− Ecological notes: found in broad-leaved and carr wood- 7.vii.1997, all D.J. Carter & C.W. Plant leg. (PPCM). land; damp and wet heathland; wetlands especially Bucklebury Common, Berkshire, 1 male, 28.iv.1997 fens, also basin mires, saltmarsh and reed beds, bogs, (reared), Storey leg. (PPCM). Burnham, Buckingham- and damp meadows; grassland with pingo pools. vi–x. shire, 3 males, 7.iv.−22.iv.1932; 2 males, 9 females, 21.iv.−11.vi.1934 (all reared), all Basden leg. (NMS). SCATOPSCIARA EDWARDS, 1927 Burnham Beeches, Buckinghamshire, 1 male, 3 − SCATOPSCIARA (SCATOPSCIARA) ATOMARIA females, 3.v. 21.vi.1931; 1 male, 8.vi.1931, all Basden − (ZETTERSTEDT, 1851) leg. (BMNH, NMS); 1 male, 28.v. 19.vi.1996; 1 male, 5.viii.−31.viii.1996; 2 males, 9.x.−26.xi.1996, all Ismay Synonyms: falsaria (Winnertz, 1867); hybrida (Win- leg. (PPCM). Canonteign Falls, Devon, 2 females, nertz, 1867); mundula (Winnertz, 1867); pagana (Win- 22.vi.2002, J.E. Smith leg. (HRI). Catfield, Norfolk, 1 nertz, 1867); pratinicola (Winnertz, 1867); silvestris male, 12.x.1988; 1 male, 13.x.−13.xii.1988, all Foster (Frey, 1936); soluta (Winnertz, 1867); vivida (Win- & Procter leg. (PBLN). Chigwell Row Wood, Chigwell nertz, 1867). Row, Essex, 1 female, 20.vi.2002; 1 male, 3 females, Literature: Edwards (1925a): 541 [as Sciara tilicola, 28.vi.2002; 1 female, 15.ix.2002; 1 female, 23.ix.2002; misidentification]; Basden (1933): 44; Edwards 1 female, 26.ix.2002; 1 female, 8.x.2002; 1 male, 5 (1934b): 34 [all as Sciara vivida]; Edwards (1938): 201 females, 16.x.2002; 2 females, 22.x.2002, all Ismay &

Schulten leg. (DEI, UMO). Clondalkin, Dublin, 1 stone, Leicestershire, 1 female, 24.viii.2003, Menzel male, 14.iii.−16.iii.1982, J.P. & M.A. O’Connor leg. leg. (DEI). Kinrive Wood, Highland Region, 1 male, (NMID). Coombe Abbey Country Park, Coventry, 15.vi.1976, Chandler leg. (PPCM). Knebworth, Hert- Warwickshire, 1 female, 28.viii.2003, J.E. Smith leg. fordshire, 1 male, iv.1922; 1 male, 13.v.1923; 1 male, (HRI). Corker Pass, Clare, 1 male, 23.v.1985, J.P. viii.1936, all Edwards leg. (BMNH). Knole Park, Kent, O’Connor leg. (NMID). Corrimony, Highland Region, 1 1 male, 4.vi.1977, Chandler leg. (PPCM). Lackham male, 28.v.2002, Chandler leg. (PPCM). Cors Caron, Park, Wiltshire, 1 male, 5.viii.2002, Chandler leg. Ceredigion, 1 male, 5.x.1987, Holmes, Boyce & Reed (PPCM). Leckford Estate, Hampshire, 1 male, leg. (PPWM). Crickhowell, Powys, 1 male, 1.x.1936, 13.iii.1974, Stubbs leg. (PPCM). Letchworth, Hert- Edwards leg. (BMNH). Culbin Sands, Highland fordshire, 1 male, v.1917; 1 male, x.1917, all Edwards Region, 1 male, 5.vii.−7.vii.1936, Coe leg. (BMNH); 1 leg. (BMNH). Little Kildrummie, Highland Region, 7 male, 27.v.2002, Chandler leg. (PPCM). Ditchley, males, 14.vi.1984, Chandler leg. (PPCM). Loch Fleet, Oxfordshire, 1 male, v.1937, Edwards leg. (BMNH). Highland Region, 1 male, 26.v.2002, Chandler leg. Dropmore, Buckinghamshire, 1 male, 8.iv.1937 (PPCM). Loch Park, Moray, 4 males, 12.viii.− (reared), Basden leg. (BMNH). Eday, Orkney, 1 male, 14.viii.1937, Coe & Edwards leg. (BMNH). Loch Ran- 1.vi.−6.vi.1988, Laurence leg. (PBLN). Epping Forest, noch, Perth & Kinross, 1 male, 27.viii.1995, Chandler Essex, 3 males, 1.iii.−22.iii.1999, Dagley leg. (PPCM). leg. (PPCM). Lockhaugh Bank Wood, Derwent Valley, Fair Isle, Shetland, 1 male, 20.xi.2000, Laurence leg. Tyne & Wear, 1 male, 24.vii.−13.viii.1990, Jessop leg. (PBLN). Fairfield Experimental Horticulture Station, (PBLN). Lopham Fen, Norfolk, 2 males, 11.x.1990, Kirkham, Lancashire, 1 male, no date, Hussey leg. Withers leg. (PPWM). Lye Hill, Oxfordshire, 1 male, (HRI). Falls of Tarnash, Moray, 1 male, 1.vii.1936, Coe 20.ii.1915; 2 males, 5.ii.1916, all Hamm leg. (BMNH). leg. (BMNH). Farnham Royal, Buckinghamshire, 1 Mayford, Surrey, 1 male, 21 females, 21.iv.−28.iv.1934 male, 3 females, 31.ii.−11.v.1931; 1 male, 9.v.1931; (reared), Basden leg. (NMS). Medmenham, Bucking- 1 male, 1 female, 21.v.−24.v.1931; 1 male, 4.ix.1933 hamshire, 1 male, 7.vi.1931; 1 male, 1 female, (all reared), all Basden leg. (BMNH, NMS). Fisher’s 19.x.1931; 4 males, 3 females, 19.x.−1.xi.1931; 1 male, Wood, Bromley, Greater London, 1 male, 2.viii.1966, 25.x.1931; 1 female, 25.ii.1932; 1 male, 5.vi.1934; 27 Chandler leg. (PPCM). Flitwick Moor, Bedfordshire, males, 26 females, 16.i.−31.iii.1931–34 (all reared), all 1 male, 13.vii.1978, J.H. Cole leg. (PJCB). Folly Farm, Basden leg. (BMNH, NMS). Mills Marsh, Norfolk, 1 Avon, 1 male, 28.ix.2001, Gibbs leg. (PDGB). Forres, male, 11.viii.1988, Foster & Procter leg. (PBLN). Moray, 1 male, 4.vii.1936, Coe leg. (BMNH). Gamlin- Monkfield Wood, Cambridgeshire, 1 male, x.1998, gay Wood, Cambridgeshire, 1 male, 22.v.1992, J.H. C.W. Plant leg. (HRI). Murrough, Wicklow, 1 male, Cole leg. (PJCB). Gearagh, Cork, 1 male, 20.iv.1990; v.1989, Blackith leg. (PPWM). Nethy Bridge, High- 1 male, 19.v.1990; 1 male, 9.vi.1990; 1 male, land Region, 3 males, 16.vii.1936, Coe leg. (BMNH). 12.vi.1990, all de Courcy Williams leg. (DEI, NMID). New Malden, Greater London, 1 male, 15.vi.1932, Coe Glasshouse Crops Research Station, Littlehampton, leg. (BMNH). Newbridge Fen, Kildare, 1 male, 8.viii.− West Sussex, 1 male, 13.viii.1956; 2 males, 10.ix.1956; 19.viii.1998, Good leg. (PPWM). Nettlebed, Oxford- 1 male, 17.ix.1956; 2 males, 13.i.1958; 1 male, shire, 1 male, 9.v.1998, Chandler leg. (PPCM). Nûg, 25.iii.1958; 1 male, 29.ii.1960; 1 male, 7.iii.1960, Conwy, 1 male, 3.viii.1988, Holmes, Boyce & Reed leg. all Hussey leg. (HRI). Grassholm, Pembrokeshire, 1 (PPWM). Offley Holes, Hertfordshire, 1 male, iv.1936, male, 25.viii.1934, Edwards leg. (BMNH). Gutcher Edwards leg. (BMNH). Oxford University Parks, Yell, Shetland, 2 males, 13.x.−16.x.1992, Laurence leg. Oxford, Oxfordshire, 1 male, 1 female, 8.vi.1915, (PBLN). Gwaun Ystrad Caron, Ceredigion, 1 male, Hamm leg. (BMNH). Radwell, Hertfordshire, 2 males, 24.vii.1987, Holmes, Boyce & Reed leg. (PPWM). 21.x.1917, all Edwards leg. [in part labelled as ‘sp. no. Harpenden, Hertfordshire, 2 males, 7.x.1934, C.B. 5′] (BMNH, ZFMK). RAF Barnham, Suffolk, 1 male, Williams leg. (BMNH). Hartslock, Oxfordshire, 1 22.vi.1996, Chandler leg. (PPCM). Reedham, Norfolk, male, 6.v.2000, Raper leg. (PDGB). Haytor, Devon, 1 1 male, 7.xii.1988–17.iii.1989, Foster & Procter female, 23.vi.2002, J.E. Smith leg. (HRI). Hickling, leg. (PBLN). Rhôs-y-gad, Isle of Anglesey, 1 male, Norfolk, 1 male, v.1988; 1 male, vi.1988; 28 males, 5.x.1988, Holmes, Boyce & Reed leg. (PPWM). Ryton x.1988, all Foster & Procter leg. (PBLN). Highgate Wood, Bubbenhall, Coventry, West Midlands, 1 Wood, Greater London, 1 male, 1998–99, Hackett leg. female, 12.x.2001, J.E. Smith leg. (HRI). Salisbury (PPCM). Hogley Bog, Oxfordshire, 2 males, 3.iii.1923 Plain Training Area, Warminster, Wiltshire, 1 male, (reared), Hamm leg. (BMNH). How Hill, Norfolk, 1 4.vii.2003, Ismay & Schulten leg. (UMO). Scadbury male, 1.x.1989, Withers leg. (PPWM). Insh Marshes, Park, Greater London, 2 males, 1 female, 3.xi.1984, Highland Region, 1 male, 30.v.2002, Chandler leg. Chandler leg. (PPCM). Sedlescombe, East Sussex, 1 (PPCM). Janet’s Foss Wood, North Yorkshire, 2 males, male, xii.1994; 1 male, 24.iii.1996, all Roper leg. vii.1980, Disney leg. (PPWM). Kelham Bridge, Raven- (PPRS). Selsdon, Greater London, 1 male, 26.i.1937,

Coe leg. (BMNH). Shefford, Bedfordshire, 1 male, SCATOPSCIARA (SCATOPSCIARA) BUCERA 17.xi.1917, Edwards leg. (BMNH). Sidmouth, Devon, RUDZINSKI, 1994 1 male, 10.v.1936, Edwards leg. (BMNH). Spey Bay, New to British list. Moray, 12 males, 2.vii.1936, Coe leg. (BMNH). Sutton Park, Birmingham, Birmingham, West Midlands, 1 Material examined: Burnham Beeches, Buckingham- female, 7.iv.2003, J.E. Smith leg. (HRI). Temple, Berk- shire, 1 male, 7.vi.−21.vi.1995, Ismay leg. (PPCM). shire, 3 males, 6 females, 12.ii.−28.iii.1932; 1 male, Epping Forest, Essex, 1 male, vii.1999, Dagley leg. 24.iii.1933; 1 male, 4 females, 24.ii.−19.v.1934; 1 male, (PPCM). West End Common, Esher, Surrey, 1 male, 2 females, 31.iii.−7.iv.1934 (all reared), all Basden leg. 16.vii.2002, Chandler leg. (PPCM). Whitacre Heath, (NMS). The Loke, Norwich, Norfolk, 2 males, Warwickshire, 1 male, 13.x.2001, J.E. Smith leg. 20.iv.1987; 4 males, 26.iv.1987; 2 males, 8.v.1987; 1 (HRI). − − male, 24.x. 25.x.1987; 1 male, 28.iv. 4.v.1990; 2 Ecological notes: found in ancient beech and other − males, 2.v. 6.v.1992; 1 male, 21.viii.1992; 1 male, broad-leaved woodland; willow carr with birch and − − 29.ix. 3.x.1992; 2 males, 4.xi. 6.xi.1992; 2 males, reed beds. vi–vii, x. 20.v.−23.v.1993, all Laurence leg. (PBLN). The Oaks, Cambridgeshire, 1 male, x.1998, C.W. Plant leg. (PPCM). Thornham Magna, Suffolk, 1 male, SCATOPSCIARA (SCATOPSCIARA) CALAMOPHILA 14.iii.1987, Withers leg. (PPWM). Tomintoul, Moray, FREY, 1948 1 male, 8.viii.−11.viii.1937, Coe leg. (BMNH). Literature: Laurence (1994): 118 [in part as Scatops- Trawsfynydd, Gwynedd, 1 male, 5.viii.1988; 1 ciara vivida]; Laurence (1996): 87 [as Scatopsciara male, 14.x.1988, all Holmes, Boyce & Reed leg. tenax, misidentiﬁcation]. (PPWM). Udale Bay, Highland Region, 1 male, 29.v.2002, Chandler leg. (PPCM). Vidlin, Shetland, New to British and Irish list. 1 male, 18.vi.−21.vi.1993, Laurence leg. (PBLN). Material examined: Chigwell Row Wood, Chigwell Wellesbourne, Warwickshire, 12 males, 2 females, Row, Essex, 1 male, 1 female, 23.v.−27.v.2002; 2 17.viii.−31.viii.2000; 1 male, viii.2000; 1 female, 19.i.− females, 18.vi.2002; 2 females, 8.x.2002; 1 female, 15.ii.2001; 1 female, 28.iii.−26.iv.2001; 1 male, 16.x.2002; 1 female, 22.x.2002, all Ismay & 27.vi.−5.vii.2001; 1 male, 13.ix.−4.x.2001, all J.E. Schulten leg. (DEI, UMO). Epping Forest, Essex, 1 Smith leg. (HRI). Welwyn, Hertfordshire, 1 male, male, 11.ix.1998; 3 males, viii.1999, all Dagley leg. 20.ii.1928; 1 male, 21.vii.−24.vii.1934, all Edwards (PPCM). Gearagh, Cork, 1 male, 7.iv.1990, de leg. (BMNH). Wennington Wood, Cambridgeshire, 1 Courcy Williams leg. (NMID). North Ronaldsay, male, 4.x.1975, J.H. Cole leg. (PJCB). West Harling Orkney, 3 males, 10.ix.1989, Laurence leg. (PBLN). Common, Norfolk, 1 male, 1.iv.1988; 2 males, Ryton Wood, Bubbenhall, Coventry, West Midlands, 30.iii.1991; 1 male, 23.iv.1991, all Withers leg. 1 male, 25.viii.2002, Menzel leg. (DEI). Sedle- (PPWM). Weston, Hertfordshire, 2 males, 4.x.1931, scombe, East Sussex, 1 male, 2.vi.1999, Roper leg. Edwards leg. (BMNH). Whitnash brook, Sydenham, (PPRS). Stallode Wash, Suffolk, 10 males, 24.vi.− Leamington Spa, Warwickshire, 1 male, 2 females, 8.vii.1988, Foster & Procter leg. (PBLN). Stronsay, 4.ix.2003, J.E. Smith leg. (HRI). Wicken Fen, Cam- Orkney, 1 male, 21.v.1988, Laurence leg. (PBLN). bridgeshire, 1 male, 26.iv.1916, Edwards leg. (BMNH). Temple, Berkshire, 1 male, 1.v.1934 (reared), Bas- Windbury, Devon, 1 male, 7.vii.1986, Chandler leg. den leg. (BMNH). The Loke, Norwich, Norfolk, 1 (PPCM). Windsor Forest, Berkshire, 4 males, 2 male, 8.v.1987; 1 male, 18.vi.1987; 1 male, females, 3.ix.1977, Chandler leg. (PPCM). Wolford 15.vii.1987; 1 male, 10.viii.1987; 1 male, 20.ii.1988; Wood, Moreton-in-Marsh, Warwickshire, 3 females, 1 male, 19.vii.1988; 1 male, 15.v.−18.v.1992; 5 11.x.2001, J.E. Smith leg. (HRI). Yell, Shetland, 4 males, 18.v.−23.v.1992; 1 male, v.1992; 1 male, males, 3.ix.−10.ix.1990, Laurence leg. (PBLN). 26.vi.−30.vi.1992; 1 male, 21.viii.1992; 1 male, 24.ix.−3.x.1992; 1 male, 24.x.−25.x.1987; 2 males, Ecological notes: very common in all types of wood- 27.vi.−30.vi.1993, all Laurence leg. (PBLN). Wind- land; heathland; moorland on granite; wetlands sor Forest, Berkshire, 1 male, iv.1933 (reared), including fens, mires, bogs, Cladium beds with tufa, Donisthorpe leg. (BMNH). sedge beds, water meadows and riverside marshes; unimproved calcareous grassland; grounds of horticul- Ecological notes: found in ancient oak and beech forest tural research station; on ivy; parkland; gardens; and other broad-leaved woodland; ancient woodland rocky island with seabird colonies. Reared from nests with oak, hazel and bramble; heathland; marshland; of coot, common shrew, ﬁeld mouse, hedgehog, mole, garden; grassland at woodland fringe; near shore. moorhen, rabbit, thrush and wasp; the fungus Laeti- Reared from spruce bark and rotten horse chestnut porus sulphureus and lime galls. i–xii. wood. ii, iv–x.

SCATOPSCIARA (SCATOPSCIARA) EDWARDSI coei]; Menzel (1998): 22; Menzel & Mohrig (2000): FREEMAN, 1983 490. Literature: Freeman (1983a): 167; Freeman (1983b): Material examined: Aylesbeare Common, Devon, 1 40; Menzel (1998): 22. male, 25.x.2000, C.W. Plant leg. (PPCM). Balmoral − Material examined: Baldock, Hertfordshire, 1 male, Forest, Aberdeenshire, 1 male, 1.viii. 4.viii.1937, Coe viii.1918, Edwards leg. (BMNH). Balmoral Forest, leg. (BMNH). Ben Lawers, Perth & Kinross, 1 male, Aberdeenshire, 1 male, 1.viii.−4.viii.1937, Coe leg. 1.vi.1932, Edwards leg. [paratype of Scatopsciara coei] (BMNH). Ditchley, Oxfordshire, 1 male, v.1937, (BMNH). Boat of Garten, Highland Region, 1 male, Edwards leg. (BMNH). How Hill, Norfolk, 1 male, v.1934, Edwards leg. [paratype of Scatopsciara coei] 8.iv.1989, Withers leg. (PPWM). Hoxne, River (BMNH). Brampton Wood, Cambridgeshire, 2 males, Waveney, Suffolk, 1 male, 1.iv.1990, Withers leg. 27.v.1987, J.H. Cole leg. (PJCB). Epping Forest, Essex, (PPWM). Market Weston Fen, Suffolk, 1 male, 1 male, 25.x.2001, Chandler leg. (PPCM). Glendal- 12.v.1991, Withers leg. (PPWM). Searles Farm, ough, Wicklow, 1 male, 24.iv.1989, J.P. & M.A. O’Con- − Reading, Berkshire, 1 male, 26.vi.2003, Chandler nor leg. (NMID). Glyn Ceiriog, Clwyd, 2 males, 26.vi. leg. (PPCM). Shepreth, Cambridgeshire, 5 males, 6 7.vii.1995, Brock leg. (HRI). Goyt Valley, Derbyshire, 2 − females, 23.ii.1918 [paratypes of Scatopsciara males, 18.vii. 20.vii.1939, Edwards leg. (BMNH). edwardsi]; 2 males, ii.1918 [holotype and paratype of Holme Fen, Cambridgeshire, 2 males, 27.v.1987, J.H. Scatopsciara edwardsi] (all reared), all Edwards leg. Cole leg. (PJCB). King’s Somborne, Hampshire, 1 (BMNH). West Harling Common, Norfolk, 1 male, male, viii.1936, Edwards leg. [paratype of Scatopsci- 30.iii.1991, Withers leg. (PPWM). ara coei] (BMNH). Loch Pityoulish, Highland Region, 1 male, 3.vii.1933, Coe leg. [holotype of Scatopsciara Ecological notes: found in damp broad-leaved wood- coei] (BMNH). Lochan an Daimh, Perth & Kinross, 1 land; conifer plantations; peat hollows and fen; fen male, 29.v.1998, Chandler leg. (PPCM). New Forest, with sedge pools; river margin; water meadow. Reared Hampshire, 1 male, 18.x.1937, Edwards leg. [paratype from unnamed fungus and Auricularia mesenterica. of Scatopsciara coei] (BMNH). Norwich, Norfolk, 1 ii–vi, viii, xi. male, 3.vii.1983, Irwin leg. (PPWM). Sutton Park, Birmingham, West Midlands, 1 male, 7.iv.2003, J.E. Smith leg. (HRI). SCATOPSCIARA (SCATOPSCIARA) FLUVIATILIFORMIS MOHRIG & MAMAEV, 1987 Ecological notes: found in broad-leaved and mixed woodland; conifer plantations; stream with alder; Synonym: germana Rudzinski, 1994. heath and parkland; wetlands; open marsh by pool; Literature: Laurence (1997a): 85 [as Scatopsiara mul- scrub and Sphagnum bog; open montane habitats; tispina, misidentification]. garden. iv–viii, x. New to British list. SCATOPSCIARA (SCATOPSCIARA) LATIPTERA Material examined: Bradfield Wood, Suffolk, 4 males, RUDZINSKI, 1995 6.vii.−11.vii.1990, Laurence leg. (PBLN). Burnham Beeches, Buckinghamshire, 1 male, 18.v.1990, Chan- Literature: Laurence (1996): 87 [as Scatopsciara dler leg. (PPCM). Epping Forest, Essex, 2 males, tenax, misidentification]. v.1999, Dagley leg. (PPCM). Morrone Birkwood, Aber- New to British list. deenshire, 1 male, 31.v.1998, Chandler leg. (NMS). Theale, Berkshire, 1 male, 6.viii.2003, Chandler leg. Material examined: The Loke, Norwich, Norfolk, 1 (PPCM). male, 5.vii.1987, Laurence leg. (PBLN). Ecological notes: found in ancient beech and other Ecological notes: found in garden. vii. broad-leaved woodland; birch woodland with calcareous seepages; carr woodland. v, viii. SCATOPSCIARA (SCATOPSCIARA) MULTISPINA (BUKOWSKI & LENGERSDORF, 1936)

SCATOPSCIARA (SCATOPSCIARA) FLUVIATILIS Synonym: barbula Mohrig & Mamaev, 1985. (LENGERSDORF, 1940) Literature: Freeman (1983b): 39; Menzel (1998): 22; Synonyms: coei Freeman, 1983; pulchra (Lengersdorf, Chandler (2001): 201; Chandler et al. (2002): 124. 1940); robusticornis (Frey, 1948). Material examined: Annahala bog, Gearagh, Cork, 1 Literature: Freeman (1983a): 167; Freeman (1983b): male, 10.viii.1990, de Courcy Williams leg. (DEI). Bal- 40; O’Connor & Ashe (1991): 29 [all as Scatopsciara dock, Hertfordshire, 1 male, v.1918, Edwards leg.

