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University Microfilms International 300 N. Zeeb Road Ann Arbor, Ml 48106 8526272 Welbourn, Warren Calvin, Jr. PHYLOGENETIC STUDIES OF TROMBIDIOID MITES The Ohio Stale University Ph.D. University Microfilms International300 N. Zeeb Road, Ann Arbor, Ml 48106 Copyright 1985 by Welbourn, Warren Calvin, Jr. All Rights Reserved PLEASE NOTE: In all cases this material has been filmed in the best possible way from the available copy. Problems encountered with this document have been identified here with a check mark V . 1. Glossy photographs or pages_____ 2. Colored illustrations, paper or print_______ 3. Photographs with dark background_____ 4. Illustrations are poor copy______ 5. Pages with black marks, not original copy______ 6. Print shows through as there is text on both sides of p_______ ag e 7. Indistinct, broken or small print on several pages\/ 8. Print exceeds margin requirements______ 9. Tightly bound copy with print lost in spine_______ 10. Computer printout pages with indistinct print_______ 11. Page(s)____________lacking when material received, and not available from school or author. 12. Page(s)____________seem to be missing in numbering only as text follows. 13. Two pages num bered . Text follows. 14. Curling and wrinkled pages______ 15. Dissertation contains pages with print at a slant, filmed as received__________ 16. Other___________________________________________________________________________ University Microfilms International PHYLOGENETIC STUDIES OF TROMBIDIOID MITES DISSERTATION Presented in Partial Fulfillment of the Requirements for the Degree Doctor of Philosophy in the Graduate School of The Ohio State University By Warren Calvin Welbourn Jr, B.A., M.A. sjc >je 3jc >jc Sjt The Ohio State University 1985 Reading Committee: Approved By Donald E. Johnston Susan W. Fisher Norman F. Johnson Adviser Department of Entomology Copyright by Warren C. Welbourn Jr 1985 ACKNOWLEDGEMENTS I wish to thank my advisor, Dr. Donald E. Johnston, for his advice, assistance and patience during this study. Very special thanks go to my wife and family for their continuing support. Funds for my 1984 trip to the British Museum (Natural History), London, Museum National D’Histoire Naturelle, Paris, Museum D ’Histoire Naturelle, Geneva and Istituto Spermentale per la Zoologia Agrara (Berlese Collection), Florence and related field work in England and Italy was supported by a National Science Foundation Dissertation Grant (BSR-8401206). Travel from Florence, Italy to University "Al. I. Cuza." in Iasi, Romania was partially supported by an Ohio State University Graduate Student Alumni Research Award. My visit to the Smithsonian Institution in April, 1981 was supported by a Visiting Researcher Grant from the Smithsonian Institution. I would like to acknowledge the assistance of Ms. Anne Baker, Dr. Don Macfarlane, Mr. Keith Hyatt and Mr. Keiren Martin in the Arachnid Section, of the British Museum (Natural History), London, for their assistance in 1982 and 1984 and arranging for loans of important specimens. I also want to thank Mrs. Judith Marshall (Entomology) for her assistance in my examination of the Orthopteran collections for mites. I am especially grateful to Anne Baker and Paul Hillyard for arranging a very successful spring 1984 collecting trip to south Wales. I also want to thank Clive and Diane Bowman for their hospitality during my two visits to London. I want to acknowledge Dr. Yves Coineau, Director of the Laboratory of Arthropods, Paris for his assistance while in Paris, as well as Mr. Michel Naudo for his patience and assistance as well as arranging for the loan of important specimens. I am especially grateful to Dr. Pierre Robaux for opening his laboratory to me and for the stimulating discussions on trombidioid mites, in addition to his hospitality. I am also grateful to Dr. Bernard Hauser of the Geneva Natural History Museum for his assistance in locating specimens in the Vercammen-Grandjean collection. I am indebted to Dr. Fausta Pegazano for her assistance in making ray visit to the Berlese Collection successful. Dr. Roberto Nannelli is appreciated for his assistance and taking the time to take me collecting at several of Berlese's collecting localities in the mountains around Florence. I wish to gratefully thank Dr. Libertina Solomon for providing space in her laboratory and locating trombidioid specimens in the Feider Collection. I am especially grateful for her family's hospitality while I was in Iasi. The 1982 field work in California and Oregon was supported by a small grant from The Ohio State University to Dr. Donald E. Johnston. I thank Dr. G. W. Krantz for opening his laboratory to us, and Dr. David Walter for his assistance in Corvallis. I acknowledge Drs. G. W. Wharton and G. R. Needham of the Acarology Laboratory for their assistance, interest and support. I am especially grateful to Dr. Edward Baker for his continuing assistance in locating missing specimens and lost literature. I also thank Drs. Evert E. Lindquist (Agriculture Canada) and John B. Kethley (Field Museum Natural of History, Chicago) for their consultation and support. I am especially indebted to the many people who loaned and/or donated specimens for this research. In addition to the persons mentioned above I thank Dr. B. Feldman-Muhsam (Hebrew University Hadassah Medical School, Jerusalem), Dr. Torbjorn Kornestedt (Naturhistoriska Rimsrauset, Stockholm), Dr. David Lee (South Australia Museum), Drs. Ian Smith and Evert Lindquist (Agriculture Canada, Ottawa), Dr. L. van der Hamraen (Rijksrauseura voor Natuurlijke Historie, Leiden), Drs. R. G. Beer and G. W. Byers (Snow Entomological Museum, University of Kansas), Dr. H. Levi (Museum of Comparative Zoology, Harvard University), Dr. Norman Platnick (American Museum of Natural History, New York), Dr. Gary Mullen (Auburn University, Auburn), Dr. Barry M. OConnor (Museum of Zoology, University of Michigan, Ann Arbor), Dr. Donald Tuttle (University of Arizona Experiment Station, Yuma), Dr. John Moser (USDA, Forest Service, Southern Forest Experiment Station, Pineville), Dr. John B. Kethley (Field Musuera of Natural History, Chicago), Dr. Wayne Moss (Philadelphia Academy of Natural Sciences, Philadelphia), Dr. Harold Denmark (Florida Department of Agriculture and Consumer Services, Division of Entomology, Gainsville). To the numerous other people who contributed directly or indirectly to this reseach I am very appreciative. VITA December 19, 1946 ......... Born - Pasadena, California 1966.............................A.A. - Zoology Citrus College Azusa, California 1969........................... B.A. - Biology California State University, Long Beach, California 1972........................... M.A. - Biology California State University Long Beach, California 1971-1978 ...................... Teacher Albuquerque Public Schools Albuquerque, New Mexico 1978-1985 ...................... Collection Manager Acarology Laboratory Department of Entomology The Ohio State University Columbus, Ohio PUBLICATIONS Loomis, R. B. and W. C. Welbourn 1969. A new species of Hannemania (Acarina, Trombiculidae) from Bufo punctatus of western North America, with comments on Hannemania hylae Ewing. Bull. So. Calif. Acad. Sci. 69:160-168. Welbourn, W. C. and R. B. Loomis 1970. Three new species of Hannemania (Acarina, Trombiculidae) from amphibians of western Mexico. Bull. So. Calif. Acad. Sci. 69: 65-73. Welbourn, W. C. 1972. Studies of two species of Hannemania (Acarina, Trombiculidae) on amphibians in western North America, (abstract) Masters Abstracts 10 No. 4. Welbourn, W. C.