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The Natural History, Distribution, and Phenotypic Variation of Cave-Dwelling Spring Salamanders, Gyrinophilus Spp
Marshall University Marshall Digital Scholar Theses, Dissertations and Capstones 2005 The aN tural History, Distribution, and Phenotypic Variation of Cave-dwelling Spring Salamanders, Gyrinophilus spp. Cope (Plethodontidae), in West Virginia Michael Steven Osbourn Follow this and additional works at: http://mds.marshall.edu/etd Part of the Aquaculture and Fisheries Commons, and the Ecology and Evolutionary Biology Commons Recommended Citation Osbourn, Michael Steven, "The aN tural History, Distribution, and Phenotypic Variation of Cave-dwelling Spring Salamanders, Gyrinophilus spp. Cope (Plethodontidae), in West Virginia" (2005). Theses, Dissertations and Capstones. Paper 735. This Thesis is brought to you for free and open access by Marshall Digital Scholar. It has been accepted for inclusion in Theses, Dissertations and Capstones by an authorized administrator of Marshall Digital Scholar. For more information, please contact [email protected]. The Natural History, Distribution, and Phenotypic Variation of Cave-dwelling Spring Salamanders, Gyrinophilus spp. Cope (Plethodontidae), in West Virginia. Thesis submitted to The Graduate College of Marshall University In partial fulfillment of the Requirements for the degree of Master of Science Biological Sciences By Michael Steven Osbourn Thomas K. Pauley, Committee Chairperson Daniel K. Evans, PhD Thomas G. Jones, PhD Marshall University May 2005 Abstract The Natural History, Distribution, and Phenotypic Variation of Cave-dwelling Spring Salamanders, Gyrinophilus spp. Cope (Plethodontidae), in West Virginia. Michael S. Osbourn There are over 4000 documented caves in West Virginia, potentially providing refuge and habitat for a diversity of amphibians and reptiles. Spring Salamanders, Gyrinophilus porphyriticus, are among the most frequently encountered amphibians in caves. Surveys of 25 caves provided expanded distribution records and insight into ecology and diet of G. -
Federal Register/Vol. 76, No. 55/Tuesday, March 22, 2011
Federal Register / Vol. 76, No. 55 / Tuesday, March 22, 2011 / Proposed Rules 15919 before the test, per the regulation. provided by the roof crush standard. safety argument. Instead, its arguments Furthermore, it is not apparent how the NTEA stated that its statistics show that are based primarily on overstated modifications generally made by a final- the vast majority of multi-stage vehicles certification risk. As such, we believe stage manufacturer will create are rated above 6,000 pounds. NTEA that this rule should continue to include compliance difficulties with FMVSS No. noted that FMVSS No. 216a excludes those vehicles with an intact, compliant 216a. Moreover, as we explained in the trucks other than ones built on chassis- roof structure, whether they are multi-stage certification rulemaking, if cabs (and incomplete vehicles with a delivered to the dealership or the final- final-stage manufacturers identify full exterior van body), meaning that the stage manufacturer. particular areas where compliance with agency excluded approximately one- FMVSS No. 216a is a problem, they, or third of multi-stage vehicles with a IV. Conclusion NTEA on behalf of its members, can GVWR of 6001 pounds to 10,000 For the reasons discussed above, we petition for a temporary exemption pounds. NTEA also said that chassis deny the petition for reconsideration under 49 CFR part 555.78 with a GVWR of over 10,000 pounds submitted by NTEA. In our Further Response, we stated constitute 94.5 percent of the entire Authority: 49 U.S.C. 322, 30111, 30115, that in analyzing the 2006 GMT–355 market of chassis rated above 6,000 30166 and 30177; delegation of authority at IVD, which is for a body-on-frame pounds. -
Tennessee-Alabama-Georgia (TAG) Cave Teaching and Learning Module
Tennessee-Alabama-Georgia (TAG) Cave Teaching and Learning Module K. Denise Kendall, Ph.D. Matthew L. Niemiller, Ph.D. Annette S. Engel, Ph.D. Funding provided by 1 Dear Educator, We are pleased to present you with a TAG (Tennessee – Alabama – Georgia) cave-themed teaching and learning module. This module aims to engage Kindergarten through 5th grade students in subterranean biology, while fostering awareness and positive attitudes toward cave biodiversity. We have chosen cave fauna for this teaching module because students have a fascination with atypical organisms and environments. Moreover, little attention has been given to subterranean biodiversity in public outreach programs. Many students will likely be intrigued by the unique fauna and composition of subterranean landscapes. Therefore, we hope these lessons enable teachers to introduce students to the unique organisms and habitat below their feet. The module presents students with background information and outlines lessons that aim to reinforce and discover aspects of the content. Lessons in this module