BULLETIN OF THE CALIFORNIA SURVEY VOLUME 1, NO. 6

THE OF CALIFORNIA (: )

BY PAUL D. HURD, JR. (Division of Entomology and Parasitology, University of California, Berkeley)

UNIVERSITY OF CALIFORNIA PRESS BERKELEY AND LOS ANGELES 1952 BULLETIN OF THE CALIFORNIA INSECT SURVEY

Editors: E. 0. Essig, S. B. Freeborn, E. G. Linsley, R. L. Usinger

Volume 1, No. 6 pp. 141-152, plates 12 and 13

Submitted by Editors, January 14, 1952

Issued June 27, 1952

Price, 25 cents

UNIVERSITY OF CALIFORNIA PRESS

BERKELEY AND LOS ANGELES

CALIFORNIA

CAMBRIDGE UNIVERSITY PRESS

LONDON, ENGLAND

PRINTED BY OFFSET IN THE UNITED STATES OF AMERICA THE SCOLIIDAE OF CALIFORNIA (Hymenoptera Aculeata)

The members of the family Scoliidae are peared to be incidental to the acquisition of among the larger and more conspicuous of nectar on the part of the . our fauna. Colorationally the sexes are quite dissimilar, the males being more somber than The wasps are frequently encountered in con- their generally larger and more brightly mac- siderable numbers congregating on the flowers of ulated females. The Scoliidae may be recog- various plants in search of nectar. The flowers nized by the very oblique second transverse of Asclepias, Baccharis, Chilopsis, Erigonum, cubitus of the forewing (Plate 12, figs. 2- 5). Lepidospartua, Pectis, Prosopis, and Tamarix seem Additionally, the apical third of the forewing to be among the more attractive floral hosts upon is virtually without venation (Plate 12, figs. which the California species have been collected. 2 - 5). The males possess, at the apex of the abdomen, a retractile trident, a character which The family, an austral intrusive element, is is sufficiently diagnostic to separate them from represented in California by eight species of other scolioid wasps. three genera. Five of these species, three of which are members of the genus , are appar- The species of this family are parasitic ently restricted to the deserts or more arid re- upon the larvae of the . According gions of the state. The two species of Campsom- to Bradley (1945:l) the females do not construct erzs are quite widely distributed in the austral a burrow of their own, but attach their eggs to regions of California, although pilipes is appar- the scarab larvae and prepare a crude cell around ently not present on the California deserts, ex- them leaving the paralyzed grub in situ. Rau and cept perhaps marginally. Two species of the Rau (1918) and Rau (1932) have discussed certain genus Carnpsoscolia are present in California and aspects of the biology of the eastern subspecies are allopatric in their distribution, one of the of’. The Raus report having observed species, flarnrnicoma, being found on the deserts the wasps in a restricted habitat near. St. Louis, is geographically replaced in the coastal and Missouri wherein there was a considerable accum- more northern austral regions of the state by ulation of stable waste. Examination of this alcione. material revealed the presence of many larvae of the scarab, (Linnaeus) -- the sus- The present report is based primarily upon pected host. However, field observations, as the material contained in the following insti- well as rearing attempts, failed to confirm the tutional collections: California Acidemy of suspected host-parasite relationship. The court- Sciences (C.A.S.), California Insect Survey ship dance, mating, and the sleeping behaviour of (C.I.S.), University of California, at Davis this wasp have been described by Rau (1932:59-62) (U.C.D. 1, at Los Angeles (U.C.L.A. ), and at Linsley (1946:27) has reported Carnpsomeris tol- Riverside (U.C.R.). To the individuals in teca (as plumipes) pollinating alfalfa in the charge of these collections as well as those Blythe region of California. Pollination was persons acknowledged elsewhere in the paper I accomplished by tripping the flowers mechanically would like to express my sincere appreciation when alighting or taking flight from a raceme; for the privilege of examining the material in however, pollination of the alfalfa flowers ap- their care.

141 Key to the genera of California Scoliidae

1. Forewing with two recurrent nervures (Plate 12, figs. 2, 3) ...... 2 Forewing with one recurrent nervure (Plate 12, figs. 4, 5) ...... Scolia

2(1) Forewing with two submarginal cells (Plate 12, fig. 3) ...... Canpsorneris Forewing with three submarginal cells (Plate 12, fig. 2) ...... Campsoscolia

Key to the California species of Scolia

1. Forewing with two submarginal cells (Plate 12, fig. 4) ...... 2 Forewing with three submarginal cells (Plate 12, fig. 5)...... ardens ( p. 151)

2(1) Abdominal tergites maculated with yellow; thorax bi- or tri- colorous, never wholly black ...... otornita (p.143) Abdominal tergites not maculated with yellow; thorax wholly black ... dub a haernatodes (p.144)

