CHAPTER I I I Sistematic DESCRIPTION
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CHAPTER III SiSTEMATIC DESCRIPTION CIUPTER III systematic description Section I : Mortonlceratinae Introduction : The ammonoid species described by Stoliczka under the group of Ammonites crlstatl were sub divided by him into species 1, Ammonites blanfordianus, 2, Ammonites inflatus, condollianus. ootatoorensis and corruptus, 3, Amnionites propinquus. 4. Ammonites obesus, 5. Ammonites se rratocarlnatus. 6, Ammonite s subtricarinatus. These were transferred by later workers e.g. Kossmat to Schloenbachia and its subgenera as : Schloenbachla (Tropitoldes) . S. (Prlnotropis) . S. (Peroniceras). S, (Munerlceras) etc. According to present ideas the above mentioned species have undergone further transfers and except those of the Inflata group, are placed under different subfamilies, viz. Peroniceratlnae, Mune rice rat inae, etc. These species are, therefore, not considered under the present section; but those of the Inflata group which are now found to belong to the sub family Mortonlceratinae are dealt with here. The species which represent this subfamily in the present collection, viz. Mortoniceras (Mortoniceras) inflatum (Sow.); M. (M.) rostratum (Sow.), Mortonioeras (Peiradoceras) indicum spo nov., M. (Purnovarites) perinflatum Spath., M. (p.) - 17 - - 18 - subquadratum Spath, M. (d .) sp. Indet; ?Mortonicera8 (?Mortoniceras) deshpandel sp, nov., Prohysteroceras (Goodhallltes) stoliczkal sp. nov. are dealt with here., Of these eight species Mortonlceras (Mortonlceras) inflatum was described by Stoliczka and Kossmat from these rocks. Three species viz. Mortoniceras (M.) rostratum (Sow.), M. (Durnovarites) perlnflatum Spath., M. (D .) subquadratum Spath. are species already known and are recorded from outside South Indian basin; they are now recorded here for the first time. The remaining species described here are new to science. The subfamily Mortoniceratinae has an interesting history, being used in the first instance by Spath (1925) to comprise species of texanum group of Senonian. Workers have differed on the position of Ammonites vespertinus Morton, the genotype of Mortoniceras, and its exact horizon was also doubtful. Adkins in 1928, however, established its horizon as Albian, and hence it was grouped with other related ammonoids which were included in Inflacticeratidae of Spath (1925) by some, and by others in Pervinquieridae of Spath (l926). Later, Spath (1932) proposed a new name, Texanites for the Mortoniceras like Senonian forms of the texanum group which were included in Peroniceratidae Hyatt. - 19 - This confusion regarding Mortonlceras was partly due to the doubtful or wrongly assigned horizon to Ammonites vespertlnus Morton, and partly due to the homeomorphy between the Alblan and Senonlan forms. According to the taxonomic standards then current this group of fossils was as mentioned earlier, once placed under Schloenbachla by Kossmat (1897) and many others. Wright (1957) while diagnosing this subfamily remarks " As the synonymies of the subfamily and its genera show, both the nomenclature and scale of classification of the group have been in doubt. A large number of species with much the same basic characters vary in the combination of nature of whorl section, number and shape of tubercles, and strength and direction of ribs ......... (p. L405), but time may justify recognition of some genera and subgenera here reduced to synonyms ” . The material at our disposal, however, at present is insufficient to make an attempt at any thing of such a classification; hence for the present we have adopted the various genera and subgenera as accepted in the Treatise . Phylum : Mollusca Class : Cephalopoda Order ; Ammonoidea Sub-order : Ammonitlna - 20 - Super family : Acanthocerataceae Ityatt, 1900 Family : Brancoceratldae Spath, 1933 Sub family ; Mortoniceratinae Spath, 1925 Genus : Mortonleeraa Meek, 1876 Sub-genus : Deiradoceras Van Hoepen, 1931 Mortoniceras (Deiradoceras) indicum sp, nov. PI. I, Figs. 1, 3 Material : One specimen, Holotype Ut l/70 Dimensions; Breadth of whorl section : 58 mm Height of whorl section ; 62 mm B / H =0.93 Description : The present material is fragmentary only the body chamber being preserved. The shell is in general compressed with height almost equal to or a little greater than the breadth of the whorl section. The sides are slightly convex. The venter is fairly broad with a strong and high median keel. The ribs are single, broad, low, prorsiradiate and weak. Each rib has very weakly developed bulloid umbilical tubercle, only one row of ventrolateral tubercles which are strong and clavate and almost vertical (i.e. parallel to the plane of coiling). There are no lateral tubercles. - 21 - Remarks ; Though the present material is only