focus primarily on habitat formation, biodiversity, evolution, and system flows in subterranean landscapes. We intend for this module to be a guide, and, thus, we have included baseline material and activity plans. Teachers are welcome to use the lessons in any order they wish, use portions of lessons, and may modify the lessons as they please. Furthermore, educators may share these lessons with other school districts and teachers; however, please do not receive monetary gain for lessons in the module. Funding for the TAG Cave module has been graciously provided by the Cave Conservation Foundation, a non-profit 501(c)3 organization dedicated to promoting and facilitating the conservation, management, and knowledge of cave and karst resources. -
Distribution and Relative Abundance of Tennessee Cave Salamanders (Gyrinophilus Palleucus and Gyrinophilus Gulolineatus) with an Emphasis on Tennessee Populations
Herpetological Conservation and Biology 3(1):1-20. Submitted: 18 September 2007; Accepted: 28 December 2007 DISTRIBUTION AND RELATIVE ABUNDANCE OF TENNESSEE CAVE SALAMANDERS (GYRINOPHILUS PALLEUCUS AND GYRINOPHILUS GULOLINEATUS) WITH AN EMPHASIS ON TENNESSEE POPULATIONS 1,2 1,3 BRIAN T. MILLER AND MATTHEW L. NIEMILLER 1 Department of Biology, Middle Tennessee State University, Murfreesboro, Tennessee 37132, USA 2 Corresponding author, e-mail: [email protected] 3 Present Address: Department of Ecology and Evolutionary Biology, University of Tennessee, Knoxville, Tennessee 37996, USA, e-mail: [email protected] Abstract.—The Tennessee Cave Salamander complex (Gyrinophilus palleucus and G. gulolineatus) consists of three obligate cave-dwelling taxa inhabiting subterranean waters of east and central Tennessee, north Alabama, and northwest Georgia. Although ranges of these taxa are poorly understood, their populations are reportedly small and declining. The IUCN lists G. gulolineatus as "Endangered" and G. p. necturoides as "Vulnerable"; whereas, NatureServe lists G. gulolineatus (G1) and G. p. necturoides (G2G3T1) as Critically Imperiled. To better determine the distribution and relative abundance of extant populations, we searched 113 cave streams in middle and east Tennessee, seven in northwest Georgia, 13 in north Alabama and two in southern Kentucky. We found 1183 salamanders, including 63 G. gulolineatus, 681 G. palleucus, and 439 G. porphyriticus (Spring Salamanders), during 229 surveys of 135 caves. Gyrinophilus palleucus and G. gulolineatus were observed in more caves (30) than G. porphyriticus (17 caves). Members of the complex were found at 52% (12 of 23) of historic caves and at 16% (18 of 110) of non-historic caves. We extended the known distribution of G. -
Standard Common and Current Scientific Names for North American Amphibians, Turtles, Reptiles & Crocodilians
STANDARD COMMON AND CURRENT SCIENTIFIC NAMES FOR NORTH AMERICAN AMPHIBIANS, TURTLES, REPTILES & CROCODILIANS Sixth Edition Joseph T. Collins TraVis W. TAGGart The Center for North American Herpetology THE CEN T ER FOR NOR T H AMERI ca N HERPE T OLOGY www.cnah.org Joseph T. Collins, Director The Center for North American Herpetology 1502 Medinah Circle Lawrence, Kansas 66047 (785) 393-4757 Single copies of this publication are available gratis from The Center for North American Herpetology, 1502 Medinah Circle, Lawrence, Kansas 66047 USA; within the United States and Canada, please send a self-addressed 7x10-inch manila envelope with sufficient U.S. first class postage affixed for four ounces. Individuals outside the United States and Canada should contact CNAH via email before requesting a copy. A list of previous editions of this title is printed on the inside back cover. THE CEN T ER FOR NOR T H AMERI ca N HERPE T OLOGY BO A RD OF DIRE ct ORS Joseph T. Collins Suzanne L. Collins Kansas Biological Survey The Center for The University of Kansas North American Herpetology 2021 Constant Avenue 1502 Medinah Circle Lawrence, Kansas 66047 Lawrence, Kansas 66047 Kelly J. Irwin James L. Knight Arkansas Game & Fish South Carolina Commission State Museum 915 East Sevier Street P. O. Box 100107 Benton, Arkansas 72015 Columbia, South Carolina 29202 Walter E. Meshaka, Jr. Robert Powell Section of Zoology Department of Biology State Museum of Pennsylvania Avila University 300 North Street 11901 Wornall Road Harrisburg, Pennsylvania 17120 Kansas City, Missouri 64145 Travis W. Taggart Sternberg Museum of Natural History Fort Hays State University 3000 Sternberg Drive Hays, Kansas 67601 Front cover images of an Eastern Collared Lizard (Crotaphytus collaris) and Cajun Chorus Frog (Pseudacris fouquettei) by Suzanne L. -
Endangered and Threatened Wildlife and Plants
