FRAGMENTA PALAEONTOLOGICA HUNGARICA 22, BUDAPEST, 2004

Contributions to the Jurassic of Kachchh, western India VIII. The gastropod fauna. Part II: Discohelicidae, Neritomorpha,

by János SZABÓ & Anand K. JAITLY

Abstract — Twenty-seven species of gastropods have been identified and systematically described from the Middle to early Late Jurassic of Kachchh (western India). Most of them have been already known from the deposits of the epicontinental seas around the western part of the Tethys but a few of them are new: Neritoma kachchhensis n. sp., Globularia khadirensis n. sp., Coelodiscus sadharaensis n. sp. and Pseudamaura hadibadhang- ensis n. sp.

Keywords — Jurassic, , systematics, Kachchh, India.

SZABÓ,J.&JAITLY, A. K.: Contributions to the Jurassic of Kachchh, western India VIII. The gastropod fauna. Part II: Discohelicidae, Neritomorpha, Caenogastropoda. — Fragmenta Palaeontologica Hungarica, 22: 9–26.

Introduction This contribution is the third one in a series of papers, gastropod genera (Bhujnerita, Neritidae), which seem to be which are based on the collections of Jurassic from endemic in the “Ethiopian Faunal Province”. localities of Kachchh (State Gujarat, western India). The Paralelly, another team has been also working on the type specimens of this paper are deposited at the Banaras Jurassic gastropods of the same area; they published their Hindu University, Varanasi (India); complementary mater- results some months earlier (DAS et al. 1999) than JAITLY ial is kept in the collections of the Würzburg University. et al. (2000). Therefore some of the new species, found in A sketch of the geology and stratigraphy of the localities Kachchh, have already synonyms. In the following list, are added to the first part of the systematical results revised names of the species, published from Kachchh, is (JAITLY et al. 2000). Another paper (JAITLY &SZABÓ given (bold letters indicate valid names; junior synonyms 2002) wishes to draw attention to one of the few Jurassic immediately follow the senior ones in square brackets).

1. Pleurotomaria sp. 29. Biarmatoidella tagorei (DAS,BARDHAN &LAHIRI, 1999) 2. Anodomaria latigradata JAITLY,SZABÓ &FÜRSICH, 2000 [Riselloidea pileiformis JAITLY,SZABÓ &FÜRSICH, 2000] 3. Anodomaria depressa JAITLY,SZABÓ &FÜRSICH, 2000 30. Helicanthus chanda DAS,BARDHAN &LAHIRI, 1999 4. Anodomaria? sp. 31. Metriomphalus (Metriomphalus) cf. lyelli (D’ARCHIAC, 1843) 5. Obornella granulata (J. SOWERBY, 1818) 32. Amphitrochus angulatus (HUDLESTON, 1894) 6. Obornella aff. granulata (J. SOWERBY, 1818) [Colpomphalus jumarense DAS,BARDHAN &LAHIRI, 1999] 7. Obornella wuertembergensis (SIEBERER, 1908) 33. Amphitrochus acutispira JAITLY,SZABÓ &FÜRSICH, 2000 8. Leptomaria fraga (J. A. EUDES-DESLONGCHAMPS, 1849) 34. Amphitrochus orbis JAITLY,SZABÓ &FÜRSICH, 2000 9. Leptomaria pseudoumbilicata JAITLY,SZABÓ &FÜRSICH, 2000 35. Neritopsis (Neritopsis) benoisti COSSMANN, 1900 10. Leptomaria simplex JAITLY,SZABÓ &FÜRSICH, 2000 [Neritopsis (Neritopsis) patchamensis DAS,BARDHAN &LAHIRI, 1999] 11. Bathrotomaria kachchhensis JAITLY,SZABÓ &FÜRSICH, 2000 36. Neritopsis (Hayamiella) sankhamata DAS,BARDHAN &LAHIRI, 1999 12. Bathrotomaria siebereri JAITLY,SZABÓ &FÜRSICH, 2000 37. Neritopsis sp. 13. Bathrotomaria millepunctata (J. A. EUDES-DESLONGCHAMPS, 1849) 38. Hayamia subsulcosa (D’ORBIGNY, 1850) 14. Bathrotomaria sp. [Hayamia mitra DAS,BARDHAN &LAHIRI, 1999)] 15. Pyrgotrochus? sp. 39. Hayamia bajocensis (D’ORBIGNY, 1852) 16. Emarginula karuna DAS,BARDHAN &LAHIRI, 1999 40. Neridomus involuta (LYCETT, 1863) [Emarginula (Tauschia) jumarensis JAITLY,SZABÓ &FÜRSICH, 2000] 41. Neridomus sp. 17. Discohelix cf. sapho (D’ORBIGNY, 1853) 42. Neritoma kachchhensis n. sp. 18. Proconulus jadavpuriensis DAS,BARDHAN &LAHIRI, 1999 43. Bhujnerita bhattii JAITLY &SZABÓ, 2002 [Proconulus (Proconulus) bhujensis JAITLY,SZABÓ &FÜRSICH, 2000] 44. remtsaensis COX, 1969 19. Proconulus (Proconulus) haboensis JAITLY,SZABÓ &FÜRSICH, 2000 45. Pseudomelania calloviensis (HEBERT &DESLONGCHAMPS,1860) 20. Proconulus (Proconulus) wagadensis JAITLY,SZABÓ &FÜRSICH, 2000 46. Pseudomelania ? sp. 21. Proconulus (Laeviconulus?) sp. 47. Bourgetia saemanni (OPPEL, 1856) 22. Calliostoma? sp. 48. Ampullospira (Ampullospira) brevispira J.–C. FISCHER,2001 23. Ataphrus (Ataphrus) kaladongarensis JAITLY,SZABÓ &FÜRSICH, 2000 49. Ampullospira (Pictavia) lorierei (D’ORBIGNY, 1852) 24. Adeorbisina? sp. 50. Ampullospira (Pictavia) bajocensis (D’ORBIGNY, 1852) 25. Eucyclus laevijugatus (QUENSTEDT, 1884) 51. Ampullospira (Pictavia) tanganyicensis COX, 1965 26. Onkospira? kutchensis DAS,BARDHAN &LAHIRI, 1999 52. Ampullospira (Pictavia)cf. pyramidata (MORRIS &LYCETT, 1851) [Onkospira? praecursor JAITLY,SZABÓ &FÜRSICH, 2000] 53. Ampullospira (Pictavia) aff. tanganyicensis Cox, 1965 27. Onkospira? aff. kutchensis DAS,BARDHAN &LAHIRI, 1999 54. Ampullospira (Pictavia) sp. [Onkospira? aff. praecursor JAITLY,SZABÓ &FÜRSICH, 2000] 55. Globularia spathi DAS,BARDHAN &LAHIRI, 1999 28. Biarmatoidella biarmata (MÜNSTER, 1844) 56. Globularia khadirensis n. sp. [Riselloidea elongata DAS,BARDHAN &LAHIRI, 1999] 57. Globularia michelini (D’ARCHIAC, 1843)

Fragmenta Palaeontologica Hungarica 22, 2004 10 SZABÓ &JAITLY 58. Globularia ? aff. michelini (D’ARCHIAC, 1843) 62. Lamelliphorus cf. ornatissimus (D’ORBIGNY, 1853) 59. Naricopsina cornelia (LAUBE, 1868) 63. Coelodiscus sadharaensis n. sp. 60. Megatylotus sp. 64. Kosmomphalus reticulatus FISCHER, 2001 61. Pseudamaura hadibadhangensis n. sp.

