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New Crustacean Records from the Late Campanian of the Gschliefgraben

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New Crustacean Records from the Late Campanian of the Gschliefgraben ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Beiträge zur Paläontologie Jahr/Year: 1999 Band/Volume: 24 Autor(en)/Author(s): Fraaye Rene H. B., Summesberger Herbert Artikel/Article: New crustacean records from the Late Campanian of the Gschliefgraben (Cretaceous, Austria) 1-6 ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien Beitr. Paläont., 24:1-6, Wien 1999 New crustacean records from the Late Campanian of the Gschliefgraben (Cretaceous, Austria) by René H.B. F raaye 1 & Herbert Summesberger * 2 F raaye , R.H.B & S ummesberger , H., 1999. New Crustacean records from the Late Campanian of the Gschliefgraben (Cretaceous, Austria). — Beitr. Palàont., 24:1-6, 1 Plate, Wien. Abstract Cenomanian rudist limestones of NE Italy (Collins & D ieni , 1995). Three species of crustacean, two malacostracans and In 1882 von A mmon described the first fossil isopods one cirripede, are described from the Late Campanian from Austria from the Oligocène (H essler , 1969). The Polyplocum Zone of the Gschliefgraben, Gmunden oldest Austrian isopods were described by B achmayer (Austria). Palaega huetteri n. sp. is the first Cretaceous (1949) from late Jurassic limestones of Ernstbrunn. isopod to be reported from Austria. A specimen of the Two years earlier, B achmayer (1947) had reported on scalpellid cirripede A rcoscalpellum maximum sulcatum two new isopods from the Miocene of Deutsch- (J. de C. S owerby , 1829) preserved in the body cham­ Altenburg. In 1950, Tauber added another Miocene ber of the large heteromorph Pseudoxybeloceras isopod species from the Vienna Basin. Palaega huetteri (Schlueterella) pseudoarmatum (S chlüter , 1872) is n. sp. is the first Cretaceous isopod from central Eu­ considered to represent stomach contents of the am­ rope. No late Cretaceous cirripedes have been described monite. The co-occurrence of the giant isopod Palaega previously from Austria. For more detailed stratigraphic huetteri n. sp. and the presumable swimming crab and palaeontological information of the Gschliefgraben IXanthilites sp. is indicative of a relatively deep area we refer to Prey (1983), Kennedy & S ummesber ­ palaeoenvironment. ger (1984), Christensen (1998), K ennedy & S ummes ­ berger (this volume), Jagt (this volume) and Trôger , Introduction S ummesberger & S koumal (this volume). Conventions All studied specimens are depos­ Records of Cretaceous crustaceans from Austria are ited in the Naturhistorisches Museum Wien, Austria rather scanty. In 1931, Glaessner mentioned a crusta­ (NHMW registration numbers, ex Hiitter collection). cean fauna from a presumed late Jurassic limestone from the non-alpine Waschberg belt, near Klement. A Systematic Palaeontology recent re-examination by W right (1997) of this small crab faunule revealed it to be of Cenomanian age com­ Class Cirripedia B urmeister , 1834 prising Rathbunopon obesum (V an Straelen , 1944), Order Thoracica D arwin , 1851 Pithonoton cenomanense (W right & Collins , 1972), Suborder Lepadomorpha Pilsbry , 1916 Palaeodromites incertus (B ell, 1863) and Diaulax Family Scalpellidae Pilsbry , 1916 oweni (B ell, 1863). A chela of Dromiopsis sp. was re­ Subfamily Arcoscalpellinae Zevina , 1978 corded by G ross (1984) from presumed allochthonous Santonian limestones. Morphologically, this specimen Genus Arcoscalpellum H oek, 1907 is almost identical with the Dromiopsis chelae known Arcoscalpellum maximum sulcatum from the Campanian of northern Germany. W ank (J. de C. S owerby , 1829) (1986) mentioned the presence of the crab Cyomocarci- nus [sic] cf. angustifrons from the Santonian of Klein (Plate 1, Fig. A, C) St. Paul (Carinthia). This specimen is in need of re­ 1829 Pollicipes sulcatus J. de C. Sowerby , p. 221, pi. 606, examination; it most probably belongs to the genus fig. 2. Graptocarcinus, an identification which is supported 1935 Scalpellum (Arcoscalpellum ) maximum var. sulcatum by M üller (1998) which has been recorded from the (J. de C. Sowerby ); Withers , p. 253, pi. 32, figs. 5- 8. 1953 Scalpellum (Arcoscalpellum) maximum var. sulcatum Oertijdmuseum, De Groene Poort, Bosscheweg 80, (J.de C. Sowerby ); Carlsson , p. 20, pi. 5, fig. 1. NL-5283 WB, Boxtel, The Netherlands 2 Naturhistorisches Museum, Burgring7, A-1014 Wien, Material: The body segments of what appears to Austria be a single specimen is preserved in a portion of the ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien 2 Beitr. Paläont. 