Phylum Arthropoda, Subphylum Trilobitomorpha: Trilobites
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Sequence of Post-Moult Exoskeleton Hardening Preserved in a Trilobite Mass Moult Assemblage from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco
Editors' choice Sequence of post-moult exoskeleton hardening preserved in a trilobite mass moult assemblage from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco HARRIET B. DRAGE, THIJS R.A. VANDENBROUCKE, PETER VAN ROY, and ALLISON C. DALEY Drage, H.B., Vandenbroucke, T.R.A., Van Roy, P., and Daley, A.C. 2019. Sequence of post-moult exoskeleton hardening preserved in a trilobite mass moult assemblage from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco. Acta Palaeontologica Polonica 64 (2): 261–273. Euarthropods have a tough exoskeleton that provides crucial protection from predation and parasitism. However, this is restrictive to growth and must be periodically moulted. The moulting sequence is well-known from extant arthropods, consisting of: (i) the long inter-moult stage, in which no changes occur to the hardened exoskeleton; (ii) the pre-moult stage where the old exoskeleton is detached and the new one secreted; (iii) exuviation, when the old exoskeleton is moulted; and (iv) the post-moult stage during which the new exoskeleton starts as soft, thin, and partially compressed and gradually hardens to the robust exoskeleton of the inter-moult stage. Trilobite fossils typically consist of inter-moult carcasses or moulted exuviae, but specimens preserving the post-moult stage are rare. Here we describe nine specimens assigned to Symphysurus ebbestadi representing the first group of contemporaneous fossils collected that preserve all key stages of the moulting process in one taxon, including the post-moult stage. They were collected from a single lens in the Tremadocian part of the Fezouata Shale Formation, Morocco. Based on cephalic displacement and comparison to other trilobite moults, one specimen appears to represent a moulted exoskeleton. -
The Case of the Diminutive Trilobite Flexicalymene Retrorsa Minuens from the Cincinnatian Series (Upper Ordovician), Cincinnati Region
EVOLUTION & DEVELOPMENT 9:5, 483–498 (2007) Evaluating paedomorphic heterochrony in trilobites: the case of the diminutive trilobite Flexicalymene retrorsa minuens from the Cincinnatian Series (Upper Ordovician), Cincinnati region Brenda R. Hundaa,Ã and Nigel C. Hughesb aCincinnati Museum Center, 1301 Western Avenue, Cincinnati, OH 45203, USA bDepartment of Earth Sciences, University of California, Riverside, CA 92521, USA ÃAuthor for correspondence (email: [email protected]) SUMMARY Flexicalymene retrorsa minuens from the upper- rate of progress along a common ontogenetic trajectory with most 3 m of the Waynesville Formation of the Cincinnatian respect to size, coupled with growth cessation at a small size, Series (Upper Ordovician) of North America lived ‘‘sequential’’ progenesis, or non-uniform changes in the rate of approximately 445 Ma and exhibited marked reduction in progress along a shared ontogenetic trajectory with respect to maximum size relative to its stratigraphically subjacent sister size, can also be rejected. Rather, differences between these subspecies, Flexicalymene retrorsa retrorsa. Phylogenetic subspecies are more consistent with localized changes in analysis is consistent with the notion that F. retrorsa retrorsa rates of character development than with a global hetero- was the ancestor of F. retrorsa minuens. F. retrorsa minuens chronic modification of the ancestral ontogeny. The evolution has been claimed to differ from F. retrorsa retrorsa ‘‘in size of F. retrorsa minuens from F. retrorsa retrorsa was largely alone,’’ and thus presents a plausible example of global dominated by modifications of the development of characters paedomorphic evolution in trilobites. Despite strong similarity already evident in the ancestral ontogeny, not by the origin of in the overall form of the two subspecies, F. -