Abbreviations to measurements — H = total height; HL = height of last whorl; HP = height of peristome; D = diameter of last whorl; WP = width of peristome; AA = apical angle; AL = coiling angle of last whorl (if different from apical angle). Asterisks indicate reconstructed measurements taken on more (**) or less (*) damaged shell parts.

Class Gastropoda CUVIER, 1797 Subclass Prosobranchia MILNE EDWARDS, 1848 Order ?Vetigastropoda SALVINI-PLÄWEN, 1980 Superfamily ?Cirroidea COSSMANN, 1916 Family Discohelicidae SCHRÖDER, 1995 Genus Discohelix DUNKER, 1848

Discohelix cf. sapho (D’ORBIGNY, 1853) (Plate I: 1–2)

1853: Straparollus Sappho — D’ORBIGNY, p. 315, Pl. 323, Figs 9–13. 1997: Discohelix sapho (D’ORBIGNY, 1853) — FISCHER &WEBER, p. 124, Pl. 22, Figs 6a–c. Material — A single, badly damaged specimen. Measurements H HL HP D WP AA AL BHU2001I 30 Plate I: 1–2 4.2* 4.2* 4.2* 9* 2.2* 228° Description — Biconcave, sinistral, discoidal shell of permits only an open identification because the morpho- small size. Whorl cross-section symmetrically trapezoidal logy of the nodules are hardly visible. They appear as undu- with feebly convex spiral and umbilical sides and more lation of the inner mould and the shell cross section of the convex outer side on spire whorls. Last whorl cross-section outer face. Beside this, the spiral angle (concavity of spire) tends to be circular. Both outer angulations bear weak seems somewhat greater than in the holotype of D. sapho carina. Ornament consists of riblet like growth-lines on (D’ORBIGNY); name corrected by FISCHER &WEBER (1997). outer face, and tubercles and nodes on outer angulations Discohelix sapho (D’ORBIGNY, 1853) has been recorded (delimiting outer face). Tubercles gradually strengthen and from the Oxfordian in France. nodes develop on last whorl. Distribution in Kachchh — Upper part of Callovian Remarks — The available, poorly preserved specimen to Oxfordian Chari Formation, Ler.

Order Neritopsina COX &KNIGHT, 1960 Superfamily Neritopsioidea RAFINESQUE, 1815 Family Neritopsidae GRAY, 1847 Genus Neritopsis GRATELOUP, 1832 Subgenus Neritopsis GRATELOUP, 1832

Neritopsis (Neritopsis) benoisti COSSMANN, 1900 (Plate I: 11–12, 17–21)

1900: Neritopsis Benoisti nov. sp. — COSSMANN, p. 567, pl. XVII, figs. 18, 23. 1918: Neritopsis Benoisti COSSMANN —ROLLIER, p. 23. 1964: Neritopsis benoisti COSSMANN — J.–C. FISCHER, p. 62, pl. II. figs 12a–b. 1999: Neritopsis (Neritopsis) patchamensis sp. nov. — DAS,BARDHAN &LAHIRI, p. 278, Figure 7: 3–5. Material — Seven, more or less incomplete specimens. Measurements H HL HP D WP AA AL BHU2001I 8 Plate I: 17–19 11.5 11.0 9.5 12.0 9.0 120° BHU2001I 9 Plate I: 20–21 11.5** 10.0** 122° BHU2001I 10 Plate I: 11–12 13.5 13.0 11.0 18.5 13.5* 133°* Description — Rather small species of the genus with whorl surface evenly arched. Sutural part and columellar lip low-spire shell, consisting of rapidly expanding whorls, of peristome partly preserved on one specimen. Frag- separated by widely canaliculate suture. Narrow, feebly ments, as well as sculpture, indicate nearly circular shape of convex ramp below sutural furrow; remaining part of peristome almost in one plane, being strongly prosocline

Fragmenta Palaeontologica Hungarica 22, 2004 Jurassic gastropods from Kachchh 11 and almost tangential to base. Peristome fragments have has sparser collabral ribs than N. benoisti, moreover, sharp outer edge; columellar lip looks like keel. Adapically alternating stronger and weaker spiral cords cross the ribs. narrowing furrow forms false umbilicus along columellar The density of the stronger ones is similar to that of the lip. Ornament consists of marked, cancellate elements of collabral ribs. The Callovian Neritopsis (Neritopsis) guerrei dense spiral cords and sparser, somewhat stronger collabral HEBERT &DESLONGCHAMPS, 1860 is another species of ribs. Number of cords between suture and inner lip falls in high resemblance but the three figured specimens (Pl. I, range of 14 to 16 on latest segment of shells. Ribs run from Figs 4 a–d) show such high variability that a totally reliable suture to suture, or to columella on last whorl. Crossings distinction method can not be given. Its collabral ribs seem of network form low spines. Growth-lines dense and sparser than in N. benoisti but they are denser (and weaker) delicate. on the last whorl of the specimen displayed in HEBERT & Remarks — All morphological characters indicate DESLONGCHAMPS’ (1860) Plate I, Fig. 4 c. The spiral cords identity with Neritopsis benoisti COSSMANN, 1900 on one in N. guerrei are every case much weaker than the collabral hand, and with Neritopsis (Neritopsis) patchamensis DAS,BAR- ribs but the spiral elements are almost as strong in N. DHAN &LAHIRI, 1999 on the other hand; consequently the benoisti as the collabral ones. N. guerrei has at least two well latter name has to be qualified as a junior synonym. distinguishable generations of spiral ornament but the Neritopsis (Neritopsis) cossmanni ROLLIER, 1918 (= N. spiral cords in N. benoisti are almost uniform. (N. guerrei deslongchampsi COSSMANN, 1885, being jun. homonym to N. seems so highly variable that one may think the figured deslongchampsi BEAUDOUIN, 1869) is most resembling to specimens belong to more than a single species). Neritopsis (Neritopsis) benoisti COSSMANN, 1900 amongst the Distribution in Kachchh — Coral Bed of Jhurio similarly aged (Bathonian) species. However, N. cossmanni Formation (Bathonian), Jumara Dome.