(BMNH). Bayfordbury Estate, Hertford, Hertford- leg. (PBLN). Rothamsted, Hertfordshire, 24 males, shire, 1 male, 17.vi.1975, J.H. Cole leg. (PJCB). 1950, Laurence leg. (PBLN). Rotherfield Park, Hamp- Berwick-upon-Tweed, Northumberland, 6 males, shire, 1 male, 3.vii.2002, Chandler leg. (PPCM). Sedle- iv.1999; 1 male, vii.1999; 1 male, 1.x.1999; 2 males, scombe, East Sussex, 1 male, 5.iv.1997, Roper leg. 4.xi.1999, all Laurence leg. (PBLN). Blaen y Werg- (PPRS). Stepaside, Wicklow, 1 male, 8.v.1989, Withers loedd, Conwy, 1 male, 4.viii.1988, Holmes, Boyce & leg. (PPWM). The Loke, Norwich, Norfolk, 2 males, Reed leg. (PPWM). Boundless Copse, Surrey, 1 male, 20.iv.1987; 1 male, 23.iv.1987; 5 males, 26.iv.1987; 26.vii.2002, Chandler leg. (PPCM). Braedownie, 1 male, 27.iv.1987; 1 male, 9.v.1987; 14 males, Angus, 1 male, 24.vii.1937, Coe leg. (BMNH). Buck- 5.vii.1987; 2 males, 10.viii.1987; 2 males, 15.ix.1987; 1 ingham Palace Garden, Greater London, 1 male, male, 24.x.−25.x.1987; 6 males, 21.iv.1988; 1 male, 26.vi.1998, Chandler leg. (PPCM). California Country 15.viii.−16.viii.1988; 1 male, 13.vii.−19.vii.1989; 2 Park, Berkshire, 3 males, 6.viii.1999, Chandler leg. males, 20.v.−23.v.1993, all Laurence leg. (PBLN). Wall- (NMS, PPCM). Cambridge, Cambridgeshire, 2 males, ington, Hertfordshire, 1 male, 13.v.1934, Edwards 3.xi.1903, Jenkinson leg. (BMNH). Cleddon Bog, Mon- leg. (BMNH). Wellesbourne, Warwickshire, 1 male, 1 mouthshire, 1 male, 10.vi.1988, Holmes, Boyce & Reed female, 28.iii.−26.iv.2001; 1 male, 27.iv.−5.vii.2001, all leg. (PPWM). Craufurdland Water, East Ayrshire, 1 J.E. Smith leg. (HRI). Welwyn, Hertfordshire, 1 male, male, 3.vii.1995, Chandler leg. (PPCM). Cuckoo Wood, 21.vii.−24.vii.1934, Edwards leg. (BMNH). Willes Downe, Greater London, 1 male, 20.vii.1985, Chandler Meadow, Leamington Spa, Warwickshire, 1 male, leg. (NMS). Dinton Pastures Country Park, Berkshire, 27.viii.2002, Menzel leg. (DEI). No locality details, ? 1 male, 7.xi.1974, Chandler leg. (PPCM). Epping For- England, 1 male, no date, Laurence leg. (PBLN). est, Essex, 1 male, 21.vi.1998; 1 male, x.1999, all Dag- Ecological notes: common in broad-leaved, mixed and ley leg. (PPCM). Fritham, New Forest, Hampshire, 1 carr woodland; gravel pits; meadow hedge; common in male, 17.vi.1953, Colyer leg. (BMNH). Gearagh, Cork, damp meadows; sedge bed; Molinia bog; parkland 1 male, 7.iv.1990, de Courcy Williams leg. (NMID). with old trees; grounds of horticultural research sta- Glasshouse Crops Research Station, Littlehampton, tion; in sparse marginal vegetation around pond; gar- West Sussex, 2 males, 10.viii.1956; 3 males, dens; on ivy. iii–xi. 13.viii.1956; 1 male, 20.viii.1956; 1 male, 22.viii.1956; 1 male, 2.xi.1956, all Hussey leg. (HRI). Grafham Water, Savages Creek, Cambridgeshire, 1 male, SCATOPSCIARA (SCATOPSCIARA) NACTA 19.vi.1988, J.H. Cole leg. (PJCB). Harpford Common, (JOHANNSEN, 1912) Devon, 1 male, 30.vi.−26.vii.2000, C.W. Plant leg. Literature: Freeman (1983b): 39 [as Scatopsciara (PPCM). Hatchett’s Pond, New Forest, Hampshire, 1 tenax, misidentification]; Menzel (1998): 22. male, 12.vii.1990, J.H. Cole leg. (PJCB). Hatherton, Staffordshire, 2 males, 22.viii.1983, Wyatt leg. Material examined: Barkham, Berkshire, 2 males, 1 (BMNH). Leckford Estate, Hampshire, 2 males, female, ix.1922 (reared), Donisthorpe leg. (BMNH). 4.vi.1972; 1 male, 1.ix.1999, all Chandler leg. (NMS, Letchworth, Hertfordshire, 1 male, vi.-vii.1936, PPCM). Lockhart Mill, South Lanarkshire, 1 male, Edwards leg. (BMNH). New Forest, Hampshire, 3 4.viii.1995, Chandler leg. (PPCM). Lockhaugh Bank males xi.1915 (reared), Sharp leg. (BMNH). Radwell, Wood, Derwent Valley, Tyne & Wear, 1 male, 26.vii.− Hertfordshire, 1 male, 21.x.1917, Edwards leg. 13.viii.1990, Jessop leg. (PBLN). Monkfield Wood, (BMNH). Cambridgeshire, 1 male, x.1998, C.W. Plant leg. (HRI). Ecological notes: found in broad-leaved woodland; on Newbold Comyn Park, Leamington Spa, Warwick- ivy. Reared from fungus and under pine bark. vi–vii, shire, 2 males, 15.vii.2001, J.E. Smith leg. (HRI). New- ix–xi. bridge Fen, Kildare, 1 male, 8.viii.−19.viii.1998, Withers leg. (PPWM). Oakley Inclosure, New Forest, Hampshire, 1 male, 17.vi.1953, Colyer leg. (BMNH). SCATOPSCIARA (SCATOPSCIARA) NANA Oxford University Museum, Oxford, Oxfordshire, 1 (WINNERTZ, 1871) male, 27.x.1922, Hamm leg. (BMNH). Pontarffinant Literature: Freeman (1983b): 40; Laurence (1996): 87; Wood, Carmarthen, 1 male, 12.x.1995, McLean leg. Laurence (1997b): 228; Laurence & James (1997): 166 (PPCM). Potter’s Bar, Hertfordshire, 1 male, 4.v.1963, [as Scatopsciara nana and Scatopsciara vitripennis, Vane-Wright et al. leg. (BMNH). Powerscourt, Wick- misidentification]; Menzel (1998): 22; Withers (2002): low, 1 male, 10.vii.1971, Chandler leg. (PPCM). Rad- 115. well, Hertfordshire, 2 males, 1 female, 21.x.1917, Edwards leg. (BMNH). Rake Beck, North Yorkshire, 1 Material examined: Alconbury, Cambridgeshire, 1 male, 9.vi.1976, J.H. Cole leg. (PJCB). Reedham, male, 9.v.1978, J.H. Cole leg. (PJCB). Blaen y Werg- Norfolk, 2 males, 26.viii.−27.x.1988, Foster & Procter loedd, Conwy, 2 males, 13.x.1988, Holmes, Boyce &

Reed leg. (PPWM). Bradfield Wood, Suffolk, 1 male, meadows; unimproved calcareous grassland; open 11.ix.−16.ix.1990, Laurence leg. (PBLN). Cadair Idris, montane habitats; near shore; garden. Reared from Gwynedd, 3 males, 4.ix.1993, Laurence leg. (PBLN). coot nest. ii–x. Cefn Grwhyd, Neath/Port Talbot, 1 male, 12.vii.1989, Holmes, Boyce & Reed leg. (PPWM). Cors Caranod, Ceredigion, 1 male, 23.vii.1987, Holmes, Boyce & Reed SCATOPSCIARA (SCATOPSCIARA) NEGLECTA MENZEL & leg. (PPWM). Gatewen Marsh, Denbighshire, 1 male, MOHRIG, 1998 3.viii.1988, Holmes, Boyce & Reed leg. (PPWM). Literature: ? Walker (1848): 109; ? Walker (1856): 55 Gors Llwyn, Neath/Port Talbot, 1 male, 14.vii.1989, [all as Sciara pusilla; misidentification]; Freeman Holmes, Boyce & Reed leg. (PPWM). Gwaun Ystrad (1983b): 40 [as Scatopsciara pusilla, misidentifica- Caron, Ceredigion, 2 males, 14.x.1986, Holmes, Boyce tion]; Laurence (1994): 118; Laurence (1997a): 85 [all & Reed leg. (PPWM). Hafod Garregog, Gwynedd, 1 in part as Scatopsciara pusilla]; Menzel (1998): 22. male, 13.x.1988, Holmes, Boyce & Reed leg. (PPWM). New to Irish list. Harpenden, Hertfordshire, 1 male, 13.v.1934; 1 male, 7.x.1934, all C.B. Williams leg. (BMNH). Haytor, Material examined: Alvecote Pools, Staffordshire, 1 Devon, 1 female, 23.vi.2002, J.E. Smith leg. (HRI). male, 13.x.2001, Chandler leg. (HRI). Badger Wood, Hengistbury Head, Hampshire, 1 male, 3.vi.1938, Cambridgeshire, 1 male, x.1998, C.W. Plant leg. Edwards leg. (BMNH). Hickling Broad, Norfolk, 2 (PPCM). Bix Bottom, Oxfordshire, 1 male, 23.ix.2000, males, viii.1932, Edwards leg. (BMNH). Kergord, Raper leg. (PDGB). Bradfield Wood, Suffolk, 1 male, Shetland, 1 male, 26.vii.−28.vii.1996, Laurence leg. 16.viii.−22.viii.1991, Laurence leg. (PBLN). Bromley, (PBLN). Killaun Bog, Offaly, 2 males, ix.2000, Withers Greater London, 1 male, 2.x.1966, Chandler leg. leg. (PPWM). Lackham Park, Wiltshire, 1 male, (PPCM). Burnham Beeches, Buckinghamshire, 2 2.vii.2002, Chandler leg. (PPCM). Leckford Estate, males, 15.ix.−4.x.1995; 1 male, 8.vii.−27.vii.1996, all Hampshire, 1 male, 18.iii.1971, Stubbs leg. (PPCM). Ismay leg. (PPCM); 1 male, 21.iv.1998, Chandler leg. Letchworth, Hertfordshire, 2 males, 2 females, ii.1918, (PPCM). California Country Park, Berkshire, 1 male, Edwards leg. (BMNH). Loch Callater, Aberdeenshire, 6.viii.1999; 1 male, 29.vi.2002, all Chandler leg. 1 male, 28.vii.−29.vii.1937, Coe leg. (BMNH). Magor (PPCM). Clovelly, Devon, 1 male, 8.v.1936, Edwards Marsh, Monmouthshire, 1 male, 21.vii.1988, Holmes, leg. (BMNH). Delamere, Cheshire, 2 males, 23.v.1924, Boyce & Reed leg. (PPWM). Malham Tarn, North Britten leg. (BMNH). Dinton Pastures Country Park, Yorkshire, 2 males, 11.ix.−14.ix.1981, Disney leg. Berkshire, 1 male, 3.x.1993, Chandler leg. (PPCM). (PPWM). Newbold Comyn Park, Leamington Spa, Epping Forest, Essex, 1 male, viii.1999; 1 male, Warwickshire, 1 male, 29.vii.2001, J.E. Smith leg. ix.1999, all Dagley leg. (PPCM). Gearagh, Cork, 1 (HRI). Newbridge Fen, Kildare, 2 males, 8.viii.− male, 7.vi.1990, de Courcy Williams leg. (NMID). Glen 19.viii.1998, Good leg. (PPWM). Newmarket, Cam- Affric, Highland Region, 8 males, v.1992 (reared), bridgeshire, 2 males, 25.vii.1893, Verrall leg. (BMNH). MacGowan leg. (NMS). Hartslock, Oxfordshire, 1 North Ronaldsay, Orkney, 1 male, 3.ix.1989; 1 male, male, 6.v.2000, Raper leg. (PDGB). Hickling, Norfolk, 9.x.1989, all Laurence leg. (PBLN). Orphir, Orkney, 2 males, 1988, Foster & Procter leg. (PBLN). Hobb’s 1 male, 26.viii.1988, Laurence leg. (PBLN). Oxford Quarry, Gloucestershire, 1 male, 4.ix.2002, Gibbs leg. University Museum, Oxford, Oxfordshire, 1 male, (PDGB). Lackham Park, Wiltshire, 1 male, 2.vii.2002, 20.iv.1921, Hamm leg. (BMNH). Redgrave Lake, Suf- Chandler leg. (PPCM). Lancaut Nature Reserve, folk, 1 male, 24.v.1987, Withers leg. (PPWM). Salis- Gloucestershire, 1 male, 5.ix.2002, Gibbs leg. (PDGB). bury Plain Training Area, Warminster, Wiltshire, 5 Lion Wood, Redditch, Worcestershire, 1 male, males, 29.v.2003; 2 males, 4.vii.2003, all Ismay & 10.iv.2003, J.E. Smith leg. (HRI). Lockhaugh Bank Schulten leg. (DEI, UMO). The Loke, Norwich, Nor- Wood, Derwent Valley, Tyne & Wear, 1 male, 26.vii.− folk, 2 males, 23.iv.1987; 3 males, 9.v.1987; 1 male, 13.viii.1990, Jessop leg. (PBLN). Mid Quoich, Aber- 5.vii.1987; 1 male, 1.iv.1988; 1 male, 5.vi.−13.vi.1989; deenshire, 1 male, 22.ix.2000, Godfrey leg. (PPCM). 1 male, 4.vi.−6.xi.1992, all Laurence leg. (PBLN). Moccas Park, Herefordshire, 1 male, 11.vi.2002, God- Trefeiddan Moor, Pembrokeshire, 1 male, 21.vii.1987, frey leg. (PPCM). Paramoor Wood, Cornwall, 1 male, Holmes, Boyce & Reed leg. (PPWM). Wellesbourne, 7.vii.1983, Chandler leg. (PPCM). Sheep Leas, West Warwickshire, 1 male, 24.v.−27.vi.2001, J.E. Smith leg. Horsley, Surrey, 1 male, 15.vi.1966 (reared), Chandler (HRI). leg. (PPCM). Sutton Park, Birmingham, West Mid- lands, 1 male, 7.iv.2003, J.E. Smith leg. (HRI). The Ecological notes: found in broad-leaved, mixed and Loke, Norwich, Norfolk, 1 male, 15.viii.-.16.viii.1988; 1 carr woodland; wet heathland; moorland on granite; male, 29.ix.−3.x.1992, all Laurence leg. (PBLN). War- wetlands including fens, reed and sedge beds, Cla- boys Wood, Cambridgeshire, 4 males, v.1931, Fryer dium beds with tufa, mires, bog, lakeshore and water leg. (BMNH). West End Common, Esher, Surrey, 2

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 BRITISH SCIARID FAUNA 125 males, 19.x.2001; 1 male, 17.vii.2002, all Chandler leg. Literature: Laurence (1994): 116 [as Scatopsciara sub- (PPCM). Weston, Hertfordshire, 1 male, ix.1931, vivida]; Menzel (1998): 22; Menzel & Mohrig (2000): Edwards leg. (BMNH). Whitacre Heath, Warwick- 500. shire, 1 male, 13.x.2001, Chandler leg. (HRI). Material examined: 60 males. Catfield, Norfolk, 1 Ecological notes: very common in all types of wood- male, 13.x.−13.xii.1988, Foster & Procter leg. land; heath and parkland; wooded river bank; old [paratype of Scatopsciara subvivida] (PBLN). Hick- hedges; sedge bed; water meadows; old sand and ling, Norfolk, 12 males, 8.viii.−22.viii.1988, Foster gravel workings; calcareous grassland; gardens. & Procter leg. (PBLN); 1 male, x.1988, Laurence Reared from the fungi Phaeolus schweinitzii on pine leg. [paratypes of Scatopsciara subvivida] (PBLN). and Trametes versicolor. iv–x. Reedham, Norfolk, 1 male, 27.vi.−11.vii.1988; 18 males, 27.x.1988, all Foster & Procter leg. [paratypes of Scatopsciara subvivida] (PBLN). Stallode Wash, SCATOPSCIARA (SCATOPSCIARA) PUSILLA Suffolk, 16 males, 10.vi.−24.vi.1988; 3 males, 10.vi.− (MEIGEN, 1818) 26.vi.1988; 8 males, 24.vi.−8.vii.1988, all Foster & Synonyms: paludicicola (Lengersdorf, 1940); pavida Procter leg. [holotype and paratypes of Scatopsciara (Winnertz, 1867); pusilliformis Mohrig & Mamaev, subvivida] (DEI, PBLN). 1986; zygoneuroides Frey, 1948. Ecological notes: found in carr woodland; fen; marsh- Literature: Laurence (1994): 118; Laurence (1997a): 85 land; sedge beds. vi–viii, x–xii. [all in part]; Menzel (1998): 22; Withers (2002): 115. Taxonomic note: In Laurence’s collection there are Material examined: Bradfield Wood, Suffolk, 1 male, 19 males that were labelled by the author as 16.viii.−22.viii.1988, Laurence leg. (PBLN). Bure ‘PARATYPE, Scatopsciara subvivida’ with the dates Marshes, Norfolk, 1 male, 15.x.1983, Foster & Procter 10.vi.−24.vi.1988 [2 males (DEI); 14 males (PBLN)] leg. (PPWM). Catfield, Norfolk, 1 male, 12.x.1988, Fos- and 10.vi.−26.vi.1988 [1 male (DEI); 2 males (PBLN)]. ter & Procter leg. (PBLN). Coombe Abbey Country In the original description, however, only 9 paratypes Park, Coventry, Warwickshire, 2 males, 28.viii.2003, from Stallode Wash, Suffolk [Laurence (1994): 117 [as Menzel leg. (DEI). Cors Graianog, Gwynedd, 1 male, ‘9 males, similar data’]] were recorded. It seems pos- 7.x.1988, Holmes, Boyce & Reed leg. (PPWM). Flatford sible that there was a printing error and that it should Mill, Suffolk, 1 male, 13.viii.1981, no collector detail have been 19, not 9! Therefore, the total type series for (PPWM). Gearagh, Cork, 1 male, 7.vi.1990, de Courcy Scatopsciara subvivida stands as the holotype and 59 Williams leg. (NMID). Hickling Broad, Norfolk, 1 paratypes. male, 27.v.1991, Withers leg. (PPWM). Malham Tarn, − North Yorkshire, 1 male, 11.ix. 18.ix.1981, Disney leg. SCATOPSCIARA (SCATOPSCIARA) SUBCILIATA (PPWM). Murrough, Wicklow, 1 male, 8.v.1989, With- TUOMIKOSKI, 1960 ers leg. (PPWM). Searles Farm, Reading, Berkshire, 1 Literature: Freeman (1983b): 37 [in part as Bradysia male, 6.viii.2003, Chandler leg. (PPCM). Sedlescombe, regularis, misidentification]. East Sussex, 1 male, 23.iii.1996, Roper leg. (PPRS). Strumpshaw, Norfolk, 1 male, 13.vi.−11.vii.1988, Fos- New to British list. ter & Procter leg. (PBLN). Vicarage Bog, Wrexham, 1 Material examined: Aviemore, Highland Region, 4 male, 12.x.1988, Holmes, Boyce & Reed leg. (PPWM). males, 1 female, v.1934, Edwards leg. (BMNH). Wheatfen Broad, Norfolk, 1 male, 19.v.1990, Withers leg. (PPWM). Whitnash brook, Sydenham, Leaming- Ecological notes: probably from the Spey valley but ton Spa, Warwickshire, 1 male, 4.ix.2003, J.E. Smith precise habitat not recorded. v. leg. (HRI). Ynys Eidiol, Ceredigion, 1 male, 9.x.1987, Holmes, Boyce & Reed leg. (PPWM). SCATOPSCIARA (SCATOPSCIARA) TRICUSPIDATA Ecological notes: found in broad-leaved woodland (WINNERTZ, 1867) especially carr; grazed wet heathland; wetlands Synonyms: degenerans (Frey, 1948); invalida (Win- including bogs, basin mire, Sphagnum kettlehole; nertz, 1867); leucoptera Frey (1948); tenax (Winnertz, Phragmites and sedge bed; damp meadow; grassland 1867). at woodland fringe. iii, v–x. Literature: Edwards (1925a): 541 [as Sciara]; Chan- dler (1978): 209; Freeman (1983b): 39; Menzel (1998): SCATOPSCIARA (SCATOPSCIARA) SUBCALAMOPHILA 22. MENZEL & MOHRIG, 1991 Material examined: Bell Heath, Worcestershire, 2 Synonym: subvivida Laurence, 1994. males, 13.ix.1922, Edwards leg. (BMNH). Brampton

Wood, Huntingdonshire, 1 male, 11.iv.1977, J.H. Cole 12.x.−13.xii.1988, Foster & Procter leg. (PBLN). Char- leg. (PJCB). Burnham Beeches, Buckinghamshire, 1 terhouse Alder Holt, Surrey, 1 female, 2.vii.1977, male, 15.ix.−4.x.1995; 2 males, 8.vii.−27.vii.1996, all Chandler leg. (PPCM). Chigwell Row Wood, Chigwell Ismay leg. (PPCM). Knole Park, Kent, 1 male, Row, Essex, 1 female, 26.ix.2002; 1 female, 16.x.2002, 10.x.1971; 1 male, 28.xii.1971 (all reared), all Chan- all Ismay & Schulten leg. (DEI, UMO). Cors Erddrein- dler leg. (PPCM). Turville Heath, Buckinghamshire, 1 iog, Isle of Anglesey, 1 male, 27.vii.1988, Holmes, male, 20.vii.2002, Chandler leg. (PPCM). Boyce & Reed leg. (PPWM). Cors Gyfelog, Gwynedd, 1 male, 26.vii.1988, Holmes, Boyce & Reed leg. Ecological notes: found in ancient beech and other (PPWM). Credenhill Park Wood, Herefordshire, 1 broad-leaved woodland; parkland; fallen trunk cov- male, 11.x.1998, Chandler leg. (PPCM). Crianlarich, ered with fungus; on large dead oak. Reared from Stirling, 1 male, 12.vi.1976, Chandler leg. (NMS). Den under bark of dead beech log. iv, vii, ix–x, xii. of Pitlurg, Aberdeenshire, 1 male, 17.vii.1936, Coe leg. (BMNH). Dickleburgh, Norfolk, 2 males, 7.v.1988; 1 male, 12.ii.−26.ii.1989, all Withers leg. (PPWM). SCATOPSCIARA (SCATOPSCIARA) VITRIPENNIS Dinton Pastures Country Park, Berkshire, 1 male, (MEIGEN, 1818) 9.vi.1994; 1 male, 1 female, 7.xi.1994, all Chandler leg. Synonyms: aucta (Winnertz, 1867); basaliseta (Yang & (PPCM). East Tisted, Hampshire, 1 male, 28.viii.1972, Zhang, 1987); coracina (Zetterstedt, 1851); intermista Chandler leg. (PPCM). Epping Forest, Essex, 1 female, (Winnertz, 1867); nitidula (Zetterstedt, 1851); 20.vii.1926, Edwards leg. (BMNH); 2 males, quinquelineata (Macquart, 1834); superba (Winnertz, 28.vi.2000, Chandler leg. (PPCM). Falls of Tarnash, 1867). Moray, 1 female, 1.vii.1936, Coe leg. (BMNH). Fferam Paradwys, Isle of Anglesey, 1 male, 5.vii.1987, Chan- Literature: ? Walker (1848): 105; ? Walker (1856): 53 dler leg. (PPCM). Freshwater East, Pembrokeshire, 1 [all as Sciara vitripennis]; Edwards (1925a): 539; male, 18.vii.1986, Chandler leg. (NMS). Gateheugh, Basden (1933): 44 [all as Sciara quinquelineata]; by River Tweed, Scottish Borders, 1 male, 15.vii.1988, Chandler (1978): 209; Freeman (1983b): 39 [all as Chandler leg. (NMS). Gearagh, Cork, 1 male, Scatopsciara quinquelineata]; Freeman (1987): 203; 18.ix.1990, de Courcy Williams leg. (NMID). Glass- Laurence (1994): 118; Laurence (1996): 87; Laurence house Crops Research Station, Littlehampton, West (1997b): 228; Chandler et al. (1996): 259; Menzel Sussex, 1 male, 10.ix.1956; 2 males, 9.xi.1956; 1 male, (1998): 22; Chandler (2001): 201. 23.xi.1956; 2 males, 17.xii.1956; 1 male, no date, Material examined: Aberffraw, Isle of Anglesey, 1 all Hussey leg. (HRI). Grafham Water, Savage’s male, 9.vii.1987, Chandler leg. (PPCM). Aversley Creek, Cambridgeshire, 1 female, 30.v.1988, J.H. Cole Wood, Cambridgeshire, 1 female, 24.iv.1988, J.H. Cole leg. (PJCB). Greywell Fen, Hampshire, 1 male, leg. (PJCB). Badger Wood, Cambridgeshire, 1 male, 29.viii.2002, Chandler leg. (PPCM). Gwenfro, Isle of ix.1998, C.W. Plant leg. (PPCM). Baldock, Hertford- Anglesey, 2 males, 14.vi.1988; 1 male, 27.vii.1988, all shire, 2 females, ix.-x.1917; 1 male, v.1918, all Holmes, Boyce & Reed leg. (PPWM). Hellingly, East Edwards leg. (BMNH). Becky Woods, Manaton, Sussex, 3 males, 29.viii.1935, Low leg. (BMNH). Hick- Devon, 1 female, 23.vi.2002, J.E. Smith leg. (HRI). ling, Norfolk, 1 male, x.1988; 1 male, v.1989, all Foster Bedgebury Pinetum, Kent, 1 female, 27.x.1968, Chan- & Procter leg. (PBLN). Holme Fen, Cambridgeshire, dler leg. (PPCM). Black Park, Buckinghamshire, 3 4 males, 1 female, 27.iv.1987, J.H. Cole leg. (PJCB). males, 15.vi.−16.vi.1934 (reared), Basden leg. (NMS). Holt Island, St. Ives, Cambridgeshire, 1 female, Blaen Cilieni, Powys, 1 male, 19.vii.1989, Holmes, 18.viii.2002, J.H. Cole leg. (PJCB). Hook Farm, Bro- Boyce & Reed leg. (PPWM). Blaen y Wergloedd, mley, Greater London, 1 female, 10.vii.1966 (reared), Conwy, 1 male, 13.x.1988, Holmes, Boyce & Reed leg. Chandler leg. (PPCM). Hoxne, River Waveney, Suffolk, (PPWM). Blaise Wood, Avon, 1 female, 4.vii.1985, 2 males, 16.vii.1988, Withers leg. (PPWM). Kingsbury Chandler leg. (PPCM). Braedownie, Angus, 1 male, 4 Water Park, Warwickshire, 1 male, 13.x.2001, Chan- females, 24.vii.1937, Coe leg. (BMNH). Braemar, dler leg. (HRI). Lackham Park, Wiltshire, 1 male, Aberdeenshire, 1 female, 19.vi.−30.vi.1938, Coe leg. 14.x.2002, Chandler leg. (PPCM). Lady’s Wood, Cam- (BMNH). Brampton, Cambridgeshire, 1 male, bridgeshire, 1 male, 3.vi.1989, J.H. Cole leg. (PJCB). 22.ii.1975, J.H. Cole leg. (PJCB). Brancaster, Norfolk, Leckford Estate, Hampshire, 1 male, 25.viii.1998, 3 males, 2.ix.−16.ix.1988, Foster & Procter leg. Chandler leg. (PPCM). Letchworth, Hertfordshire, 1 (PBLN). Castlemartin Corse, Pembrokeshire, 1 male, female, x.1917; 1 male, vi.1918, all Edwards leg. 22.vii.1987; 1 male, 24.vii.1987; 1 male, 3.x.− (BMNH). Llyn Hafodol, Isle of Anglesey, 1 male, 14.x.1987, all Holmes, Boyce & Reed leg. (PPWM). 11.vii.1987, Chandler leg. (NMS). Loch Eye, Highland Catacol, Arran, North Ayrshire, 1 female, ix.1920, J. Region, 1 male, 26.v.2002, Chandler leg. (PPCM). Loch Waterston leg. (BMNH). Catﬁeld, Norfolk, 1 male, Fleet, Highland Region, 1 male, 26.v.2002, Chandler