Instituto De Geociências Revisão Sistemática E
UNIVERSIDADE DE SÃO PAULO INSTITUTO DE GEOCIÊNCIAS REVISÃO SISTEMÁTICA E PALEOBIOGEOGRÁFICA DE TRILOBITAS PHACOPIDA (HOMALONOTIDAE E CALMONIIDAE) DO DEVONIANO DAS BACIAS DO PARNAÍBA E AMAZONAS, BRASIL Felipe van Enck Meira Tese apresentada ao Programa de Pós- Graduação em Geoquímica e Geotectônica do Instituto de Geociências da Universidade de São Paulo, como parte dos requisitos para a obtenção do título de doutor. Orientadora: Prof. Dra. Juliana de Moraes Leme TESE DE DOUTORAMENTO Programa de Pós-graduação em Geoquímica e Geotectônica São Paulo 2016 “Kites rise highest against the wind - not with it.” - Winston Churchill Agradecimentos Agradeço a Deus, por sempre iluminar o caminho durante esses anos de altos e baixos do Doutorado. Agradeço a Ele também por enviar dois verdadeiros anjos da guarda à minha vida – minha esposa Angela Faleiros van Enck Meira e meu filho, Thomas Faleiros van Enck Meira. Agradeço a meus pais, José Carlos e Sylvia, e à minha irmã, Patrícia, pelo apoio durante a jornada, ainda que à distância, por vezes. Sou grato aos meus sogros, Jair e Lucia, que sempre foram como verdadeiros pais e conselheiros. À FAPESP (Processo n° 2012/07075-3), pelo suporte financeiro, sem o qual o Doutorado não seria viável. À minha orientadora, Drª. Juliana de Moraes Leme, pela orientação, discussões e esclarecimentos pertinentes ao projeto. Ao Doutorando Fabio Carbonaro e ao Dr. Renato Ghilardi (UNESP-Bauru), pela parceria e por discussões importantes na realização do trabalho. À Drª. Niède Guidon (FUMDHAM), pelo empréstimo de fósseis da região de São João Vermelho (Piauí), estudados aqui. Sou grato às seguintes pessoas, por permitirem meu acesso às instituições para visita de acervos, e por sua disponibilidade, atenção e ajuda durante minha permanência: Bushra Hussaini (AMNH), Flávia Alessandra Figueiredo, Mônica de Medina Coeli, Dr. -
Smithsonian Miscellaneous Collections Volume 148, Number 3
—r SMITHSONIAN MISCELLANEOUS COLLECTIONS VOLUME 148, NUMBER 3 UPPER CAMBRIAN TRILOBITE FAUNAS OF NORTHEASTERN TENNESSEE (With 21 Plates) By FRANCO RASETTI The Johns Hopkins University Baltimore, Md. (Publication 4598) CITY OF WASHINGTON PUBLISHED BY THE SMITHSONIAN INSTITUTION June 10, 1965 ,v, . SMITHSONIAN MISCELLANEOUS COLLECTIONS VOLUME 148, NUMBER 3 l^psfarrl| 3F«ttb UPPER CAMBRIAN TRILOBITE FAUNAS OF NORTHEASTERN TENNESSEE (With 21 Plates) By FRANCO RASETTI The Johns Hopkins University Baltimore, Md. (Publication 4598) CITY OF WASHINGTON PUBLISHED BY THE SMITHSONIAN INSTITUTION June 10, 1965 <.%'^ Q^fii^ CONNECTICUT PRINTERS, INC. HARTFORD, CONN., U.S.A. CONTENTS Part I. Stratigraphy and Faunas Pj^ge Acknowledgments 3 Descriptions of Localities and Sections 3 General statement 3 Hawkins County 4 Hamblen County 10 Grainger County 12 Jefferson County I5 Union County 19 Claiborne County 21 Knox County 22 Monroe County 23 Purchase Ridge, Scott Coimty, Virginia 24 Index of Localities 25 Fauna of the Cedaria Zone 26 Fauna of the Crepicephalus Zone 28 Fauna of the Aphelaspis Zone 30 Part II. Systematic Paleontology General Statement 38 Descriptions of Trilobite Genera and Species Order AGNOSTIDA 38 Family AGNOSTIDAE 38 Order CORYNEXOCHIDA 39 Family DORYPYGIDAE 39 Order PTYCHOPARHDA 40 Family LONCHOCEPHALIDAE 40 Family CATILLICEPHALIDAE 44 Family CREPICEPHALIDAE 45 Family TRI CREPICEPHALIDAE 54 Family ASAPHISCIDAE 55 Family KINGSTONIIDAE 60 Family MENOMONIIDAE 61 Family NORWOODIIDAE 64 Family CEDARIIDAE 69 Family ELVINIIDAE 71 Family PTEROCEPHALIIDAE 72, PTYCHOPARHDA of uncertain affinities 102 Undetermined trilobites 112 References 115 Explanation of Plates 118 UPPER CAMBRIAN TRILOBITE FAUNAS OF NORTHEASTERN TENNESSEE By FRANCO RASETTI The Johns Hopkins University PART I. STRATIGRAPHY AND FAUNAS The purpose of this paper is both to describe the fossils and to present them in their proper stratigraphic setting. -