Neritopsis sp. (Plate I: 13–16) Material — Single specimen. Measurements H HL HP D WP AA AL BHU2001I 4 Plate I: 13–16 11.0* 10.5* 9.5* 12.0 8.5 133° Description — Small, depressed specimen with wide however, its elements are much denser and equally strong subsutural ramp, being abaxially bounded by angulation dislike in Neritopsis sp. On these differences, the two in earlier growth phases. Angulation vanishing on last species can be distinguished. whorl of available specimen. Peristome not completely The juvenile part of Neritopsis (Hayamiella) sankhamala preserved, its sutural corner strongly thickened; aperture DAS,BARDHAN &LAHIRI, 1999 shows a comparable suborbicular. Rather broad (?false) umbilicus separates sculpture but both the collabral and spiral elements are almost vertical columellar lip from base. Four strong spi- denser (like in N. benoisti), and the shape is more globular ral carinae, weaker spiral cords and sparse collabral ribs than that of Neritopsis sp. result in marked network ornament. Distribution in Kachchh — Coral Bed of Jhurio Remarks — In the studied material, Neritopsis benoisti Formation (Bathonian), Jumara Dome. COSSMANN, 1900 has also a marked cancellate ornament,

Genus Hayamia KASE, 1980

Hayamia bajocensis (D’ORBIGNY, 1852) (Plate I: 22–25)

1852: Neritopsis bajocensis D’ORBIGNY, 1847 — D’ORBIGNY, p. 223, Pl. 300, Figs 8–10. 1852: Neritopsis baugeriana D’ORBIGNY, 1852 — D’ORBIGNY, p. 224, Pl. 300, Figs 11–13. 1868: Neritopsis bajocensis D’ORBIGNY, 1850 — LAUBE, p. 6, Pl. 1, Fig. 9. 1894: Neritopsis bajocensis D’ORBIGNY, 1850 — HUDLESTON, p. 340, Pl. 27, Figs 9 a–c. 1997: Neritopsis bajocensis D’ORBIGNY, 1852 — FISCHER &WEBER, p. 86, Pl. 17, Figs 14 a–c, 15a–c. Material — Six specimens, three of them poorly preserved. Measurements H HL HP D WP AA AL BHU2001I 11 Plate I: 23–25 29.0* 28.0* 24.0* 26.0* 19.5* 115° BHU2001I 12 Plate I: 22 21.0** 20.0** 16.0** 21.5** 18.5** 125° Description — Rather large, thick-walled, globular Aperture slightly ovate axially, peristome does not lie in shells of relatively high spire (shell height bigger than single plane; upper part of outer lip more prosocline than diameter). Whorls and base evenly arched, except narrow, remaining parts. Columellar lip separated from base by subsutural concave belt where shell upward bending. rather wide and deep furrow (pseudoumbilicus).

Fragmenta Palaeontologica Hungarica 22, 2004 12 SZABÓ &JAITLY Evenly spaced, sparse spiral cords cover whorls and and this opinion was subsequently accepted also by COSS- base with four to six weaker threads between their closest MANN (1885) and FISCHER &WEBER (1997). pairs. Regularly repeating, rounded, suture to suture ribs In the remarks of designation of Hayamia, KASE (1980, cross them on spire whorls. During growth, ribs change p. 303.) called the attention to a figured in situ operculum into wide, low ridges or undulations, which usually fade of “Neritopsis” bajocensis D’ORBIGNY (in HUDLESTON 1896) away around periphery on last whorl. that indicates belonging to the genus Hayamia KASE, 1980. Remarks — Though the Kachchh specimens are In stable Europe, “Neritopsis” bajocensis has been found rather poorly preserved, they are well identifiable. The in the Bajocian (Humphresianum Zone) to Bathonian only significant difference from D’ORBIGNY’s specimens (Retrocostatum Zone) interval (FISCHER &WEBER is the wider and deeper basal furrow along the columellar 1997). lip (compare to FISCHER &WEBER 1997, Pl. 17, Fig. 15c). Distribution in Kachchh — Lower Callovian part of The valid date of the species description is 1852 by Chari Formation, Jumara Dome; ?Bathonian, S of Cheriya FISCHER &WEBER (1997). LAUBE (1868) unified D’OR- Bet, Khadir Island and Bathonian of Sponge Limestone BIGNY’s two species, Neritopsis bajocensis and N. baugeriana, Member, Patcham Formation, Jumara Dome.

Hayamia subsulcosa (D’ORBIGNY, 1850) (Plate I: 26)

1843: Neritopsis sulcosa BROCH. var. — D’ARCHIAC, p. 377, pl. XXVIII, figs 10, 10a. (non BROCCHI) 1850: Stomatia subsulcosa D’ORBIGNY — D’ORBIGNY, p. 301. 1851: Neritopsis sulcosa (D’ARCHIAC) — MORRIS &LYCETT, p. 59, Pl. XI, Fig. 12 1853: Stomatia subsulcosa D’ORBIGNY — D’ORBIGNY, p. 372, Pl. 339, Figs 4–5. 1885: Neritopsis sulcosa D’ARCHIAC —COSSMANN, p. 159, Pl. III, figs 15–17. 1950: Neritopsis subsulcosa (D’ORBIGNY)—COX &ARKELL, p. 64. 1969: Neritopsis subsulcosa (D’ORBIGNY) — J.–C. FISCHER, p. 134, Pl. XV, Figs 12 a–c 1915: Neritopsis sulcosa D’ARCH. — ROLLIER, p. 22. 1997: Neritopsis subsulcosa (D’ORBIGNY) — J.–C. FISCHER &WEBER, p. 149. 1999: Hayamia mitra sp. nov. — DAS,BARDHAN &LAHIRI, p. 281, Fig. 8: 1–3. Material — Three inner moulds with shell portions. Measurements H HL HP D WP AA AL BHU2001I 15 Plate I: 26 20.0 19.0 14.5 20.0 14.5 135° 120° BHU2001I 16 (not figured) 17.0 16.5 13.0 21.0 11.5 140°* 130° Description — Low spired, thick-walled shell of who found characteristic operculum in situ in a Hayamia rapidly expanding whorls (shell diameter bigger than “mitra” specimen. height). Suture narrowly canaliculate, no distinct ramp Most similar species is the Toarcian Neritopsis abbas ROL- below it. Adapical and basal parts of whorls less but LIER, 1919 [the name was given to HUDLESTON’s N. philea peripheral part more strongly arched. Last whorl feebly variety (1894, p. 341, Pl. XXVII, Figs 11a–c). Due to its deviating downward. Ornament of whorls and base con- larger and higher (H/D value) shell, this species is distin- sists of dense, finely granulate spiral cords with threads guishable from Hayamia subsulcosa (D’ORBIGNY). in their interspaces. Delicate growth-lines cause corruga- Hayamia subsulcosa (D’ORBIGNY, 1850) is Middle to tion of spiral ornamental elements. Late Bathonian in “stable” Europe Remarks — No operculum has been found in the Distribution in Kachchh — Lower Callovian Chari available specimens; the generic identification follows Formation of Jumara Dome and Kamaghuna section of KASE (1980, p. 303) opinion, verified by DAS et al., 1999 Jhura Dome.