24, Wien 1999 relatively large body chamber of the ammonite Pseu- with axial keel on the anterior half with tripartite end­ doxybeloceras (Schlueterella) pseudoarmatum ing in the centre; pleotelson and five partially preserved (S chlüter , 1872) (NHMW 1998z29/ll). Preserved are pleonites are covered with fine, more or less uniform the carina, both terga and one scutum. dense punctae. Pleotelson slightly wider (c.3.1 cm) than Description For a detailed description of this long (c. 2.8 cm); posterior border with 17 spines, me­ taxon reference is made to W ithers (1935). The dian spine very large (c. 3 mm), others smaller (c. 0.5 Gschliefgraben specimen most closely resembles the mm) and more or less of equal size; anterior border specimens from the late Campanian of Norwich, Eng­ irregularly covered with numerous small incisions. land. Pleonites subequal in width and length, with posteriorly Remarks Examples of late Cretaceous cirripedes pointed pleurae. Smooth ventral side of partially pre­ preserved within the body chambers of relatively large served right uropod. ammonites have been recorded by several authors (e.g. Etymology This species is named after Fritsch & K afka , 1887, E rnst , 1967; Collins , 1986; Mag. Herbert H otter , who donated not only the three O ekentorp , 1989; H auschke , 1994). In all of these specimens described herein but also his complete cases only cirripedes of the family Stramentidae W ith­ Gschliefgraben collection to the Museum of Natural ers , 1920 were involved. History, Vienna. Although the majority of these cirripedes have been Remarks Palaega huetteri n.sp. differs from all found inside or attached to the bodychambers of am­ species known to date in having 17 spines on, and a monites, they have mostly been interpreted as hitch­ tripartite subdivision of, the pleotelson. Cenozoic forms hikers on living ammonites or as epibionts on/in empty have only 11 or fewer spines. Of all Cretaceous forms, ammonite shells (benthic islands). O ekentorp (1989) the overall morphology, ornament and especially was the only author who considered the possibility, that pleotelson morphology of P. huetteri n. sp. most closely the lepadomorph cirripedes could represent stomach resembles P. guadalupensis from the Coniacian-Cam- contents of the ammonites. The Gschliefgraben panian of Texas, USA (Rathbun , 1935; W ieder & cirripede is situated in the posterior half of the body Feldmann , 1992). chamber, as do the majority of the above mentioned Palaega guadalupensis differs in having 21 small teeth examples. Jäger & F raaye (1997) reported portions on the posterior border of the pleotelson. Recent of chelipeds and abdominal segments of decapod crus­ Palaega species occur at depths between 70 and 2140 taceans as ammonite stomach contents. The majority metres, with an acme between 100 and 800 metres of these remains have been found in the posterior half (K arasawa , N obuhara & M atsuoka , 1992). of the body chambers. This fact and O ekentorp ’s re­ The occurrence of P. huetteri n.sp. in the present faunule ported size-relationship (larger cirripedes are found in thus corresponds well to a relatively deep and open larger body chambers and smaller cirripedes in smaller marine palaeoenvironment. ones) are seen as evidence in favour of the view that most “in ammonite” cirripedes are in fact ammonite Order Decapoda Latreille , 1803 stomach contents. Infraorder Brachyura L atreille , 1803 The genus Arcoscalpellum is common in the late Cre­ Section Brachyrhynchia B orradaille , 1907 taceous of Europe. Arcoscalpellum maximum sulcatum Superfamily Xanthoidea D ana , 1852 is known from the Campanian and Maastrichtian of Family Xanthidae D ana , 1852 Denmark, England, France, Germany (W ithers , 1935), ? Xanthilites sp. Sweden (Carlsson , 1953), Belgium (Jagt & Collins , (Plate 1, Fig. B) 1989) and the Czech Republic (Fritsch & Kafka , 1887) Material The only known specimen (NHMW Class Malacostraca L atreille , 1806 1998z42/l) is the incomplete ventral part of a brach- Order Isopoda L atreille , 1817 yuran decapod. Suborder Flabellifera G.O. Sars , 1882 Description: Although the ventral part of the Family Cirolanidae D ana , 1852 specimen is very well preserved it lacks the carapace, Genus Palaega W oodward , 1870 which must have been wider than long as deduced from the preserved ventral parts (abdomen, stemites, legs Palaega huetteri n.sp. and mouth parts). Left chela almost twice as large as (Plate 1, Fig. D) right one. Both chelae uniformly covered with fine Material Holotype and sole specimen (NHMW pustules. Elongated fingers with curved smooth tips 1998z42/2), an external mould lacking the anterior part, and cutting edge darker coloured. Fixed finger gently from the late Campanian (Polyplocum Zone) of curving down- and inwards with a longitudinal shal­ Gschliefgraben, Gmunden (Austria). low groove extending from the tip to the middle of Description: Large-sized palaegid; inner side of three cusps. Longer dactylus
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