Available Generic Names for Trilobites
AVAILABLE GENERIC NAMES FOR TRILOBITES P.A. JELL AND J.M. ADRAIN Jell, P.A. & Adrain, J.M. 30 8 2002: Available generic names for trilobites. Memoirs of the Queensland Museum 48(2): 331-553. Brisbane. ISSN0079-8835. Aconsolidated list of available generic names introduced since the beginning of the binomial nomenclature system for trilobites is presented for the first time. Each entry is accompanied by the author and date of availability, by the name of the type species, by a lithostratigraphic or biostratigraphic and geographic reference for the type species, by a family assignment and by an age indication of the type species at the Period level (e.g. MCAM, LDEV). A second listing of these names is taxonomically arranged in families with the families listed alphabetically, higher level classification being outside the scope of this work. We also provide a list of names that have apparently been applied to trilobites but which remain nomina nuda within the ICZN definition. Peter A. Jell, Queensland Museum, PO Box 3300, South Brisbane, Queensland 4101, Australia; Jonathan M. Adrain, Department of Geoscience, 121 Trowbridge Hall, Univ- ersity of Iowa, Iowa City, Iowa 52242, USA; 1 August 2002. p Trilobites, generic names, checklist. Trilobite fossils attracted the attention of could find. This list was copied on an early spirit humans in different parts of the world from the stencil machine to some 20 or more trilobite very beginning, probably even prehistoric times. workers around the world, principally those who In the 1700s various European natural historians would author the 1959 Treatise edition. Weller began systematic study of living and fossil also drew on this compilation for his Presidential organisms including trilobites. -
Copertina Guida Ai TRILOBITI V3 Esterno
Enrico Bonino nato in provincia di Bergamo nel 1966, Enrico si è laureato in Geologia presso il Dipartimento di Scienze della Terra dell'Università di Genova. Attualmente risiede in Belgio dove svolge attività come specialista nel settore dei Sistemi di Informazione Geografica e analisi di immagini digitali. Curatore scientifico del Museo Back to the Past, ha pubblicato numerosi volumi di paleontologia in lingua italiana e inglese, collaborando inoltre all’elaborazione di testi e pubblicazioni scientifiche a livello nazonale e internazionale. Oltre alla passione per questa classe di artropodi, i suoi interessi sono orientati alle forme di vita vissute nel Precambriano, stromatoliti, e fossilizzazioni tipo konservat-lagerstätte. Carlo Kier nato a Milano nel 1961, Carlo si è laureato in Legge, ed è attualmente presidente della catena di alberghi Azul Hotel. Risiede a Cancun, Messico, dove si dedica ad attività legate all'ambiente marino. All'età di 16 anni, ha iniziato una lunga collaborazione con il Museo di Storia Naturale di Milano, ed è a partire dal 1970 che prese inizio la vera passione per i trilobiti, dando avvio a quella che oggi è diventata una delle collezioni paleontologiche più importanti al mondo. La sua instancabile attività di ricerca sul terreno in varie parti del globo e la collaborazione con professionisti del settore, ha permesso la descrizione di nuove specie di trilobiti ed artropodi. Una forte determinazione e la costruzione di un nuovo complesso alberghiero (AZUL Sensatori) hanno infine concretizzzato la realizzazione -