Superfamily Neritoidea RAFINESQUE, 1815 Family Neritidae RAFINESQUE, 1815 Genus Neridomus MORRIS &LYCETT, 1851

Neridomus involuta (LYCETT, 1863) (Plate I: 27–29)

1863: Nerita involuta — LYCETT, p. 20, Pl. XXXI, Fig. 15. 1950: Neridomus involuta LYCETT — COX &ARKELL, p. 66. Material — One specimen; apex somewhat eroded. Measurements H HL HP D WP AA AL BHU2001I 14 Plate I: 27–29 20.5 20.5 18.5 21.5 13.0 170°* 151°

Fragmenta Palaeontologica Hungarica 22, 2004 Jurassic gastropods from Kachchh 13 Description — Medium size, slightly depressed, ovate Remarks — The state of preservation does not permit shell with extremely low spire that consists of about two to see clearly the morphology of the early shell. Shape of flattened whorls, separated by flush suture. Wide subsutu- the last one and a half whorl and some remnants of earlier ral ramp on last whorl without distinct outer rim. No outer parts suggest the juvenile shell is sunken into the penulti- lip part preserved; parietal lip with large but thin callus, mate whorl. With the depressed spire, Neridomus involuta delimited from base by shallow groove. Shell-wall seems has a unique shape. The visible morphological elements unusually thin, unlike in other species of genus Neridomus. suggest its identity with LYCETT’s species. Growth-lines rather marked, their strength irregularly Neridomus involuta is Bathonian in Europe. changes. Traces of original shell pattern (dark dots and Distribution in Kachchh — Lower Yellow Flagstone spots) preserved on spire and adapical part of last whorl. Member (Bathonian), Pachchhmai Pir, Pachchham Island.

Neridomus sp. (Plate I: 30–32) Material — Four damaged specimens. Measurements H HL HP D WP AA AL BHU2001I 1 Plate I: 30–32 13.5 13.0 10.0 13.0* 10.0* 132° 112° BHU2001I 2 not figured 13.0 12.5 11.5* 12.5 9.0 130° 115° Description — Small-medium size, ovate shell with cally much younger (Middle Oxfordian) Neridomus mais widely cyrtoconical spire outline. Suture in marked inci- (BUVIGNIER, 1843) but it has a higher spire and a stronger, sion, no subsutural ramp present. Shell wall prominently swelling-like parietal callus that is truncated by a marked thick. Parietal callosity belt rather narrow but its thickness sinus at the short columella. considerable; sutural peristome corner also infilled by Distribution in Kachchh — Hadibhadang Shale parietal callus. Member (Bathonian), Khadir Island and ?Bajocian/Batho- Remarks — A comparable species is the stratigraphi- nian, Kuar Bet.

Genus Neritoma MORRIS, 1849

Neritoma kachchhensis n. sp. (Plate I: 33–37) Holotype — BHU2001I 46 (Plate I: 33–37) Type locality — Khadir Island, Kachchh, western India. Type strata — Hadibadhang Shale Member of Khadir Formation, Bathonian. Species name — From the name of Kachchh district, Gujarat. Diagnosis — Slightly depressed shell with low spire; rather flush suture; wide, feebly convex ramp; peristome sinus and sinus belt above periphery, on lower edge of ramp. Material — Two damaged specimens with repaired shell parts. Measurements H HL HP D WP AA AL BHU2001I 2 not figured 13.0 12.5 11.5* 12.5 9.0 130° BHU2001I 3 Plate I: 33–37 14.5 13.5 13.0 15.0 11.5 140° Description — Subglobular-ovate shell with flush elements are different in the Kachchh species: spire is suture. Shell-wall moderately thick. Outer lip subcircular, just feebly protruding and the position of the periphery parietal lip almost straight, and having only narrow callous gradually shifts below the sinus (sinus belt). belt. Growth-lines strong and arranged into irregular, The spire region is rather similar to that of N. ridge-like bundles. Marked peristomal sinus that gradually bisinuata (BUVIGNIER, 1842) from the West European gets fairly above periphery from its initially peripheral Oxfordian. However, this species differs in having also position; more exactly: position of periphery shifts down- peripheral sinus, and in the shape of the last whorl, ward from onset of last whorl; sinus-belt well visible on which downward deviates from the normal coiling near whole last whorl. Depth of sinuses on growth lines the last peristome while increasingly exposes the pen- increasing towards peristome. ultimate whorl. Remarks — The type species of the genus (Nerita Distribution in Kachchh — Hadibhadang Shale sinuosa J. SOWERBY, 1818) has a peripheral peristome Member, Khadir Formation, Bathonian, Khadir Island and sinus and a rather high spire. These morphological ?Bajocian–Bathonian, Kuar Bet.

Fragmenta Palaeontologica Hungarica 22, 2004 14 SZABÓ &JAITLY Order Caenogastropoda COX, 1959 Superfamily Pseudomelanioidea FISCHER, 1886 Family FISCHER, 1886 Genus Pseudomelania PICTET &CAMPICHE, 1862

Pseudomelania remtsaensis COX, 1969 (Plate II: 1–5)

1969: Pseudomelania (Oonia) remtsaensis sp. nov. — COX, p. 249, Pl. I, Figs 13 a–b. Material — One hundred and eight specimens in different ontogenetic stages and preservation. Measurements H HL HP D WP AA AL BHU2001I 22 Plate II: 2–3 47.0 28.5 20.0 22.5 17.0 40° 35° BHU2001I 5 Plate II: 4–5 38.5 9.5 9.5 15.5 8.5 45° 38° BHU2001I 13 Plate II: 1 18.5 11.5 8.0 8.5 5.5 41° 34° Description — Medium sized species of the genus. COX (1969) regarded this species as member of Oonia Shells thick-walled, moderately turriculate and consisting GEMMELLARO, 1878 probably because of the cyrto- of few rather high and moderately convex whorls. Juve- conical juvenile shell. However, true Oonia, has cyrto- nile whorls cyrtoconical with blunt apex. Suture shallow- conical outline (complete shell) and much lower spire canaliculate. Peristome entire, outer lip simple and evenly (related to the last whorl). A careful revision of Pseudo- arched, inner lip wide V-shaped; columellar lip heavy but melaniidae is needed to clarify the relation and distinction of parietal lip appears as a callus of moderate thickness. the genera of this family. Remarks — The morphology of most available speci- The North African (Tunisian) occurrence is Early mens well corresponds to that of the holotype. However, Callovian. some specimens show considerable difference in the Distribution in Kachchh — Chari Formation (Am- dimensions of the last whorl, which is more expanded phiastrea Bed), Dhrang section, northern Habo Dome than the earlier ones, exhibiting normal growth (e.g. Plate and Raimalro Limestone Member (Bathonian), South of I: 4–5). Gadhada Village, Khadir Island.