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 BRITISH SCIARID FAUNA 127 leg. (PPCM). Lopham Fen, Norfolk, 1 male, 26.vi.1988, Ecological notes: very common in broad-leaved and Withers leg. (PPWM). Lough Gill, Castlegregory, carr woodland; also found in mixed woodland; arbore- Kerry, 2 males, 13.vii.1990, Withers leg. (PPWM). tum of conifers; heathland including Breckland; Malahide Castle, Dublin, 1 female, 17.xi.1984, Chan- wetlands including floodplain and calcareous fens, dler leg. (PPCM). Malham Tarn, North Yorkshire, floating fen, reed and sedge beds, river and lake mar- 1 male, 11.ix.−14.ix.1981, Disney leg. (PPWM). gins, Cladium with tufa, bog and water meadows; old Monkfield Wood, Cambridgeshire, 2 males, viii.1998; hedges; gravel pits; open glen with stream; grazed 2 males, x.1998, all C.W. Plant leg. (HRI, PPCM). grassland; sand dunes; grounds of horticultural Nethy Bridge, Highland Region, 1 male, 2 females, research station; parkland; gardens. At ivy and 16.vii.1936, Coe leg. (BMNH). Newbridge Fen, Kil- guelder rose flowers. Reared from nests of rook and dare, 1 male, 8.viii.−19.viii.1998, Good leg. (PPWM). coot and from the fungus Pleurotus cornucopiae on Old Slade Wood, Buckinghamshire, 1 female, pine stump. i–xii. 3.vii.1973, Chandler leg. (PPCM). Orphir, Orkney, 1 male, 28.vii.1988, Laurence leg. (PBLN). Pingle Wood, Warboys, Cambridgeshire, 1 male, 1 female, SCATOPSCIARA (XENOPYGINA) CURVILINEA 25.vi.1988, J.H. Cole leg. (PJCB). Porthcawl, Brid- (LENGERSDORF, 1934) gend, 1 female, 18.v.1906, Yerbury leg. (BMNH). Pul- Synonym: ventrospina Mohrig & Mamaev, 1983. ham St. Mary, Norfolk, 4 males, 31.iii.1989, Withers New to British list. leg. (PPWM). Radwell, Hertfordshire, 4 males, 5 females, 21.x.1917; 1 male, 31.x.1917; 2 females, Material examined: Sheffield Bottom, Theale, Berk- 11.v.1918; 1 male, v.1918, all Edwards leg. (BMNH). shire, 1 male, 31.viii.2003; 1 male, 26.ix.2003, all Reedham, Norfolk, 2 males, 26.viii.−27.x.1988, Foster Chandler leg. (PPCM). & Procter leg. (PBLN). Rhôs-y-gad, Isle of Anglesey, 1 Ecological notes: found in broad-leaved woodland. viii, male, 27.vii.1988, Holmes, Boyce & Reed leg. (PPWM). ix. Rotherfield Park, Hampshire, 1 male, 3.vii.2002, Chandler leg. (PPCM). Rye Water, Kildare, 1 male, 9.viii.1981, J.P. & M.A. O’Connor leg. (NMID). Sandy SCATOPSCIARA (XENOPYGINA) DENTIFERA (FREY, 1936) Warren, Bedfordshire, 1 male, 17.ix.1996, J.H. Cole Synonyms: aculea Mohrig, 1985; siccata Mohrig & leg. (PJCB). Sedlescombe, East Sussex, 1 male, Kauschke, 1997. 25.iii.2000, Roper leg. (PPRS). Stallode Wash, Suffolk, 2 males, 10.vi.−24.vi.1988; 8 males, 30.viii.− New to British list. 13.ix.1988, all Foster & Procter leg. (PBLN). Stoke Material examined: Blaen Cilieni, Powys, 1 male, Poges, Buckinghamshire, 1 female, 29.x.1967, Chan- 2.x.1989, Holmes, Boyce & Reed leg. (PPWM). Illtyd dler leg. (PPCM). Temple, Berkshire, 7 females, Pools, Powys, 1 male, 2.x.1989, Holmes, Boyce & Reed 30.i.−31.i.1932; 13 females, 2.ii.−31.iii.1932; 1 female, leg. (PPWM). Pant-y-Sais, Neath/Port Talbot, 1 male, 12.ii.1932 (all reared), all Basden leg. (BMNH, NMS). 13.vii.1989, Holmes, Boyce & Reed leg. (PPWM). The Loke, Norwich, Norfolk, 1 male, 5.vii.1987; 1 Rhôs-y-gad, Isle of Anglesey, 1 male, 27.vii.1988, male, 5.vi.−13.vi.1989, all Laurence leg. (PBLN). The Holmes, Boyce & Reed leg. (PPWM). Oaks, Cambridgeshire, 2 males, ix.1998, C.W. Plant leg. (NMS). Tomies Wood, Killarney, Kerry, 1 male, Ecological notes: found on Carex paniculata and 15.ix.1981, J.P. O’Connor leg. (NMID). Tomintoul, Molinia heath; calcareous fen; Juncus basin mire; Moray, 15 females, 8.viii.−11.viii.1937; 1 male, 1 Phragmites and Molinia bog. vii, x. female, viii.1937, all Coe leg. (BMNH). Tregardock Cliff, Cornwall, 1 male, 8.vii.1983, Chandler leg. (NMS). Vidlin, Shetland, 1 male, 18.vi.−21.vi.1993, SCATOPSCIARA (XENOPYGINA) FRITZI MOHRIG & Laurence leg. (PBLN). Wangford, Suffolk, 1 male, MENZEL, 1992 12.vii.−13.vii.1974, Rogers, Fitton & Day leg. (BMNH). Synonym: semantica (Laurence, 1994). Wellesbourne, Warwickshire, 2 males, 6.iv.−13.iv.2000; Literature: Laurence (1994): 114 [as Bradysia seman- 2 males, 17.viii.−31.viii.2000; 2 females, 19.i.− tica]; Menzel (1998): 22; Menzel & Mohrig (2000): 505. 15.ii.2001, all J.E. Smith leg. (HRI). Wicken Fen, Cambridgeshire, 1 male, 1 female, 26.iv.1916, Material examined: Brancaster, Norfolk, 4 males, Edwards leg. (BMNH). Woodwalton Fen, Cam- 17.vi.−1.vii.1988; 2 males, 1.vii.−15.vii.1988; 2 males, bridgeshire, 1 male, vi.1939; 1 male, 24.ix.1940, all 2.ix.1988; 7 males, 2.ix.−16.ix.1988, all Foster & Edwards leg. (BMNH). Wytham Woods, Wytham, Procter leg. [holotype and paratypes of Bradysia Oxfordshire, 1 female, 21.viii.2002, J.E. Smith leg. semantica] (DEI, PBLN); 1 male, 28.x.1988, Foster & (HRI). Procter leg. (PBLN). Brandon Marsh, Ryton-on-

Dunsmore, West Midlands, 1 male, 22.viii.2002, Men- New to British list. zel leg. (DEI). Burnham Beeches, Buckinghamshire, 1 Material examined: Llyn Pencraig, Gwynedd, 2 males, male, 1.vi.−8.vi.1999, Ismay leg. (PPCM). Castlemar- 6.vii.1988, Holmes, Boyce & Reed leg. (DEI, PPWM). tin Corse, Pembrokeshire, 1 male, 22.vii.1987; 1 male, 14.x.1987, all Holmes, Boyce & Reed leg. (PPWM). Ecological notes: found in wetland with Molinia and Catﬁeld, Norfolk, 1 male, 7.vi.−21.vi.1988, Foster & Phragmites. vii. Procter leg. [paratype of Bradysia semantica] (PBLN). Taxonomic note: This species has been transferred Cefn Bryn, Swansea/Abertawe, 1 male, 4.x.1989, from Bradysia to the subgenus Xenopygina Frey on Holmes, Boyce & Reed leg. (PPWM). Chardeon, the basis of several Scatopsciara characteristics that Gwynedd, 1 male, 11.vi.1988, Holmes, Boyce & Reed suggest a close relationship to the species complex of leg. (PPWM). Cors-y-Llyn, Powys, 1 male, 17.vii.1989, Sc. paradoxa (Frey), Sc. simillima (Tuomikoski) and Holmes, Boyce & Reed leg. (PPWM). Foulden, Norfolk, Sc. weiperti Menzel & Mohrig [redescription in Menzel 1 male, 10.vi.−8.vii.1988, Foster & Procter leg. & Heller (2005)]. These include different length of [paratype of Bradysia semantica] (PBLN). Heming- middle and posterior tibial spurs [one spur on middle ford, Cambridgeshire, 1 male, ix.1938, Edwards leg. leg more distinctly reduced than one on posterior leg]. (BMNH). Hickling, Norfolk, 1 male, 23.vi.−7.vii.1988; The wide-compact and trapezoid tegmen, the shape of 11 males, 8.viii.−13.viii.1988; 2 males, 22.viii.1988, all the aedeagus [stronger-shorter genital apodeme with Foster & Procter leg. [including paratypes of Bradysia wide-funnel-shaped base], the widely rounded apex of semantica] (DEI, PBLN). Lackham Park, Wiltshire, the gonostyle with dense bristles, strong spines and 1 male, 13.ix.2002, Chandler leg. (PPCM). Middle missing apical tooth, the narrow comb on the anterior Harling, Norfolk, 1 male, 28.vi.−12.vii.1988; 5 tibia, the absence of spines on the anterior tibiae as males, 1.ix.−15.ix.1988, all Foster & Procter leg. well as the many bristles on the basal segment of the [paratypes of Bradysia semantica] (PBLN). palpi and missing sensory pit are also typical for the Scarning, Norfolk, 10 males, 8.vi.−22.vi.1988, Foster subgenus Xenopygina. & Procter leg. [paratypes of Bradysia semantica] (PBLN). Searles Farm, Reading, Berkshire, 1 male, 9.vii.2003; 1 male, 22.vii.2003, all Chandler leg. SCATOPSCIARA (XENOPYGINA) SIMILLIMA − (PPCM). Sutton, Norfolk, 3 males, 9.vi. 23.vi.1988; 6 (TUOMIKOSKI, 1960) males, 4.viii.−18.viii.1988, all Foster & Procter leg. Synonym: pectinata (Mohrig & Menzel, 1990). [paratypes of Bradysia semantica] (PBLN). Theale, Berkshire, 1 male, 26.vi.2003, Chandler leg. (PPCM). Literature: Laurence (1994): 113 [as Bradysia pecti- Tocil Wood, Warwick University Campus, Coventry, nata]; Menzel (1998): 22. Warwickshire, 1 male, 28.viii.2003, J.E. Smith leg. Material examined: Comin Esgair Maen, Ceredigion, (HRI). Walberswick, Suffolk, 1 male, 1.vi.−15.vi.1988, 1 male, 17.ix.1986, Holmes, Boyce & Reed leg. Foster & Procter leg. [paratype of Bradysia semantica] (PPWM). Cors Barfog, Gwynedd, 1 male, 2.x.1988, (PBLN). Wheatfen Broad, Norfolk, 1 male, 8.vi.1986; 1 Holmes, Boyce & Reed leg. (PPWM). Cors Caron, Cere- male, no date, all Withers leg. (PPWM). Whitnash digion, 1 male, 15.vi.1987; 1 male, 27.vii.1987, all brook, Sydenham, Leamington Spa, Warwickshire, 1 Holmes, Boyce & Reed leg. (PPWM). Cors Farlais, male, 4.ix.2003, J.E. Smith leg. (HRI). Carmarthen, 1 male, 13.vii.1989, Holmes, Boyce & Ecological notes: found in broad-leaved and carr wood- Reed leg. (PPWM). Cors Fochno, Ceredigion, 1 male, land; humid heathland; wetlands including fen, reed 24.vii.1989, Holmes, Boyce & Reed leg. (PPWM). Cors and sedge beds, acidic mire, wet meadows and salt- Gyfelog, Gwynedd, 1 male, 5.x.1988, Holmes, Boyce & marsh; former gravel workings. vi–x. Reed leg. (PPWM). Llanbrynmair Moor, Powys, 1 male, 1.viii.1988, Holmes, Boyce & Reed leg. (PPWM). Taxonomic note: As well as the listed types for Rhôs Cilcennin, Ceredigion, 4 males, 1.x.1986, Bradysia semantica from Laurence (1994: 114), 2 fur- Holmes, Boyce & Reed leg. (PPWM). Rhôsgyll Fawr, ther examples from the location ‘Hickling, Norfolk’ Gwynedd, 1 male, 7.x.1988, Holmes, Boyce & Reed leg. with the dates 23.vi.−7.vii.1988 [1] and 22.viii.1988 [1] (PPWM). Strumpshaw, Norfolk, 3 males, 13.vi.− were found in Laurence’s collection in another box. 11.vii.1988, Foster & Procter leg. (PBLN). Thompson These do not belong to the type series for Bradysia Common, Thompson, Norfolk, 6 males, 13.ix.− semantica. 27.ix.1988, Foster & Procter leg. (DEI, PBLN). Wal- berswick, Suffolk, 3 males, 1.vi.−15.vi.1988, Foster & SCATOPSCIARA (XENOPYGINA) GABYAE (HELLER, 1998) Procter leg. (PBLN). Literature: Menzel & Heller (2005): 353 [combination Ecological notes: found in one site comprising broad- in Scatopsciara (Xenopygina)]. leaved woodland and grassland with pingo pools; oth-

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 BRITISH SCIARID FAUNA 129 erwise wetlands including valley and floodplain fens; gion, 3 males, 15.vii.1987, Holmes, Boyce & Reed leg. reed beds, ungrazed basin mire, blanket bog and (PPWM). Rhôs Pwllygawnen, Carmarthen, 2 males, raised bogs, saltmarsh and damp meadow. vi–x. 11.vii.1989, Holmes, Boyce & Reed leg. (PPWM). Rhôsgoch, Powys, 1 male, 10.x.1989, Holmes, Boyce & Reed leg. (PPWM). Rhôs-y-Gad, Isle of Anglesey, 1 SCATOPSCIARA (XENOPYGINA) WEIPERTI MENZEL & male, 5.x.1988, Holmes, Boyce & Reed leg. (PPWM). MOHRIG, 1991 Roydon Common, Norfolk, 3 males, 8.vi.−22.vi.1988, Literature: Laurence (1994): 117; Menzel (1998): 22. Foster & Procter leg. (PBLN). Strumpshaw, Norfolk, 1 male, 13.vi.−11.vii.1988, Foster & Procter leg. (PBLN). New to Irish list. Walberswick, Suffolk, 2 males, 1.vi.−15.vi.1988, Foster Material examined: Blaen y Wergloedd, Conwy, 2 & Procter leg. (DEI). Woodbastwick, Norfolk, 6 males, males, 4.viii.1988, Holmes, Boyce & Reed leg. 6.vi.−20.vi.1988, Foster & Procter leg. (PBLN). (PPWM). Brampton Wood, Cambridgeshire, 1 male, Ecological notes: found in broad-leaved and carr wood- 13.v.1989, J.H. Cole leg. (PJCB). Brancaster, Norfolk, land; grazed wet heath; wetlands especially fens, also 9 males, 2.ix.−16.ix.1988; 5 males, 28.x.1988, all Fos- sedge beds, mires, bogs and wet meadows; grazed ter & Procter leg. (PBLN). Catfield, Norfolk, 2 males, grassland; saltmarsh and reed beds; glacial pool. v–x. 27.ix.−18.x.1988, Foster & Procter leg. (PBLN). Cefn Grwhyd, Neath/Port Talbot, 1 male, 12.vii.1989, Holmes, Boyce & Reed leg. (PPWM). Colwyn Brook, SCHWENCKFELDINA FREY, 1942 Powys, 1 male, 20.vi.1989, Holmes, Boyce & Reed leg. (PPWM). Comin Esgair Maen, Ceredigion, 1 male, SCHWENCKFELDINA CARBONARIA (MEIGEN, 1830) 23.vii.1987, Holmes, Boyce & Reed leg. (PPWM). Cors Synonyms: frauenfeldi (Winnertz, 1867); illepida Caranod, Ceredigion, 1 male, 10.ix.1986, Holmes, (Winnertz, 1867); indigena (Winnertz, 1867); pilosa Boyce & Reed leg. (PPWM). Cors Erddreiniog, Isle Antonova, 1975. of Anglesey, 4 males, 27.vii.1987, Holmes, Boyce & Literature: ? Verrall (1886): 180; Edwards (1925a): 537 Reed leg. (PPWM). Cors Farlais, Carmarthen, 1 male, [all as Sciara]; Edwards (1938): 201 [as Sciara 13.vii.1989, Holmes, Boyce & Reed leg. (PPWM). Cors (Neosciara)]; Grensted (1944): 202; Laurence (1952): Geirch, Gwynedd, 3 males, 28.vii.1988; 1 male, 90 [all as Sciara]; Kidd (1953): 137; Parmenter (1960): 21.x.1988, all Holmes, Boyce & Reed leg. (PPWM). 71 [all as Sciara (Bradysia)]; Freeman (1983b): 21; Cors Graianog, Gwynedd, 1 male, 30.vi.1988; 1 male, Laurence (1994): 118; Chandler et al. (1996): 258; Lau- 28.vii.1988, all Holmes, Boyce & Reed leg. (PPWM). rence (1996): 86; Laurence (1997a): 85; Laurence Cors Gyfelog, Gwynedd, 1 male, 5.x.1988, Holmes, (1997b): 228; Menzel (1998): 22; Chandler (2001): 201 Boyce & Reed leg. (PPWM). Cors Llyn Farch a Llyn [as Schwenkfeldina; recte Schwenckfeldina]. Fanod, Ceredigion, 1 male, 10.ix.1986, Holmes, Boyce & Reed leg. (PPWM). East Harling, Norfolk, 1 male, Material examined: Acton, Greater London, 2 males, 14.vi.−28.vi.1988, Foster & Procter leg. (PBLN). Fair- no date [19th century], no collector details (BMNH). wood Common, Swansea/Abertawe, 1 male, 4.x.1989, Alice Holt Forest, Hampshire, 2 males, 23.viii.1999, Holmes, Boyce & Reed leg. (PPWM). Foulden, Norfolk, Chandler leg. (NMS). Annahala bog, Gearagh, 1 male, 10.vi.−8.vii.1988, Foster & Procter leg. Cork, 1 male, 6.iv.1990; 2 males, 10.v.1990; 4 males, (PBLN). Hickling Stubb Mill, Norfolk, 1 male, v.1988; 10.viii.1990; 1 male, 25.viii.1990; 6 males, 24.ix.1990, 1 male, vii.1988; 1 male, x.1988, all Laurence leg. all de Courcy Williams leg. (DEI, NMID). Arrochar, (PBLN); 1 male, 23.vii.−27.vii.1988; 1 male, 8.viii.− Argyll & Bute, 1 male, 26.v.1905, King leg. (NMS). 22.viii.1988, all Foster & Procter leg. (PBLN). Illtyd Ayot Green, Hertfordshire, 2 males, 15.v.1949, Colyer Pools, Powys, 1 male, 19.vii.1989, Holmes, Boyce & leg. (BMNH). Bagley, Oxfordshire, 1 male, 19.v.1928; 1 Reed leg. (PPWM). Lakenheath, Suffolk, 1 male, male, 12.v.1938, all Hamm leg. (UMO). Baldock, Hert- 30.viii.−13.ix.1988, Foster & Procter leg. (PBLN). fordshire, 2 males, 1 female, v.1918, Edwards leg. Langstone, Monmouthshire, 1 male, 12.viii.1988, (BMNH). Ballynalacken, Clare, 2 males, 1 female, Holmes, Boyce & Reed leg. (PPWM). Llanon, Swansea/ 31.viii.1959, Richards leg. (BMNH). Bayfordbury Abertawe, 1 male, 29.vi.1989, Holmes, Boyce & Reed Estate, Hertford, Hertfordshire, 1 male, 2.v.1975; 2 leg. (PPWM). Llyn Ystumllyn, Gwynedd, 2 males, males, 20.v.1975, all J.H. Cole leg. (PJCB). Bearsted, 5.x.1988, Holmes, Boyce & Reed leg. (PPWM). Middle Kent, 1 female, v.1896, Green leg. (BMNH). Bedding- Harling, Norfolk, 1 male, 28.vi.−11.vii.1988, Foster & ton, Greater London, 1 male, 19.v.1935, Parmenter Procter leg. (PBLN). Newbridge Fen, Kildare, 1 male, leg. (BMNH). Bembridge, Isle of Wight, 1 male, 2 8.viii.−19.viii.1998, Withers leg. (PPWM). Pentwd females, 29.viii.−7.ix.1932, Coe leg. (BMNH). Binfield, Marshes, Ceredigion, 2 males, 29.vii.1987, Holmes, Berkshire, 1 male, 13.v.1923, Dudgeon leg. (BMNH). Boyce & Reed leg. (PPWM). Rhôs Llawr Cwrt, Ceredi- Binnegar Quarry, Dorset, 1 male, 15.viii.2002, Gibbs