Key to the California species of

1. Antennae thirteen segmented; abdomen com- glabrous, except on upper margin ...... posed of seven visible segments (males) .. 2 tolteca (p. 144) Antennae twelve segmented; abdomen com- posed of six visible segments (females) . . 3

2(1) Pronoturn with a yellow spot near the hum- eral angle; antennocellar groove present, deeply incised, extending from anterior ocellus nearly to base of interantennal crest, bounded on either side with an impunctate glabrous tubercle ...... pilipes (p. 14.4) Pronotum with a broad yellow transverse band extending to or near the tegulae; antennocellar groove absent, or if pres- ent, short, terminating in a pit, the swelling on each side, if at all evi- dent, weak and punctate throughout ..... tolteca (p.i44j 3(1) Abdomen maculated with yellow bands; pos- terior face of propodeum punctate throughout; densely clothed with long hairs ...... pilipes (p.144) b Abdomen maculated with orange bands; pos- Fig. 1 - a, antenna of Campsoscolia octonacuiata; terior face of propodeum impunctate, b, antenna of Caapsoscolia alcione.

Key to the California species of Campsoscolia

1. Antennae thirteen segmented; abdomen cm- Antennae twelve segmented; abdomen com- posed of seven visible segments(ma1es) .. 2 posed of six visible segments(fema1es) .. 4

142 THE SCOLIIDAE OF CALIFORNIA 143

2(1) Antennae clavate, apical flagellar seg- the level of the face; inner hind tib- ments noticeably enlarged (fig. lb) ..... 3 ial spur not spatulate, scarcely if at Antennae not clavate, apical flagellar all expanded at its apex ...... 5 segment not enlarged (fig. la) ...... Interantennal area scarcely raised above octonaculata I texensis (p. 143 the level of the face; inner hind tib- ial spur spatulate ...... 3(2) Abdominal tergites with basal and apical octoutaculata texensis (p. 145) bands entirely black ...... alcione (p.146) Abdominal tergites with basal and apical 5(4) Abdominal tergites with basal and apical bands ferruginous or predominantly so bands entirely black ...;.. alcione (p.146) flapuzicoma (p. 146) Abdominal tergites with basal and apical bands ferruginous ...... f Zammicoma (p. 146) 4(1) Interantennal area markedly raised above

Discussion of Species

Scdia (Triscolia) ardens Smith' Scolia otomita Saussure2

Scolia ardens Smith, 1855, Cat. Hym. Brit. Mus., Scolia otomita Saussure, 1858, Ann. SOC.Ent. -3;112. France, ser. 3, 6: 223. Type 6.

Geographic range: Geographic range: Mex. , Tex., N. Mex., Ariz. , and Calif. Mexico, N. Mex., Ariz., Nev., and Calif.

California records : California records: RIVERSIDE CO.: Cottonwood Sprine, 6, IV-18- IMPERIAL 0.:Palo Verde, 6, VIII-9-46 35 (C. M. Dammers, U.C.R. 1. Whitewater, 2 6, Y, (P. D. Hurd, Jr. , G.I.S. 1; 2 6, Y, VIII-20-46 M-24-33 (C. M. Dammers, U.C.R.); Y, IX-28-33 (W. F. Barr, C.I.S. 1: 4 6, 9 9, VIII-20-46 (P. D. (C. M. Dammers, U.C.R.); 126, X-10-33 (C. M. Dam- Hurd, Jr., C.1.S.); 2 6, 8 9, VIII-20-46 (J. W. mers, U.C.R.); 17 6, 8 $, VIII-1-34 (C. M. Dam- MacSwain, C.I.S. ); 6,VIII-22-46 (W. F. Barr, C. mers, U.C.R.). Whitewater Canyon, 2 6, $, IX-11- 1.S.); 2 6, VIII-22-46 (P. D. Hurd, Jr., C.I.S.); 35 (P. H. Timberlake, U.C.R.). Y, VIII-27-46 (W. F. Barr, C.I.S.); 6, Y, VIII-27 46 (P. D. Hurd, Jr., C.I.S. 1. SAN BERNAFDINO CO.: Twentynine Palms, 6, 4 Y, RIVERSIDE CO.: Mission Canyon, 6,X-1-32 VIII-3-33 (P. H. Timberlake, C.I.S., U.C.R. 1; Y, (P. H. Timberlake, U.C.R.). Palm Springs, 0, VIII-4-33 (P. H. Timberlake, U.C.R.). VIII-28-34 (P. H. Timberlake, U.C.R. 1. Andreas SAN DIEGO CO.: San Felipe Creek, 3 6, IX-9- Canyon, 2 6, X-27-34 (P. H. Timberlake, U.C.R.). 38 (P. H. Timberlake, U.C.R.). Blythe, 9 2, X-16-39 (C. M. Danuners, U.C.R.); 6, 5 $, VIII-20-46 (.I.W. MacSwain, C.I.S.). u, Discussion: Y, X-15-47 (P. H. Timberlake, U.C.R.). This species is easily distinguished from the SAN -IN0 CD.: Mexican Well, Y, 1X-10- other members of the genus known to occur in Cal- 34 (C. M. DaPmers, U.C.R.). Twentynine Palms, ifornia by the three submarginal cells of the 3 6, $, IX-5-36 (P. H. Timberlake, U.C.R.); Y, forewing. The wings are heavily infuscated with VIII-9-46 (P. H. Timberlake, U.C.R.). black and are brilliantly. violaceous, especially SAN DIMX) CO.: San Felipe Creek, 6, VIII-8- in bright light. The body is black, except for 35 (H. S. Gentry, U.G.D.); 14 Y, M-10-38 (P. H. those abdominal segments posterior to the second Timberlake, U.G.R.); 2 6,X-11-38 (P. €I.Timber- which are reddish-brown. lake, U.G.R. ).