Pseudomelania calloviensis (HÉBERT &DESLONGCHAMPS, 1860) (Plate II: 9–10)

1860: Eulima calloviensis — HÉBERT &DESLONGCHAMPS, p. 35, Pl. 7: Figs 8a–b. 1909: Pseudomelania (Hudlestoniella) calloviensis (HÉBERT &DESLONGCHAMPS)— COSSMANN, p. 95, Pl. 2: Fig. 19. 1968: Pseudomelania sp. cf. P. calloviensis (HÉBERT &DESLONGCHAMPS) — REINER, p. 173, Pl. I: 4–5. 1980: Pseudomelania sp. cf./(aff.) trochiformis (PIETTE, 1855) — HIRSCH, p. 140, 146, Pl. 9: Figs 13–16. Material — A single specimen; apical part and peristome lacking. Measurements H HL HP D WP AA AL BHU2001I 64 Plate II: 9–10 12.5* 8.5* 9* 21° Description — Moderately high-turriculate, small 1980 — Makhtesh Hatira). Elements of REINER’s (1968) species of Pseudomelania. Rather thin-walled shell consists thorough comparison to the European specimens could of relatively high number of whorls. Juvenile whorls slightly be repeated here, however, the authors of the present paper convex but subsequent ones flat or only feebly convex. Rem- think that the differences between the European and the nants of juvenile shell suggest cyrtoconical outline but African specimens (2–3° wider spiral angle of the former subsequent shell part simply high conical. Whorls sepa- ones) are within the frame of a moderate specific vari- rated by flush suture. Periphery rounded angular, base ability (the differences may origin also from the changing anomphalous and convex. No peristome part preserved. spiral angle of the cyrtoconical apical region). Remarks — The specimen is conspecific with some Distribution in Kachchh — Kaladongar, Pachchham Sinai forms (REINER 1968 — Hamaktesh Hagadol; HIRSCH Island, Goradongar Formation, Bathonian.

Pseudomelania ? sp. (Plate I: 46–47)

Material — Two damaged, shelly specimens. Measurements H HL HP D WP AA AL BHU2001I 24 Plate I: 46–47 12.5 9.0 6.5 8.0* 5.5* 55°

Fragmenta Palaeontologica Hungarica 22, 2004 Jurassic gastropods from Kachchh 15 Description — Low spired, small member of genus their opisthocyrt part visible on whorls. Pseudomelania. Spire medium widely conical and composed Remarks — This species has some characters that are of moderately convex whorls that bend upward along not typical in the Pseudomelaniidae but remind also rissoid upper edge. Suture distinct but not deeply cut. Base convex shells (shape of growth lines, rudimentary siphonal outlet). and anomphalous. Aperture pyriform, preserved parts of Further specimens are needed to reach a more reliable peristome (inner lip, basal lip) unthickened. Shallow, identification. wide sinus present on basal lip generating a low ridge on Distribution in Kachchh — Gadhada Formation the base around columella during growth. (Lower Callovian), N of Mouwana village, Mouwana Dome. Growth lines parasigmoidal between suture and columella;

Genus Bourgetia TERQUEM & JOURDY, 1870

Bourgetia saemanni (OPPEL, 1856) (Plate II: 6–8) 1818: Melania striata SOWERBY — J. SOWERBY, p.101, Pl. 47. 1854: Phasianella striata (SOW.) — D’ORBIGNY, p.322, Pl. 324, Fig. 15; Pl. 325, Fig. 1. 1856: Phasianella sämanni OPPEL — OPPEL, p. 387. 1870: Bourgetia striata (SOW.) — TERQUEM &JOURDY, p. 51, Pl. 2, Figs 21–23. 1938: Bourgetia saemanni (OPPEL) — COX, p. 60. 1965: Bourgetia saemanni (OPPEL) — COX, p. 152, Pl. 25, Figs 8-9. 1968: Bourgetia “saemanni” (OPPEL) — REINER, p. 175, pl. I: 12–13. 1997: Bourgetia saemanni (OPPEL) — FISCHER &WEBER, p. 127, Pl. 3, Fig. 1. 2001: Bourgetia saemanni (OPPEL) — FISCHER, p. 79. Material — Two damaged, shelly specimens. Measurements H HL HP D WP AA AL BHU2001I 20 Plate II: 6–7 35.5* 26.0* 18.0 18.5 12.0 30° BHU2001I 21 Plate II: 8 53.5* 36.5 22.0 25.0 18.0 16° Description — High littorinform and moderately variable, persistent and widespread species. Therefore turriculate shells suggest high variability of shapes. Whorls both Kachchh specimens can be regarded as Bourgetia moderately convex with shallow-canaliculate suture. Peri- saemanni (OPPEL, 1856) in spite of the stratigraphically phery rounded, base convex and anomphalous. Apparent younger one (Plate II: 8) has a much slenderer shell than peristome pyriform. No true peristome part preserved. the older one, having a more “typical” morphology. Whorls and base covered by nearly equally spaced spiral Bourgetia saemanni is Bajocian to Oxfordian in Europe, striae, crossed only by fine, sigmoidal growth lines. Callovian in Near East, Bajocian in Arabia and Callovian Remarks — B. saemanni (OPPEL, 1856) (spec. name to Kimmeridgeian in East Africa (FISCHER 2001). corrected by COX 1938) is the first valid name to the type Distribution in Kachchh — (20) Lower Chari species of Bourgetia TERQUEM & JOURDY, 1870. The Formation (?Callovian), Jumara Dome; (21) Upper Chari authors of the synonymy composed a concept of a rather Formation (?Oxfordian), Fakirwari.