Federal Register / Vol. 76, No. 55 / Tuesday, March 22, 2011 / Proposed Rules 15919 before the test, per the regulation. provided by the roof crush standard. safety argument. Instead, its arguments Furthermore, it is not apparent how the NTEA stated that its statistics show that are based primarily on overstated modifications generally made by a final- the vast majority of multi-stage vehicles certification risk. As such, we believe stage manufacturer will create are rated above 6,000 pounds. NTEA that this rule should continue to include compliance difficulties with FMVSS No. noted that FMVSS No. 216a excludes those vehicles with an intact, compliant 216a. Moreover, as we explained in the trucks other than ones built on chassis- roof structure, whether they are multi-stage certification rulemaking, if cabs (and incomplete vehicles with a delivered to the dealership or the final- final-stage manufacturers identify full exterior van body), meaning that the stage manufacturer. particular areas where compliance with agency excluded approximately one- FMVSS No. 216a is a problem, they, or third of multi-stage vehicles with a IV. Conclusion NTEA on behalf of its members, can GVWR of 6001 pounds to 10,000 For the reasons discussed above, we petition for a temporary exemption pounds. NTEA also said that chassis deny the petition for reconsideration under 49 CFR part 555.78 with a GVWR of over 10,000 pounds submitted by NTEA. In our Further Response, we stated constitute 94.5 percent of the entire Authority: 49 U.S.C. 322, 30111, 30115, that in analyzing the 2006 GMT–355 market of chassis rated above 6,000 30166 and 30177; delegation of authority at IVD, which is for a body-on-frame pounds. -
Resources and Biodiversity in Tennessee Caves Annette S
W 453-D Landowner’s Guide to Biological Resources and Biodiversity in Tennessee Caves Annette S. Engel, Department of Earth and Planetary Sciences, University of Tennessee Matthew L. Niemiller, Department of Biological Sciences, University of Alabama in Huntsville Editor: Eric Drumm, Department of Biosystems Engineering and Soil Science, University of Tennessee Caves and associated subterranean habitats represent one of the most unforgiving and challenging environments on earth, even though significant and diverse fauna are found in these habitats. More than 25 percent of all known caves in the United States are located in Tennessee, Alabama and Georgia (TAG). More than 1,138 cave-restricted animal species have been described in the United States alone, representing 112 families and 239 genera (Hobbs 2012). Tennessee ranks second behind Texas for the most obligate subterranean species in the United States. This guide summarizes what it is like to live in a cave, describes the types of life that may be encountered in Tennessee caves, and provides an overview of how understanding more about cave biodiversity can help with conservation and management of caves and karst in the state. Other information about Tennessee caves and sinkholes can be found in the series of publications, W 453-A: Guide to Caves and Sinkholes in Tennessee.pdf The Cave Habitat Figure 1. Entrance to a cave in Putnam Co., Tennessee (photo by M.L. Niemiller). The cave habitat consists of three major ecological zones: the entrance (Figure 1), twilight zone and dark zone. Each zone has specific light and other abiotic conditions. The biology of each zone is adapted to live within those conditions. -
Appalachian Salamander Cons
PROCEEDINGS OF THE APPALACHIAN SALAMANDER CONSERVATION WORKSHOP - 30–31 MAY 2008 CONSERVATION & RESEARCH CENTER, SMITHSONIAN’S NATIONAL ZOOLOGICAL PARK, FRONT ROYAL, VIRGINIA, USA Hosted by Smithsonian’s National Zoological Park, facilitated by the IUCN/SSC Conservation Breeding Specialist Group A contribution of the IUCN/SSC Conservation Breeding Specialist Group © Copyright 2008 CBSG IUCN encourages meetings, workshops and other fora for the consideration and analysis of issues related to conservation, and believes that reports of these meetings are most useful when broadly disseminated. The opinions and views expressed by the authors may not necessarily reflect the formal policies of IUCN, its Commissions, its Secretariat or its members. The designation of geographical entities in this book, and the presentation of the material, do not imply the expression of any opinion whatsoever on the part of IUCN concerning the legal status of any country, territory, or area, or of its authorities, or concerning the delimitation of its frontiers or boundaries. Gratwicke, B (ed). 2008. Proceedings of the Appalachian Salamander Conservation Workshop. IUCN/SSC Conservation Breeding Specialist Group: Apple Valley, MN. To order additional copies of Proceedings of the Appalachian Salamander Conservation Workshop, contact the CBSG office: [email protected], 001-952-997-9800, www.cbsg.org. EXECUTIVE SUMMARY Salamanders, along with many other amphibian species have been declining in recent years. The IUCN lists 47% of the world’s salamanders threatened or endangered, yet few people know that the Appalachian region of the United States is home to 14% of the world’s 535 salamander species, making it an extraordinary salamander biodiversity hotspot, and a priority region for salamander conservation. -
Download the Parcas Criteria and Implementation Plan Here!