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 130 F. MENZEL ET AL. leg. (PDGB). Bix Bottom, Oxfordshire, 1 male, Five Oaks, Devon, 1 male, 16.v.1954, Spooner leg. 23.ix.2000, Raper leg. (PDGB). Black Park, Bucking- (BMNH). Forest of Dean, Gloucestershire, 2 males, 1 hamshire, 1 male, 9.ix.1944, Richards leg. (BMNH). female, 27.v.1962, K.G.V. Smith leg. (BMNH). Gal- Blessington, shore of Lacken Reservoir, Wicklow, 1 way’s Bridge, Killarney, Kerry, 1 female, 8.ix.1981, J.P. male, 15.ix.1966, Chandler leg. (PPCM). Bolt Wood, O’Connor leg. (NMID). Gearagh, Cork, 1 male, Stockerston, Leicestershire, 2 males, 24.viii.2003, 19.v.1990; 1 male, 26.ix.1990; 5 males, 10.x.1990, all Menzel & J.E. Smith leg. (DEI, HRI). Bookham Com- de Courcy Williams leg. (DEI, NMID). George Green, mon, Surrey, 1 female, 12.v.1946, Parmenter leg. Buckinghamshire, 1 male, 1.v.1943, Richards leg. (BMNH). Bournemouth, Dorset, 1 male, 26.viii.− (BMNH). Glasshouse Crops Research Station, Little- 12.ix.1991, George leg. (PPCM). Brampton Wood, hampton, West Sussex, 1 male, 10.ix.1956, Hussey leg. Cambridgeshire, 1 male, 14.v.1966; 1 female, (HRI). Glen Lednock, Perth & Kinross, 1 female, 25.v.1987, all J.H. Cole leg. (PJCB). Brandon Marsh, 21.ix.1944, Kevan leg. (NMS). Glendurgan, Cornwall, Ryton-on-Dunsmore, West Midlands, 9 males, 1 1 male, 3.x.1983, Chandler leg. (PPWM). Gomshall, female, 22.viii.2002, Menzel & J.E. Smith leg. (DEI, Surrey, 1 male, 14.v.1939, Parmenter leg. (BMNH). HRI). Brodick, Arran, North Ayrshire, 1 male, 22.v.− Goring-on-Thames, Oxfordshire, 1 male, 1 female, 25.v.1919, Edwards leg. (BMNH). Bromley, Greater 30.iv.1959; 1 female, 11.v.1962, all J.H. Cole leg. London, 1 female, 20.v.1966; 1 male, 2.ix.1966, all (PJCB). Hackhurst Farm, Gomshall, Surrey, 1 male, Chandler leg. (NMS, PPCM). Brown’s Folly, Avon, 1 27.viii.1966, Chandler leg. (PPCM). Hartslock, male, 24.viii.2000, Gibbs leg. (PDGB). Burledge Hill, Oxfordshire, 1 male, 14.v.2000, Raper leg. (PDGB). Avon, 1 male, 23.vi.2001, Gibbs leg. (PDGB). Burren, Hayling Island, Hampshire, 1 male, 21.iv.1953 near Caher River, Clare, 1 male, 7.vi.−8.vi.1970, Vane- (reared), no collector detail (BMNH); 6 males, 1 Wright leg. (BMNH). Byﬂeet, Surrey, 1 male, female, 21.iv.1963, Vane-Wright leg. (BMNH). Hay 15.ix.1920, Coe leg. (BMNH). Cadair Idris, Gwynedd, Wood, Baddesley Clinton, Solihull, West Midlands, 4 1 male, 4.ix.1993, Laurence leg. (PBLN). Caher, Bur- males, 25.viii.2002, Menzel leg. (DEI). Henﬁeld, West ren, Clare, 2 males, 7.v.−8.v.1970, Vane-Wright leg. Sussex, 3 females, 14.v.1989, J.H. Cole leg. (PJCB). (BMNH). Castlefreke, Cork, 1 male, 12.v.1989, With- Hickling, Norfolk, 1 female, v.1988; 2 males, v.1989, all ers leg. (PPWM). Castlehill, Dumfries & Galloway, 1 Foster & Procter leg. (PBLN); 1 male, 27.v.1991, With- female, 22.ix.1979, Duncan leg. (NMS). Catacol, ers leg. (PPWM). Hitchin, Hertfordshire, 3 females, Arran, North Ayrshire, 1 male, ix.1920, J. Waterston vi.1918, Edwards leg. (BMNH). Hobb’s Quarry, Glouc- leg. (BMNH). Chedworth, Gloucestershire, 1 male, estershire, 1 male, 4.ix.2002, Gibbs leg. (PDGB). Hod 5.vi.1936, Edwards leg. (BMNH). Chesil Beach, Dor- Hill, Dorset, 1 male, 1 female, 22.v.1968, Edwards leg. set, 1 male, 12.ix.1968, Vane-Wright leg. (BMNH). (BMNH). How Hill, Norfolk, 1 male, 27.v.1989; 1 male, Coed-y-Brenin, Gwynedd, 1 male, 18.ix.1968, Vane- 13.viii.1989, all Withers leg. (PPWM). Hunstanton, Wright leg. (BMNH). Collinpark Wood, Gloucester- Norfolk, 1 female, 16.v.1937, Parmenter leg. (BMNH). shire, 1 male, 3.ix.2002, Gibbs leg. (PDGB). Comrie, Inishowen, Donegal, 1 female, ix.1928, Latham leg. Perth & Kinross, 1 female, 14.ix.−23.ix.1940, Kevan (NMS). Kelham Bridge, Ravenstone, Leicestershire, 5 leg. (NMS). Coombe Abbey Country Park, Coventry, males, 1 female, 24.viii.2003, Menzel & J.E. Smith leg. Warwickshire, 56 males, 28.viii.2003, Menzel & J.E. (DEI, HRI). Killaun Bog, Offaly, 3 males, ix.2000, Smith leg. (DEI, HRI). Corsemalzie, Dumfries & Gal- Withers leg. (PPWM). Kirtling, Cambridgeshire, 3 loway, 1 female, v.1899, Gordon leg. (NMS). Cothill, males, 1 female, 9.v.1920, Collin leg. (UMO). Kneb- Oxfordshire, 2 males, 6.v.1934, Collin leg. (UMO); 2 worth, Hertfordshire, 1 male, 7.ix.−9.ix.1923, Edwards males, 28.viii.1989, Chandler leg. (NMS). Cripplegate, leg. (BMNH). Knole Park, Kent, 2 males, 20.v.1967; 2 Greater London, 1 male, 17.v.1947, Parmenter males, 12.ix.1971, all Chandler leg. (NMS, PPCM). leg. (BMNH). Daneway, Gloucestershire, 1 male, Lancelot Clark Storth Nature Reserve, Cumbria, 2 12.x.1979, Chandler leg. (NMS). Dinton Pastures males, 27.viii.1965, J.H. Cole leg. (PJCB). Lea Mead- Country Park, Berkshire, 1 male, 1.v.1993; 1 male, ows, Newtown Linford, Leicestershire, 66 males, 8.viii.1993; 1 female, 13.viii.1995; 1 male, 1 female, 5 females, 24.viii.2003, Menzel & J.E. Smith leg. 13.vii.1998, all Chandler leg. (NMS, PPCM). East (DEI, HRI). Leaves Green, Greater London, 1 male, Harptree Combe, Somerset, 1 female, 6.vii.1985, 18.v.1968, Chandler leg. (NMS). Leckford Estate, Chandler leg. (PPCM). Ebbor Gorge, Somerset, 1 Hampshire, 3 males, 1 female, 19.v.−20.v.1971; 1 male, male, 3.vii.1989, Chandler leg. (NMS). Emerson’s 3.vi.1972, all Stubbs leg. (PPCM); 1 male, 26.viii.1972; Green, Avon, 1 male, 12.v.2001, Gibbs leg. (PDGB). 1 male, 1977 (reared), Chandler leg. (PPCM). Lelant, Emo Park, Laois, 1 male, 11.vii.1971, Chandler leg. Cornwall, 1 male, 1 female, 16.viii.1912, Yerbury leg. (PPCM). Felbridge, Surrey, 1 male, 26.v.1974, Chan- (BMNH). Letchworth, Hertfordshire, 1 male, 1 female, dler leg. (PPCM). Fisher’s Wood, Bromley, Greater v.1917; 1 male, vi.1917, all Edwards leg. (BMNH). Lit- London, 1 female, 10.v.1966, Chandler leg. (NMS). tle Wenallt, Monmouthshire, 1 male, 9.vii.1980, Chan-

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 BRITISH SCIARID FAUNA 131 dler leg. (NMS). Llandre, Ceredigion, 1 male, 1 female, Scragh Bog, Westmeath, 1 male, 10.v.1989, Withers 31.viii.1943, Richards leg. (BMNH). Loch Earn, Perth leg. (PPWM). Selkirk, Scottish Borders, 1 female, & Kinross, 1 female, 23.ix.1940, Kevan leg. (NMS). 12.ix.1962, Pelham-Clinton leg. (NMS). Sheep Leas, Lough Luogh, Clare, 1 male, 31.v.1984, J.P. O’Connor West Horsley, Surrey, 1 female, 21.v.1966, Chandler leg. (NMID). London, Greater London, 1 female, leg. (PPCM). Shrubland Park, Suffolk, 1 male, iii.1955, Buxton leg. (BMNH). Long Range, Killarney, 27.viii.1920, Morley leg. (BMNH). Slough, Berkshire, Kerry, 1 male, 14.ix.1981, J.P. O’Connor leg. (NMID). 1 female, 11.v.1933; 1 male, 20.viii.1936; 1 male, Lopham Fen, Norfolk, 1 male, 20.v.1990, Withers leg. 2.ix.1938; 1 male, 11.v.1942; 1 female, 31.viii.1942, all (PPWM). Lough Bunny, Clare, 1 male, 21.v.1985, J.P. Richards leg. (BMNH). Snailbeach, Shropshire, 4 & M.A. O’Connor leg. (NMID). Mereworth Wood, Kent, males, 17.iv.1908, Verrall leg. (UMO). 1 male, 8.v.1921, 1 male, 29.viii.1966, Chandler leg. (NMS). Mersea, Edwards leg. (BMNH). St. David’s Airﬁeld, Pem- Essex, 1 male, x.1909, no collector detail (BMNH). brokeshire, 1 male, 29.ix.1987, Holmes, Boyce & Middle Hope, Avon, 1 male, 9.v.2001, Gibbs leg. Reed leg. (PPWM). Stamford, Lincolnshire, 1 male, (PDGB). Mitcham Common, Greater London, 1 male, 17.v.1937, Parmenter leg. (BMNH). Stoke Common, 1 female, 19.v.1937, Parmenter leg. (BMNH). Monu- Buckinghamshire, 1 male, 9.ix.1967, Chandler leg. ment Field, Epping Forest, Essex, 1 male, 1 female, (NMS). Stoke Row, Oxfordshire, 1 female, 3.vi.1962, 19.iv.−2.v.1999, Dagley leg. (HRI). Narborough Bog, Burtt leg. (BMNH). Stoke Wood, Herefordshire, 2 Whetstone, Leicestershire, 11 males, 24.viii.2003, males, 5.iv.1905; 2 males, 5.ix.1905, all Verrall leg. Menzel & J.E. Smith leg. (DEI, HRI). Natural History (BMNH, UMO). Stokeford Heath, Dorset, 1 male, Museum, South Kensington, Greater London, 1 male, 7.v.2001, Gibbs leg. (PDGB). Tarrington, Hereford- 16.v.1916, Edwards leg. (BMNH). New Barnet, shire, 1 male, 7.ix.1905, Verrall leg. (UMO). Terenure, Greater London, 1 female, 27.viii.1964, K.G.V. Smith Dublin City, Dublin, 1 male, 7.v.1989, Ashe leg. leg. (BMNH). New Forest, Hampshire, 3 males, (NMID). The Loke, Norwich, 1 male, 7.ix.1987; 2 16.v.1968, Parmenter leg. (BMNH). Newbridge males, 13.v.1988; 1 male, 16.viii.1988; 2 males, 5.vi.− Fen, Kildare, 1 male, 8.viii.−19.viii.1998, Good leg. 13.vi.1989; 1 male, 2.v.−6.v.1992; 2 males, 4.v.− (PPWM). Newmarket, Cambridgeshire, 7 males, 2 6.v.1993; 1 male, 30.vii.−2.viii.1993; 3 males, 27.viii.− females, 19.v.1884; 1 male, 1 female, 22.viii.1894; 2 1.ix.1997, all Laurence leg. (PBLN). Thornham males, 31.v.1908, all Verrall leg. (BMNH, UMO). Magna, Suffolk, 1 male, 29.viii.1988, Withers leg. Northwood, Greater London, 1 female, 12.v.1921, Aus- (PPWM). Thundersley Glen, Essex, 1 male, 30.iv.1986, ten leg. (BMNH). Oxford University Museum, Oxford, Withers leg. (PPWM). Tilgate Forest, West Sussex, 1 Oxfordshire, 1 female, 13.v.1921; 1 female, 28.v.1924, male, 28.v.1939, Parmenter leg. (BMNH). Tocil Wood, all Hamm leg. (UMO). Patcham, East Sussex, 1 male, Warwick University Campus, Coventry, Warwickshire, 1 female, 15.v.1935; 1 male, 24.v.1936, all Parmenter 1 male, 28.viii.2003, J.E. Smith leg. (HRI). Tomies leg. (BMNH). Pencarreg, Carmarthen, 1 male, Wood, Killarney, Kerry, 1 male, 15.ix.1981, J.P. O’Con- 28.ix.1974, Chandler leg. (NMS). Pennington, Hamp- nor leg. (NMID). Totton, Hampshire, 1 female, no date, shire, 4 males, 12.v.1964, Colyer leg. (BMNH). Ply- Vardy leg. (BMNH). Upton, Chester, Cheshire, 1 male, mouth, Devon, 2 males, 1 female, 14.v.1954, Spooner 21.v.1958, Colyer leg. (BMNH). Upton Meadows, leg. (BMNH). Porthcawl, Bridgend, 1 male, 18.v.1906, Slough, Berkshire, 1 female, 7.v.1942, Richards leg. Yerbury leg. (BMNH). Purn Hill, Avon, 1 male, (BMNH). Walney, Cumbria, 1 male, 20.viii.1992, 21.viii.2001, Gibbs leg. (PDGB). Putsborough, Devon, Birkett leg. (PPCM). Wellesbourne, Warwickshire, 3 1 male, 10.viii.−16.viii.1946, d’Assis-Fonseca leg. males, 1999; 5 males, 2 females, 10.v.−24.v.2001, all (BMNH). Radwell, Hertfordshire, 1 male, viii.1918; J.E. Smith leg. (HRI). West Malling, Kent, 1 male, 1 male, viii.1919, all Edwards leg. (BMNH). Rhôs 29.viii.1966, Chandler leg. (PPCM). West Mersea, Rhydd, Ceredigion, 1 male, 25.ix.1986, Holmes, Boyce Essex, 1 male, 1 female, 24.vii.1909, Wesche leg. & Reed leg. (PPWM). River Teme, Bromﬁeld, Shrop- (BMNH). Weston Big Wood, Avon, 1 male, 1.ix.2001, shire, 1 male, 24.v.2002, Pont leg. (HRI). Rodborough Gibbs leg. (PDGB). Weston, Hertfordshire, 1 male, Forest, Gloucestershire, 3 males, 1.v.−2.v.1944, Bain- 4.x.1931, Edwards leg. (BMNH). Wexham Place Wood, bridge-Fletcher leg. (BMNH). Rostrevor, Down, 1 Buckinghamshire, 1 male, 7.vi.1967, Chandler leg. female, 16.ix.1925, Johnson leg. (NMID). Ryton Wood, (NMS). Wheatfen Broad, Norfolk, 1 male, 11.vii.1987, Bubbenhall, Coventry, West Midlands, 10 males, Withers leg. (PPWM). Whitnash brook, Sydenham, 25.viii.2002, Menzel & J.E. Smith leg. (DEI, HRI). Leamington Spa, Warwickshire, 1 male, 4.ix.2003, J.E. Salisbury, Wiltshire, 1 female, 26.v.1972, Dear leg. Smith leg. (HRI). Wicken Fen, Cambridgeshire, 1 (BMNH). Sally Gap, Wicklow, 1 male, 9.ix.1999, With- male, 8.vii.1989, Chandler leg. (NMS); 1 male, ers leg. (PPWM). Sandbanks, Hampshire, 2 females, 12.v.1990, Withers leg. (PPWM). Willes Meadow, 13.v.1951, Colyer leg. (BMNH). Savernake Forest, Leamington Spa, Warwickshire, 4 males, 1 female, Wiltshire, 1 male, 24.ix.1972, Chandler leg. (PPCM). 27.viii.2002, Menzel & J.E. Smith leg. (DEI, HRI).

Windermere, Cumbria, 1 female, vi.1929, Edwards female, 18.ix.1905, all A.E.J. Carter leg. (UMO). leg. (BMNH). Windsor Forest, Berkshire, 1 male, Archers Wood, Cambridgeshire, 1 male, 1.viii.1986, 8.ix.1980, Chandler leg. (PPCM). Wisley, Surrey, 1 J.H. Cole leg. (PJCB). Baldock, Hertfordshire, 1 male, 10.v.1923, Dudgeon leg. (BMNH). Wood Green, female, 8.vii.1937, Edwards leg. (BMNH). Ban-y-Gor Greater London, 1 male, 12.v.1946; 1 female, Wood, Gloucestershire, 1 male, 12.ix.2001, Gibbs leg. 23.viii.1947; 3 females, 14.v.1948; 1 female, 3.ix.1948; (PDGB). Barton Broad, Norfolk, 2 females, 5.viii.1906, 1 female, 26.v.1951, all Colyer leg. (BMNH). Wood- Rothschild leg. (BMNH). Barton Mills, Suffolk, 4 bastwick, Norfolk, 1 male, viii.1932, Edwards leg. females, 9.viii.1901; 1 female, 10.viii.1901, all Hamm (BMNH). Woodditton Wood, Cambridgeshire, 1 male, leg. (UMO). Bayswater, Oxfordshire, 2 females, 20.v.1928, Collin leg. (UMO). Woodwalton Fen, Cam- 14.vii.1917, Hamm leg. (UMO). Beaulieu, New Forest, bridgeshire, 1 male, 4.ix.1971; 1 male, 16.v.1982, all Hampshire, 1 female, 13.vii.1895, no collector detail J.H. Cole leg. (PJCB). Woolacombe Bay, Devon, 2 (BMNH). Bishop Wood, North Yorkshire, 1 female, males, 14.viii.1945, Richards leg. (BMNH). Wychwood, 20.vii.1997, Chandler leg. (PPCM). Bix Bottom, Oxfordshire, 1 male, 12.ix.1998, Chandler leg. (NMS). Oxfordshire, 1 female, 6.vii.2002, Chandler leg. Wytham Woods, Wytham, Oxfordshire, 10 males, (PPCM). Blackford Hill, City of Edinburgh, 6 males, 6 21.viii.2002, Menzel & J.E. Smith leg. (DEI, HRI). females, 2.vii.2002 (reared), Bland leg. (NMS). Bod- min, Cornwall, 1 female, 18.viii.1936; 1 female, Ecological notes: very common in all types of wood- 20.viii.1936, all Parmenter leg. (BMNH). Bonhill, land; sandy and wet heathland; wetlands including West Dunbartonshire, 1 male, 3 females, 22.vii.1905; fens, reed and sedge beds, Cladium beds with tufa, 1 male, 6.viii.1905, all Malloch leg. (NMS). Bookham basin mire, quaking and raised bogs, butterbur Common, Surrey, 1 female, 26.viii.1934, Parmenter swamp, lake and river margins and wet meadows; leg. (BMNH). Boughton Fen, Norfolk, 2 males, 10.vi.- former gravel workings; open glen with stream; ix.1980, Foster & Procter leg. (PBLN). Braco, Glen meadow hedges and ditches; pasture; common on cal- Eagles, Perth & Kinross, 2 females, 13.viii.1948, Jack- careous and neutral grassland; open montane habi- son leg. (NMS). Budshead Wood, Devon, 1 female, tats; grikes; coastal shingle; grounds of horticultural 1.vii.1889, Yerbury leg. (BMNH). Burghfield Common, research station; gardens; cabbage field. At hawthorn Berkshire, 1 female, 11.vii.1970, Chandler leg. (NMS). flowers; larvae in group at base of grass stems, adults Burnham Beeches, Buckinghamshire, 2 females, in large swarm. Reared from the fungus Peziza carnea 5.vii.−20.vii.1995, Ismay leg. (HRI). Bushey Heath, and poplar log, in laboratory. iii–xi. Hertfordshire, 4 females, 22.vii.1912, Edwards leg. (BMNH). Burrows, no county, 2 females, vi.1912, no SCIARA MEIGEN, 1803 collector detail (BMNH). Carrigower, Wicklow, 1 female, 8.viii.1979, Speight leg. (PPCM). Castlebar SCIARA FLAVIMANA ZETTERSTEDT, 1851 Lough, Mayo, 1 female, 31.vii.1911, Grimshaw leg. Synonyms: fulgens Winnertz, 1867; mannii Winnertz, (NMS). Catfield, Norfolk, 1 male, 1 female, 9.viii.1980, 1867. Shaw leg. (NMS). Chippenham Fen, Cambridgeshire, Literature: Freeman (1983b): 19; Menzel (1998): 22. 1 male, 2.viii.1988, Chandler leg. (PPCM). Church Stretton, Shropshire, 1 female, 24.vii.1920, Edwards Material examined: Nairn, Highland Region, 1 female, leg. (BMNH). Cors Erddreiniog, Isle of Anglesey, 1 10.viii.1904, King leg. (BMNH). male, 27.vii.1988, Holmes, Boyce & Reed leg. (PPWM). Ecological notes: no details of habitat recorded. viii. Cors Goch, Isle of Anglesey, 1 male, 18.vii.1989, Holmes, Boyce & Reed leg. (PPWM). Cosford Mill, Surrey, 1 female, 28.vi.1970, Chandler leg. (NMS). SCIARA HEMEROBIOIDES (SCOPOLI, 1763) Cothill, Oxfordshire, 1 male, 1 female, 13.vii.1985, Synonyms: lateralis Meigen, 1818; thomae (Linnaeus, Chandler leg. (NMS). Croft Pascoe Pool, Cornwall, 1 1767); valida Winnertz, 1867. female, 2.x.1983, Chandler leg. (NMS). Crowborough, East Sussex, 1 male, 2 females, 22.viii.1911, Jackson Literature: Haliday (1833): 148; Walker (1848): 104; leg. (NMS). Crownhill Park, Plymouth, Devon, 1 Walker (1856): 52; Edwards (1925a): 535; Grensted female, 15.vii.1889, Yerbury leg. (BMNH). Dinas (1944): 202; Parmenter (1950): 110; Perry & Stubbs Head, Pembrokeshire, 1 female, 11.ix.1943, Grensted (1978): 71; Freeman (1983b): 18; Skidmore et al. leg. (UMO). Dinton Pastures Country Park, Berk- (1987): 119; Laurence (1994): 118 [all as Sciara shire, 1 male, 1 female, 26.vi.1993; 1 female, thomae]; Menzel (1998): 22. 25.vi.1994, all Chandler leg. (PPCM). Dover, Kent, 1 Material examined: Aberffraw, Isle of Anglesey, 1 female, viii.1900, no collector detail (BMNH). East female, 9.vii.1987, Chandler leg. (PPCM). Aberfoyle, Bergholt, Suffolk, 1 female, 22.vii.−29.vii.1970, Lane Stirling, 1 female, 21.v.1896; 1 female, 11.ix.1905; 1 leg. (BMNH). Ebbor Rocks, Somerset, 1 male,