Superfamily Naticoidea FORBES, 1838 Family Ampullinidae COSSMANN, 1918 Genus Ampullospira G. F. HARRIS, 1897 Subgenus Ampullospira G. F. HARRIS, 1897

Ampullospira (Ampullospira) brevispira J.–C. FISCHER, 2001 (Plate II: 20–21)

2001: Ampullospira (A.) brevispira nov. sp. — J.–C. Fischer, p. 89, Pl. 2, figs 13–14. Material — Two specimens with damaged peristome. Measurements H HL HP D WP AA AL BHU2001I 36 Plate II: 20–21 15.0 13.0 10.0 12.5 11.0 107° 85° Description — Medium size, low spired specimen, bad to see umbilicus (if present). Aperture semicircular; consists of six whorls; protoconch unknown. Subsutu- peristome remnants simple, suggesting continuous deve- rally, distinct ramp appears on early juvenile shell and lopment; parietal part appears as narrow callosity. Orna- lasts to adult aperture. Outer limit of ramp strongly arched ment not visible. convexity; keel like angulation absent. Ramp feebly con- Remarks — The size and the shape of this species vex. Base strongly convex, preservation of central part too resemble to those of Ampullospira (Pictavia) lorierei (D’OR-

Fragmenta Palaeontologica Hungarica 22, 2004 16 SZABÓ &JAITLY BIGNY, 1852). The only marked differences are in the temporaneous, and the occurrences suggest similar/same morphology of the subsutural ramp. Possibly, the spire paleobiotopes, the identification needs special attention. morphology provides further tools to a reliable distinc- Distribution in Kachchh — Gora Doongar Yellow tion, but the available specimen number has been low to Flagstone Member (Bathonian), Sadhara village, Pachchham observe the variability. Because the two species are con- Island.

Subgenus Pictavia COSSMANN, 1925

Ampullospira (Pictavia) lorierei (D’ORBIGNY, 1852) (Plate II: 24–25, 31–32)

1852: Natica Lorierei — D’ORBIGNY, p. 190, Pl. CCLXXXIX, Figs 6–7. 1885: Ampullina Lorierei D’ORBIGNY — COSSMANN, P. 144, Pl. XVI, Figs 36–37. cf. 1914: Ampullina sp. — DIETRICH, p. 125, pl. 11, figs 15a–b 1968: Pictavia cf. P. lorierei (D’ORBIGNY) — REINER, p. 187, Pl. II, Fig. 9. 1969: Ampullospira (Pictivia) lorierei (D’ORBIGNY) — J.–C. FISCHER, p. 191, Pl. XVIII, Figs 29–31 1997: Ampullospira (Pictavia) lorierei (D’ORBIGNY) — J.–C. FISCHER &WEBER, p. 70, Pl. 17, Fig. 9. 2001: Ampullospira (Pictavia) lorierei (D’ORBIGNY) — J.–C. FISCHER, p. 90, Pl. 2, Fig. 16. Material — Two+?three specimens. Measurements H HL HP D WP AA AL BHU2001I 40 31–32 16.5 14.5 12.5 14.0 11.0 95° 85° BHU2001I 41 no photo 14.0 11.5 12.5 14.0 11.0 BHU2001I 35 24–25 24.0** 20.5** 18.0** 21.0** 15.5** 101° 84° Description — Medium size, low littoriniform species. network ornament, visible under magnification. Largest specimen consists of six whorls; protoconch un- Remarks — COSSMANN (1885) and FISCHER (1969) known. Subsuturally, keel like angulation appears on early revised the originals of the earlier authors; their synony- juvenile shell and lasts to adult aperture. Between keel and mies add further details to the history of species interpreta- nearest suture, rather deep, narrow canal developed. Con- tion. Among the published forms, showing considerable cavity of canal gradually decreases with growth while width variability, FISCHER (2001) presented a specimen from Ara- increases. From penultimate whorl, canal becomes narrow bia that has the closest morphology to the Kachchh finds. ramp that remains feebly concave and seems to slope to axis The species is Bathonian in western Europe (in France), every case. Base strongly convex, narrow umbilicus visible Bathonian to Middle Callovian in Arabia. centrally. Aperture half circle shaped; peristome simple and Distribution in Kachchh — Amphiastrea Coral Bed, continuous but parietal part appears as thin callus, slightly Chari Formation, Habo Dome; Lower Callovian Chari For- extending anteriorly onto base. Growth-lines feebly proso- mation, Jara Dome and Gora Doongar Yellow Flagstone cline, delicate spiral threads cross them that results very fine Member (Bathonian), Sadhara village, Pachchham Island.

Ampullospira (Pictavia) bajocensis (D’ORBIGNY, 1852) (Plate I: 40–43)

1852: Natica Bajocensis D’ORBIGNY — p.189, pl. 289, figs 1–3. 1852: Natica Pictaviensis D’ORBIGNY — p. 191, pl. 289, figs 8–10 1868: Natica Bajocensis D’ORBIGNY —LAUBE, p.4, Tab. I, Fig. 5. 1892: Natica Bajocensis D’ORBIGNY —HUDLESTON, p. 263, Pl. XX, 13a, 13b. 1997: Ampullospira (Pictavia) bajocensis (D’ORBIGNY, 1852) — FISCHER &WEBER, p. 70, 71, Pl. 17, figs 7, 8. 2003: Pictavia bajocensis (D’ORBIGNY, 1852) — GRÜNDEL, p.76, Pl. 9, Figs 5–10. Material — Ten shelly specimens, more or less damaged. Measurements H HL HP D WP AA AL BHU2001I 26 Plate I: 42–43 25* 17.0* 13.0* 13.5 9.0 61° 45° BHU2001I 27 Plate I: 40–41 20* 15.0 12.5 12.0 10.0 63° 53° Description — Rather high spired, relatively slender, considerably wider spiral angles than the Kachchh speci- phasianelliform shells of slightly cyrtoconical outline. mens. In this feature, the latter ones resemble also A. (P.) Narrow but distinct, concave subsutural ramp present calypso (D’ORBIGNY, 1852) that has been established on from early whorls (suture canaliculate). Aperture elongate, stratigraphically younger (Oxfordian) form. The Bathonian peristome remnants suggest short columellar lip; no trace (BHU2001I 27) specimen has less but the Early Callovian of peristome modification observable. (BHU2001I 26) shows more resemblance to A. (P.) calypso. Remarks — The holotype of Ampullospira (Pictavia) However, the subsutural “ramp” is more marked in A. (P.) bajocensis, selected by FISCHER &WEBER (1997), has bajociensis (like in our specimens) that provides a feebly gradate

Fragmenta Palaeontologica Hungarica 22, 2004 Jurassic gastropods from Kachchh 17 outline, while this morphological feature hardly visible in A. Distribution in Kachchh — Gora Doongar Yellow (P.) calypso. Amongst the figured forms, the shape of the Flagstone Member (Bathonian), Sadhara village, Pach- Kachchh finds is most similar to the Bajocian (Parkinsoni chham Island; Lower Callovian Chari Formation, Kama- Zone) specimen figured by HUDLESTON (1892) in Plate ghuna section, Jhura Dome and Lower Callovian Chari XX, Fig. 13b, a “narrower variety, approaching N. Calypso”. Formation of Jumara Dome.