Model Criteria and Implementation Guidance for a Priority Amphibian and Reptile Conservation Area (PARCA) System in the U.S.A. V.4 October 2012 PARCA Task Team Prepared by Ron Sutherland and Phillip deMaynadier In collaboration with National PARC’s "Priority Amphibian and Reptile Conservation Areas" Task Team: Co-Leads: Ron Sutherland , Wildlands Network Phillip deMaynadier, Maine Dept. of Inland Fisheries & Wildlife Team Members: Margaret Trani Griep, U.S. Forest Service Southern Region Audrey Hatch, formerly Oregon Dept. of Fish and Wildlife Randy Jennings, Western New Mexico University Karen Kinkead , Iowa Dept. of Natural Resources Priya Nanjappa, Assoc. of Fish & Wildlife Agencies Cover Photographs: Blanding’s Turtle (Emydoidea blandingii) – J. Mays Coastal Tailed Frog (Ascaphus truei) – B. McCreary Eastern Indigo Snake (Drymarchon couperi) – J.D. Willson Blue-spotted Salamander (Ambystoma laterale) – L. Kenney Reticulate Gila Monster (Heloderma s. suspectum) – K. Stohlgren Suggested citation: R. Sutherland and P. deMaynadier. 2012. Model Criteria and Implementation Guidance for a Priority Amphibian and Reptile Conservation Area (PARCA) System in the USA. Partners in Amphibian and Reptile Conservation, Technical Publication PARCA-1. 28 pp. 2 V.4 October 2012 PARCA Task Team Contents: Acknowledgments ………………………………………………………………………….....................…...3 Introduction…………..……. ……………………………………………………………………………………3 Criteria for PARCA Selection…………………… …………………………………………………………5 Criteria Overview and PARCA Scale.............……………………………………………………….…..8 -
FEIS Citation Retrieval System Keywords
FEIS Citation Retrieval System Keywords 29,958 entries as KEYWORD (PARENT) Descriptive phrase AB (CANADA) Alberta ABEESC (PLANTS) Abelmoschus esculentus, okra ABEGRA (PLANTS) Abelia × grandiflora [chinensis × uniflora], glossy abelia ABERT'S SQUIRREL (MAMMALS) Sciurus alberti ABERT'S TOWHEE (BIRDS) Pipilo aberti ABIABI (BRYOPHYTES) Abietinella abietina, abietinella moss ABIALB (PLANTS) Abies alba, European silver fir ABIAMA (PLANTS) Abies amabilis, Pacific silver fir ABIBAL (PLANTS) Abies balsamea, balsam fir ABIBIF (PLANTS) Abies bifolia, subalpine fir ABIBRA (PLANTS) Abies bracteata, bristlecone fir ABICON (PLANTS) Abies concolor, white fir ABICONC (ABICON) Abies concolor var. concolor, white fir ABICONL (ABICON) Abies concolor var. lowiana, Rocky Mountain white fir ABIDUR (PLANTS) Abies durangensis, Coahuila fir ABIES SPP. (PLANTS) firs ABIETINELLA SPP. (BRYOPHYTES) Abietinella spp., mosses ABIFIR (PLANTS) Abies firma, Japanese fir ABIFRA (PLANTS) Abies fraseri, Fraser fir ABIGRA (PLANTS) Abies grandis, grand fir ABIHOL (PLANTS) Abies holophylla, Manchurian fir ABIHOM (PLANTS) Abies homolepis, Nikko fir ABILAS (PLANTS) Abies lasiocarpa, subalpine fir ABILASA (ABILAS) Abies lasiocarpa var. arizonica, corkbark fir ABILASB (ABILAS) Abies lasiocarpa var. bifolia, subalpine fir ABILASL (ABILAS) Abies lasiocarpa var. lasiocarpa, subalpine fir ABILOW (PLANTS) Abies lowiana, Rocky Mountain white fir ABIMAG (PLANTS) Abies magnifica, California red fir ABIMAGM (ABIMAG) Abies magnifica var. magnifica, California red fir ABIMAGS (ABIMAG) Abies -
And Spring Salamander (G