Ardens is apparently restricted in its dis- Discussion: tribution within California to the Mojava and Col- Previously unrecorded from California, this orado Deserts. species appears to be restricted in its occur- rence within the state to the Mojave and Colorado Deserts. It has been observed congregating in fairly large numbers upon the flowers of Pectis

p appo s a ~

'Scolia feruida Burmeister (preoccupied) end S. nonte- 2Scol!a fulutuentris Bartlett is a synonym of this zwae Seussure ere synonyms of this species. species. 144 BULLETIN OF THE CALIFORNIA INSECT SURVEY

Scolia dubia haematodes Burmeister' MONTEREY CO.: Carmel, 6, IV-20-30 (L. S. Slevin, C.A.S.). Carmel Valley, 6,111-29-30 Scolia haematodes Burmeister, 1853, Abhandl. (C. W. Chancey, U.C.R. ). Naturf. Gesell. Halle, 1(4):33. Type 9. NAPA CO.: Sage Canyon, 6, IV-25-48 (L. W. Isaak, U.C.D.). Monticello, 6, IV-10-49 (IV-10-49 Geographic range: (R. C. Bechtel, U.C.D. ). Conn Lake, V-30-50 Mex., Tex., N. Mex., Ariz., and Calif. (R. S. Beal. C.I.S.). ORANGE CO.: Newport Bay, 6, VIII-6-41 (P. D. California records: Hurd, Jr., C.I.S.). Bartlett (1912:321) recorded this species RIVERSIDE CO. : Riverside, 6, IX-10133 from California; however, the present writer has (P. H. Timberlake, U.C.R.). Hemet, Y, VI-6-45 not found any representatives of this species (J. W. MacSwain, C.I.S. ); 6, VIII-22-46 (J. W. from the area under consideration. MacSwain, C.I.S.); 6,VII-5-50 (J. W. MacSwain, C.I.S.). Keen Camp, 6 Y, VI-13-39 (E. G. Linsley, C.I.S.); Keen Camq, 42. E., 6, V-17-39 (B. Campsomeris pilipes (Saussure) Brookman, C.I.S. ). Hemet Reservoir, 2 9, VI-13-39 (E. S. Ross. C.1-S.1: ..6. V-22-39 4J. H. Dorman. Elis pilipes Saussure, 1858, Ann. SOC.Ent. France, C.I.S.); 3, V-23-40 (F. H. Rindge, C.I.S.). Ser. 3, 6:245, 246. Type Y. Vandevanter Flat, Y, VI-12-39 (B. Brookman, C.I.S.). Herkey Creek, y, VI-24-34 (P. H. Timber- Geographic range : lake, U.C.R.); #!,VI-8-34 (P. H. Timberlake, Tex., N. Mex., Ariz., Calif., Nev., Utah, U.C.R.); 2, V-20-39 (B. Brookman, C.1.S.); 2 6, Colo., Kans., *Neb., Wyo., and S.D. 2, V-22-39 (E. G. Linsley, C.1.S.); 6, V-27-39 (E. G. Linsley, C.I.S.); 2, VI-3-39 (E. G. Lins- California records: ley, C.I.S.); 9, VI-1-40 (C. D. Michener, C.I.S.); AIAMEtM 00.: Livermore, 3, 111-1931 (E. S. 4 y, VI-4-40 (C. D. Michener, C.1.S.); 9, VI-13-40 Ross, C.A.S. 1. (H. T. Reynolds, C.1.S.); 3 9, VI-14-40 (C. D. EUIl'E €0.: Butte Creek, nr. Chico, 24 6, Michener, C.I.S.); 9, VI-14-40 (F. H. Rindge, IV-25-22 (E. P. Van Duzee, C.A.S. 1. C.I.S. 1. Whitewater, 9, VIII-1-34 (P. H. Timber- CALAVERAS 0.:Mokelumne Hill, 6,April lake, U.C.R.); 6, 111-13-39 (C. M. Dammers, (F. E. Blaisdell, C.A.S.). U.C.R:). San Andreas Canyon, 2 6, 2 Y, IV-25-50 CONIRA C@TA CO.: Antioch, 6, V-1-32 (C.I. (C. D. MacNeill). S.); 3 Y, V-15-32 (C.1.S.); 9 VI-8-33 (M. A. SACRAMENTO CO.: Fair Oaks, 2 Y, V-11-47 Cazier, C.I.S.); 2 9, VI-12-35 (E. P. Van Duzee, (A. T. McClay, U.C.D.). C.A.S.); #!,VI-2-36 (M. A. Fhburg, C.I.S.); 2 Y, SAN BERNARDINO CO.: Lee Lake, 9, VI-2-33 VII-4-36 (R. M. Bohart, U.C.D.); Y, VII-1937 (C. M. Dammers, U.C.R.). Victorville, 4 9, (E. S. Ross, C.A.S.); 2 6, IV-9-49 (J. E. Gilla- V-24-36 (C. M. Dammers, U.C.R. 1. Cajon Pass, Y, spy, C.I.S.); 2 6, Y, V-21-49 (P. D. Hurd, Jr.. VI-24-49 (L. W. Isaak, U.C.D.). Deep Creek, 3 6, C.I.S.) 6, same data (L. W. Quate, C.1.S.); 6, IV-26-36 (K. A. Caard. U.C.L.A.). IV-29-50 (P. D. Hurd, Jr. , C.I.S. 1; 2 6, same SAN DIECO CO.: Warner Springs, 6,V-9-36 date (J. W. MacSwain, C.I.S.); 4 6, V-8-50 (R. S. (P. H. Timberlake, U.C.R.). Ekal, C.I.S.). TULARE CO. : Exeter, Y, VIII-25-38 (C.I.S.). EL DORADO CO.: Chile Bar, 2 $!, VII-5-48 Three Rivers, 3 mi. W., 2 6, IX-1-39 (P. H. Tim- (P. D. Hurd, Jr., C.1.S.); #!, same data (J. W. berlake. U.C.R. ). MacSwain, C. I.S. 1. VENTURA Co.: Saticoy, y, V-14-26 (S. F. FRESNO CO.: Coalinga, di, 111-10-31 (E. P. Flanders, U.C.R. ). Van Duzee, C.A.S.). Selma, 46 Y, VI-19-50 (R. C. YOLO CO.: Putah (:anyon, 2 6, V-1950 (J: C. Bechtel, U.C.D. ). Hall, U.C.D. 1. KERN CO.: Mojave Desert, 2 6, V-9-37 (N. F. Hardman, C.I.S. 1: 6, IV-23-39 (G. Fi Smith, C. I.S.). Kernville, 6, IV-26150 (E. I. Schlinger, U1C.D. ). Campsomeris tolteca (Saussure)4 Bakersfield, 6, IV-15-49 (W. E. Hazeltine, C.I.S.). LOS ANGELES CO.: Westwood Hills, 3 6, VI-2-35 (E. G. Linsley. C.I.S., U.C.R.). Little Elis tolteca Saussure, 1857, Rev. et Mag. Zool., -8Rock 2 6, IV-11-36 (G. E. & R. M. Bohart, U.C.D., ser. 2, 9:282. Type Y. UCLA). Palmdale, 3 6, IV-11-36 (G. E. & R. M. Bohart, U.C.D.). b,6, IV-15-36 (E. G. Linsley, Geographic range : C.I.S.). Castaic, 3 Y, VII-4-50 (U.C.L.A.). Haiti, Mex., Tex., Ariz., Lower Calif., and Elizabeth Lake, 6,V-6-50 (B. Lazaroff, U.C.L.A.). Calif.

