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Evolutionary Morphology of the Rabbit Skull
Evolutionary morphology of the rabbit skull Brian Kraatz1 and Emma Sherratt2 1 Department of Anatomy, Western University of Health Sciences, United States 2 Department of Evolution, Ecology and Genetics, Research School of Biology, The Australian National University, Canberra, ACT, Australia ABSTRACT The skull of leporids (rabbits and hares) is highly transformed, typified by pronounced arching of the dorsal skull and ventral flexion of the facial region (i.e., facial tilt). Previous studies show that locomotor behavior influences aspects of cranial shape in leporids, and here we use an extensive 3D geometric morphometrics dataset to further explore what influences leporid cranial diversity. Facial tilt angle, a trait that strongly correlates with locomotor mode, significantly predicts the cranial shape variation captured by the primary axis of cranial shape space, and describes a small proportion (13.2%) of overall cranial shape variation in the clade. However, locomotor mode does not correlate with overall cranial shape variation in the clade, because there are two district morphologies of generalist species, and saltators and cursorial species have similar morphologies. Cranial shape changes due to phyletic size change (evolutionary allometry) also describes a small proportion (12.5%) of cranial shape variation in the clade, but this is largely driven by the smallest living leporid, the pygmy rabbit (Brachylagus idahoensis). By integrating phylogenetic history with our geometric morphometric data, we show that the leporid cranium exhibits weak phylogenetic signal and substantial homoplasy. Though these results make it difficult to reconstruct what the `ancestral' leporid skull looked like, the fossil records suggest that dorsal arching and facial tilt could have occurred before the origin of the crown group. -
Field Guide Mammals of Ladakh ¾-Hðgå-ÅÛ-Hýh-ºiô-;Ým-Mû-Ç+Ô¼-¾-Zçàz-Çeômü
Field Guide Mammals of Ladakh ¾-hÐGÅ-ÅÛ-hÝh-ºIô-;Ým-mÛ-Ç+ô¼-¾-zÇÀz-Çeômü Tahir Shawl Jigmet Takpa Phuntsog Tashi Yamini Panchaksharam 2 FOREWORD Ladakh is one of the most wonderful places on earth with unique biodiversity. I have the privilege of forwarding the fi eld guide on mammals of Ladakh which is part of a series of bilingual (English and Ladakhi) fi eld guides developed by WWF-India. It is not just because of my involvement in the conservation issues of the state of Jammu & Kashmir, but I am impressed with the Ladakhi version of the Field Guide. As the Field Guide has been specially produced for the local youth, I hope that the Guide will help in conserving the unique mammal species of Ladakh. I also hope that the Guide will become a companion for every nature lover visiting Ladakh. I commend the efforts of the authors in bringing out this unique publication. A K Srivastava, IFS Chief Wildlife Warden, Govt. of Jammu & Kashmir 3 ÇSôm-zXôhü ¾-hÐGÅ-mÛ-ºWÛG-dïm-mP-¾-ÆôG-VGÅ-Ço-±ôGÅ-»ôh-źÛ-GmÅ-Å-h¤ÛGÅ-zž-ŸÛG-»Ûm-môGü ¾-hÐGÅ-ÅÛ-Å-GmÅ-;Ým-¾-»ôh-qºÛ-Åï¤Å-Tm-±P-¤ºÛ-MãÅ-‚Å-q-ºhÛ-¾-ÇSôm-zXôh-‚ô-‚Å- qôºÛ-PºÛ-¾Å-ºGm-»Ûm-môGü ºÛ-zô-P-¼P-W¤-¤Þ-;-ÁÛ-¤Û¼-¼Û-¼P-zŸÛm-D¤-ÆâP-Bôz-hP- ºƒï¾-»ôh-¤Dm-qôÅ-‚Å-¼ï-¤m-q-ºÛ-zô-¾-hÐGÅ-ÅÛ-Ç+h-hï-mP-P-»ôh-‚Å-qôº-È-¾Å-bï-»P- zÁh- »ôPÅü Åï¤Å-Tm-±P-¤ºÛ-MãÅ-‚ô-‚Å-qô-h¤ÛGÅ-zž-¾ÛÅ-GŸôm-mÝ-;Ým-¾-wm-‚Å-¾-ºwÛP-yï-»Ûm- môG ºô-zôºÛ-;-mÅ-¾-hÐGÅ-ÅÛ-h¤ÛGÅ-zž-Tm-mÛ-Åï¤Å-Tm-ÆâP-BôzÅ-¾-wm-qºÛ-¼Û-zô-»Ûm- hôm-m-®ôGÅ-¾ü ¼P-zŸÛm-D¤Å-¾-ºfh-qô-»ôh-¤Dm-±P-¤-¾ºP-wm-fôGÅ-qºÛ-¼ï-z-»Ûmü ºhÛ-®ßGÅ-ºô-zM¾-¤²h-hï-ºƒÛ-¤Dm-mÛ-ºhÛ-hqï-V-zô-q¼-¾-zMz-Çeï-Çtï¾-hGôÅ-»Ûm-môG Íï-;ï-ÁÙÛ-¶Å-b-z-ͺÛ-Íïw-ÍôÅ- mGÅ-±ôGÅ-Åï¤Å-Tm-ÆâP-Bôz-Çkï-DG-GÛ-hqôm-qô-G®ô-zô-W¤- ¤Þ-;ÁÛ-¤Û¼-GŸÝP.ü 4 5 ACKNOWLEDGEMENTS The fi eld guide is the result of exhaustive work by a large number of people. -
World Distribution of the European Rabbit (Oryctolagus Cuniculus)
1 The Evolution, Domestication and World Distribution of the European Rabbit (Oryctolagus cuniculus) Luca Fontanesi1*, Valerio Joe Utzeri1 and Anisa Ribani1 1Department of Agricultural and Food Sciences, Division of Animal Sciences, University of Bologna, Italy 1.1 The Order Lagomorpha to assure essential vitamin uptake, the digestion of the vegetarian diet and water reintroduction The European rabbit (Oryctolagus cuniculus, (Hörnicke, 1981). Linnaeus 1758) is a mammal belonging to the The order Lagomorpha was recognized as a order Lagomorpha. distinct order within the class Mammalia in Lagomorphs are such a distinct group of 1912, separated from the order Rodentia within mammalian herbivores that the very word ‘lago- which lagomorphs were originally placed (Gidely, morph’ is a circular reference meaning ‘hare- 1912; Landry, 1999). Lagomorphs are, however, shaped’ (Chapman and Flux, 1990; Fontanesi considered to be closely related to the rodents et al., 2016). A unique anatomical feature that from which they diverged about 62–100 million characterizes lagomorphs is the presence of years ago (Mya), and together they constitute small peg-like teeth immediately behind the up- the clade Glires (Chuan-Kuei et al., 1987; Benton per-front incisors. For this feature, lagomorphs and Donoghue, 2007). Lagomorphs, rodents and are also known as Duplicidentata. Therefore, primates are placed in the major mammalian instead of four incisor teeth characteristic of clade of the Euarchontoglires (O’Leary et al., 2013). rodents (also known as Simplicidentata), lago- Modern lagomorphs might be evolved from morphs have six. The additional pair is reduced the ancestral lineage from which derived the in size. Another anatomical characteristic of the †Mimotonidae and †Eurymilydae sister taxa, animals of this order is the presence of an elong- following the Cretaceous-Paleogene (K-Pg) bound- ated rostrum of the skull, reinforced by a lattice- ary around 65 Mya (Averianov, 1994; Meng et al., work of bone, which is a fenestration to reduce 2003; Asher et al., 2005; López-Martínez, 2008). -
Violations in the Name of Conservation “What Crime Had I Committed by Putting My Feet on the Land That I Own?”