31.viii.1948, Cowley leg. (BMNH). Exeter, Devon, 1 Brunetti leg. (BMNH). Ports Down, Hampshire, 3 female, 17.vii.1891, Verrall leg. (UMO). Felden, Hert- females, 29.vii.1957, Clark leg. (BMNH). Puttenham, fordshire, 1 female, 16.viii. 1893, A. Piffard leg. Surrey, 1 male, 31.vii.1950, Parmenter leg. (BMNH). (BMNH). Fferam Paradwys, Isle of Anglesey, 1 male, Red Wharf, Isle of Anglesey, 1 female, 15.viii.1948, 5.vii.1987, Chandler leg. (NMS). Folkestone, Kent, 1 Grensted leg. (UMO). Redlake Cottage Meadow, Corn- male, 1 female, 1866, A. Piffard leg. (BMNH). Forest of wall, 1 female, 12.vii.2000, Chandler leg. (PPCM). Dean, Gloucestershire, 1 male, 21.vii.1897, Verrall leg. Rowridge Valley, Isle of Wight, 1 female, 2.vii.1980, (UMO). Freshfield, Merseyside, 1 male, 1 female, Chandler leg. (PPCM). Savernake Forest, Wiltshire, 19.ix.1970, Hutson leg. (BMNH). Glensanda, Morv- 1 female, 24.vii.1973, Chandler leg. (NMS). Send, ern, Highland Region, 4 females, 10.ix.1961, Pelham- Surrey, 1 male, 1 female, 27.vi.1948, Parmenter leg. Clinton leg. (NMS). Goring Heath, Oxfordshire, 1 (BMNH). Shapwick Heath, Somerset, 1 female, female, 14.vii.1965, Burtt leg. (BMNH). Goring-on- 28.viii.1942, Cowley leg. (BMNH). Silchester Com- Thames, Oxfordshire, 1 male, 2 females, 11.vii.1957, mon, Hampshire, 1 female, 22.viii.1963, Burtt leg. J.H. Cole leg. (PJCB). Gurnard’s Head, Cornwall, (BMNH). Slapton Ley Nature Reserve, Devon, 2 4 females, 1.viii.−8.viii.1936, Hale-Carpenter leg. females, 18.vii.−19.vii.1964, Chandler leg. (PPCM). (UMO). Hackhurst Wood, Surrey, 1 female, Southrepps Common, Norfolk, 1 female, 8.vii.1993, 27.viii.1966, Chandler leg. (PPCM). Hartcliff Combe, J.H. Cole leg. (PJCB); 1 female, 10.vii.1993, Chandler Dorset, 3 females, 30.vi.1998, Chandler leg. (NMS, leg. (PPCM). St. Erth, Cornwall, 1 female, 8.viii.1935, PPCM). Hatfield Forest, Hertfordshire, 1 female, Parmenter leg. (BMNH). Sutton Fen, Cambridgeshire, 24.vii.1960, Vardy leg. (BMNH). Hobb’s Quarry, 1 female, 12.vii.1983, Chambers leg. (BMNH). Sway- Gloucestershire, 1 male, 4.ix.2002, Gibbs leg. (PDGB). thling, Hampshire, 1 female, 25.vi.1989, Chandler leg. Hogley, Oxfordshire, 2 males, 1 female, 4.vii.1914, (NMS). Tamerton Folliatt, Devon, 1 female, 8.viii. Hamm leg. (UMO). Humbie, East Lothian, 1 female, 1889, Yerbury leg. (BMNH). Thatcham, Berkshire, 1 13.vii.1988, J.H. Cole leg. (PJCB). Killarney, Kerry, 2 female, 19.vii.1964, J.H. Cole leg. (PJCB). The Keld, females, 23.viii.1902, King leg. (NMS). Leap, Cork, 1 Driffield, North Yorkshire, 1 female, 15.vii.1996, female, 15.vii.1976, Chainey leg. (BMNH). Leckford Chandler leg. (PPCM). Theale, Berkshire, 2 males, 10 Estate, Hampshire, 1 female, 25.vii.1998, Chandler females, 26.vi.2003, Chandler leg. (HRI). Thompson leg. (NMS). Lelant, Cornwall, 2 males, 3.viii.1941; 1 Common, Norfolk, 1 male, 4.vii.1993, J.H. Cole leg. male, 28.viii.1947, all Parmenter leg. (BMNH). Llan- (PJCB). Thursley Common, Surrey, 1 female, deilo, Carmarthen, 3 males, 1 female, 26.viii.− 13.vii.1968, Chandler leg. (PPCM). Tintagel, Corn- 2.ix.1967, Elstow leg. (PPCM). Llyn Hafodol, Angle- wall, 2 females, 2.viii.−15.viii.1947, Benson leg. sey, 1 female, 11.vii.1987, Chandler leg. (NMS). (BMNH). Torc Falls, Killarney, Kerry, 1 male, Long Crichel, Dorset, 1 female, 2.viii.1959, Colyer 11.vii.1990, Withers leg. (PPWM). Torrs, Dumfries & leg. (BMNH). Lough Derragh, Longford, 1 male, Galloway, 1 female, 4.viii.1981, Duncan leg. (NMS). 15.vii.1990, Withers leg. (PPWM). Lundy Island, Tresco, Scilly Isles, Cornwall, 1 female, 16.vii.1927, Devon, 1 female, 20–22.vii.1972, Hutson leg. (BMNH). Richards leg. (BMNH). Tring, Hertfordshire, 1 female, Lyndhurst Road, New Forest, Hampshire, 1 female, 26.vii.1031, Edwards leg. (BMNH). Tubney, Oxford- 29.vii.1894, Yerbury leg. (BMNH). Matley Bog, New shire, 1 female, 1.viii.1910; 1 female, vii.1912; 1 Forest, Hampshire, 3 females, 12.viii.−13.viii.1903, female, 2.ix.1916, all Hamm leg. (UMO). Twitchen, Hamm leg. (UMO). Meathop, Cumbria, 1 female, Devon, 4 females, 28.vii.−29.vii.1909, no collector 17.vii.1945, no collector detail (UMO). Merton, detail (UMO). Upware, Cambridgeshire, 1 female, Greater London, 3 females, 27.vii.1899, Walsingham 15.vii.1875, Verrall leg. (BMNH). Uxbridge, Greater leg. (BMNH). Mevagissey, Cornwall, 1 female, London, 1 female, vii.1924, Edwards leg. (BMNH). 13.ix.1941, Grensted leg. (UMO). Minsmere, Suffolk, 1 Weston Big Wood, Avon, 1 male, 1.ix.2001, Gibbs leg. male, 5.viii.−27.ix.2001, Edwards leg. (PDGB). Myross (PDGB). Wheatfen, Surlingham, Norfolk, 2 females, Wood, Leap, Cork, 1 male, 11.vii.1977, Chainey leg. 27.vi.1950, Biggs leg. (NMS). Wheatfen Broad, Nor- (BMNH). Newton Abbot, Devon, 1 female, 9.viii.1901; folk, 1 male, 13.vii.1986, Withers leg. (PPWM). 2 females, 8.viii.1902, Hamm leg. (UMO). Oughton Wicken Fen, Cambridgeshire, 1 female, 11.viii.1901, Head, Hertfordshire, 1 female, 2.vii.1966, Colyer leg. no collector detail (UMO). Wisley, Surrey, 1 female, (BMNH). Ovington, Hampshire, 1 male, 2 females, 5.x.1922; 1 female, 19.x.1922, Dudgeon leg. (BMNH). 14.vii.1990, Chandler leg. (NMS, PPCM). Penzance, Wisley Common, Surrey, 3 females, 7.vii.1968, Chan- Cornwall, 1 male, 5 females, 11.vii.1871, Verrall leg. dler leg. (PPCM). Woodwalton Fen, Cambridgeshire, 1 (BMNH, UMO). Piddletrenthide, Dorset, 1 female, female, 6.vii.1970, J.H. Cole leg. (PJCB). Woking, Sur- 6.viii.1941, Richards leg. (BMNH). Pitlochry, Perth & rey, 2 females, 15.viii.1891, Beaumont leg. (BMNH). Kinross, 1 female, 2.ix.1892, Beaumont leg. (BMNH). Woolacombe, Devon, 1 female, 14.viii.1945, Richards Plymstock, Plymouth, Devon, 3 females, 30.viii.1901, leg. (BMNH). Wyre Forest, Worcestershire, 2 males,

14.ix.1922, Edwards leg. (BMNH). Yelverton, Devon, 1 Ecological notes: found in broad-leaved woodland female, 11.viii.1976, no collector detail (BMNH). No especially carr; wetlands especially fens, reed and locality details [? Holywood, Down], 2 males, 2 sedge beds, ombrotrophic and basin mires, Molinia females, no date, Haliday leg. (NMID). and quaking bogs, saltmarsh and water meadows; heath grassland and scrub; grassland with pingo Ecological notes: very common in broad-leaved, mixed pools; rough pasture with ditches. vi–x. and carr woodland; heathland; wetlands including fens, bogs, reed and Cladium beds; calcareous grassland; grassland with pingo pools; ditches; old hedges; SCIARA LACKSCHEWITZI (LENGERSDORF, 1934) gravel pits; in mixed vegetation by coastal lake; dunes; garden. At flowers of water dropwort, wild parsnip and Synonym: mendax Tuomikoski, 1960. hogweed. v–x. Literature: Menzel (1998): 22. Material examined: Cors Gyfelog, Gwynedd, 1 male, 30.vi.1988, Holmes, Boyce & Reed leg. (PPWM). SCIARA HUMERALIS ZETTERSTEDT, 1851 Moorthwaite, Cumbria, 1 male, 20.vi.1988, no collec- Synonyms: analis var. bezzii Del Guercio, 1905; tor detail (PPWM). armata Winnertz, 1867; hamatilis Yang, Zhang & Yang, 1993. Ecological notes: found on site comprising floodplain fen, Sphagnum, Carex limosa and Phragmites. vi. Literature: Freeman (1983b): 19; Laurence (1994): 118; Menzel (1998): 22; Withers (2002): 115. Material examined: Aberithon Turbary, Powys, 1 SCIARA NURSEI FREEMAN, 1983 RESTIT. male, 3.vii.1989, Holmes, Boyce & Reed leg. (PPWM). Synonyms: marginata Mohrig & Krivosheina, 1983 Blackheath, Albury, Surrey, 1 male, 2.vii.1968, Chan- [preocc.]; ulrichi Menzel & Mohrig, 1998 syn. nov. dler leg. (PPCM). Chilbolton Common, Hampshire, 4 Literature: Freeman (1983a): 161; Freeman (1983b): males, 6.vii.1991, Chandler leg. (NMS, PPCM). Cors 18; Laurence (1994): 118 [all as Sciara nursei]; Menzel Gyfelog, Gwynedd, 1 male, 26.vii.1988, Holmes, Boyce (1998): 22; Menzel & Mohrig (2000): 523 [all as Sciara & Reed leg. (PPWM). Gors Llwyn, Neath/Port Talbot, ulrichi and Sciara flavomarginata, misidentification 1 male, 12.vii.1989; 1 male, 4.x.1989, all Holmes, and incorrectly synomymized from Sciara nursei to Boyce & Reed leg. (PPWM). Greywell Fen, Hampshire, Sciara flavomarginata]; Chandler et al. (2002): 124 1 male, 11.viii.1999; 1 male, 20.vi.2002; 2 males, 1 [Sciara ulrichi]. female, 29.viii.2002, all Chandler leg. (HRI, PPCM). Horning Ferry, Norfolk, 1 male, 1 female, 21.viii.1932, Material examined: Burnham Beeches, Buckingham- Collin leg. (UMO). Leckford Estate, Hampshire, 1 shire, 2 males, 1 female, 21.vi.−5.vii.1995; 4 males, male, 29.vi.1974; 1 male, 14.viii.1983, all Chandler 20.vii.−2.viii.1995; 11 males, 2.viii.−16.viii.1995, all leg. (NMS, PPCM). Llyn Ystumllyn, Gwynedd, 1 male, Ismay leg. (PPCM). Cambridge, Cambridgeshire, 1 5.x.1988, Holmes, Boyce & Reed leg. (PPWM). Lough female, 4.vii.1903; 1 female, 13.viii.1903, all Jenkin- Derragh, Longford, 2 males, 15.vii.1990, Withers leg. son leg. [paratypes of Sciara nursei] (BMNH). Cat- (PPWM). New Forest, Hampshire, 1 male, 16.vii.1907, field, Norfolk, 3 males, 29.ix.−12.x.1988, Foster & King leg. (BMNH). Redmoor, Cornwall, 1 male, Procter leg. (PBLN). Colt Park Wood, North Yorkshire, 8.x.2001, J.H. Cole leg. (PJCB). Rempstone Heath, 1 male, 27.viii.−30.viii.1981, Disney leg. (PPWM). Dorset, 2 males, 9.ix.1906, Verrall leg. (UMO). Rhôs Daneway, Gloucestershire, 1 male, 12.x.1979, Chan- Cruglas, Carmarthen, 1 male, 29.vi.1989; 1 male, dler leg. (PPCM). Epping Forest, Essex, 1 male, 13.vii.1989, all Holmes, Boyce & Reed leg. (PPWM). 27.vi.2000, J.H. Cole leg. (PJCB). Lea Meadows, New- Rhôs Rhydd, Ceredigion, 1 male, 8.x.1987, Holmes, town Linford, Leicestershire, 2 males, 24.viii.2003, Boyce & Reed leg. (PPWM). Scragh Bog, Westmeath, 1 J.E. Smith leg. (HRI). Letchworth, Hertfordshire, 2 male, 15.vii.1990, Withers leg. (PPWM). Thompson females, vi.-vii.1936, Edwards leg. [paratypes of Sci- Common, Norfolk, 3 males, 16.vii.1989, Chandler leg. ara nursei] (BMNH). Moy, Tyrone, 2 males, 24.ix.− (NMS, PPCM). Upton, Norfolk, 2 males, 5.vii.1934, 30.ix.1984, Boston leg. (PPCM). Reedham, Norfolk, 1 Collin leg. (UMO). Walberswick, Suffolk, 4 males, male, 26.viii.−27.x.1988, Foster & Procter leg. (PBLN). 15.vi.−29.vi.1988, Foster & Procter leg. (PBLM, Ryton Wood, Bubbenhall, Coventry, West Midlands, 1 PPCM, PPWM). Winnall Moor, Hampshire, 1 male, male, 25.viii.2002, J.E. Smith leg. (HRI). Sea Mere, 25.vi.1989, Chandler leg. (NMS). Woodbastwick, Nor- Norfolk, 2 males, 6.vii.−2.ix.1988, Foster & Procter leg. folk, 22 males, 2 females, viii.1932, Edwards leg. (PBLN). Seaford, East Sussex, 2 males, vi.1934 (BMNH). Woolcombe Meadow, Dorset, 2 males, (reared), Basden leg. [paratypes of Sciara nursei] 3.vii.1998, Chandler leg. (NMS, PPCM). (BMNH). St. Mary’s Nature Reserve, East Ham,

Greater London, 1 male, 15.vii.−21.vii.1987, C.W. wing membrane, and a large area of yellow laterally Plant leg. (PPCM). Tunbridge Wells, Kent, 1 male, on the mesonotum [as well as 2 paler lateral bands]. vii.1922, Nurse leg. [holotype of Sciara nursei] Moreover, Sciara flavomarginata has no lobe-like (BMNH). Turville Heath, Buckinghamshire, 4 males, swelling on the ventral inner surface of the gonostyle 20.vii.2002, Chandler leg. (PPCM). Wheatfen Broad, and no spine on its apex. By contrast with Sciara Norfolk, 1 male, 13.vii.1986, Withers leg. (PPWM). nursei, there are 7–8 spines that are arranged in a Woodwalton Fen, Cambridgeshire, 1 male, vi.1939, strict row in the upper third of the gonostyle and are of Coe leg. (BMNH). different lengths, distinctly increasing in length and strength from above to below. Ecological notes: common in broad-leaved, mixed and carr woodland; wetlands including fens, sedge beds and wet meadows. Reared from nest of meadow pipit. SCIARA RUFICAUDA MEIGEN, 1818 vi–x. Synonyms: boleti Winnertz, 1867; mamaevi Antonova, Taxonomic note: After studying the original descrip- 1978; vigilax Winnertz, 1867. tions by Freeman and Mohrig et al., the name ‘nursei Literature: Edwards (1925a): 536; Freeman (1983b): Freeman, 1983’ was placed as a synonym of Sciara fla- 19; Menzel (1998): 22. vomarginata Mohrig & Mamaev, 1982, in the recent revision of Palaearctic Sciaridae (Menzel & Mohrig, Material examined: Brockenhurst, Hampshire, 1 2000). This synonymy was based among other things male, 31.vii.1910, King leg. (BMNH). on the illustrations by Freeman, which were prepared Ecological notes: probably found in broad-leaved from the single preserved gonostyle on the hypopy- woodland. vii. gium of the holotype [compare Freeman (1983a): fig. 1; Freeman (1983b): fig. 11]. In these figures, Freeman showed 6 strong, subapical spines on the inner side of SCYTHROPOCHROA ENDERLEIN, 1911 the gonostyle, which are stated to be arranged in a row. However, re-examination of the holotype has SCYTHROPOCHROA QUERCICOLA (WINNERTZ, 1869) shown that the position of the spines is the result of Synonym: thomae var. nevadensis (Strobl, 1900). the way in which the gonostyle was mounted on the Literature: Freeman (1990): 52; Menzel (1998): 23. slide, and it is clear that the spines are not all in one plane. Furthermore, Freeman has drawn 2 very fine Material examined: Brampton Wood, Cambridgeshire, spinose bristles at the apex of the gonostyle that are 2 males, 3.iv.1975; 4 males, 3 females, 3.iv.−7.iv.1975 not actually developed in this form [there is only one (all reared), all J.H. Cole leg. (BMNH, PJCB, PPCM). short, strong ventral spine, which is placed on the dis- Ecological notes: Found in ancient broad-leaved wood- tinctly demarcated tip of the gonostyle]. This taxo- land on clay. Reared from rotten log. iv. nomic conclusion has widespread nomenclatural ramifications. All specimens with the character combination of ‘inner surface of gonostyle with subapical SCYTHROPOCHROA RADIALIS LENGERSDORF, 1926 lobe-like swelling bearing 4–6 subequal, similarly Literature: Freeman (1983b): 22; Chandler (1987): strong and relatively closely grouped spines; ventral 204; Freeman (1987): 197; Menzel (1998): 23; Chan- apex of the gonostyle with 1–2 distinctly shorter dler et al. (2002): 124. spines among the covering of hairs; anal area of the wing membrane [below CuA2] without or with only Material examined: Ashtead Common, Surrey, 2 very few macrotrichia; mesonotum dark brown to males, 6.vii.1994, Brock leg. (HRI). Beinn Eighe, black, at most paler dark yellow at base’ belong to a Highland Region, 1 male, 10.vi.1984, J.H. Cole leg. relatively small Sciara species with the name-complex (PJCB). Burnham Beeches, Buckinghamshire, 1 male, ‘marginata–nursei–ulrichi’. The oldest name ‘margin- 7.vi.−21.vi.1995; 1 male, 21.vi.−5.vii.1995; 1 male, ata Mohrig & Krivosheina, 1983’ is a junior primary 20.vii.−2.viii.1995; 3 males, 1 female, 8.vi.−18.vi.1999, homonym and is therefore not available. Conse- all Ismay leg. (HRI); 1 male, 2 females, 10.v.1998 quently, the name ‘Sciara nursei Freeman, 1983’ is the (reared), Chandler leg. (NMS, PPCM). California valid name for this species, and the later name ‘ulrichi Country Park, Berkshire, 1 male, 1 female, Menzel & Mohrig, 1998’ must be placed in synonymy 19.vi.1998, Chandler leg. (PPCM). Chippenham Fen, with it. Cambridgeshire, 1 male, 2.vii.1985, Chandler leg. Only the holotype of Sciara flavomarginata Mohrig (PPCM). Comrie, Perth & Kinross, 1 male, 20.vi.1996 & Mamaev from the Far East of Russia is known. It is (reared), MacGowan leg. (NMS). Conon Island, High- characterized by a distinctly longer body length, a land Region, 1 male, 18.vi.1976, Chandler leg. (NMS). denser covering of macrotrichia on the anal area of the Edford Wood, Somerset, 1 female, 2.viii.1985, Chan-

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 136 F. MENZEL ET AL. dler leg. (PPCM). Epping Forest, Essex, 1 male, Material examined: Brodick, Arran, North Ayrshire, 2 14.vi.−20.vi.1995, Dagley leg. (HRI); 5 males, males, 22.v.−25.v.1919, Edwards leg. [holotype and 26.vi.2000, J.H. Cole leg. (PJCB); 1 male, 28.vi.2000, paratype of Sciara scotica] (BMNH). Upper Quoich, Chandler leg. (PPCM). Grantown-on-Spey, Highland Aberdeenshire, 3 males, 21.vi.2000, Godfrey leg. (HRI, Region, 4 males, 16.vi.1989, Chandler leg. (NMS, PPCM). PPCM). How Hill, Norfolk, 1 male, 27.v.1989, Withers Ecological notes: found in broad-leaved and ancient leg. (PPWM). Knole Park, Kent, 4 males, 23.vi.1979, pine woodland. v–vi. Chandler leg. (NMS, PPCM). Loch Chon/Loch Ard, Stirling, 1 female, 22.vi.1999, Chandler leg. (PPCM). Loch Katrine, Stirling, 1 male, 15.vi.1992, J.H. Cole TRICHOSIA (MOUFFETINA) PULCHRICORNIS − leg. (PJCB). Moy, Tyrone, 1 male, 24.ix. 30.ix.1984, (EDWARDS, 1925) Boston leg. (PPCM). Nethy Bridge, Highland Region, Literature: Edwards (1925b): 224 [as Plastosciara]; 1 female, 6.vii.1922; 1 female, 12.viii.1922, all King Freeman (1983a): 169; Freeman (1983b): 22 [all as leg. (BMNH). Old Buckenham, Norfolk, 1 male, Mouffetina]; Menzel (1998): 23; Menzel & Mohrig 11.vi.1993, Chandler leg. (PPCM). Old Slade, Buck- (1997a): 30; Menzel & Mohrig (2000): 562. inghamshire, 2 females, 7.v.1972 (reared), Chandler leg. (NMS, PPCM). Peasmarsh, Hurtwood, Surrey, 9 Material examined: Ashley Hill, Berkshire, 4 males, males, 18 females, 17.ii.−27.iii.1975 (reared), Rich- iv.1999 (reared), Sims leg. (NMS, PPCM). Burnham ards leg. (BMNH). Sandy Warren, Bedfordshire, 1 Beeches, Buckinghamshire, 1 male, 7.vi.−21.vi.1995; 1 male, 10.vii.1996; 2 males; 4.vi.1998; 2 males, male, 1 female, 28.v.−14.vi.1996, all Ismay leg. (HRI, 24.vi.1998, all J.H. Cole leg. (PJCB). Santon Down- PPCM). Dupplin Loch, Perth & Kinross, 1 male, ham, Suffolk, 1 male, 5.viii.1970, no collector detail 15.vi.1992, Chandler leg. (PPCM). Epping Forest, (BMNH). Sedlescombe, East Sussex, 1 male, no date, Essex, 1 female, 24.v.1999, Chandler leg. (PPCM). Roper leg. (PPCM). The Loke, Norwich, Norfolk, 1 Greywell Fen, Hampshire, 1 female, 20.vi.2002, male, 17.vi.1987, Laurence leg. (PBLN). West Harling Chandler leg. (PPCM). Letchworth, Hertfordshire, 5 Common, Norfolk, 2 males, 20.iv.−12.v.1981 (reared), females, 12.vi.1925, Edwards leg. [lectotype and Withers leg. (PPWM). Whitmoor Vale, Hampshire, 1 paralectotypes of Plastosciara pulchricornis] (BMNH, male, 12.vii.1990, Merriﬁeld leg. (PPCM). Woolcombe ZFMK). Mark Ash Wood, New Forest, Hampshire, 1 Wood, Dorset, 1 male, 3.vii.1998, Chandler leg. female, 17.vi.1990, Chandler leg. (PPCM). Newburn, (NMS). Northumberland, 12 males, 3 females, 25.vi.1937; 2 males, 5 females, 8.vii.−10.vii.1937 (reared), all Ecological notes: common in all types of woodland; Topham leg. (BMNH). heathland; wetlands including fens and water meadows; grassland at woodland fringe; acid grassland and Ecological notes: found in ancient broad-leaved, mixed wet woodland; parkland. Reared from rotten elm, and carr woodland; in house. Reared from rotten wood. birch and beech wood; also found on dead pines. iv–ix. iv–vii.