4Eli3 dives Provancher and E Lis 4-cincta Provancher 3Elis anericana Saussure is a synonym of this species. are synonyms of this species. THE SCOLIIDAE OF CALIFORNIA 145 California records: Springs, 2 9, V-14-41 (E. C. Van Dyke, C.A.S.). CONTRA COSTA CO.: Antioch, Y, V-8-32 Whitewater, 9, VIII-9-50 (J. W. MacSwain, (C.I.S. 1; 6, 5 Y, V-15-32 (C. I.S. 1; 3 Y, VIII- C.I.S.); ?, same data (J. D. Paschke, C.I.S.). 22-33 (C.I.S.); Y, IX-23-33 (B. J. Hall, U.C.R.); Hemet, 2 y, VII-15-45 (J. W. MacSwain, C.I.S.); Y, M-1935 (E. S. ROSS, C.A.S.); 2 2, IX-12-36 XVIII-4-46(J. W. MacSwain, C.I.S. 1; Y. (E. G. Linsley, U.C.R.); 2 6, IV-11-37 (E. C. VIII-26-46 (J. W. MacSwain, C.I.S.); 6, VIII-28- Van Dyke, C.A.S.); 3 6,IV-18-37 (E. C. Van Dyke, 46 (J. W. MacSwain, C.I.S. 1. kt,6. _S. C.A.S.); 2 Y, VII-15-37 (E. C. Van Dyke, C.A.S.); W., VIII-7-46 (J. W. MacSwain, C.I.S.). 9, VIII-28-37 (T. H. G. Aitken, C.I.S. 1; 9, Springs, 6, IV-1-46 (C. L. Fox, C.A.S. 1; 3 9, VI-15-41 (C.1.S.); 6,VIII-10-41 (E. C. Van Dyke, 111-26-32 (P. H. Timberlake, U.C.R.); 9,. C.A.S.); 6,X-16-48 (C. D. MacNeill); ?, X-24-48 V-11-33 (P. H. Timberlake, U.C.R.). Ripley, 1 P. D. Hurd, Jr., C.I.S. 1; 9, X-21-39 (C. D. Mac- -mi. S., 9, VII-10-46 (J. W. MacSwain, C.I.S.). Neill); ?, IV-29-50 (P. D. Hurd, Jr., C.I.S.); Blythe, 3 ?, M-25-32 (C. M. Darners, U.C.R. 1; 6, 2 Y, V-8-50 (R. S. Ekal, C.I.S.). 9 Y, X-16-34 (C. M. Dannners, U.C.R.); Y, FRESNO CO.: Coalinga, 6, 111-19-31 (E. P. VII-29-35 (C. M. Darners, U.C.R.); Y. IX-1-35 Van Duzee, C.A.S.). Alcade Canyon, E. Coalinga, (P. H. Timberlake, U.C.R.); 3 6,2 y, XI-2-36 6, IV-2-32 (H. Fincher, U.C.D. ). Firebaugh, $, (C. M. Dammers, U.C.R.); 6 b, VII-15-38 (P. H. 111-24-49 (R. F. Smith, C.I.S.); 2 6,VIII-16-49 Timberlake, U.C.R.); 6,Y, VI-23-45 (E. G. (A. D. Telford, C.I.S.). Selma, 9, VI-19-50 Linsley, C.I.S. 1; 3.6, same data (J. W. Mac- (R. C. Bechtel, U.C.D.). Raisin City, $, X-15-51 Swain, C.I.S.; 2 6, VI-25-45 (E. G. Linsley, (E. G. Linsley, C. I.S. 1. C.I.S.); 2 y, VII-27-46 (P. D. Hurd, Jr., IMPERIAL CO.: Potholes, Y, IV-7-23 (E. P. C.I.S. 1; ?, VIII-13-46 (P. D. Hurd, Jr., C. I.S. 1; Van Duzee. C.A.S.). Holtville, Y, VI-1936 (E. S. Y. VIII-20-46 (J. W. MacSwain, C.I.S.); 6, ROSS, C.A.S.); 6, VI-20-39 (E. S. ROSS, C.A.S.); VII-6-47 (W. F. Barr, C-1.S.); 3 Y, IV-27-49 6, VI-23-40 (W. F. Barr, C.I.S.). Palo Verde, 6, (C. D. MacNeill). 