VIOLATIONS IN THE NAME OF CONSERVATION “WHAT CRIME HAD I COMMITTED BY PUTTING MY FEET ON THE LAND THAT I OWN?” Amnesty International is a movement of 10 million people which mobilizes the humanity in everyone and campaigns for change so we can all enjoy our human rights. Our vision is of a world where those in power keep their promises, respect international law and are held to account. We are independent of any government, political ideology, economic interest or religion and are funded mainly by our membership and individual donations. We believe that acting in solidarity and compassion with people everywhere can change our societies for the better. © Amnesty International 2021 Except where otherwise noted, content in this document is licensed under a Creative Commons Cover photo: Illustration by Colin Foo (attribution, non-commercial, no derivatives, international 4.0) licence. Photo: Chitwan National Park, Nepal. © Jacek Kadaj via Getty Images https://creativecommons.org/licenses/by-nc-nd/4.0/legalcode For more information please visit the permissions page on our website: www.amnesty.org Where material is attributed to a copyright owner other than Amnesty International this material is not subject to the Creative Commons licence. First published in 2021 by Amnesty International Ltd Peter Benenson House, 1 Easton Street London WC1X 0DW, UK Index: ASA 31/4536/2021 Original language: English amnesty.org CONTENTS 1. EXECUTIVE SUMMARY 5 1.1 PROTECTING ANIMALS, EVICTING PEOPLE 5 1.2 ANCESTRAL HOMELANDS HAVE BECOME NATIONAL PARKS 6 1.3 HUMAN RIGHTS VIOLATIONS BY THE NEPAL ARMY 6 1.4 EVICTION IS NOT THE ANSWER 6 1.5 CONSULTATIVE, DURABLE SOLUTIONS ARE A MUST 7 1.6 LIMITED POLITICAL WILL 8 2. -
Factors Influencing Snowshoe Hare (Lepus Americanus) in Michigan
Factors influencing snowshoe hare (Lepus americanus) in Michigan By David Michael Burt A THESIS Submitted to Michigan State University in partial fulfillment of the requirements for the degree of Fisheries and Wildlife – Master of Science 2014 ABSTRACT Factors influencing snowshoe hare (Lepus americanus) in Michigan By David Michael Burt The goal of my thesis was to identify the climatic and vegetation factors that influence snowshoe hare occupancy in Michigan. My objectives were to: 1) quantify snowshoe hare occupancy as related to climate change and 2) quantify the relationships between patch-level snowshoe hare occupancy and land cover change over time, current land cover, habitat structure, and mesocarnivore presence. In Chapter 1, I determined the most efficient transect configurations for winter track surveys and found that transects 150m in length with 100 or 75m spacing, or 125m in length with 75m spacing provided reliable estimates of hare occupancy. In Chapter 2, I researched climatic variables that are potentially linked to snowshoe hare population performance and assessed whether those factors affected the localized extinction of snowshoe hares. I found that localized extinction was influenced by maximum temperature from May 15 – January 19; as temperature increased the likelihood of localized extinction increased. I also found that the total number of days with measurable snow on the ground affected localized extinction; as number of days with snow on the ground decreased the likelihood of localized extinction increased. In Chapter 3, I evaluated whether land cover change over time, current land cover, and habitat factors affected snowshoe hare occupancy and habitat use. I found that land cover change over time did not affect hare occupancy. -
Appendix Lagomorph Species: Geographical Distribution and Conservation Status
Appendix Lagomorph Species: Geographical Distribution and Conservation Status PAULO C. ALVES1* AND KLAUS HACKLÄNDER2 Lagomorph taxonomy is traditionally controversy, and as a consequence the number of species varies according to different publications. Although this can be due to the conservative characteristic of some morphological and genetic traits, like general shape and number of chromosomes, the scarce knowledge on several species is probably the main reason for this controversy. Also, some species have been discovered only recently, and from others we miss any information since they have been first described (mainly in pikas). We struggled with this difficulty during the work on this book, and decide to include a list of lagomorph species (Table 1). As a reference, we used the recent list published by Hoffmann and Smith (2005) in the “Mammals of the world” (Wilson and Reeder, 2005). However, to make an updated list, we include some significant published data (Friedmann and Daly 2004) and the contribu- tions and comments of some lagomorph specialist, namely Andrew Smith, John Litvaitis, Terrence Robinson, Andrew Smith, Franz Suchentrunk, and from the Mexican lagomorph association, AMCELA. We also include sum- mary information about the geographical range of all species and the current IUCN conservation status. Inevitably, this list still contains some incorrect information. However, a permanently updated lagomorph list will be pro- vided via the World Lagomorph Society (www.worldlagomorphsociety.org). 1 CIBIO, Centro de Investigaça˜o em Biodiversidade e Recursos Genéticos and Faculdade de Ciˆencias, Universidade do Porto, Campus Agrário de Vaira˜o 4485-661 – Vaira˜o, Portugal 2 Institute of Wildlife Biology and Game Management, University of Natural Resources and Applied Life Sciences, Gregor-Mendel-Str. -
