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Novitates PUBLISHED by the AMERICAN MUSEUM of NATURAL HISTORY CENTRAL PARK WEST at 79TH STREET, NEW YORK, N.Y AMERICAN MUSEUM Novitates PUBLISHED BY THE AMERICAN MUSEUM OF NATURAL HISTORY CENTRAL PARK WEST AT 79TH STREET, NEW YORK, N.Y. 10024 Number 3161, 18 pp., 17 figures, 3 tables April 10, 1996 New Records of Acanthoceratid Ammonoids from the Upper Cenomanian of South Dakota W. J. KENNEDY,1 W. A. COBBAN,2 AND N. H. LANDMAN3 ABSTRACT The upper Cenomanian Metoicoceras mos- thoceras proteus vascoceratoides Wright and Ken- byense zone of the Belle Fourche area in Butte nedy, 1987, and Yezoites n. sp. These species are County, South Dakota, has yielded a number of similar to those reported from the M. mosbyense ammonoid species, some of them new, that show zone in southwestern New Mexico and suggest a both U.S. Westem Interior and northwest Euro- possible influx offorms from the south, migrating pean affinities, including Calycoceras (Calycocer- via the Gulf Coast region, well before the better- as) boreale n. sp., C. (C.) sp., C. (C.) cf. dromense known interchange of Western Interior and more (Thomel, 1972), Calycoceras (Gentoniceras) sp., cosmopolitan taxa in the succeeding Sciponoceras Hamites cimarronensis (Kauffman and Powell, gracile zone. 1977), Neocardioceras transiens n. sp., Protacan- INTRODUCTION The bulk of the ammonoid species known demic species (fig. 1). This is in marked con- from the upper Cenomanian Metoicoceras trast to the predominantly cosmopolitan as- mosbyense zone in the northern part of the semblage known in the succeeding Scipono- Western Interior ofthe United States are en- ceras gracile zone (for a description of this I Curator of the Geological Collections, University Museum, Parks Road, Oxford, OXI 3PW, United Kingdom. 2 Research Scientist Emeritus, U.S. Geological Survey, Box 25046, Mail Stop 919, Federal Center, Denver, CO 80225. 3 Curator and Chairman, Department of Invertebrates, American Museum of Natural History. Copyright C American Museum of Natural History 1996 ISSN 0003-0082 / Price $2.50 2 AMERICAN MUSEUM NOVITATES NO. 3161 zoites n. sp. are not described in the system- SUBSTAGE ZONE atic section of this paper, but will be treated elsewhere.) Nigericeras scott! Of these species, Protacanthoceras proteus vascoceratoides also occurs in northwest Eu- Neocardloceras juddll rope (Wright and Kennedy, 1987). Hamites Burroceras clydense UPPER cimarronensis occurs elsewhere in the West- Sciponoceras gracile ern Interior Seaway including Oklahoma, CENOMANIAN Metolcoceras mosbyense Colorado, and Texas (Kauffman and Powell, Dunveganoceras problematicum 1977). The other species cited are related to Dunveganoceras pond! forms previously reported from New Mexico and northwest Europe. The exact age relationship between the am- Fig. 1. Upper Cenomanian ammonite zona- monoids in South Dakota and those in New tion in the central and northern parts of the U.S. Mexico is difficult to resolve. It is possible Western Interior [modified from Cobban, 1988, that the ammonoids in South Dakota occur table 2] higher up in the Metoicoceras mosbyense zone than those in the south and, therefore, the two occurrences are not exactly equivalent in latter fauna, see Cobban and Scott, 1972). age. Nevertheless, we suggest that the am- Several northwest European and more cos- monoids in South Dakota may have migrated mopolitan species are known from the M. from the