Ampullospira (Pictavia) tanganyicensis COX, 1965 (Plate I: 48–49)

1965: Pictavia tanganyicensis sp. nov. — COX, p. 167, Pl. 28, Figs 6a–c, 7a–b. Material — Twenty-three, more or less damaged, shelly specimens Measurements H HL HP D WP AA AL BHU2001I 23 Plate I: 48–49 12.5 9.0 7.0* 7.0 5.5 62° 44° Description — Small, cyrtoconical adult shells, con- Growth lines delicate, feebly prosocline; crossed by sisting of about six feebly convex whorls. Apex obtuse, fine spiral lines (visible under magnification). protoconch dome like, details not observable. Suture canali- Remarks — The East African specimens are smaller culate, narrow, concave ramp follows it on upper edge of but their morphology does not show too much difference. whorls from third volution. Last whorl almost flat between The last whorls of the larger Kachchh specimens are of sharply rounded ramp angle and broadly rounded-angular smaller spiral angle, but in the same growth stage, the meas- periphery. Base convex with narrow umbilical (?) fissure urements are similar to those of the East African ones. along columellar lip. Aperture (and probably whole peri- In East Africa, the species occurred in ?Bajocian strata. stome) ovate. Parietal lip appears as thin callosity, columel- Distribution — Gadhada Formation (Lower Callo- lar lip strong and having sharp edge along basal fissure. vian), north of Mouwana village, Mouwana Dome.

Ampullospira (Pictavia) cf. pyramidata (MORRIS &LYCETT, 1851) (Plate II: 26, 27–28, 40–41) cf. 1851: Natica (Euspira) pyramidata n. sp. — MORRIS &LYCETT, 1851, p. 46, Pl. VI, Figs 8, 8a. Material — Three worn specimens. The specimens on Plate II: 26 and 27–28 are somewhat deformed. Measurements H HL HP D WP AA AL BHU2001I 38 (Plate II: 40–41) 9.5 7.0 6.5 7.5 4.5 77° 77° BHU2001I 33 (Plate II: 27–28) 14.1 11.3 9.0 11.5** 7.5** Description — Rather high spired naticiform–litto- poor preservation of the base and the peristome makes riniform shell with strongly convex whorls, having rather the identification tentetive. wide, concave subsutural ramp with marked outer angu- The marked angulation below the suture and the simple lation, providing gradate outline to shell. Remaining part conical outline of the shell distinguishes A. cf. pyramidata of whorls below ramp angle evenly arched. Base convex, from the other species of the studied material. peristome badly damaged on all specimens. Distribution in Kachchh — Coral Bed Jhurio Remarks — The visible elements of the shell mor- Formation, Jumara Dome; Gadhada Formation (Lower phology suggest belonging to A. pyramidata, however, the Callovian), N of Mouwana village, Mouwana Dome.

Ampullospira (Pictavia) aff. tanganyicensis COX, 1965 (Plate I: 38–39) aff. 1965: Pictavia tanganyicensis sp. nov. — COX, p. 167, Pl. 28, Figs 6a–c, 7a–b. Material — Single, poorly preserved specimen. Measurements H HL HP D WP AA AL BHU2001I 31 Plate I: 38–39 14.5 10.5 7.5 9.5 5.5 63° 48° Description — Small, nearly trochiform shell with lacking or extremely thin. Base somewhat flattened and slightly cyrtoconical outline. Whorls feebly convex, last having narrow (false?) umbilicus. one flattened, and periphery subangular. Suture runs in Remarks — The wider ramp, having sharply angular moderately deep incision; narrow, almost horizontal (ab- outer rim, distinguishes this species from the similarly sized axially sloping) shoulder follows it with sharply angular Ampullospira (P.) tanganyicensis (above) and A. (P.) sp. (below). outer rim. Width of shoulder considerably increasing on Distribution in Kachchh — ?Bajocian/Bathonian, last whorl. Aperture and peristome elongate; parietal lip Kuar Bet and Coral Bed, Jhurio Formation, Jumara Dome. Fragmenta Palaeontologica Hungarica 22, 2004 18 SZABÓ &JAITLY Ampullospira (Pictavia) sp. (Plate I: 44–45) Material — Most of the 17 specimens are poorly preserved. Measurements H HL HP D WP AA AL BHU2001I 18 Plate I: 44–45 14.5* 10.0* 6.5* 8.5 63° 44° BHU2001I 19 not figured 14.2* 10.0 6.5 8.5 5.3 65° 41° Description — Small, phasianelliform shells with slightly The main differences are: A. (P.) tanganyicensis has lower cyrtoconical outline. Whorls slightly convex, periphery whorls (number of whorls is equal in the adult speci- rounded. Suture runs in moderately deep incision. Upper mens of the two species in spite of the difference in belt of whorls forms narrow but marked shoulder with the total height); Ampullospira (Pictavia) sp. has more rounded-angular outer rim. Aperture and peristome elong- slender shells; the subsutural ramp is more distinct than ate; parietal lip lacking or extremely thin. Base somewhat in A. (P.) tanganyicensis. (Distinction from Ampullospira flattened and having narrow (false?) umbilicus. (Pictavia) aff. tanganyicensis above.) Remarks — The shells resemble those of the simi- Distribution in Kachchh — Gadhada Formation larly aged Ampullospira (Pictavia) tanganyicensis COX, 1965. (Lower Callovian), Umrapur section, Khadir Island.

Genus Globularia SWAINSON, 1840

Globularia khadirensis n. sp. (Plate II: 17–19, 22–23) Holotype — BHU2001I 46 (Plate II: 17–19) Type locality — Khadir Island, Kachchh, western India. Type strata — Hadibadhang Shale Member of Khadir Formation, Bathonian. Species name — From the name of Khadir Island. Diagnosis — Axially elongate shells with short, coeloconical spire; feebly convex, rather wide subsutural ramp, developed on last two whorls but disappearing near last peristome; subconical base with feebly concave outline near columellar lip; parietal lip present as callosity, inner lip reflecting over (hiding) possible umbilicus. Material — Two specimens. Measurements H HL HP D WP AA AL BHU2001I 46 Plate II: 17–19 18.5 15.5 12.5 14.5 11.5 75° 99° BHU2001I 47 Plate II: 22–23 20.5* 18.0* 16.0* 16.5 13.0 73° 110° Description — Medium size species of axially slightly basal depression. Outer lip not preserved. elongate shells. Spire low, coeloconical and composed of Remarks — Globularia eparcyensis (D’ARCHIAC, 1843) few (3–4) convex whorls. Protoconch large, subglobular but and Globularia subumbilicata (D’ARCHIAC, 1843) resemble details not preserved. Suture slightly impressed; almost Globularia khadirensis n. sp. because of their coeloconical horizontal, feebly convex and rather wide shoulder follows spire. However, the subglobular last whorl distinguishes it below that causes subgradate outline of spire. Ramp them from Globularia khadirensis n. sp., having subconical slopes out of axis and turns downward without angulation. body whorl. Strongly convex belt of whorl surface gives outer boundary Two young specimens of Ampullospira besairiei COX, of ramp; this convexity much stronger than on remain- 1965 (Pl. 28, Figs 10, 11a–b) from Madagascar and East ing whorl surface but changes uniform near last peri- Africa are comparable to the specimens of Globularia khadir- stome therefore ramp becomes narrow belt close to ensis n. sp. However, the holotype of A. besairiei (COX suture. Base subconical, with narrow, axial furrow along 1965, Pl. 28, Figs 12a–b, from Tanganyika) and another columellar lip, indicative of (false?) umbilicus. Aperture larger specimen seem strongly different because their ovate, peristome continuous. Inner lip consists of parietal wide subsutural ramp exists also near the last peristome. callosity with feebly opisthocline–opisthocyrt edge and Distribution in Kachchh — Hadibhadang Shale columellar lip, being relatively long and bending over Member, Khadir Island (Bathonian).