Lagomorphs: Pikas, Rabbits, and Hares of the World
LAGOMORPHS 1709048_int_cc2015.indd 1 15/9/2017 15:59 1709048_int_cc2015.indd 2 15/9/2017 15:59 Lagomorphs Pikas, Rabbits, and Hares of the World edited by Andrew T. Smith Charlotte H. Johnston Paulo C. Alves Klaus Hackländer JOHNS HOPKINS UNIVERSITY PRESS | baltimore 1709048_int_cc2015.indd 3 15/9/2017 15:59 © 2018 Johns Hopkins University Press All rights reserved. Published 2018 Printed in China on acid- free paper 9 8 7 6 5 4 3 2 1 Johns Hopkins University Press 2715 North Charles Street Baltimore, Maryland 21218-4363 www .press .jhu .edu Library of Congress Cataloging-in-Publication Data Names: Smith, Andrew T., 1946–, editor. Title: Lagomorphs : pikas, rabbits, and hares of the world / edited by Andrew T. Smith, Charlotte H. Johnston, Paulo C. Alves, Klaus Hackländer. Description: Baltimore : Johns Hopkins University Press, 2018. | Includes bibliographical references and index. Identifiers: LCCN 2017004268| ISBN 9781421423401 (hardcover) | ISBN 1421423405 (hardcover) | ISBN 9781421423418 (electronic) | ISBN 1421423413 (electronic) Subjects: LCSH: Lagomorpha. | BISAC: SCIENCE / Life Sciences / Biology / General. | SCIENCE / Life Sciences / Zoology / Mammals. | SCIENCE / Reference. Classification: LCC QL737.L3 L35 2018 | DDC 599.32—dc23 LC record available at https://lccn.loc.gov/2017004268 A catalog record for this book is available from the British Library. Frontispiece, top to bottom: courtesy Behzad Farahanchi, courtesy David E. Brown, and © Alessandro Calabrese. Special discounts are available for bulk purchases of this book. For more information, please contact Special Sales at 410-516-6936 or specialsales @press .jhu .edu. Johns Hopkins University Press uses environmentally friendly book materials, including recycled text paper that is composed of at least 30 percent post- consumer waste, whenever possible. -
Groundwater Research in NEPAL for Tiger Conservation
GROUNDWATER RESEARCH IN NEPAL FOR TIGER CONSERVATION A reconnaissance study to groundwater dynamics in an alluvial mega-fan in Bardiya National Park (Terai), focusing on the interaction between groundwater and the Karnali river. Author: Hanne. Berghuis. MSc. Thesis. Program: Earth, Surface and Water at Utrecht University. 1st Supervisor: Prof. Dr. Jasper Griffioen. 2nd Supervisor: Dr. Derek Karssenberg. Date: 28-06-2019. Student No.: 6190987. Contact: [email protected]. Photo credits: Esther Leystra (2019). Nepal: Bardiya National Park. Acknowledgement I’d like to thank my supervisor Jasper Griffioen for the opportunity to hydrologically explore Bardiya. His enthusiasm for the project was inspiring and his close involvement was very motivating. My friend Ewa van Kooten introduced me to this project. Together we travelled to Nepal for three months. Thanks to her I enjoyed every single day of our time in Bardiya. She often came up with new ideas for field measurements, creative ways to fabricate field equipment or interpretations for unexpected observations. I am grateful for the Himalayan Tiger Foundation (HTF), who took the initial initiative for hydrological research in Bardiya. I very much appreciate their efforts for the conservation of the wild tiger. During the meetings in the Netherlands and around the campfire in Bardiya with the members and co of HTF, I have learned and laughed a lot. Moreover, I like to thank them for getting us in touch with the National Trust for Nature Conservation (NTNC). The staff of NTNC heartily welcomed us in Bardiya and at their office. They made us feel like a part of the NTNC-family by letting us join their festivals, dinners and campfires. -
Review Paper Status of Tiger and Its Conservation Efforts in Nepal
International Journal of Global Science Research ISSN: 2348-8344 (Online) Vol. 7, Issue 1, April 2020, pp. 1277-1283 DOI: 10.26540/ijgsr.v7.i1.2020.149 Available Online at www.ijgsr.com © Copyright 2014 | ijgsr.com | All Rights Reserved Review Paper Status of tiger and its conservation efforts in Nepal: A review Nabina Dhakal1*, Sami Shrestha2 and Jiban Shrestha3 1Agriculture and Forestry University, Rampur, Chitwan, Nepal 2Institute of Forestry, Tribhuvan University, Hariyokharka, Pokhara-15, Nepal 3Nepal Agricultural Research Council, Agriculture Botany Division, Khumaltar, Lalitpur, Nepal *Corresponding author email: [email protected] Received: 01/03/2020 Revised: 14/03/2020 Accepted: 30/03/2020 Abstract: Tiger (Panthera tigris) is one of INTRODUCTION the most endangered wildlife species in Tiger (Panthera tigris) is considered