111-7-47 (E. G. Linsley, C.I.S.). Palo Verde, SAWENTO CO.: Sacramento, 1 Y, VI-15-48 -3 mi. S-., Y, IV-8-49 (P. D. Hurd, Jr., C.I.S.). (C. D. MacNeill). Westmorland, 2 9, V-6-39 (U.C.L.A.); 6, IV-16-39 SAN BERNARDINO CO:: Needles, 2 6, VII-4-21 (U.C.L.A. ). (J. A. Kusche, C.A.S. ). h,2 6, XI-5-33 (C. KERN CO.: Arvin, $, 111-31-37 (G. L. M. Darners, U.C.R. 1; d, IV-24-50 (U.C.L.A. 1. Smith, C.I.S. 1. Bakersfield, Y, VII-1938 (C.I.S. 1. Mojave River at Apple Valley, 2 6, Y, VI-29-40 Bakersfield, 15.E., 6,VIII-4-46 (F. A. (J. W. MacSwain, C.I.S. 1:.. ?, same data (P. H. Ehrenford, C.I.S.). Shafter, 9, VI-7-40 (C. G. Timberlake, U.C.R.). Morongo, 2 6, IX-29-34 (P. Lewis, C.I.S. ); 6, VI-25-40 (C. G. Lewis, C.I.S. 1. H. Timberlake, U.C.R.); $, IX-26-44 (P. H. Tim- Kern Park, 6,VII-7-46 (F. A. Ehrenford, C.I.S.). berlake, U.C.R.). Morongo Valley, 6, VIII-20-36 Kern Canyon, $, VII-28-46 (F. A. Ehrenford, (P. H. Timberlake, -Orville, 0, C.I.S.). Kern River County Park, 4 9, VI-27-49 IV-18-33 (C. M. Dammers, U.C.R.); ?, IV-17-36 L. W. Isaak, U.C.D.). Devils Den, Y, IV-8-51 (C. M. Dammers, U.C.R.); 2 9, VIII-17-36 (C. M. (J. W. MacSwain, C.I.S.). Dammrs, U.C.R.); 2 $, IX-28-38 (P. H. Timberlake, LOS ANGELES CO.: Claremont, 2 6, VI-1-28 U.C.R. 1. (E. L. Kessel, C.A.S.). Huntington Park, 9, SAN DIEGO CO.: Jacumba, 6,X-3-25 (J. D. IV-28-50 (R. L. Langston, C.I.S.). Gunder, C.A.S.). Coronado Beach, 9, IX-26-35 MWCED CO.: Cressy, 4 6,VII-21-48 (R. P. (L. S. Slevin, C.A.S.). San Felipe Creek, Y, Allen, C.A.S. ). DOS Palos, 2 6,9, IX-9-48 IX-2-35 (F. T. Thorne. U.C.R. 1; ?, IX-7-38 (P. H. (R. F. Smith, C.1.S.); 6,VI-21-50 (C. D. Mac- Timberlake, U.C.R. 1. oak Grove, ?, VI-6-40 Neill). Dos Palos, 7 mi. E., 6, VIII-16-49 (R. F. (F. H. Rindge, C.I.S.). Borego Valley, 2 d', Y, Smith, C.I.S.). 111-30-? (W. A. MacDonald, U.C.L.A.). ORANGE CO.: Anaheim, Y, V-14-27 (E. L. Kes- TULARE CO.: Lindsay, 2 6, 9, IX-6-32 (P. sel, C.A.S. 1. Seal Beach, 5 6, 21 Y, VIII-25-38 H. Timberlake, U.C.R.). Wood Lake, 6, IV-23-47 P. H. Timberlake, U.C.R. 1- Newport Bay, 6, 9, (N. W. Frasier, C.I.S.). VII-17-41 (P. D. Hurd, Jr., C.I.S. ); 4 6, 5 Y, VENTURA CO.: Santa Susana, 7 ?, VIII-13-32 VII-25-41 (P. D. Hurd, Jr., C.I.S.). (C.1.S.); 0, VIII-15-32 (C.I.S. ); Y, VIII-23-32 RIVERSIDE CO.: Riverside, 32 8, 18 y, (C.I.S. 1. Saticoy, 0, V-19-36 (U.C.R. 1. February-October 1929-1938 (P. H. Timberlake, U.C.R. 1.. Soboba Hot Springs, 9, 11-22-36 (P. H. 9, Timberlake, U.C.R.). Ribbonwood, 3 V-20-39 Campsoscolia octomaculata texensis (Saussure) a (C.I.S.). Fairmont Lake, 3 6,VII-13-36 (W. C. ko,6, VIII-2-48 (P. H. Reeves, C.I.S.). Elis texensis Saussure. 1858, Ann. SOC. Ent. France, Timberlake, U.C.R.). Palm Desert, 9, IV-12-50 ser. 3, 6_:224. Type 6. (P. D. Hurd, Jr., C.I.S. 1. Banning, 2 8, VII- 16-50 (J. C. Hall, U.C.D.); 3 6, 9. same data '~coliaregina Cresson, ~co~iaflavosignata Cresson, (W. C. Marshall, U.C.D.). Cathedral City, 9, Scolia consors Cresson (nec Saussure), Elis (Frielis) 1. zonaria Cresson. and Blis Lueina Cresson are synonyms VII-20-50 (L. W. Isaak, U.C.D. Gilman Hot of this subspecies. 146 BULLETIN OF THE CALIFORNIA INSECT SURVEY