TRICHODAPUS MOHRIG & MENZEL, 1997 TRICHOSIA (TRICHOSIA) ACROTRICHA TUOMIKOSKI, 1960 TRICHODAPUS RHENANUS (FRITZ, 1982) New to British list. Synonym: schoengeisingensis (Rudzinski, 1994). Material examined: New Close Wood, Brandon, War- New to British list. wickshire, 1 male, 28.viii.2003, J.E. Smith leg. (HRI). Material examined: Trefonen Marsh, Shropshire, 1 Ecological notes: found in broad-leaved woodland with male, 23.v.1994, J.H. Cole leg. (PJCB). pedunculate oak, birch and hazel. viii. Ecological notes: found by shaded stream in pasture grazed by cattle. v. TRICHOSIA (TRICHOSIA) BASDENI FREEMAN, 1983 Literature: Freeman (1983a): 161; Freeman (1983b): TRICHOSIA WINNERTZ, 1867 20 [all as Trichosia (Leptosciarella)]; Menzel (1998): TRICHOSIA (BAEOSCIARA) SCOTICA (EDWARDS, 1925) 23; Menzel & Mohrig (1997a): 29; Menzel & Mohrig (2000): 554. Literature: Edwards (1925a): 536 [as Sciara]; Free- man (1983a): 169 [as Trichosia]; Freeman (1983b): 20 Material examined: Black Park, Buckinghamshire, 1 [as Trichosia (Trichosia)]; Menzel & Mohrig (1997a): male, 27.vi.1941, Richards leg. (BMNH). Burnham 37; Menzel (1998): 23. Beeches, Buckinghamshire, 3 males, 23.v.−7.vi.1995,

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 BRITISH SCIARID FAUNA 137 all Ismay leg. (HRI, PPCM). Farley Down, Hampshire, Meadows, Newtown Linford, Leicestershire, 1 male, 1 male [paratype of Trichosia basdeni], 1 female, 24.viii.2003, Menzel leg. (DEI). Salisbury Plain Train- 20.v.1936, Edwards leg. (BMNH). Farnham Royal, ing Area, Warminster, Wiltshire, 1 male, 30.v.2003, Buckinghamshire, 2 males, 27.v.1934 [paratypes of Ismay & Schulten leg. (DEI). Trichosia basdeni, but dates as 21.v.−24.v and locality Ecological notes: found in ancient beech woodland; as Temple in Freeman (1983a)]; 1 male, 28.v.1934 (all wet meadow with mixed woodland with alder, birch, reared), all Basden leg. (BMNH); 6 males, 21.v.− hazel, spruce and fir; unimproved calcareous grass- 26.v.1934 (reared), Basden leg. (NMS). Frilford Heath, land. v, viii, x–xi. Oxfordshire, 1 male, 25.v.−15.vi.1991, Porter leg. (PPCM). Lundy Island, Devon, 1 female, 26.vii.− 27.vii.1972, Lane leg. (BMNH). Milham Ford School, Oxford, Oxfordshire, 1 male, 18.v.2001; 1 male, 1 TRICHOSIA (TRICHOSIA) CONFUSA MENZEL & female, 29.v.−4.vi.2002, all Webb leg. (PPCM). New- MOHRIG, 1997 market, Cambridgeshire, 1 female, 18.ix.1888, Verrall Literature: Edwards (1925a): 536 [as Sciara trochant- leg. (UMO). Stokeford Heath, Dorset, 1 male, erata, misidentification]; Freeman (1983b): 20; O’Con- 20.v.2001, Gibbs leg. (PDGB). Temple, Berkshire, 1 nor & Ashe (1991): 30; Laurence (1994): 118 [all as male, 26.v.1934 [holotype of Trichosia basdeni]; 1 Trichosia trochanterata, misidentification]; Menzel female, 28.v.1934; 1 female, 2.vi.1934; 1 female, (1998): 23; Chandler (2001): 201; Withers (2002): 115. 9.vi.1934 [paratypes of Trichosia basdeni but not same date as male as in Freeman (1983a)] (all reared), all Material examined: Badger Wood, Cambridgeshire, 1 Basden leg. (BMNH); 3 females, 26.v.−4.vi.1934 male, vi.1998, C.W. Plant leg. (NMS). Balmoral Forest, (reared), Basden leg. (NMS). The Loke, Norwich, Aberdeenshire, 1 male, 19.viii.−30.viii.1938, Coe leg. Norfolk, 1 male, 5.vii.1987; 2 males, 10.viii.1987; 1 (BMNH). Benacre Broad, Suffolk, 1 male, 28.v.1973, male, 5.v.1988; 3 males, 5.vi.−13.vi.1989; 1 male, Hutson leg. (NMS). Binfield, Berkshire, 1 male, 21.viii.−24.viii.1989; 1 male, 28.iv.−4.v.1990; 1 male, 6.v.1923, Dudgeon leg. (BMNH). Black Wood, Ran- 2.v.−6.v.1992; 14 males, 15.v.−18.v.1992; 9 males, 15.v.− noch, Perth & Kinross, 1 male, 11.vii.1988, Chandler 25.v.1992; 1 male, 18.v.−23.v.1992; 8 males, 21.v.− leg. (NMS). Bolderwood, Hampshire, 1 male, 23.v.1993, all Laurence leg. (DEI, PBLN). The Oaks, 29.vii.1953, Colyer leg. (BMNH). Boundless Copse, Cambridgeshire, 1 male, ix.1998, C.W. Plant leg. Surrey, 1 male, 26.vii.2002, Chandler leg. (PPCM). (PPCM). Tunbridge Wells, Kent, 1 female, 3.vi.1986, Brampton Wood, Cambridgeshire, 1 male, 9.v.1987, Verrall leg. (UMO). Wellesbourne, Warwickshire, 1 J.H. Cole leg. (PJCB). Brodick, Arran, North Ayrshire, male, 24.v.−27.vi.2001, J.E. Smith leg. (HRI). No local- 1 male, 22.v.−25.v.1919, Edwards leg. (BMNH). Bro- ity details, England, 1 male, 3.v.1924 (reared), Hamm mley, Greater London, 1 male, 24.vi.1964, Chandler leg. (UMO). leg. (PPCM). Buckingham Palace Garden, Greater London, 1 male, 15.viii.−5.ix.1995, Chandler leg. Ecological notes: found in broad-leaved and mixed (PPCM). Burnham Beeches, Buckinghamshire, 1 woodland; school grounds with old poplars; calcareous male, 8.vii.−22.vii.1996, Ismay leg. (PPCM); 1 male, grassland; sandy heathland; garden. Reared from 20.iv.1998, Chandler leg. (PPCM). California Country nests of blackbird, thrush, decoy nest. v–ix Park, Berkshire, 2 males, 15.vi.1996; 1 male, Taxonomic note: The type series of Trichosia basdeni 22.v.1999, all Chandler leg. (PPCM). Chippenham according to Freeman (1983a: 163) ought to be 4 males Fen, Cambridgeshire, 1 male, 6.v.1934, J.H. Cole and 3 females. However, he incorrectly transcribed leg. (PJCB). Chobham Common, Surrey, 1 male, some of the data. The specimens from Farnham Royal, 5.vii.1984, Chandler leg. (PPCM). Coldborough Park, Buckinghamshire, 2 males, 27.v.1934 (paratypes) Herefordshire, 1 male, 31.v.1972, Chandler leg. were given as Temple, Berkshire, 2 males, 21.v.−24.v (PPCM). Cothill, Oxfordshire, 1 male, 8.vi.1975, and the specimens from Temple, Berkshire, 1 male, Chandler leg. (PPCM). Craigmore Wood, Aberfoyle, 26.v.1934 (holotype); 1 female, 28.v.1934; 1 female, Stirling, 1 male, 21.vi.1999, Chandler leg. (NMS). Cul- 2.vi.1934; 1 female, 9.vi.1934 (all paratypes) were bin Forest, Highland Region, 1 male, 27.v.2002, Chan- given as 3 females, 26.v.1934; 2 males, 21.v.−24.v.1934. dler leg. (PPCM). Dinton Pastures Country Park, Berkshire, 1 male, 17.vi.1995, Chandler leg. (PPCM). Drumguish, Highland Region, 1 male, 17.viii.1986, TRICHOSIA (TRICHOSIA) BOREALIS (FREY, 1942) Chandler leg. (NMS). Easter Fearn, Highland Region, 3 males, 11.vi.1984, Chandler leg. (PPWM). Epping New to British list. Thicks, Epping Forest, Essex, 1 male, 12.vii.− Material examined: Burnham Beeches, Buckingham- 18.vii.1999, Dagley leg. (HRI); 1 male, 26.vi.2000, shire, 1 male, 4.x.−15.xi.1995, Ismay leg. (PPCM). Lea J.H. Cole leg. (PJCB). Fish House Moss, Cumbria, 1

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 138 F. MENZEL ET AL. male, 16.vi.1999, Chandler leg. (NMS). Folly Farm, Ecological notes: very common in all types of wood- Avon, 3 males, 25.vii.2001, Gibbs leg. (PDGB). Glen- land; fen; water meadows by river; gravel pits; old dalough, Wicklow, 1 male, 10.vi.1985, Chandler hedges; calcareous grassland and scrub; dunes; gar- leg. (PPCM). Glentanar, Aberdeenshire, 2 males, dens; in house. Reared from rotten wood. iv–x. 13.vii.1977, Chandler leg. (NMS). Grantown-on-Spey, Highland Region, 1 male, 15.vi.1984, Chandler leg. (PPWM). Grondra Great Wood, Monmouthshire, 1 TRICHOSIA FLAVICOXA TUOMIKOSKI, 1960 male, 9.vii.1986, Chandler leg. (PPCM). Hartslock, New to British and Irish list. Oxfordshire, 1 male, 10.v.−19.v.2000; 1 male, 14.v.2000, all Raper leg. (HRI, PDGB). High Dam, Material examined: Burnham Beeches, Bucking- − − Cumbria, 1 male, 15.vi.1999, Chandler leg. (NMS). hamshire, 3 males, 23.v. 7.vi.1995; 1 male, 27.x. − − Kinrive Wood, Highland Region, 1 male, 15.vi.1976, 7.xi.1995; 4 males, 28.v. 1.vii.1996; 2 males, 17.v. − Chandler leg. (NMS). Knebworth, Hertfordshire, 1 25.v.1999; 4 males, 1 female, 1.vi. 8.vi.1999, all Ismay male, vi.1922, Edwards leg. (BMNH). Knocksink leg. (HRI, PPCM). Frithsden Beeches, Hertfordshire, Wood, Wicklow, 1 male, 17.vi.1985, Chandler leg. 1 male, 6.vi.1997, Chandler leg. (PPCM). Glasdrum, (PPWM). Knole Park, Kent, 1 male, 16.v.1971; 1 male, Argyll & Bute, 1 male, 12.vi.1976, Stubbs leg. 23.vi.1979, all Chandler leg. (NMS). Leckford Estate, (PPCM). Glen Coiltie, Highland Region, 1 male, Hampshire, 1 male, 8.v.1976, Chandler leg. (PPCM). 28.v.2002, Chandler leg. (PPCM). Loch Meiklie, High- Letchworth, Hertfordshire, 1 male, vii.1917; 2 males, land Region, 1 male, 13.vi.1984, Chandler leg. (NMS). iv.1918; 2 males, v.1924; 1 male, v.1925, all Edwards Moccas Park, Herefordshire, 1 male, 11.vi.2002, God- leg. (BMNH). Loch Garten, Highland Region, 1 male, frey leg. (PPCM). River Flesk, Killarney, Kerry, 1 15.vi.1984, Chandler leg. (NMS). Loch Katrine Dam, male, 8.vii.1989, Ashe leg. (NMID). Rotherfield Park, Stirling, 1 male, 21.vi.1999, Chandler leg. (PPCM). Hampshire, 1 male, 3.vii.2002, Chandler leg. (PPCM). Loch Tummel, Perth & Kinross, 1 male, 30.vi.1937, Selborne Common, Hampshire, 1 male, 28.v.1988, Collin leg. (BMNH). Lough Ballynafid, Westmeath, Chandler leg. (PPCM). The Lythes, Selborne, Hamp- 1 male, 16.vi.1985, Chandler leg. (PPWM). Monk’s shire, 1 male, 28.v.1988, Chandler leg. (PPCM). Wind- Wood, Cambridgeshire, 1 male, 29.v.1972; 1 male, sor Forest, Berkshire, 1 male, 15.v.1991, Chandler leg. 12.vi.1972; 1 male, 18.vi.1978, all J.H. Cole leg. (PPCM). (PJCB). New Forest, Hampshire, 1 male, 28.v.1948, Ecological notes: found in alder and other broad- Colyer leg. (UMO). Oxford, Oxfordshire, 1 male, leaved woodland; ancient oak and beech forest; park- 13.v.1925; 1 male, 30.v.1925, all Hamm leg. (UMO). land. v–vii, x–xi. Oversley Wood, Warwickshire, 1 male, x.1969 (reared), Evans leg. (BMNH). Pett’s Wood, Greater London, 1 male, 25.v.1983, Freeman leg. (BMNH). Pingle Cut- TRICHOSIA (TRICHOSIA) GLABRA (MEIGEN, 1830) ting, Warboys, Cambridgeshire, 1 male, 26.v.1985, Literature: Edwards (1925a): 537 [as Sciara]; Free- J.H. Cole leg. (PJCB). Pitmaduthy, Highland Region, 1 man (1983b): 20; Freeman (1990): 51 [all as Trichosia male, 19.vii.1991, Stubbs leg. (PPCM). Purn Hill, (Leptosciarella)]; Menzel (1998): 23. Avon, 1 male, 24.v.2001, Gibbs leg. (PDGB). Redfield, Bristol, Avon, 1 male, 2.v.2001, Gibbs leg. (PDGB). Material examined: Badger Wood, Cambridgeshire, 1 Ridley Wood, Hampshire, 1 male, 4.v.1964, Colyer leg. male, 1.v.−31.v.1998, C.W. Plant leg. (PPCM). Burn- (BMNH). Rothiemurchus, Highland Region, 1 male, ham Beeches, Buckinghamshire, 2 males, 28.iv.− 18.viii.1986, Chandler leg. (NMS). Savernake Forest, 10.v.1995; 1 male, 7.vi.−21.vi.1995; 1 male, 21.vi.− Wiltshire, 1 male, 3.v.1993, Chandler leg. (PPCM). 5.vii.1995; 1 male, 17.v.−25.v.1999; 1 male, 1.vi.− Scadbury Park, Greater London, 2 males, 18.vi.1983, 8.vi.1999, all Ismay leg. (HRI, PPCM). Bwlch y Ddeu- Chandler leg. (NMS, PPCM). Skinner’s Pastures, faen, Conwy, 1 male, 5.vii.1987, J.H. Cole leg. (PJCB). Cumbria, 1 male, 17.vi.1999, Chandler leg. (NMS). Chigwell Row Wood, Chigwell Row, Essex, 2 males, Teesdale, Durham, 1 male, 22.vi.1924, Edwards leg. 23.v.−27.v.2002; 10 males, 2 females, 4.vi.2002; 1 (BMNH). Terenure, Dublin City, Dublin, 1 male, female, 11.vi.2002; 2 males, 18.vi.2002; 1 male, 23.v.1990, Ashe leg. (NMID). The Loke, Norwich, Nor- 20.vi.2002; 5 males, 28.vi.2002, all Ismay & Schulten folk, 1 male, 5.vi.−13.vi.1989; 1 male, 28.v.1992, all leg. (DEI, UMO). Dulicht, Highland Region, 1 female, Laurence leg. (PBLN). The Spittles, Dorset, 1 male, 19.vii.1997, Chandler leg. (PPCM). Epping Forest, 2.vii.1998, Chandler leg. (NMS). West Walk, Hamp- Essex, 1 female, 24.v.1999, Chandler leg. (PPCM). shire, 1 male, 6.vi.1998, Chandler leg. (NMS). Whit- Hartslock, Oxfordshire, 1 male, 14.v.2000, Raper leg. barrow Scar, Cumbria, 1 male, 13.vi.1999, J.H. Cole (PDGB). Ipswich, Suffolk, 1 male, 10.v.1898 (reared), leg. (PJCB). Windsor Forest, Berkshire, 1 male, Morley leg. (BMNH). Letchworth, Hertfordshire, 2 7.vi.1977, Chandler leg. (NMS). males, 1 female, vi.1917; 1 female, 18.vi.1934, all

Edwards leg. (BMNH). Loch Assynt, Highland Region, Material examined: Aldbury Common, Hertfordshire, 1 male, 5.vi.1911, Yerbury leg. (BMNH). Monk’s 1 male, 17.vi.1997, Chandler leg. (PPCM). Alder- Soham, Suffolk, 1 male, 8.v.1914, Morley leg. (BMNH). moors, Berkshire, 1 male, 14.v.1994, Chandler leg. Monk’s Wood, Cambridgeshire, 1 male, 5.vi.1972; 1 (PPCM). Arden Hall, Cheshire, 1 male, 1 female, male, 15.vi.1972; 1 male, 31.vii.1972, all J.H. Cole leg. 2.vii.1919 (reared), Britten leg. (BMNH). Ashley Hill, (PJCB). Newmarket, Cambridgeshire, 1 female, 5.vi. Berkshire, 1 male, iv.1999 (reared), Sims leg. (NMS). 1885, Verrall leg. (UMO). Richmond Park, Greater Aviemore, River Spey, Highland Region, 1 male, London, 1 female, 6.vi.1992, Chandler leg. (PPCM). 9.vii.−13.vii.1982, Withers leg. (PPWM). Ballyhigh- The Loke, Norwich, Norfolk, 2 males, 15.v.−18.v.1992; land, Wexford, 1 male, 14.vi.1987, J.P. & M.A. O’Con- 4 males, 15.v.−25.v.1992; 1 male, 18.v.−22.v.1992, all nor leg. (NMID). Barcaldine Forest, Argyll & Bute, 1 Laurence leg. (PBLN). Wicken Fen, Cambridgeshire, 2 male, 1 female, 11.vii.1974, Chandler leg. (NMS, males, 14.v.1992, J.H. Cole leg. (PJCB). Wychwood, PPCM). Barmby Moor, East Riding of Yorkshire, 1 Oxfordshire, 1 male, 20.v.−16.vi.1989, Porter leg. male, 25.v.1998, Chandler leg. (NMS). Barnsley, (PPCM). Wytham Wood, Oxfordshire, 1 male, South Yorkshire, 27 males, 37 females, v.1924 19.xii.1967 (reared), Paviour-Smith & Elbourn leg. (reared), Bayford leg. (BMNH). Barton Mills, Suffolk, (BMNH). 1 female, 29.v.1909, Verrall leg. (UMO); 1 male, 10.v.1931; 1 male, 30.v.1935, all Collin leg. (UMO). Ecological notes: common in broad-leaved woodland; Bevills Wood, Abbot’s Ripton, Cambridgeshire, 1 ancient woodland with oak, hazel and bramble; heath- male, 31.v.1976, J.H. Cole leg. (PJCB). Birming- land (heather) with oak and holly; moorland ﬂushes; ham, West Midlands, 3 males, iv.1896, Bradley fen meadows and ditches; ancient parkland; garden. leg. (UMO). Bix Bottom, Oxfordshire, 1 male, Reared from larvae in galleries of the beetle Ischno- 9.v.1998, Chandler leg. (NMS). Bonhill, West mera caerulea. iv–vii. Dunbartonshire, 1 female, 22.vi.1905; 1 male, 18.v.1906; 1 male, 20.viii.1907; 1 male, 29.viii.1907; 1 TRICHOSIA (TRICHOSIA) JENKINSONI FREEMAN, 1987 female, 26.vi.1909, all Malloch leg. (NMS). Boundless Copse, Surrey, 1 male, 26.vii.2002, Chandler leg. Literature: Freeman (1987): 197 [as Trichosia (Leptos- (PPCM). Braelangwell, Highland Region, 1 male, ciarella)]; Menzel & Mohrig (1997a): 27; Menzel 12.vi.1984, Chandler leg. (PPWM). Brown’s Folly, (1998): 23; Menzel & Mohrig (2000): 558. Avon, 1 male, 24.viii.2000, Gibbs leg. (PDGB). Buck- Material examined: Cambridge, Cambridgeshire, 1 ingham Thick Copse, Northamptonshire, 1 female, male, 10.vii.1923, Jenkinson leg. [holotype of Tricho- 9.vi.1972, Warne leg. (NMS). Burnham Beeches, sia jenkinsoni] (BMNH). Buckinghamshire, 1 male, 17.vi.1967; 1 female, 20.iv.1968; 1 female, 13.v.1990; 1 male, 1 female, Ecological notes: possibly from garden of F. Jenkinson 10.v.−11.v.1998 (reared), all Chandler leg. (PPCM); 1 or in the University Parks. vii. male, 2 females, 23.v.−7.vi.1995; 2 females, 1.vi.− 8.vi.1999, all Ismay leg. (HRI). Butley Thicks, Suf- TRICHOSIA (TRICHOSIA) MORIO (FABRICIUS, 1794) folk, 1 male, 1 female, 1.vi.1952, d’Assis-Fonseca leg. (NMS). Cairngorm, Highland Region, 1 female, Synonyms: caudata (Walker, 1848); dziedzickii (Grze- 5.viii.1972, Chandler leg. (PPCM). California Country gorzek, 1884); edwardsi (Lengersdorf, 1930); longiven- Park, Berkshire, 2 males, 2.vi.1996, Chandler leg. tris (Zetterstedt, 1851); mikii (Grzegorzek, 1884); (PPCM). Callander, Stirling, 1 female, 27.v.1916, sznablii (Grzegorzek, 1884). A.E.J. Carter leg. (UMO). Canonteign Falls, Devon, 1 Literature: ? Haliday (1833): 148 [as Sciara]; Walker male, 22.vi.2002, J.E. Smith leg. (HRI). Charlbury, (1848): ? 104 and 106; Walker (1856): ? 52 and 53 [all Oxfordshire, 1 male, 4.vii.1936, Hobby leg. (UMO). as ? Sciara morio and Sciara caudata]; Verrall (1912): Chippenham Fen, Cambridgeshire, 1 male, 1 female, 20; Edwards (1925a): 535 [all as Sciara longiventris]; 8.viii.1940, Collin leg. (UMO). Cliveden, Bucking- Edwards (1925c): 228 [as Sciara caudata]; Edwards hamshire, 1 female, 3.vii.1984, Chandler leg. (NMS). (1926b): 142 [as Sciara longiventris]; Lengersdorf Coed y Rhygen, Gwynedd, 1 male, 14.vii.1976, J.H. (1930): 29 [as Lycoria trochanterata] and 67 [as Lyco- Cole leg. (PJCB). Coombe Dingle, Avon, 1 male, ria edwardsi]; Kidd (1953): 137 [as Sciara (Sciara) 26.vi.1949, d’Assis-Fonseca leg. (NMS). Cothill, caudata]; Brindle (1963): 83 [as Sciara longiventris]; Oxfordshire, 1 male, 28.vii.1989, Chandler leg. Perry & Stubbs (1978): 71; Freeman (1983b): 20; Lau- (NMS). Craigston, Barra, Western Isles, 1 female, rence (1997b): 228 [all as Trichosia caudata]; Menzel 23.vi.1980, A.R. Waterston leg. (NMS). Crickhowell, & Mohrig (1997a): 19; Menzel (1998): 23; Coldwell Powys, 1 male, v.1937, Edwards leg. (BMNH). Culbin (1999): 25 [as Trichosia caudata]; Menzel & Mohrig Forest, Highland Region, 1 male, 1 female, 27.v.2002, (2000): 558. Chandler leg. (PPCM). Culbin Sands, Highland