Herpetological Conservation and Biology 5(1):32-43. Submitted: 17 September 2009; Accepted: 22 February 2010. CONSERVATION STATUS AND HABITAT USE OF THE WEST VIRGINIA SPRING SALAMANDER (GYRINOPHILUS SUBTERRANEUS) AND SPRING SALAMANDER (G. PORPHYRITICUS) IN GENERAL DAVIS CAVE, GREENBRIER CO., WEST VIRGINIA 1 2 3 4 MATTHEW L. NIEMILLER , MICHAEL S. OSBOURN , DANTE B. FENOLIO , THOMAS K. PAULEY , 5 6 BRIAN T. MILLER , AND JOHN R. HOLSINGER 1Department of Ecology and Evolutionary Biology, University of Tennessee, Knoxville, Tennessee 37996, USA, e-mail: [email protected] 2Division of Biological Sciences, University of Missouri, Columbia, Missouri 65211, USA, e-mail: [email protected] 3Center for Conservation Research, Atlanta Botanical Garden, 1345 Piedmont Ave. NE, Atlanta, Georgia 30309, USA, e-mail: [email protected] 4Department of Biology, Marshall University, Huntington, West Virginia 25755, USA, e-mail: [email protected] 5Department of Biology, Middle Tennessee State University, Murfreesboro, Tennessee 37132, USA, e-mail: [email protected] 6Department of Biological Sciences, Old Dominion University, Norfolk, Virginia 23529, USA, E-mail: [email protected] Abstract.—The West Virginia Spring Salamander (Gyrinophilus subterraneus) is one of four obligate cave-dwelling species of plethodontid salamanders found east of the Mississippi River in the United States. This species is endemic to a single cave system; General Davis Cave, in Greenbrier Co., West Virginia, where it is syntopic with the closely-related Spring Salamander (G. porphyriticus). Accordingly, the West Virginia Spring Salamander is a species of critical conservation concern. Because of its conservation status and lack of data regarding the ecology and life history, particularly about population trends, we present data on relative abundance of and habitat use by the West Virginia Spring Salamander during a 33-year period from 1975–2008. -
A New Maximum Body Size Record for the Berry Cave Salamander
A peer-reviewed open-access journal Subterranean Biology 28:A new29–38 maximum (2018) body size record for the Berry Cave Salamander... 29 doi: 10.3897/subtbiol.28.30506 SHORT COMMUNICATION Subterranean Published by http://subtbiol.pensoft.net The International Society Biology for Subterranean Biology A new maximum body size record for the Berry Cave Salamander (Gyrinophilus gulolineatus) and genus Gyrinophilus (Caudata, Plethodontidae) with a comment on body size in plethodontid salamanders Nicholas S. Gladstone1, Evin T. Carter2, K. Denise Kendall Niemiller3, Lindsey E. Hayter4, Matthew L. Niemiller3 1 Department of Earth and Planetary Sciences, University of Tennessee, Knoxville, Tennessee 37916, USA 2 Department of Ecology and Evolutionary Biology, University of Tennessee, Knoxville, Tennessee 37916, USA 3 Department of Biological Sciences, The University of Alabama in Huntsville, Huntsville, Alabama 35899, USA 4 Admiral Veterinary Hospital, 204 North Watt Road, Knoxville, Tennessee 37934, USA Corresponding author: Matthew L. Niemiller ([email protected]) Academic editor: O. Moldovan | Received 12 October 2018 | Accepted 23 October 2018 | Published 16 November 2018 http://zoobank.org/2BCA9CF1-52C7-47B6-BDFB-25DCF9EA487A Citation: Gladstone NS, Carter ET, Niemiller KDK, Hayter LE, Niemiller ML (2018) A new maximum body size record for the Berry Cave Salamander (Gyrinophilus gulolineatus) and genus Gyrinophilus (Caudata, Plethodontidae) with a comment on body size in plethodontid salamanders. Subterranean Biology 28: 29–38. https://doi.org/10.3897/ subtbiol.28.30506 Abstract Lungless salamanders in the family Plethodontidae exhibit an impressive array of life history strategies and occur in a diversity of habitats, including caves. However, relationships between life history, habitat, and body size remain largely unresolved.