Descargar Trabajo En Formato
Upper Cambrian – Arenig stratigraphy and biostratigraphy in the Purmamarca area, Jujuy Province, NW Argentina Guillermo F. Aceñolaza1 and Sergio M. Nieva1 1 INSUGEO – Conicet/Facultad de Ciencias Naturales e I.M.L. Universidad Nacional de Tucumán, Miguel Lillo 205, 4000 Tucumán. E-mail: [email protected] Palabras clave: Estratigrafía. Bioestratigrafía. Ordovícico. Purmamarca. Jujuy. Argentina. Key words: Stratigraphy. Biostratigraphy. Ordovician. Purmamarca. Jujuy. Argentina. Introduction The region of Purmamarca is a classical locality for geological studies in the argentine literature. Many authors have recognized and analyzed the Ordovician strata and its paleontological heritage (eg. Keidel, 1917; Kobayashi,1936, 1937; Harrington, 1938; De Ferraris, 1940; Harrington and Leanza, 1957; Ramos et al., 1967 and Tortello, 1990). However, strata are several times sliced by longitudinal faults of submeridional trend, cropping out in narrow belts alternating with Precambrian/Cambrian, Cambrian and Mesozoic units. As long ago stated by Harrington (1957), no continuous section is displayed, recording isolated Ordovician blocks of different stratigraphic position. Several Lower Ordovician units crop out in the Purmamarca area. The Casa Colorada Formation (Uppermost Cambrian – Lower Tremadocian) is widely represented by the eastern flank of the Quebrada de Humahuaca when meeting the Quebrada de Purmamarca. The Coquena and Cieneguillas shales (Upper Tremadocian), and the Sepulturas Formation (Arenig) are nicely displayed in the tributaries of the Quebrada de Purmamarca. Finally, it is worthwhile to mention that a paleontological collection from this South American locality was given to the renowned Japanese paleontologist Teiichi Kobayashi in the first decades of the XX century (Kobayashi, 1936). Within the material was a new trilobite genus, defined by him as Jujuyaspis(after Jujuy province). -
Enrollment and Coaptations in Some Species of the Ordovician Trilobite Genus Placoparia
Enrollment and coaptations in some sp eeies of the Ordovician trilobite genus Placoparia JEAN-LOUIS HENRY AND EUAN N.K. CLARKSON Henry, J.-L. & Clarkson, E.N.K. 1974 12 15: Enrollment and coaptations in some species of the Ordovician trilobite genus Placoparia. Fossils and Strata, No. 4, pp. 87-95. PIs. 1-3. Oslo ISSN 0300-949l. ISBN 82-00-04963-9. The species Pl. (Placoparia) cambriensis, Pl. (Coplacoparia ) tournemini and Pl. (Coplacoparia ) borni, described by Hammann (1971) from the Spanish Ordovician, are found in Brittany in formations attributed to the Llanvirn and the LIandeilo. The lateral borders of the librigenae of Pl. (Coplacoparia) tournemini and Pl. (Coplacoparia ) borni bear depressions into which the distal ends of the thoracic pleurae and the tips of the first pair of pygidial ribs come and fit during enrollment. These depressions, or coaptative structures sensu Cuenot (1919), are also to be observed in Pl. (Placoparia) zippei and Pl. (Hawleia) grandis, from the Ordovician of Bohemia. However, in the species tournemini and borni, additional depressions appear on the anterior cephalic border, and the coaptations evolve towards an increasing complexity. From Pl. (Placoparia) cambriensis, an ancestrai form with a wide geographic distribution, two distinct populations se em to have individu· alised by a process of allopatric speciation; one of these, probably neotenous, is represented by Pl. (Coplacoparia ) tournemini (Massif Armoricain and Iberian Peninsula), the other by Pl. (Placoparia ) zippei (Bohemia). Geo graphic isolation may be indirectly responsible for the appearance of new coaptative devices. jean-Louis Henry, Laboratoire de Paleontologie et de Stratigraphie, Institut de Geologie de l'Universite, B.P. -