Region, 1 female, 5.vii.−7.vii.1936, Coe leg. (BMNH). 26.vi.1909, Adams leg. [lectotype of Lycoria edwardsi] Denny Wood, New Forest, Hampshire, 1 male, 22.v.− (ZFMK). Nightingale Wood, Hampshire, 1 male, 26.v.1987, Townsend leg. (BMNH). Denton Covert, 18.ix.1975, Chandler leg. (PPCM). Old Slade, Buck- Holme Fen, Cambridgeshire, 1 male, 4.ix.1977, J.H. inghamshire, 1 male, 2 females, 9.iv.1972 (reared), Cole leg. (PJCB). Derwent Valley, Tyne & Wear, 1 Chandler leg. (NMS, PPCM). Oversley Wood, War- male, 14.vii.−13.viii.1990, Jessop leg. (PBLN). Dickle- wickshire, 4 males, x.1969 (reared), Evans leg. burgh, Norfolk, 1 male, 18.vii.−5.viii.1989, Withers (BMNH). Pass of Killiecrankie, Perth & Kinross, 1 leg. (PPWM). Dinton Pastures Country Park, Berk- male, 7.vii.1975, Chandler leg. (NMS). Pot Riding shire, 1 male, 5.ix.1993, Chandler leg. (PPCM). Dun- Wood, Doncaster, South Yorkshire, 1 male, stall Wood, Shoreham, Kent, 3 males, 7.vi.1968, 2.viii.1992, Godfrey leg. (PPCM). RAF Barnham, Chandler leg. (NMS, PPCM). Eday, Orkney, 1 male, Suffolk, 1 male, 28.vi.1995, J.H. Cole leg. (PJCB). 3.vi.−5.vi.1988, Laurence leg. (PBLN). Edinburgh, Rosemarkie, Highland Region, 1 female, 21.ix.1989, City of Edinburgh, 1 female, 7.vi.1959, A.R. Water- Chandler leg. (PPCM). Rowardennan Forest, Lochan ston leg. (NMS). Ellerton Wood, Devon, 1 female, Maoil Dhuinne, Stirling, 1 male, 17.vi.1992, J.H. Cole 2.vi.2000, Chandler leg. (PPCM). Epping Forest, leg. (PJCB). Sand, Shetland, 1 male, 5.vii.−6.vii.1992, Essex, 1 male, 27.x.1997, Chandler leg. (PPCM); 1 Laurence leg. (PBLN). Savernake Forest, Wiltshire, male, 17.v.−23.v.1999, Dagley leg. (HRI). Farnham 1 male, 28.vi.1971, Chandler leg. (NMS). Scadbury Royal, Buckinghamshire, 1 female, 27.viii.1972, Park, Greater London, 1 male, 1 female, 18.vi.1983, Chandler leg. (PPCM). Farnley, Leeds, West York- Chandler leg. (NMS). Searles Farm, Reading, Berk- shire, 1 female, vii.1918, Cheetham leg. (BMNH). shire, 1 male, 26.vi.2003, Chandler leg. (HRI). Sheep Folly Farm, Avon, 1 male, 25.vii.2001, Gibbs leg. Leas, West Horsley, Surrey, 1 male, 9.vi.1968, Chan- (PDGB). Freshfield, Merseyside, 2 males, 19.ix.1970, dler leg. (PPCM). Sutton, Norfolk, 2 males, 25.v.1895; Hutson leg. (BMNH). Gamlingay Wood, Cam- 1 female, 28.vi.1895; 1 female, 2.vii.1895, all Verrall bridgeshire, 1 female, 13.vi.2001, J.H. Cole leg. leg. (UMO). Sutton Coldfield, West Midlands, 1 male, (PJCB). Glendalough, Wicklow, 1 male, 20.ix.1968, 26.v.1895, Bradley leg. (BMNH). The Loke, Norwich, Chandler leg. (PPCM). Glentanar, Aberdeenshire, 1 Norfolk, 2 males, 5.v.1988; 2 males, 15.vi.−19.vi.1988; male, 23.vi.1977, Chandler leg. (PPCM). Grange, 1 male, 5.vi.−13.vi.1989; 2 males, 13.vii.−19.vii.1989; Cumbria, 1 male, 27.vii.1923, Edwards leg. (BMNH). 1 male, 27.vii.1992, all Laurence leg. (PBLN). Tresil- Grantown-on-Spey, Highland Region, 1 male, lian, Cornwall, 1 male, 7.vii.1983, Chandler leg. 31.viii.1984, Chandler leg. (NMS). Hampstead Heath, (PPWM). Vidlin, Shetland, 2 males, 16.vi.−21.vi.1993, Greater London, 1 male, 31.v.1973, Cranston leg. Laurence leg. (PBLN). Wadhurst Place, East Sussex, (BMNH). Hartslock, Oxfordshire, 1 male, 6.v.2000, 1 male, 11.v.1974, Stubbs leg. (PPCM). Walberswick, Gibbs leg. (PDGB). Holme Fen, Cambridgeshire, 1 Suffolk, 1 male, 26.v.−27.v.1973, Hutson leg. (BMNH). male, 12.viii.1977; 1 male, 25.v.1978, all J.H. Cole leg. Warboys Wood, Cambridgeshire, 1 female, 21.v.1967, (PJCB). Kingston Wood, Dorset, 1 male, 19.ix.1975, J.H. Cole leg. (PJCB). Wicken Fen, Cambridgeshire, 1 Chandler leg. (PPCM). Knole Park, Kent, 1 female, female, 14.v.1992, J.H. Cole leg. (PJCB). Wildmoor 1.x.1967; 1 male, 12.ix.1971, all Chandler leg. (NMS, Heath, Berkshire, 1 male, 15.v.1999, Chandler leg. PPCM). Leckford Estate, Hampshire, 1 male, (NMS). Windsor Forest, Berkshire, 1 male, 13.v.1967, 7.vi.1971; 1 male, 4.vi.1972, all Chandler leg. Stubbs leg. (PPCM); 1 male, 30.vi.1971; 1 male, (PPCM). Letchworth, Hertfordshire, 1 female, 27.v.1985, all Chandler leg. (NMS, PPCM). Wisley vi.1927, Edwards leg. (BMNH). Little Paxton Pits, Common, Surrey, 1 male, 15.vi.1968, Chandler leg. Cambridgeshire, 1 male, 18.v.1993, J.H. Cole leg. (NMS). Woodwalton Fen, Cambridgeshire, 1 female, (PJCB). Llandecwyn, Gwynedd, 1 male, 16.vii.1976, vii.1939, Coe leg. (BMNH); 1 male, 2 females, Chandler leg. (NMS). Loch Meiklie, Highland Region, 24.v.1940, Edwards leg. (BMNH). Wytham Woods, 1 male, 13.vi.1984, Chandler leg. (NMS). Loch na Wytham, Oxfordshire, 1 male, 1 female, 21.viii.2002, Gainmhich, Lewis, Western Isles, 1 male, 10.v.1962, Menzel & J.E. Smith leg. (DEI, HRI). No locality Broomfield & Vardy leg. (BMNH). Loch Rannoch, details, England, 1 female, no date, Walker leg. [lec- Perth & Kinross, 2 females, vi.1936, Edwards leg. totype of Sciara caudata] (BMNH). (BMNH). Lopham Fen, Norfolk, 1 male, 16.iv.1982; 1 male, 20.v.1990, all Withers leg. (PPWM). Milham Ecological notes: very common in all types of wood- Ford School, Oxford, Oxfordshire, 1 male, v.2001, land; heathland; wetlands including fen and reed Webb leg. (HRI). Mound Alderwood, Highland beds, scrub and Sphagnum bog; bog and lakeshore and Region, 1 male, 14.vi.1976, Chandler leg. (NMS). water meadows by river; gravel pits; calcareous grass- Monk Wood, Avon, 2 males, 24.v.2000, Gibbs leg. land; school grounds with old poplars; gardens; park- (PDGB). Murrough, Wicklow, 1 male, 8.v.1989, With- land; old hedges. Reared from pupa in rotten wood, ers leg. (PPWM). New Forest, Hampshire, 1 male, rotten beech and birch bark. v–x.

TRICHOSIA (TRICHOSIA) SPLENDENS WINNERTZ, 1867 Devon, 1 female, 8.vii.1992, Chandler leg. (NMS). Synonyms: maxima Strobl, 1880; winnertzi Nowicki, Glasdrum, Argyll & Bute, 2 males, 7.vii.1974, 1868. Chandler leg. (NMS). Glen Doll, Angus, 2 females, 26.iv.1998 (reared), Bland leg. (NMS). Glen Lochay, Literature: Edwards (1925a): 533 [as Trichosia hirti- Stirling, 1 male, 3.vi.−18.vi.1932, Edwards leg. pennis, misidentiﬁcation]; Freeman (1983b): 20; Men- (BMNH). Glencoe, Highland Region, 1 male, zel & Mohrig (1997a): 11; Menzel (1998): 23; Withers 13.vi.1976, Chandler leg. (NMS). Glendalough, Wick- (2002): 115. low, 1 male, 8.vii.1969; 1 male, 1 female, 10.vii.1971, Material examined: Alice Holt Forest, Hampshire, 1 all Chandler leg. (PPCM). Grange, Cumbria, 1 male, male, 23.viii.1999, Chandler leg. (PPCM). Apes Down, 3.vii.−7.viii.1923, Edwards leg. (BMNH). Holker Isle of Wight, 1 male, 31.vii.1980, Stubbs leg. (PPCM). Moss, Cumbria, 1 male, 1 female, 11.vii.−13.vii.1923, Arniston, Midlothian, 1 male, 17.vii.1906, A.E.J. Edwards leg. (BMNH). Holly Bank, Shropshire, 1 Carter leg. (UMO). Aviemore, Highland Region, 1 male, 17.vi.1992, Gibbs leg. (PDGB). Humphrey male, 9.vi.1913, Yerbury leg. (BMNH). Beaulieu River, Head, Cumbria, 1 female, 1.vii.−8.vii.1923, Edwards New Forest, Hampshire, 1 male, 22.v.1936, Edwards leg. (BMNH). Inholm’s Copse, West Sussex, 1 male, leg. (BMNH). Bedford Purlieus, Cambridgeshire, 2 26.vii.1974, Chandler leg. (PPCM). Invermoriston, males, 23.vi.1985, J.H. Cole leg. (PJCB). Bettws, Highland Region, 1 female, 20.viii.1977, Dear Conwy, 1 male, 16.vi.1887, Verrall leg. (UMO). Bix leg. (BMNH). Knebworth, Hertfordshire, 1 male, Bottom, Oxfordshire, 1 female, 6.vii.2002, Chandler 10.vi.1927, Edwards leg. (BMNH). Lisdoonvarna, leg. (PPCM). Black Wood, Rannoch, Perth & Kinross, 1 Clare, 1 female, 8.vii.1981, J.P. O’Connor leg. (NMID). male, 11.viii.1988, Chandler leg. (PPCM). Blean Wood, Loch Davan, Aberdeenshire, 1 female, 10.vii.1977, Kent, 1 female, 31.v.1965, Parmenter leg. (BMNH). Chandler leg. (NMS). Loch na Ba Ruaidhe, Highland Bohuntine, Highland Region, 1 female, 20.vii.1997, Region, 1 female, 13.vi.1984, Chandler leg. (PPCM). Chandler leg. (PPCM). Bonhill, West Dunbartonshire, Loch Tummel, Perth & Kinross, 2 males, 20.vi.1937; 2 1 male, 22.vi.1907; 1 male, 7.viii.1908; 1 male, males, 30.vi.1937, all Collin leg. (BMNH). Logie, 5.vi.1909, all Malloch leg. (NMS). Boundless Copse, Moray, 1 male, viii.1923, Jenkinson leg. (BMNH). Surrey, 1 male, 1 female, 26.vii.2002, Chandler leg. Lough Ballynaﬁd, Westmeath, 1 male, 16.vi.1985, (PPCM). Brighstone Forest, Isle of Wight, 1 male, 1 Chandler leg. (PPWM). Mark Ash Wood, New Forest, female, 1.vii.1980, Chandler leg. (PPCM). Burnham Hampshire, 1 female, viii.1936, Edwards leg. Beeches, Buckinghamshire, 1 male, 1 female, 1.vii.− (BMNH); 1 male, 4.vi.1939, Collin leg. (UMO); 1 8.vii.1996, Ismay leg. (HRI). Butley Thicks, Suffolk, 1 female, 7.vi.1989; 1 female, 17.vi.1990, all Chandler female, 1.vi.1952, d’Assis-Fonseca leg. (NMS). Califor- leg. (NMS, PPCM). Matley Bog, New Forest, Hamp- nia Country Park, Berkshire, 1 male, 15.ix.1996, shire, 1 male, 19.viii.1952, d’Assis-Fonseca leg. (NMS). Chandler leg. (PPCM). Chippenham Fen, Cam- Maulden Wood, Bedfordshire, 1 male, 5.vi.1980, J.H. bridgeshire, 1 female, 4.viii.1940, Collin leg. (UMO). Cole leg. (PJCB). Miller’s Dale, Derbyshire, 1 female, Cod Wood, Devon, 1 male, 24.v.1978, Chandler leg. 20.vi.1888, Verrall leg. (UMO). Monk’s Wood, Cam- (PPCM). Coed Caeddafydd, Gwynedd, 1 female, bridgeshire, 1 male, 28.vi.1971, Service leg. (PJCB). 8.vii.1987, Chandler leg. (PPCM). Comrie, Perth Nesbitt Dene, Durham, 1 female, 5.vii.1995, Chandler & Kinross, 1 female, 12.vii.1907, A.E.J. Carter leg. (PPCM). Nethy Bridge, Highland Region, 1 leg. (UMO). Crowborough, East Sussex, 1 male, female, 17.vii.1913, King leg. (BMNH). New Forest, 30.vii.1912; 2 females, 8.ix.1919, all Jenkinson leg. Hampshire, 1 male, vii.1905, Sharp leg. (BMNH); (BMNH). Cwm Bychan, Gwynedd, 1 male, 12.vii.1976, 1 male, 10.vii.1907, King leg. (BMNH); 1 male, Chandler leg. (NMS). Dalnaglar Castle, Perth & Kin- 28.v.1948, Colyer leg. (BMNH); 1 male, 23.vi.1965; 1 ross, 1 male, 7 females, 18.v.1998 (reared), Bland leg. male, 10.vi.1968, all Parmenter leg. (BMNH). Nichol’s (NMS). Derwentwater, Cumbria, 1 female, 14.vi.1977, Moss, Cumbria, 1 female, 19.vi.1999, Perry leg. Chandler leg. (PPCM). Dinton Pastures Country (PPCM). Peppercombe, Devon, 1 male, 14.vi.1989, J.H. Park, Berkshire, 1 female, 12.vi.1994, Chandler leg. Cole leg. (PJCB). Pitmaduthy Moss, Highland Region, (PPCM). Dolgellau, Gwynedd, 1 female, 13.vi.1887, 1 male, 16.vi.1976, Chandler leg. (NMS). Pond Wood, Verrall leg. (UMO). Epping Forest, Essex, 1 male, Chislehurst, Greater London, 1 female, 24.vi.1964, 18.viii.1998; 1 female, 26.v.1999, all Chandler leg. Chandler leg. (PPCM). Ringstead, Dorset, 1 male, (PPCM); 1 male, vi.1999; 3 males, 1 female, 5.vii.− viii.1939, Edwards leg. (BMNH). River East Lyn, 11.vii.1999, all Dagley leg. (HRI). Farley Downs, Rockford, Devon, 1 male, 16.vi.1989, J.H. Cole leg. Hampshire, 2 males, 20.v.1936, Edwards leg. (BMNH). (PJCB). Slapston Holm Wood, North Yorkshire, 1 Felbrigg Great Wood, Norfolk, 1 female, 10.vii.1993, male, 27.vi.1981, Chandler leg. (PPCM). Stoke Wood, Chandler leg. (PPCM). Ffrith, Flintshire, 1 male, Herefordshire, 1 male, 1 female, 11.vii.1909, Verrall or 7.vi.−9.vi.1919, Edwards leg. (BMNH). Fingle Bridge, Collin leg. (UMO). Stokenchurch, Oxfordshire, 1

© 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 146, 1–147 142 F. MENZEL ET AL. female, 13.vii.1907, Verrall leg. (UMO). Stronord, North Ayrshire, 1 male, 2.vi.−4.vi.1919, Edwards leg. Dumfries & Galloway, 1 male, 18.vii.1979, J.H. Cole (BMNH). Hengistbury Head, Hampshire, 1 male, leg. (PJCB). The Spittles, Dorset, 1 male, 2.vii.1998, 13.vi.1938, Edwards leg. (BMNH). Lea Meadows, Chandler leg. (NMS). Thompson Common, Norfolk, 1 Newtown Linford, Leicestershire, 1 male, 24.viii.2003, male, 17.vi.1991, Withers leg. (PPWM). Vicars Bridge, Menzel leg. (DEI). Malham Tarn, North Yorkshire, 1 Perth & Kinross, 1 male, 14.vi.1992, J.H. Cole leg. male, 29.vi.−2.vii.1981, Disney leg. (PPWM). Mur- (PJCB). Walter’s Copse, Isle of Wight, 1 female, rough, Wicklow, 3 males, 8.v.1989, Withers leg. 2.vii.1980, Chandler leg. (NMS). Welwyn, Hertford- (PPWM). Narborough Bog, Whetstone, Leicestershire, shire, 1 female, viii.1927, Edwards leg. (BMNH). With- 1 male, 24.viii.2003, Menzel leg. (DEI). Nethy Bridge, erslack, Cumbria, 2 females, 12.vii.1923, Edwards leg. Highland Region, 1 male, 16.vi.1936, Coe leg. (BMNH). Woodwalton Fen, Cambridgeshire, 2 males, (BMNH). Searles Farm, Reading, Berkshire, 1 male, 24.v.1940, Edwards leg. (BMNH). Wormsley, Oxford- 26.vi.2003, Chandler leg. (PPCM). Spey Bay, Moray, 1 shire, 1 male, 7.vii.1909, Verrall or Collin leg. (UMO). male, 2.vii.1936, Coe leg. (BMNH). Theale, Berkshire, 1 male, 9.vii.2003, Chandler leg. (PPCM). Wheatfen Ecological notes: common in all types of woodland Broad, Norfolk, 1 male, 25.vii.1989, Withers leg. especially where there is plenty of dead wood; old (PPWM). Woodwalton Fen, Cambridgeshire, 1 male, hedges; grassland with pingo pools; calcareous grass- 24.v.1940, Edwards leg. (BMNH). Wytham Woods, land; marginal vegetation by upland lake; fen; bog; Wytham, Oxfordshire, 2 males, 21.viii.2002, Menzel water meadows and gravel pits. Reared from pupae in leg. (DEI). moss, from yellow larvae in rotten alder wood. v–ix. Ecological notes: found in broad-leaved and mixed woodland especially carr; wetlands including fen, XYLOSCIARA TUOMIKOSKI, 1957 bogs, wet meadow with reed bed on peat, lakeshore; XYLOSCIARA (PROTOXYLOSCIARA) LONGIFORCEPS former gravel workings; sea cliff. v–viii. (BUKOWSKI & LENGERSDORF, 1936) Synonym: denudata (Mohrig & Menzel, 1990). XYLOSCIARA (XYLOSCIARA) LIGNICOLA New to British list. (WINNERTZ, 1867) Material examined: Epping Thicks, Epping Forest, Synonyms: belingi (Winnertz, 1869); hortulana (Bel- Essex, 1 male, iv.1999, Dagley leg. (PPCM). ing, 1873); nitidissima (Strobl, 1880). Ecological notes: found in ancient broad-leaved wood- Literature: Edwards (1938): 201 [as Sciara (Plastosci- land. iii. ara)]; Perry & Stubbs (1978): 71; Freeman (1983b): 26; Menzel (1998): 23. Material examined: Delamere, Cheshire, 2 males, XYLOSCIARA (XYLOSCIARA) BETULAE 17.iv.−26.iv.1928 (reared), Britten leg. (BMNH). Mon- TUOMIKOSKI, 1960 ument Field, Epping Forest, Essex, 1 male, iv.1999, New to British list. Dagley leg. (PPCM). Harpenden, Hertfordshire, 2 Material examined: Beinn Eighe, Highland Region, 1 males, 30.vii.1934, C.B. Williams leg. (BMNH). Letch- male, 11.iv.1952, Richards leg. (BMNH). Parham, worth, Hertfordshire, 1 male, 9.ix.1924; 1 male, West Sussex, 1 male, 13.xi.1957, Hussey leg. (HRI). 30.iv.1925, all Edwards leg. (BMNH). Longdendale, Sparrow Wood, Greater London, 1 male, 15.iv.1972, Cheshire, 5 males, 11.v.−14.v.1932 (reared), Britten Chandler leg. (PPCM). leg. (BMNH). Windsor Forest, Berkshire, 4 males, 1.v.1933 (reared), Donisthorpe leg. (BMNH). Ecological notes: found in broad-leaved and mixed woodland; bark of old birch and on the fungus Pipto- Ecological notes: found in ancient oak and beech forest porus betulinus on birch. iv, xi. and other broad-leaved woodland; conifer plantations. Reared from bark of pine, spruce, birch and oak. iv–v, vii, ix. XYLOSCIARA (XYLOSCIARA) HEPTACANTHA TUOMIKOSKI, 1957 Literature: Menzel (1998): 23; Withers (2002): 115. XYLOSCIARA (XYLOSCIARA) SPECTABILIS RUDZINSKI, 1992 Material examined: Becky Woods, Manaton, Devon, 1 Literature: Edwards (1925a): 542 [as Plastosciara pic- male, 23.vi.2002, J.E. Smith leg. (HRI). Brandon tiventris, misidentiﬁcation]. Marsh, Ryton-on-Dunsmore, West Midlands, 1 male, 22.viii.2002, Menzel leg. (DEI). Corriegills, Arran, New to British list.

Material examined: Radwell, Hertfordshire, 1 male, 1 lebury Common, Berkshire, 1 male, 20.vi.1993; 1 female, 18.iv.1919, Edwards leg. (BMNH). The Loke, male, 30.viii.1993, all Chandler leg. (PPCM). Burton Norwich, Norfolk, 2 males, 20.iv.1987; 1 male, Wood, Cumbria, 1 male, 9.x.1972, Stubbs leg. (PPCM). 1.iv.1988; 1 male, 27.v.1997, all Laurence leg. (PBLN). Caerkief Wood, Cornwall, 1 male, 3.vii.1983, Chandler leg. (NMS). Carbins Wood, Berkshire, 2 males, Ecological notes: found in garden; on a fence. iv–v. 26.vi.1994, Chandler leg. (PPCM). Castlerosse, Killar- Taxonomic note: Radwell specimens referred to by ney, Kerry, 1 male, 2.v.1981, Chandler leg. (PPCM). Edwards (1925a) were Xylosciara spectabilis rather Charlbury, Oxfordshire, 1 male, 4.vii.1936, Edwards than Xylosciara lignicola under which Freeman leg. (BMNH). Charleville Wood, Offaly, 2 males, included this misidentiﬁcation of Plastosciara pic- 15.vi.1985, Chandler leg. (PPWM). Charterhouse tiventris by Edwards. Alder Holt, Surrey, 1 male, 1 female, 16.vii.1989, Chandler leg. (NMS, PPCM). Chedworth, Gloucester- shire, 1 male, 5.vi.1936, Edwards leg. (BMNH). Chee XYLOSCIARA (XYLOSCIARA) SUBBETULAE MOHRIG & Dale, Derbyshire, 1 male, 1 female, 30.x.1991, Chan- KRIVOSHEINA, 1982 dler leg. (NMS). Chippenham Fen, Cambridgeshire, 1 Literature: Menzel (1998): 23. male, 2.vii.1980, Chandler leg. (NMS). Clovelly, Devon, 3 males, 8.v.1936, Edwards leg. (BMNH). Material examined: Hickling Broad, Norfolk, 1 male, Coombe Abbey Country Park, Coventry, Warwick- 13.viii.1989, Withers leg. (PPWM). shire, 2 males, 28.viii.2003, J.E. Smith leg. (HRI). Ecological notes: found in a site comprising alder carr Dean Wood, Devon, 1 male, 11.x.1980, Chandler leg. and sedge beds. viii. (NMS). Devil’s Punchbowl, Surrey, 1 male, 16.v.1992, Chandler leg. (PPCM). Dinton Pastures Country Park, Berkshire, 1 male, 8.v.1993; 2 males, 15.v.1993; 1 ZYGONEURA MEIGEN, 1830 male, 14.vii.1993; 1 male, 18.vi.1994; 2 females, ZYGONEURA (ALLOZYGONEURA) CALTHAE 7.xi.1994; 1 male, 28.v.1998; 1 male, 13.viii.1998; 1 TUOMIKOSKI, 1960 male, 7.x.1998; 1 male, 8.xi.1999, all Chandler leg. (NMS, PPCM). Ditchley, Oxfordshire, 1 male, v.1937, Literature: Menzel (1998): 23. Edwards leg. (BMNH). Drominahassig, Kerry, 1 male, Material examined: Colwyn Brook, Powys, 1 male, 15.v.1973, Chandler leg. (PPCM). Dromore Forest, 20.vi.1989¸ Holmes, Boyce & Reed leg. (PPWM). Kerry, 1 male, 15.x.1973, Chandler leg. (NMS). Dunamarc Falls, Cork, 1 male, 14.x.1973, Chandler Ecological notes: found on Carex acutiformis fen. leg. (NMS). Duncombe Park, North Yorkshire, 1 male, Reared from stem of marsh marigold. vi. 16.vii.1996, Chandler leg. (PPCM). Easington Beck, North Yorkshire, 1 male, 1 female, 11.x.1990, Chan- ZYGONEURA (ZYGONEURA) SCIARINA MEIGEN, 1830 dler leg. (NMS). Epping Forest, Essex, 1 male, 1 female, 12.iv.1999; 1 male, 1 female, 19.iv.1999, all Literature: Walker (1848): 103; Walker (1856): 57; Hal- Dagley leg. (NMS, PPCM). Farnborough Park, War- iday (1857): 31–33; Edwards (1925a): 533; Chandler wickshire, 1 male, 1 female, 10.x.2001, Chandler leg. (1978): 209; Freeman (1983b): 22; Freeman (1987): (HRI). Ferndown, Dorset, 1 male, 24.x.1965, Par- 197; Laurence (1994): 118; Menzel (1998): 23; Coldwell menter leg. (BMNH). Forge Valley, North Yorkshire, 1 (1999): 25. male, 13.ix.1999, Crossley leg. (PPCM). Glen of Material examined: Aldermoors, Berkshire, 1 male, Knocknarea, Sligo, 1 male, 1.x.1977, Chandler leg. 11.vi.1994, Chandler leg. (PPCM). Alvecote Pools, (NMS). Grange-over-Sands, Cumbria, 1 female, Staffordshire, 1 female, 13.x.2001, Chandler leg. xii.1939, Collin leg. (UMO). Grindon Wood, Stafford- (HRI). Ashtead, Surrey, 1 female, 8.v.1932, Coe leg. shire, 1 male, 13.x.1997, Chandler leg. (PPCM). (BMNH). Aversley Wood, Cambridgeshire, 1 male, Gweek Wood, Cornwall, 2 males, 4.vii.1983, Chandler 2.iv.1988, J.H. Cole leg. (PJCB). Bagley Wood, Oxford- leg. (NMS, PPCM). High Dam, Cumbria, 1 male, shire, 1 male, 20.v.−21.v.1937, Edwards leg. (BMNH). 15.vi.1999, Chandler leg. (NMS). Ilfracombe, Devon, 1 Baldock, Hertfordshire, 1 male, ix.-x.1917; 1 male, 1 female, 6.vii.1980, Chandler leg. (NMS). Killarney, female, v.1918, all Edwards leg. (BMNH). Ballyvour- Kerry, 1 male, 25.v.−27.v.1929, Edwards leg. (BMNH). ney, Cork, 1 male, 13.v.1989; 1 male, 10.vii.1990, all Killaun Bog, Offaly, 1 male, ix.2000, Withers leg. Withers leg. (PPWM). Ban-y-Gor Wood, Gloucester- (PPWM). Knap Mill, Devon, 1 male, 2.vi.1978, J.H. shire, 1 male, 12.ix.2002, Gibbs leg. (PDGB). Bramp- Cole leg. (PJCB). Lackham Wood, Wiltshire, 1 male, ton, Cambridgeshire, 6 males, 8 females, 25.iv.1997 18.x.2002, Chandler leg. (HRI). Lancaut Nature (reared), J.H. Cole leg. (PJCB). Brancaster, Norfolk, 1 Reserve, Gloucestershire, 1 female, 5.ix.2002, Gibbs male, 28.x.1988, Foster & Procter leg. (PBLN). Buck- leg. (PDGB). Lathkill Dale, Derbyshire, 1 female,