south, via the GulfofMexico, rather mosbyense zone in the southernmost parts of than from the north, through Hudson Bay. the Western Interior Seaway (southwestern This hypothesis is based on the presence of New Mexico), where Cobban et al. (1989) related species to the south, albeit not nec- recorded Calycoceras (Proeucalycoceras) essarily of exactly equivalent age, and the guerangeri (Spath, 1926a), Eucalycoceras complete absence of any related species in pentagonum (Jukes-Browne, 1896), Euom- this zone to the north, for example, in central phaloceras euomphalum (Sharpe, 1855), Montana (Cobban, 1952). Hamites cf. H. simplex (d'Orbigny, 1842), and Vascoceras diartianum (d'Orbigny, 1850). Eucalycoceras pentagonum is also present in GEOGRAPHIC AND STRATIGRAPHIC the Sciponoceras gracile zone in southeastern SETTING Colorado as well as in the S. gracile zone in The following localities are referred to in the Black Hills area. the text and indicated in figure 2: It is, therefore, of paleobiogeographic in- AMNH locality 3155-Greenhorn For- terest to record the presence of several am- mation, west side of the Belle Fourche Res- monoid species, one of which also occurs in ervoir in the 51/2 sec. 15, T9N, R3E, Butte northwest Europe, from the Metoicoceras County, South Dakota. mosbyense zone in the northern part of the USGS Mesozoic locality D10511 -Green- Western Interior Seaway. These ammonoids horn Formation, east bank of the Belle occur in the Greenhorn Formation ofthe Belle Fourche Reservoir in the NW/4 sec. 30, T9N, Fourche area, Butte County, South Dakota, R4E, Butte County, South Dakota. considerably farther north than the New USGS Mesozoic locality D10512-same Mexican occurrences cited above, and in- as Dl 0511, but a few meters higher in the clude Calycoceras (Calycoceras) boreale n. sp., section. C. (C.) sp., C. (C.) cf. dromense (Thomel, The stratigraphic sequence at AMNH lo- 1972), Calycoceras (Gentoniceras) sp., Ham- cality 3155 is shown in figure 3. At this lo- ites cimarronensis (Kauffman and Powell, cality, the Greenhorn Formation consists of 1977), Neocardioceras transiens n. sp., Pro- a succession of fine silty, calcareous shales, tacanthocerasproteus vascoceratoides Wright interbedded with layers of flat, ovoid, red- and Kennedy, 1987, and Yezoites n. sp. (The orange weathering concretions less than 30 two heteromorphs, H. cimarronensis and Ye- cm in diameter. The Metoicoceras mosbyense 1 996 KENNEDY ET AL.: ACANTHOCERATID AMMONOIDS 3 in millimeters, with D = diameter, W = whorl width, H = whorl height, and U = umbilical diameter measured at the umbilical wall. The angle of the body chamber (BC L) is ex- pressed in degrees. Suture terminology is that of Wedekind (1916) as expounded by Kull- mann and Wiedmann (1970). Photographs ofspecimens are natural size unless otherwise indicated. Arrows on photographs indicate the adapical end ofthe body chamber, where preserved. The orientations in which speci- mens were photographed are shown in figure 4. SYSTEMATIC PALEONTOLOGY ORDER AMMONOIDEA ZITTEL, 1884 SUBORDER AMMONITINA HYATT, 1889 Fig. 2. Map ofparts ofT9N, R3 and 4E, Butte SUPERFAMILY ACANTHOCERATACEAE County, South Dakota, showing localities men- DE GROSSOUVRE, 1894 tioned in the text. Each section is one square mile. FAMILY ACANTHOCERATIDAE DE GROSSOUVRE, 1894 zone is exposed in the lower part of this sec- SUBFAMILY ACANTHOCERATINAE tion below the Sciponoceras gracile zone. DE GROSSOUVRE, 1894 The ammonoids described