Geographic range: 1931 (E. S. Ross, C.A.S.). Tex., Kans., Colo., N. Mex., Ariz., Lower bSARIPOSA CO.: Yosemite, 9, VI-1926 (E. Calif., and Calif. 0. Essig, C.I.S. ). MERCED CO.: Dos Palos, 2 $', VIII-15-49 California records: (A. D. Telford, C.I.S. 1. IMPERIAL CO. : Palo Verde, 67, VIII-15-46 MOWC CO.: Lake City, 6, VIII-4-22 (C. L. (P. D. Hurd, Jr., C.I.S.); $', VIII-20-46 (a. F. Fox, C.A.S. 1. Barr, C. I.S. ); Y, VIII-20-46 (P. D. Hurd, Jr., ORANGE CO.: Laguna Beach, y, VII-19-21 C.I.S.); $', VIII-27-46 (P.'D. Hurd, Jr., C.I.S.). (E. 0. Essig, C.I.S. 1; 6, VI-1930 (E. L. Kessel, RIVESIDE CO.: Blythe, 17 mi. W., 9, C.A.S.). Laguna Canyon, 3 V, VI-28-29 (E. L. VIII-27-46 (P. D. Hurd, Jr., C.I.S.). Kessel, C.A.S.). Newport Bay, 3 Y, VI-25-41 (P. SAN BERNARDINO CO.: Twentynine Palms. 17 D. Hurd. Jr., C.I.S. ). mi. W., 5 6, VIII-9-42 (P. H. Timberlake, U.C.R.). RIVERSIDE CO.: Corona, 6, VII-1929 (C. I.S.). Winchester, 2 9, VII-25-46 (J. W. hiacSwain; C. I.S. 1. Temecula, 12 6,2 $, VII-4-50 (E. G. Linsley & J. W. MacSwain, C.I.S. ). Banning, d', Campsoscolia alcione (Banks) 4 Y, VII-9-50 (M. J. Stebbins, U.C.D. ); 3, Y, same data (H. N. Yokoyama, C.I.S.); 2 8, 6 Y, 7rielis alcione Banks, 1917, Bull. Mus. Comp. Zozil., same data (T. R. Haig). Hemet, $, VII-5-50 (E.G. Harvard College, g:112. Type 6. Linsley, C.I.S. ). Hemet, 6 G. G.,$, VIII-7-46 (J. VI. MacSwain, C.I.S.). Elsinore, Y, VII-4-50 Geographic range: (J. W. MacSwain, C. I.S. ). Calif., Nev., N. Mex., Utah, Colo., and Wash. SAN BERNARDINO CO.: Morongo Valley, 6, VIII-4-33 (P. H. Timberlake, U.C.R.). Yucca California records: Valley, 6, VIII-3-33 (P. H. Timberlake, U.C.R.). ALAMELIA CO. : Murrieta Caves, 6, VIII-1937 Mojave River at Apple Valley, 6, VI-29-40 (J. W. (T. H. G. Aitken, C.I.S.). Tesla, Corral Hollow, MacSwain, C.1.S.); 2y, same data (P. H. Timber- 3 6, Y, VII-16-46 (P. D. Hurd, Jr., C.I.S.); 6, lake, U.C.R.). Cajon Canyon, 2,700 ft., $, 2 9, IX-19-49 (P. D. Hurd, Jr., C. I.S. 1; 14 6, VII-21-36 (P. H. Timberlake, U.C.R.). Lone Pine IX-21-51 (W. C. Bentinck, C.I.S.); Y, X-4-51 (J. Canyon, San Gabriel Mts., 6,000 ft., VII-21-36 E. Gillaspy, C.I.S. 1. Livermore, 2 8, IV-7-37 (P. H. Timberlake, U.C.R.). Oak Glen Lodge, 3 Y, (U.C.L.A.) (F. Daggett, C.A.S. ). ALPINE CO.: Woodfords, 3 mi. N. E., 1 6, SAN JOAQUIN CO.: Tracy, !#,IX-3-46 (P. D. 13 Y, VIII-21-49 (C. D. MacNeill). Hurd, Jr., C.I.S.). CONTRA COSTA CO.: Antioch, 9, M-1-37 SIERRA CO.: Calpine, 2, VIII-27-48 (E. G. (M. A. Cazier, C.I.S.); Y, IX-18-38 (E. C. Van Linsley, C.I.S.). Dyke, C.A.S. ). STANISLAUS CO.: . Patterson, 6, $, VI-30-49 FRESNO CO.: Firebaugh, $', VII-25-49 (E. (T. F. Leigh, C.I.S. ); H'estley, 2 J, ?, VII-25-49 V. d. Bosch, C.I.S.); 6, VIII-4-47 (R. V. d. (T. F. Leigh, C.I.S.).