10.x.1991, Chandler leg. (PPCM). Leckford Estate, bridgeshire, 2 males, 1 female, 24.v.1940, Edwards leg. Hampshire, 1 female, 12.iii.−15.iii.1976 (reared); 4 (BMNH). Wychwood, Oxfordshire, 2 females, v.1937, males, 2 females, 1977 (reared); 1 male, 1.ix.1998; 1 Edwards leg. (BMNH). Wytham Woods, Wytham, male, 11.xi.1999, all Chandler leg. (NMS, PPCM). Oxfordshire, 4 males, 21.viii.2002, J.E. Smith leg. Letchworth, Hertfordshire, 1 male, 11.xi.1916, (HRI). Edwards leg. (BMNH). Llanmynach Hills, Shropshire, Ecological notes: very common in broad-leaved and 1 male, 23.v.1994, J.H. Cole leg. (PJCB). Maulden mixed woodland; common in carr woodland; also found Wood, Bedfordshire, 1 male, 23.vii.1980, J.H. Cole leg. on heathland; wetlands including fen, raised bog, (PJCB). Monk’s Wood, Cambridgeshire, 1 female, sedge beds, water meadows and gravel pits; mossy 3.vi.1972, J.H. Cole leg. (PJCB). Mulgrave Wood, rocks by waterfall; old hedges; grassland and scrub in North Yorkshire, 1 male, 23.viii.−3.ix.1937, Edwards old limestone quarry; gardens. Reared from poplar log, leg. (BMNH). Murrough, Wicklow, 1 male, 8.v.1989, borings of the beetle Hylesinus fraxini and fungus Withers leg. (PPWM). Natural History Museum, Auricularia mesenterica. iii–xi. South Kensington, Greater London, 1 male, 8.vi.1948, Ward leg. (BMNH). Nesbitt Dene, Durham, 1 male, 29.vii.1992; 1 male, 5.viii.1995, all Chandler leg. ACKNOWLEDGEMENTS (NMS, PPCM). Neuadd Fawr, Ceredigion, 1 male, We would like to thank the Deutsches Entomologis- 28.ix.1974, Chandler leg. (PPCM). North Grimston, ches Institut and Warwick HRI for allowing staff time North Yorkshire, 1 male, 16.vii.1996, Chandler leg. and use of their resources; Phil White, Dr Rosemary (PPCM). Oxford, Oxfordshire, 1 male, 2.x.1950 Collier, Dr Dave Chandler and Professor Mark Tatch- (reared), Laurence leg. (PBLN). Oxford University ell for their support and encouragement; Bianka Kat- Museum, Oxford, Oxfordshire, 1 male, 9.xi.1920; 1 nig for her technical help with the figures of the new male, 5.xii.1928, all Hamm leg. (UMO). Perrott’s species; the GCRI Trust, Percy Sladen Fund, British Brook, Gloucestershire, 1 male, 12.x.1979, Stubbs leg. Entomological and Natural History Society and the (PPCM). Radcliffe on Trent, Nottinghamshire, 1 Linnean Society for providing funding; the authorities female, 10.vii.1912, Robinson leg. (BMNH). Ranmore of the Natural History Museum, London, and of the Common, Surrey, 2 males, 12 females, 10.ii.− museums in Edinburgh, Dublin, Belfast and Oxford 17.iii.1936 (reared), Richards leg. (BMNH). Reedham, who kindly gave access to their sciarid collections and Norfolk, 1 male, 26.viii.−27.x.1988, Foster & Procter arranged loans as necessary. We are also indebted to leg. (PBLN). Reelig Glen, Moniack, Highland Region, the many private collectors who have either provided 1 male, 21.x.1999, Chandler leg. (NMS). Reydon Wood, us with data or passed material to us for determina- Suffolk, 1 male, 11.v.1986, Withers leg. (PPWM). Sal- tion. In particular, Brian Laurence, Jonathan Cole, combe, Devon, 1 male, 25.vii.1908, Verrall leg. (UMO). David Gibbs, Patrick Roper and Phil Withers who gave Scadbury Park, Greater London, 1 male, 18.vi.1983; 1 us full access to their sciarid collections and assisted male, 22.x.1983, all Chandler leg. (PPCM, PPWM). us with information on localities and habitats. We also Scotland Fen, Suffolk, 1 female, 19.x.1996, Chandler thank Professor Werner Mohrig for help with fly iden- leg. (PPCM). Selborne Common, Hampshire, 1 male, tification and Dr John Ismay and Barbara Schulten 28.v.1988, Chandler leg. (NMS). Sharpenhoe, Bedford- for reviewing the work. shire, 2 females, 16.vi.1997, Chandler leg. (NMS, PPCM). Shepreth, Cambridgeshire, 1 male, 3 females, iii.1918 (reared), Edwards leg. (BMNH). Sherrard’s REFERENCES Wood, Welwyn, Hertfordshire, 1 male, 22.x.1930, Edwards leg. (BMNH). Sidmouth, Devon, 2 males, Ashe P, O’Connor JP. 1991. Phytosciara (Prosciara) producta Tuomikoski (Diptera: Sciaridae) new to the British 10.v.1936, Edwards leg. (BMNH). The Dingle, Isle of Isles. Dipterists Digest (First Series) 8: 16–17. Anglesey, 1 male, 5.vii.1987, Chandler leg. (PPCM). Basden EB. 1933. On some species of the genus Sciara (Dipt., Theale, Berkshire, 2 males, 1 female, 7.vii.2003, Chan- Mycetophilidae) from the nests of birds and mammals. Ento- dler leg. (HRI). Tintern, Wexford, 1 male, 28.iii.1989, mologist’s monthly Magazine 69: 44. J.P. O’Connor leg. (NMID). Torc Cascade, Kerry, 1 Binns ES. 1973. Laboratory rearing, biology and chemical con- male, 9.v.1981, Chandler leg. (PPCM). Upper Lake, trol of the mushroom sciarid Lycoriella auripila (Diptera: Killarney, Kerry, 1 male, 5.v.1981, Chandler leg. Lycoriidae). Annals of Applied Biology 73: 119–126. (PPCM). Westhide Wood, Herefordshire, 1 male, Binns ES. 1975. Mushroom mycelium and compost substrates 9.x.1998, Chandler leg. (NMS). Weston Big Wood, in relation to the survival of the sciarid Lycoriella auripila. Avon, 1 male, 12.vii.2001, Gibbs leg. (PDGB). Wicken Annals of Applied Biology 80: 1–15. Fen, Cambridgeshire, 1 male, 25.iv.1993, Perry leg. Binns ES. 1980. Field and laboratory observations on the sub- (PJCB). Woodbastwick, Norfolk, 1 male, viii.1932, strates of the mushroom fungus gnat Lycoriella auripila Edwards leg. (BMNH). Woodwalton Fen, Cam- (Diptera: Sciaridae). Annals of Applied Biology 96: 143–152.

Binns ES. 1981. Sciarids as migrants. Mushroom Journal Edwards FW. 1915b. A second British species of Plastosciara. 108: 415–423. Entomologist’s monthly Magazine 51: 263–264. Brindle A. 1963. Culicidae to Hippoboscidae. In: Henson H, Edwards FW. 1922. A third new British Plastosciara ed. The insects of the Malham Tarn Area. Proceedings of the (Diptera, Sciaridae). Entomologist’s monthly Magazine 58: Leeds Philosophical and Literary Society, Scientific Section 160–161. 9(2): 83–90 Edwards FW. 1925a. XXII. British fungus-gnats (Diptera, Bury H. 1920. Diptera in South Shropshire, 1913–1920. Ento- Mycetophilidae). With a revised generic classification of the mologist’s monthly Magazine 56: 249–256. family. Transactions of the Entomological Society of London Buxton JH. 1993. Control of sciarid fly with parasitic nema- 1925 (3–4) (1924): 505–670. todes. Bulletin de 1’Organisation Internationale de Lutte Edwards FW. 1925b. Two new British Sciarinae (Diptera). Biologique et Intégrée contre les. Animaux et les Plantes Entomologist’s monthly Magazine 61: 223–224. Nuisibles, Section Régionale Ouest Paléarctique (OILB/ Edwards FW. 1925c. Sciara caudata Walk., Pezomyia vander- SROP) 16(2): 23–25. wulpi de Meij. and other Diptera (two new to the British list) Buxton, PA. 1954. British Diptera associated with fungi. 2.- reared from a rotten willow log. Entomologist’s monthly Diptera bred from Myxomycetes. Proceedings of the Royal Magazine 61: 228. Entomological Society of London (Series A) 29(10–12): 163– Edwards FW. 1926a. Another case of reduced wings in a 167. male Sciarine fly (Diptera, Mycetophilidae). Entomologist’s Chandler P. 1978. Associations with plants. In: Stubbs A, monthly Magazine 62: 111–113. Chandler P, eds. A dipterist’s handbook. The Amateur Ento- Edwards FW. 1926b. North Lancashire and South Westmor- mologist 15: 199–211 land Diptera. North Western Naturalist 1: 141–143. Chandler P. 1987. Scythropochroa radialis Lengersdorf (Dipt. Edwards FW. 1934a. A new brachypterous Sciarine fly. Ento- Sciaridae) identity of British material confirmed. Entomolo- mologist’s monthly Magazine 70: 140–141. gist’s monthly Magazine 123: 204. Edwards FW. 1934b. Diptera on Grassholm Island, Pem- Chandler P. 1998. Checklists of Insects of the British isles brokeshire. Journal of the Society for British Entomology 1: (new series). Part 1: Diptera (incorporating a list of Irish 33–34. Diptera). Handbooks for the Identification of British Insects Edwards FW. 1938. Notes on the Mycetophilidae (Dipt.) 12: i–xix and 1–234. obtained by Dr. C.B. Williams in a light-trap at Harpenden, Chandler P. 2001. Nematocera (Diptera) in Buckingham Pal- Herts. Journal of the Society for British Entomology 1: 199– ace Garden. In: Plant CW, ed. The natural history of Buck- 202. ingham Palace Garden, London, Part 2. The London Edwards FW. 1941. Notes on British fungus-gnats (Dipt., Naturalist 80 (Suppl.): 197–206. Mycetophilidae). Entomologist’s monthly Magazine 77: 21– Chandler PJ, O’Connor JP, Ashe P. 1996. Notes on the 32 and 67–82. Irish Sciaridae (Diptera), including five species new to Ire- Edwards FW, Collin JE. 1932. A revised list of the Diptera of land. Irish Naturalists’ Journal 25: 258–259. St. Kilda. Entomologist’s monthly Magazine 68: 263–266. Chandler PJ, O’Connor JP, Nash R, Withers P. 2002. Edwards FW, Williams CB. 1916. Sciara tritici, Coq. A fly Diptera new to Ireland in seventeen families. Dipterists injurious to seedlings. Annals of Applied Biology 2: 258–262. Digest (Second Series) 9: 121–136. Freeman P. 1983a. Revisionary notes on British Sciaridae Coldwell JD. 1999. Insects of the Barnsley Area. Sorby (Diptera). Entomologist’s monthly Magazine 119: 161–170. Record Special Series 12: 1–170. Freeman P. 1983b. Sciarid flies. Diptera, Sciaridae. Hand- Disney RHL. 1976. A further case of a nematoceran fly books for the Identification of British Insects 9: 1–68. (Diptera: Sciaridae) parasitised by a species of scuttle fly Freeman P. 1987. British Sciaridae (Diptera): new species (Diptera: Phoridae). Entomologist’s Gazette 27: 91–98. and records with notes on the Tuomikoski collection. Ento- Donisthorpe H. 1913. Ants and myrmecophiles on Lundy. mologist’s monthly Magazine 123: 195–204. Entomologists Record and Journal of Variation 25: 207–269. Freeman P. 1990. Notes on British Sciaridae (Diptera) with Donisthorpe H. 1924. Myrmecophilous Notes for 1923. Ento- descriptions of three new species and of a species new to mologists Record and Journal of Variation 36: 69–75. Britain. Entomologist’s monthly Magazine 126: 51–55. Donisthorpe H. 1927. The guests of British ants. Their habits Grensted LW. 1944. Records of Diptera and other insects from and life-histories. London: George Routledge & Sons Ltd. Dinas Head, Pembrokeshire. Entomologist’s monthly Maga- Dunne R, O’Connor JP. 1989. Some insects (Thysanoptera: zine 80: 201–203. Diptera) of economic importance, new to Ireland. Irish Nat- Haliday AH. 1833. Catalogue of Diptera occurring about uralists’ Journal 23: 63–65. Holywood in Downshire. Entomological Magazine 1: 147– Edwards FW. 1913a. Sexual dimorphism in a species of Sci- 180. ara. Entomologist’s monthly Magazine 49: 209–211. Haliday AH. 1857. Entomological notes. Proceedings of the Edwards FW. 1913b. Notes on the British Mycetophilidae. Dublin University Zoological and Botanical Association 4: Transactions of the Royal Entomological Society of London 31–36. 1913: 334–382. Hancock EG. 1997. Scottish insect records for 1995. Glasgow Edwards FW. 1915a. Ten new British Diptera (Nematocera). Naturalist 23: 25–29. Entomologist’s monthly Magazine 51: 164–166. Heller K. 2004. Eine Bestandsaufnahme der Sciaridae

(Diptera) Schleswig-Holsteins mit Ergänzungen und Kor- Menzel F, Mohrig W. 1997a. Revision der paläarktischen rekturen zum bisher bekannten Arteninventar. Faunistisch- Arten von Trichosia Winnertz sensu Tuomikoski, 1960 Ökologische Mitteilungen 8: 233–257. (Diptera, Sciaridae). – Teil I. Gattung Trichosia Winnertz, Hippa H, Vilkamaa P. 1994. The genus Camptochaeta gen. 1867. Studia Dipterologica 4: 3–40. nov. (Diptera, Sciaridae). Acta Zoologica Fennica 194: 1–85. Menzel F, Mohrig W. 1997b. 2.6. Family Sciaridae. In: Papp Hippa H, Vilkamaa P, Mohrig W. 2003. Phylogeny of L, Darvas B, eds. Contributions to a manual of Palaearctic Corynoptera Winnertz and related genera, with the descrip- Diptera (with special reference to flies of economic impor- tion of Claustropyga gen. nov. (Diptera, Sciaridae). Studia tance). Nematocera and Lower Brachycera 2: Budapest: Sci- Dipterologica 9: 469–511. ence Herald, pp. 51–69 Kidd LN. 1953. Some West Cheshire Diptera. Journal of the Menzel F, Mohrig W. 1998. Beiträge zur Taxonomie und Fau- Society for British Entomology 4: 137–138. nistik der paläarktischen Trauermücken (Diptera, Scia- King G. 1991. Trichosia hirtipennis (Zetterstedt) (Dipt., Scia- ridae). Teil VI – Neue Ergebnisse aus Typenuntersuchungen ridae) new to Britain. Entomologist’s monthly Magazine 127: und die daraus resultierenden taxonomisch-nomenkla- 43–44. torischen Konsequenzen. Studia Dipterologica 5: 351–378. Kloet GS, Hincks WD. 1945. A Check List of British Insects, Menzel F, Mohrig W. 2000. Revision der paläarktischen i–iix and, pp. 1–483. Arbroath: T. Buncle Ltd (private publi- Trauermücken (Diptera: Sciaridae). Studia Dipterologica 6 cation). (Suppl.): 1–761. Kloet GS, Hincks WD. 1976. A Check List of British Insects, Menzel F, Smith JE, Colauto NB. 2003. Bradysia difformis 2nd edn. (completely revised). Handbooks for the Identifica- Frey and Bradysia ocellaris (Comstock): two additional Neo- tion of British Insects 11: 1–139. tropical species of Black Fungus Gnats (Diptera: Sciaridae) Laurence BR. 1952. Unusual sexual behaviour of Sciara car- of economic importance: a redescription and review. Annals bonaria Mg. Entomologist’s Record and Journal of Variation of the Entomological Society of America 96: 448–457. 64: 90. Mohrig W, Menzel F. 1997. Revision der paläarktischen Laurence BR. 1994. Sciaridae (Dipt.) from East Anglian wet- Arten von Trichosia Winnertz sensu Tuomikoski, 1960 lands, with descriptions of new species. Entomologist’s (Diptera, Sciaridae). – Teil II. Gattungen Leptosciarella monthly Magazine 130: 105–119. Tuomikoski, 1960 und Trichodapus gen. nov. Studia Dipter- Laurence BR. 1996. Beyond the Red Data Book. Dipterists ologica 4: 41–98. Digest (Second Series) 2: 85–89. Mohrig W, Menzel F, Kozánek M. 1992. Neue Trauer- Laurence BR. 1997a. Flies from ancient coppiced woodland mücken (Diptera, Sciaridae) aus Nord-Korea und Japan. in Suffolk. Dipterists Digest (Second Series) 4: 78–92. Dipterological Research 3: 17–32. Laurence BR. 1997b. Diptera in the northern Isles of Britain. O’Connor JP, Ashe P. 1991. Ten species of Sciaridae Entomologist’s monthly Magazine 133: 225–232. (Diptera) new to Ireland. Dipterists Digest (First Series) 8: Laurence BR, James R. 1996. Corynoptera stammeri (Leng- 29–30. ersdorf) (Sciaridae) and Colobostema infumatum (Haliday) O’Toole, C. 1978. Ants, bees and wasps (aculeate (Scatopsidae) from Wales: new to the British list of Diptera. Hymenoptera). In: Stubbs A, Chandler P, eds. A Dipterist’s Entomologist’s monthly Magazine 132: 73–75. Handbook. The Amateur Entomologist 15: 157–164. Laurence BR, James R. 1997. Diptera from the mountain Parmenter L. 1950. The Diptera of Bookham Common. Lon- of Cadair Idris, Gwynedd, North Wales. Entomologist’s don Naturalist 29: 98–133. monthly Magazine 133: 165–171. Parmenter L. 1960. A further list of the Diptera of Bookham Leigh C, Ware C. 2003. The development of the flora, fauna Common. London Naturalist 39: 66–76. and environment of the Wildlife Garden at the Natural His- Perry I, Stubbs AE. 1978. Dead wood and sap runs. In: tory Museum, London. London Naturalist 82: 75–134. Stubbs A, Chandler P, eds. A Dipterist’s Handbook. The Ama- Lengersdorf F. 1928–30. 7. Lycoriidae (Sciaridae). In: Lind- teur Entomologist 15: 65–73. ner E, ed. Die Fliegen der Palaearktischen Region 2 (1): 1–22, Pethybridge NJ. 1991. The systematic use of bioassay meth- 1–37, 1–16, 1–75, 1–62, 1–71 and 1–196. Stuttgart: E. Sch- ods to determine the effectiveness of Bacillus thuringiensis weizerbart’sche-Verlagsbuchhandlung, pp. 1–71 isolates for the control of the mushroom sciarid Lycoriella Madwar S. 1937. Biology and morphology of the immature auripila (Diptera: Sciaridae). In: Maher MJ, ed. Science and stages of Mycetophilidae (Diptera: Nematocera). Publication Cultivation of Edible Fungi. Proceedings of the 13th Interna- of Royal Entomological Society of London 227: 42–51. tional Congress of the Science and Cultivation of Edible Menzel F. 1998. Sciaridae. In: Chandler P, ed. Checklists of Fungi. Rotterdam: A. A. Balkema, and Brookfield, pp. 453– Insects of the British isles (new series). Part 1: Diptera (incor- 456 porating a list of Irish Diptera). Handbooks for the Identifi- Poinar GO, Doncaster CC. 1965. The penetration of Trichius cation of British Insects 12: i-xix and 1–234, 20–24. sciarae (Bovien) (Sphaerulariidae: Aphelenchoidea) into its Menzel F, Heller K. 2005. Fünf neue Arten aus den Gattun- insect host, Bradysia paupera Tuom. (Mycetophilidae: gen Bradysia, Camptochaeta und Corynoptera (Diptera: Diptera). Nematologica 11: 73–78. Sciaridae) nebst einigen Bemerkungen zur Nomenklatur Schmitz H. 1918. Neue Beiträge zur Kenntnis der Sciariden europäischer Trauermücken. Studia Dipterologica 11: 335– mit reduzierten Maxillarpalpen. Tijdschrift Voor Entomolo- 357. gie 61: 88–111.

Skidmore P. 1978. Dung. In: Stubbs A, Chandler P, eds. A Walker F. 1848. List of the Specimens of Dipterous Insects in Dipterist’s Handbook. The Amateur Entomologist 15: 73–79. the Collection of the British Museum 1: 1–229. Skidmore P. 1991. Insects of the British cow-dung commu- Walker F. 1856. Insecta britannica. Diptera 3: 1–352. nity. Field Studies Council AIDGAP project. Occasional Pub- White PF. 1986. The effect of sciarid larvae (Lycoriella aurip- lication 21: 1–166. ila) on cropping of the cultivated mushroom (Agaricus Skidmore P, Limbert M, Eversham BC. 1987. The insects biporus). Annals of Applied Biology 109: 11–17. of Thorne Moors. The Sorby Record. A Journal of Natural White PF. 1997. The use of chemicals, antagonists, repellents History for the Sheffield Area 23 (Suppl.): 90–153. and physical barriers for the control of Lycoriella auripila Smith KGV. 1974. Changes in the British Dipterous Fauna. (Diptera: Sciaridae), a pest of the cultivated mushroom In: Hawksworth, DL, ed. The changing flora and fauna of Agaricus bisporus. Annals of Applied Biology 131: 29–42. Britain. London: Academic Press, pp. 371–391. White PF, Jarrett P. 1990. Laboratory and field tests with Symes CB. 1921. Insect pests of mushrooms. Part I. The Bacillus thuringiensis for the control of the mushroom scia- mushroom fly (Sciara praecox Meig.). Fruit-Grower 51: 142– rid Lycoriella auripila. Brighton Crop Protection Conference. 145, 188–190 and 234–236. Pests and Diseases 1990: 373–378. Theobald FV. 1892. An account of British Flies (Diptera). White PF, Smith JE. 2000. Bradysia lutaria (Winn.) (Dipt., London: Elliott Stock. Sciaridae) – a recent addition to the British fauna and a pest Tuomikoski R. 1960. Zur Kenntnis der Sciariden (Dipt.) of the cultivated mushroom (Agaricus bisporus). Entomolo- Finnlands. Annales Zoologici Societatis Zoologicae Botani- gist’s monthly Magazine 136: 165–167. cae Fennicae ‘Vanamo’ 21: 1–164. White PF, Smith JE, Menzel F. 2000. Distribution of Verrall GH. 1886. A hundred new British species of Diptera. Sciaridae (Dipt.) species infesting commercial mushroom The Entomologist’s monthly Magazine 22: 179–182. farms in Britain. Entomologist’s monthly Magazine 136: Verrall GH. 1888. A list of British Diptera. London: Pratt. 207–209. Verrall GH. 1901. A list of British Diptera, 2nd edn. Cam- Winnertz J. 1867. Beitrag zu einer Monographie der Scia- bridge: The University Press. rinen. Vienna. Verrall GH. 1912. Another hundred new British species of Withers P. 2002. Thirty-nine species of Diptera new to Ire- Diptera. The Entomologist’s monthly Magazine (Second land, including a species of Psychodidae new to the British Series) 48: 20–27. Isles. Dipterists Digest (Second Series) 9: 113–119.