fragmentary still it provides details which help to distinguish it from any of the known comparable species. It is placed in Mortoniceras (Deiradoceras) because of strong ventral keel and the nature of ribbing and the tubercles. The present species is comparable to Ammonites inflatus Sow. var. I Stoliczka (Stoliczka, 1865, p. 48, pi. 27, fig, 1, la, lb) from Utatur group at Utatur and Odium, but can be distinguished from it by much feebler ornament, absence of lateral tubercles and presence of strongly clavate ventro-lateral tubercles which are oriented vertically. The present species may be compared with M. (d .) bipunctatum Spath (Spath, 1931, p. 422-24, pi. XLV, figs. 2a, b; text fig. 145 c, 146) from Upper Gault (Orbignyi and Varicosum - sub zone) England, but differs in having stronger clavate ventrolateral tubercles and prorsiradiate, low, rounded ribs. The low rounded character of ribs is seen also in M. (D.) albense Spath (Spath, 1931, p. 424, pi. XLIII, fig. 2, pi. XLIV, fig. 4) from Orbignyi and Varicosum sub zone of England which, however, does not show sharp clavate ventro lateral tubercles which are parallel to the plane of coiling of the shell and thus is easily distinguished from the present speci es. Occurrence : White clays from Mortoniceras inflatum zone near Utatur. - 22 - Subgenus : Mortonlceras Meek, 1876 Mortonlceras (Mortonlceras) Inflatum ( Sowerby ) PI. I, Figs. 2, 4 1818 Amraonltes Inflatus Sowerby : Sowerby, p. 170, pi. 178 1868 -Amnionltes rostratus Sowerby; Stoliczka, p. 53 1897 Schloenbachla inflate (Sowerty) : Kossmat, p. 185 (parts) 1899 Schloenbachia inflata (Sowerby) : V, Popovici-Hatzeg, p.17 1930 Infla^ticeras aequatoralis (Kossmat) : Besairie, p. 630. 1930 Pervinquleria aequatoralis (Kossmat) : Spath, p. 64 1931 Mortoniceras (Pervinquleria) inflatum (Sowerby): Spath, p. 381, text fig. 128 1931 Mortoniceras (Pervinquleria) rostratum (Sowerby) : Spath, p. 399, text fig. 136 1968 Schloenbachla inflata (Sowerby) ; Sastry, Rao and Mamgain, pi. 1, figs. 1 and 2. Material : Several specimens, Plesiotype Ut 44/70 Dimensions; Width of whorl section : 78 mm Height of whorl section; 78 mm B / H =1.0 only Ftemarks : The Piesiotype is a fragment representing the|^body chamber as it does not show any suture. It is identified with the Sowerby’s species Inflatum as it shows all essential characters of the species, such as a squarish cross section with a rounded intercostal cross section, an arched venter with a rounded strong keel; simple broad, slightly prorsiradiat< - 23 - moderately strong ribs which are separated by wider inter spaces; each rib bearing a bulloid, low and elongated umbilical tubercle, a mediolateral and a strong conical ventrolateral tubercle; pointing almost vertically to the plane of colling; the ribs almost stop at the ventrolateral tubercles and give only faint indication of their oblique continuation towards the ventral keel. Stoliczka has reported this species from the Lower Utatur near Utatur. Occurrence : Gypseous clays of Mortoniceras inflatum zone near Utatur. Distribution ; This species is widely recorded from Upper Alblan of Europe, Africa, North America, South America and Pakistan (Samana Range). Mortoniceras (Mortoniceras) rostratum (Sowerby) PI. II, Figs. 3, 4 1817 Ammonites rostratus Sowerby ; Sowerby, p. 163, pi. clxxlli 1849 Ammonites rostratus Sowerby : Brown, pi. 19, fig. 10 1921 Subschloenbachia gostrata (Sowerby) : Spath, p. 284 1922 Inflactlceras (subschloenbachia) rostratum (Sowerby) : Spath, p. 100 1930 Mortoniceras (Pervinquleria) rostratum (Sowerby) : Spath, p. 400, pi. 36, fig. 6; pi. 38, fig. 4; pi. 39, figs. 4a,b; pi. 40, figs. la,b, 7a,b; pi.42, fig. 2; text figs. 130 f, 136, 137e - 24 - Material : One specimen Ut 68/70 Remarks : The present material is a fragment of the body chamber with part of the rostrum preserved but the umbilical wall is not wholly preserved. The cross section is somewhat rounded and depressed, with maximum width near the umbilical margins; venter comparatively narrow with a strong, high median keel; the anterior part of the shell is curved up and projected up Into a rostrum. Ornamentation consists of simple, recti- radiate ribs stopping at the ventrolateral tubercles separated by interspaces about 3 to 4 times wider than ribs. Each rib with a faint umbilical bulla, a slightly stronger bulloid lateral tubercle and a very strong ventrolateral tubercle. The ventrolateral tubercles on the two sides make an acute angle between themselves and pointing obliquely away from equitorial plane of the shell. Though our material shows a depressed cross- section other characters as described above even in its conditioi leads us to recognize it as Mortoniceras (Mortonioeras) rostraturn (Sowerby) (Spath, 1930, p. 400, text fig, 136) from Upper Albian (Substuderi-and dispar- sub