in this paper come from an interval 4.9 m above the base GENUS AND SUBGENUS of the measured section, in the Metoicoceras CALYCOCERAS HYATT, 1900 mosbyense zone. They occur in light-grey to TYPE SPECIES: Ammonites navicularis yellowish-grey weathering limestone concre- Mantell, 1822: 198, pl. 22, fig. 5, by desig- tions, less than 15 cm in diameter; most of nation under the Plenary Powers (ICZN these concretions are, in fact, large fragments Opinion no. 557). of ammonoid body chambers, such as those of Calycoceras (Calycoceras) boreale n. sp., Calycoceras (Calycoceras) boreale, with smaller, more complete ammonoids new species embedded inside (fig. 7A). Maeda (1991) Figures 5-9 called this mode of preservation "sheltered preservation" and described similar occur- DERIVATION OF NAME: Latin, borealis, rences from the Upper Cretaceous Yezo northern. Group of Japan. DIAGNOSIS: A coarsely ribbed species of Calycoceras (Calycoceras); on the inner CONVENTIONS whorls, umbilical and inner ventrolateral tu- bercles are approximated on a short rib, with The following abbreviations are used to in- no flank tubercles; ventrolateral shoulders dicate repositories of specimens: occupy most of the whorl height; the venter AMNH American Museum of Natural is broad, with outer ventrolateral and si- History, New York phonal clavi. BMNH British Museum (Natural History), TYPES: Holotype AMNH 44332 (fig. 51, J) London from AMNH locality 3155; paratypes AMNH USNM U.S. National Museum of Natural 44317, 44318, 44329-44331, all from History, Washington, D.C. AMNH locality 3155, and USNM 441253 and 441255, both from USGS Mesozoic lo- All dimensions of specimens are expressed cality D10512. 4 AMERICAN MUSEUM NOVITATES NO. 3161 1 -1 __ * 4- GSciponoceras gracile i-D --- 0I--r_. _ _1 _1L C=) Z I' C:: calcareous shale Fine, silty calcareous shale Red-orange weathenng concretion L 1 PI> Red-orange weathenng concretion, burrowed _= -~~-C--111--~~-~~------~~~~--'-oC= 3::ChalCalarouLight-grey weathenng concretion CL- - °1-1-1-.-- 3L-C5 - - ___ Bioturbated calcarenite ~~1-1°-1-1-1-1-1 m~~Pyrite -MetoicocerasCZ Z gsiia Rdoanewaheigcnceinmbroe F-~~ 4- Metoicoceras geslinianum Fee Z __1__1__1_L. ._..=) 3__10.- _ . 2_ r._ c c~ c._. 7 -------I--4 --1--1-1-1--- 71__1__1__1- Z~.C=() ._ _ Wrhcrsvriuu w __1__1__1_L. 1 :5,C:__ CSC_C= _ 12 Cc 6 'o-o _.eC:,-oc~ CDI,o o 4- Calycoceras (Calycoceras) boreale, C. (C.) cf. dromense, C. (C.) sp., ES1..1 1_.15E Protacanthoceras o ~~~~~~_._.1._.i~._..i. proteus vascoceratoides, Neocardioceras transiens, Hamites cimarronensis, Yezoites sp. a5 C;DzcCM)_...1__C=) CZ)1-D 1__ _ 1 _ _ 1 < Yezoites sp. Fig. 3. The stratigraphic succession in the Greenhorn Formation at AMNH locality 3155 on the west side of the Belle Fourche Reservoir. 1 996 KENNEDY ET AL.: ACANTHOCERATID AMMONOIDS 5 give rise to one or two coarse, strong second- ary ribs. One or two short intercalated ribs appear between the primary ribs both high and low on the ventrolateral shoulder; they strengthen and match the secondary ribs on ventral the venter. All ribs are straight and rectira- diate to slightly rursiradiate and bear weak, outer ventrolateral and siphonal clavi, the latter connected by a delicate siphonal ridge. Paratype AMNH 44331 (fig. 5K, L) is also completely septate; the innermost whorls, Fig. 4. Specimens are illustrated in lateral, visible within the umbilicus ofthe specimen, ventral, and apertural views.
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