PROGRAMME ABSTRACTS AGM Papers
The Palaeontological Association 63rd Annual Meeting 15th–21st December 2019 University of Valencia, Spain PROGRAMME ABSTRACTS AGM papers Palaeontological Association 6 ANNUAL MEETING ANNUAL MEETING Palaeontological Association 1 The Palaeontological Association 63rd Annual Meeting 15th–21st December 2019 University of Valencia The programme and abstracts for the 63rd Annual Meeting of the Palaeontological Association are provided after the following information and summary of the meeting. An easy-to-navigate pocket guide to the Meeting is also available to delegates. Venue The Annual Meeting will take place in the faculties of Philosophy and Philology on the Blasco Ibañez Campus of the University of Valencia. The Symposium will take place in the Salon Actos Manuel Sanchis Guarner in the Faculty of Philology. The main meeting will take place in this and a nearby lecture theatre (Salon Actos, Faculty of Philosophy). There is a Metro stop just a few metres from the campus that connects with the centre of the city in 5-10 minutes (Line 3-Facultats). Alternatively, the campus is a 20-25 minute walk from the ‘old town’. Registration Registration will be possible before and during the Symposium at the entrance to the Salon Actos in the Faculty of Philosophy. During the main meeting the registration desk will continue to be available in the Faculty of Philosophy. Oral Presentations All speakers (apart from the symposium speakers) have been allocated 15 minutes. It is therefore expected that you prepare to speak for no more than 12 minutes to allow time for questions and switching between presenters. We have a number of parallel sessions in nearby lecture theatres so timing will be especially important. -
001-012 Primeras Páginas
PUBLICACIONES DEL INSTITUTO GEOLÓGICO Y MINERO DE ESPAÑA Serie: CUADERNOS DEL MUSEO GEOMINERO. Nº 9 ADVANCES IN TRILOBITE RESEARCH ADVANCES IN TRILOBITE RESEARCH IN ADVANCES ADVANCES IN TRILOBITE RESEARCH IN ADVANCES planeta tierra Editors: I. Rábano, R. Gozalo and Ciencias de la Tierra para la Sociedad D. García-Bellido 9 788478 407590 MINISTERIO MINISTERIO DE CIENCIA DE CIENCIA E INNOVACIÓN E INNOVACIÓN ADVANCES IN TRILOBITE RESEARCH Editors: I. Rábano, R. Gozalo and D. García-Bellido Instituto Geológico y Minero de España Madrid, 2008 Serie: CUADERNOS DEL MUSEO GEOMINERO, Nº 9 INTERNATIONAL TRILOBITE CONFERENCE (4. 2008. Toledo) Advances in trilobite research: Fourth International Trilobite Conference, Toledo, June,16-24, 2008 / I. Rábano, R. Gozalo and D. García-Bellido, eds.- Madrid: Instituto Geológico y Minero de España, 2008. 448 pgs; ils; 24 cm .- (Cuadernos del Museo Geominero; 9) ISBN 978-84-7840-759-0 1. Fauna trilobites. 2. Congreso. I. Instituto Geológico y Minero de España, ed. II. Rábano,I., ed. III Gozalo, R., ed. IV. García-Bellido, D., ed. 562 All rights reserved. No part of this publication may be reproduced or transmitted in any form or by any means, electronic or mechanical, including photocopy, recording, or any information storage and retrieval system now known or to be invented, without permission in writing from the publisher. References to this volume: It is suggested that either of the following alternatives should be used for future bibliographic references to the whole or part of this volume: Rábano, I., Gozalo, R. and García-Bellido, D. (eds.) 2008. Advances in trilobite research. Cuadernos del Museo Geominero, 9. -