Bosch, C.I.S.). Kingsburg, 3 6, 3 y, VII-10-39 TULARE CO,: ~Kaweah, ~~ 3 6, VI-19-36 (F. T. (H. V. Bohart, U.C.L.A. ). Scott, U.C.R.). California Hot Springs, 6, Y, INYO CO.: Owens Valley, $!, VIII-2-36 (R. V-25-40 (C. G. Lewis, C.I.S. ). Wood Lake, 6, M. Bohart, U.C.D. ). Bishop, d, VIII-1-36 (R. V. VI-12-47 (N. W. Frazier, C.I.S.). Strathmore, Rohart, U.C.D.). Bishop, 15 mi. E, 11 6, 29 $, 4 6, 2 $2, VI-8-28 (T. Craig, C.A.S. VIII-5-48 (P. D. Ilurd, Jr..& J. W. MacSwain, VENTURA CO.: Santa Susana, Y, VIII-13- 32 c. I.S. 1. (C.I.S.). Santa Paula, 2 9, VII-237? (D. Burk I KEEN CO.: Greenfield, 8 mi. W., 2 6, (C.1.S.); $, VII-13-31 (C.1.S.). VIII-31-49 (P. H. Timberlake, U.C.R.). Walker -Pass, 8, VI-26-47 (11. E. Cott, U.C.D.). Shafter, 2 d, VI-8-40 (C. G. Lewis, C.I.S.); 6, $, VI-25- Campsoscolia jlarnmicorna (Bradley) 40 (C. G. Lewis, C.I.S. 1; ?, VIII-10-40 (C. G. Lewis, C.I.S.). Grapevine, d', 4 Y, VI-10-47 (R. Ompsomeris (Trielis) f lainmicoma Bradley, 1928, F. Smith, C.I.S.). Bakersfield, $, VIII-1938 Trans. Amer. Ent. SOC., %:199, 200, 209- (C.I.S.). Lerdo, Y, VII-17-46 (F. A. Ehrenford, 211. Type Y. C.I.S.). Arvin, 2 6', V-29-40 (C. G. Lewis, C.I.S.). &, d', VII-4-46 (F. A. Ehrenford, Geographic range: C.I.S.). Kern River County Park, 6, $, VI-27-49 Mex. (Sonora), N. Mex., Ariz., and Calif. (L. W. Isaak, U.C.D.). Poso Creek, $, VI-5-29 (E. P. Van Duzee, C.A.S. 1. California records: LASSEN CO. : Hallelujah Jct., $, VI1-13-49 IMPERIAL CO. : Holtville, 9, VI-1936 (P. D. Hurd, Jr., C.I.S.). Constantia, 4 mi. S., (E. S. Ross, C. I.S. 1. Salton Sea, 6 (C.I.S. ). 6 d, VII-2-51 (C. D. MacNeill). Westmorland, 4 9, V-24-46 (P. D. Gerhardt, LOS ANGELES CO.: Los Angeles, $, VII- U.C.R.). Palo Verde, 3 y, VIII-10-46 (B. F. THE SCOLIIDAE OF CALIFORNIA 147