as Nepal. Currently there are 235 tigers symbol of strength, mysterious and noble counted in Nepal. The areas of habitat of beings. It is the largest cat (feline) species tigers have been extended from 4502.5 km2 with pattern of dark vertical stripes on (before 2010) to 6167.12 km2 (after 2018). reddish-orange fur with a lighter underside Five national parks (Chitwan, Bardia, along with muscular body, powerful Banke, Shuklaphata and Parsa National forelimbs, large head and a tail that is about Parks), four protection forest (Brandabhar, half the length of the body. They are Khata, Baanta and Laljhadi Protection generally different in size with Forest) and one conservation area distinguishable sexual dimorphism (Krishnasaar Conservation Area) are between males and females, with females preserving tigers in Nepal. Tigers have being smaller than males. A 2016 survey been facing extinction due to poaching, loss found out 3,890 wild tigers and 5,000 of habitat from urbanization and specimens in the United States (WWF, deforestation, and depletion of prey 2018). -
Annual Report DM Last Final Copy.Indd
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Developing Conservation Governance Strategies: Holistic Management of Protected Areas in Nepal Prabhu Budhathoki
DEVELOPING CONSERVATION GOVERNANCE STRATEGIES: HOLISTIC MANAGEMENT OF PROTECTED AREAS IN NEPAL PRABHU BUDHATHOKI A thesis submitted in partial fulfilment of the requirements of the University of Greenwich for the degree of Doctor of Philosophy. April 2012 DECLARATION I certify that this work has not been accepted in substance for any degree, and is not concurrently being submitted for any degree other than that of Doctor of Philosophy being studied at the University of Greenwich. I also declare that this work is the result of my own investigations except where otherwise identified by references and that I have not plagiarized the work of others. Candidate: Prabhu Budhathoki …………………………………… Supervisor: Dr. S. Russell ……………………………………. Supervisor: Dr. M. J. McGibbon …………………………………… ii ACKNOWLEDGEMENTS This study could not have been completed without the support and encouragement from numerous individuals and institutions. I would like to take this opportunity to express my sincere gratitude to those who have contributed directly and indirectly to the accomplishment of this study. Firstly, I am deeply indebted to Dr. Bill Jordan, former President of the Care for the Wild and founder President of the Bill Jordon Wildlife Defence Fund, UK, for providing me the necessary financial support to carry out this study. Similarly, I am grateful to my principal supervisor Dr. Shaun Russell, Director, Wales Environment Research Hub, Bangor University, for arranging the fund for my study and travels. He also deserves my special gratitude for his excellent guidance and scholarly support in organising and improving the document. My special appreciation also goes to Dr. Elizabeth Hughes, late Executive Director, International Centre for Protected Landscape (ICPL), for her support, encouragement and intellectual inputs throughout my study. -
Lagomorphs: Pikas, Rabbits, and Hares of the World
LAGOMORPHS 1709048_int_cc2015.indd 1 15/9/2017 15:59 1709048_int_cc2015.indd 2 15/9/2017 15:59 Lagomorphs Pikas, Rabbits, and Hares of the World edited by Andrew T. Smith Charlotte H. Johnston Paulo C. Alves Klaus Hackländer JOHNS HOPKINS UNIVERSITY PRESS | baltimore 1709048_int_cc2015.indd 3 15/9/2017 15:59 © 2018 Johns Hopkins University Press All rights reserved. Published 2018 Printed in China on acid- free paper 9 8 7 6 5 4 3 2 1 Johns Hopkins University Press 2715 North Charles Street Baltimore, Maryland 21218-4363 www .press .jhu .edu Library of Congress Cataloging-in-Publication Data Names: Smith, Andrew T., 1946–, editor. Title: Lagomorphs : pikas, rabbits, and hares of the world / edited by Andrew T. Smith, Charlotte H. Johnston, Paulo C. Alves, Klaus Hackländer. Description: Baltimore : Johns Hopkins University Press, 2018. | Includes bibliographical references and index. Identifiers: LCCN 2017004268| ISBN 9781421423401 (hardcover) | ISBN 1421423405 (hardcover) | ISBN 9781421423418 (electronic) | ISBN 1421423413 (electronic) Subjects: LCSH: Lagomorpha. | BISAC: SCIENCE / Life Sciences / Biology / General. | SCIENCE / Life Sciences / Zoology / Mammals. | SCIENCE / Reference. Classification: LCC QL737.L3 L35 2018 | DDC 599.32—dc23 LC record available at https://lccn.loc.gov/2017004268 A catalog record for this book is available from the British Library. Frontispiece, top to bottom: courtesy Behzad Farahanchi, courtesy David E. Brown, and © Alessandro Calabrese. Special discounts are available for bulk purchases of this book. For more information, please contact Special Sales at 410-516-6936 or specialsales @press .jhu .edu. Johns Hopkins University Press uses environmentally friendly book materials, including recycled text paper that is composed of at least 30 percent post- consumer waste, whenever possible.