U.S. Government Printing Office Style Manual, 2008
U.S. Government Printing Offi ce Style Manual An official guide to the form and style of Federal Government printing 2008 PPreliminary-CD.inddreliminary-CD.indd i 33/4/09/4/09 110:18:040:18:04 AAMM Production and Distribution Notes Th is publication was typeset electronically using Helvetica and Minion Pro typefaces. It was printed using vegetable oil-based ink on recycled paper containing 30% post consumer waste. Th e GPO Style Manual will be distributed to libraries in the Federal Depository Library Program. To fi nd a depository library near you, please go to the Federal depository library directory at http://catalog.gpo.gov/fdlpdir/public.jsp. Th e electronic text of this publication is available for public use free of charge at http://www.gpoaccess.gov/stylemanual/index.html. Use of ISBN Prefi x Th is is the offi cial U.S. Government edition of this publication and is herein identifi ed to certify its authenticity. ISBN 978–0–16–081813–4 is for U.S. Government Printing Offi ce offi cial editions only. Th e Superintendent of Documents of the U.S. Government Printing Offi ce requests that any re- printed edition be labeled clearly as a copy of the authentic work, and that a new ISBN be assigned. For sale by the Superintendent of Documents, U.S. Government Printing Office Internet: bookstore.gpo.gov Phone: toll free (866) 512-1800; DC area (202) 512-1800 Fax: (202) 512-2104 Mail: Stop IDCC, Washington, DC 20402-0001 ISBN 978-0-16-081813-4 (CD) II PPreliminary-CD.inddreliminary-CD.indd iiii 33/4/09/4/09 110:18:050:18:05 AAMM THE UNITED STATES GOVERNMENT PRINTING OFFICE STYLE MANUAL IS PUBLISHED UNDER THE DIRECTION AND AUTHORITY OF THE PUBLIC PRINTER OF THE UNITED STATES Robert C. -
Scope: Munis Entomology & Zoology Publishes a Wide Variety of Papers
_____________ Mun. Ent. Zool. Vol. 4, No. 1, January 2009___________ I MUNIS ENTOMOLOGY & ZOOLOGY Ankara / Turkey II _____________ Mun. Ent. Zool. Vol. 4, No. 1, January 2009___________ Scope: Munis Entomology & Zoology publishes a wide variety of papers on all aspects of Entomology and Zoology from all of the world, including mainly studies on systematics, taxonomy, nomenclature, fauna, biogeography, biodiversity, ecology, morphology, behavior, conservation, paleobiology and other aspects are appropriate topics for papers submitted to Munis Entomology & Zoology. Submission of Manuscripts: Works published or under consideration elsewhere (including on the internet) will not be accepted. At first submission, one double spaced hard copy (text and tables) with figures (may not be original) must be sent to the Editors, Dr. Hüseyin Özdikmen for publication in MEZ. All manuscripts should be submitted as Word file or PDF file in an e-mail attachment. If electronic submission is not possible due to limitations of electronic space at the sending or receiving ends, unavailability of e-mail, etc., we will accept “hard” versions, in triplicate, accompanied by an electronic version stored in a floppy disk, a CD-ROM. Review Process: When submitting manuscripts, all authors provides the name, of at least three qualified experts (they also provide their address, subject fields and e-mails). Then, the editors send to experts to review the papers. The review process should normally be completed within 45-60 days. After reviewing papers by reviwers: Rejected papers are discarded. For accepted papers, authors are asked to modify their papers according to suggestions of the reviewers and editors. Final versions of manuscripts and figures are needed in a digital format.