Barr, C.I.S. ); d', VIII-15-46 (P. D. Hurd, Jr., X-23-51 (P. D. Hurd, C.I.S.). C.I.S.1; 2 0, VIII-20-46 (P. D. Hurd, Jr., SAN BERNPSDINO CO. : Twentynine Palms, C.I.S.); 9, same data (J. W. MacSwain, C.I.S.). d', VIII-3-33 (H. L. hlcKenzie, U.C.K.); 6, same Niland, 6, X-24-51 (P. D. Hurd, Jr., C.I.S. ). data (P. H. Timberlake, U.C.H.). Cronise, 5 J, RIVERSIDE CO. : Coachella, 9, X-13-39 VII-7-38 (P. H. Timberlake, U.C.n.). V' (E. C. Linsley, U.C.R.). Rlythe, 2 d',VII-10-46 Jct., 20 mi. N., 3 y, VI-19-49 (R. F. Fritz, (W. F. Barr, C.I.S.); 2 8, 17 $, VII-4-51 (J. w. C.I.S.). MacSwain, R. F. Smith, C.I.S. ). &, 3 9,

Selected Bibliography

Bartlett, Oscar C. cial reference to Venezuela.-I. 1912. The North American digger wasps of The genus Campsomeris. €301. Ent. the subfamily Scoliinae. Ann. Ent. Venezolana, 4: 1-36, 2 figs. SOC. Amer., 5:293-340, 3 plates. Linsley, E. Gorton Bradley, J. Chester 1946. Insect of alfalfa in 1928a. The species of Campsomeris (Hymen- California. Jour. Econ. Ent., optera-Scoliidae) of the plumipes 39: 18-29. group, inhabiting the United States, the Greater Antilles, and the Baha- Rau, Phil and Nellie Rau ma Islands. Proc. Acad. Nat. Sci. 1918. Wasp studies afield. Princeton, Phila., 80:313-337, 1 plate, 2 text Princeton University Press. xv figs. -+ 372 pp. 1928b. A revision of the New World species of Trielis. a subgenus of Campso- Rau, Phil meris. Trans. Amer. Ent. Soc., 1932. The courtship dance and sleeping 54:195-214, 1 prate, 4 text figs habits of Scolia dubia. Bull. 1945. The Scoliidae (Hymenoptera) of Brooklyn Ent. SOC.,27~59-62. northern South America, with espe-

PLATES

Campsoscolia

3 Campsomeris

4 Scolia

5 Sc o I i Q (Tr is c o I ia ) Plate 12

Forewings of Scoliid wasps. Fig. 2, Campsoscolia; Fig. 3, Campsomeris; Fig. 4, Scolia; Fig. 5, Scolia (7riscoZia). Recurrent nervures indicated in hlacb.

151 I 2

3

Plate 13

Scoliid wasps (females). 1, Scolia otomita Saus- sure; 2, Cumpsoscolia alcione (Banks); 3, Camp- someris tolteca (Saussure); 4, Scolia ardens Smith.

152