Globularia michelini (D’ARCHIAC, 1843) (Plate II: 11–13)

1843: Natica Michelini D’ARCHIAC — p. 377, pl. XXX, figs 3a–b. 1851: Natica Michelini D’ARCHIAC —MORRIS &LYCETT, p. 44, Pl. VI, figs 2a, 3–3a. 1852: Natica Michelini D’ARCHIAC — D’ORBIGNY, p. 192, Pl. 289, Figs 11–12. 1885: Ampullina Michelini D’ARCHIAC —COSSMANN, p. 131, pl. II, figs 9–10. 1892: Natica cf. Michelini D’ARCHIAC —HUDLESTON, p. 269, Pl. XX, Figs 18a–b. Fragmenta Palaeontologica Hungarica 22, 2004 Jurassic gastropods from Kachchh 19 1950: Globularia michelini (D’ARCHIAC, 1843) — COX &ARKELL, p. 83. 1969: Globularia michelini (D’ARCHIAC, 1843) — J.–C. FISCHER, p. 189, Figs 49–52. 1997: Globularia michelini (D’ARCHIAC, 1843) — J.–C. FISCHER &WEBER, p. 71. Material — Twenty-three specimens in various state of preservation. Measurements H HL HP D WP AA AL BHU2001I 55 Plate II: 11–13 56.0 52.5 46.5 44.5 37.5 103° Description — Rather large, ovate naticiform shell variability of the shell morphology (see more details in with low spire, consisting of few convex whorls. Suture COX &ARKELL 1950 and FISCHER 1969). All transitional moderately incised, base strongly convex. Aperture sub- forms between the more globular “typical” specimens and axially elongate, outer lip does not show trace of any the ovate ones (“Natica minchinhamptonensis” DE LORIOL) modification. Parietal lip forms narrow callus on base; exist. The Kachchh specimens are nearer to the latter columellar lip slightly reflected. Outer edge of whole forms, especially to the form figured by HUDLESTON inner lip well marked. (1892) as “Natica cf. Michelini D’ARCHIAC”. Remarks — This species is widespread along the Globularia michelini is Bathonian in Western Europe shelves and epicontinental seas surrounding the western and Early to Middle Callovian in Tunisia. Tethys (“stable” Europe and Africa). The earlier inter- Distribution in Kachchh — Callovian Chari Formation pretations of this species have outlined an extremely high (Amphiastrea Bed), Dhrang Section, Northern Habo Dome.

Globularia ? aff. michelini (D’ARCHIAC, 1843) (Plate II: 14–16) Material — Single inner mould specimen with some shelly portions on the last whorl. Measurements H HL HP D WP AA AL BHU2001I 56 Plate II: 14–16 29.0* 27.0* 23.0* 25.0 21.0 125°* 113° Description — Subglobular shell, consisting of few may belong also to a different genus. Morphology of whorls. Spire low, subgradate. Whorls evenly arched except the inner mould and shell remnants along the suture along columella where feeble concavity visible. Few marked, indicate possible presence of subsutural ramp on the collabral ridges and slight downvard deviation from normal shell; the shape and the measurements also well agree coiling of last preserved whorl suggest adult stage. Peristome with those of Ampullospira quennelli COX, 1965, however, in not preserved, apparent aperture subaxially ovate. lack of shell, the identification must remain doubtful. Remarks — There are figured specimens with The available shell remnants do not permit also a comparable shape in Globularia michelini, however, the reliable generic identification. inferable adult size of the discussed specimen is much Distribution in Kachchh — ?Bajocian/Bathonian, smaller that indicates affinity to a different species that Kuar Bet.

Genus Naricopsina CHELOT, 1886

Naricopsina cornelia (LAUBE, 1868) (Plate II: 37–39)

1868: Natica Cornelia — LAUBE, p. 5, Taf. I, Fig. 8. Material — A single specimen. Measurements H HL HP D WP AA AL BHU2001I 61 Plate II: 37–39 9.5 9 7 10 152° 83° Description — Few-whorled naticiform species Remarks — Naricopsina texata (LYCETT, 1863) is a with low spire. Earliest shell parts almost planispiral. similarly aged and shaped species. From this species, Axis of protoconch seems slightly different from that LAUBE distinguished Natica cornelia on the basis of the of the subsequent shell parts. Whorls of semicircular shape of the aperture, and the nature of the ornament. cross-section, separated by rather deeply canaliculate The Kachchh species well corresponds to LAUBE (1868) suture. Peristome not visible. Base convex, umbilical description and figures. region not cleanable. Shell ornamented by subregularly Naricopsina cornelia (LAUBE, 1868) is Bathonian in repeating, prosocline growth lines, suggesting simple, (“stable”) Europe. tangential peristome. Obscure spiral lines cross growth Distribution in Kachchh — Ler, Chari Formation, lines. Callovian.

Fragmenta Palaeontologica Hungarica 22, 2004 20 SZABÓ &JAITLY Genus Megatylotus FISCHER, 1885

Megatylotus sp. (Plate II: 29–30) Material — One poorly preserved specimen. Measurements H HL HP D WP AA AL BHU2001I 57 Plate II: 29–30 33.0* 29.0* 26.0* 29.0* 22.0* 90° Description — Moderately wide base, conical speci- Remarks — No similarly shaped species has been men. Whorls convex but last one becomes flattened near found in the literature of the Middle and Late Jurassic but peristome. Flattened last whorl part accompanied by the preservation is unsatisfactory to designate it a new narrow, convex, subsutural shoulder that tends to be species. This genus is characteristically Cretaceous, there- horizontal in belt close to suture. Apparent aperture tri- fore the Kachchh species could give important data to angular. Peristome not preserved, remnants suggest also the history of Megatylotus. triangular form with short, strongly arched columellar part; Disrtibution — Hadibhadang Shale Member, Khadir outer lip forms wide “V” shape in cross-section. Island, (Bathonian).

Genus Pseudamaura FISCHER, 1885

Pseudamaura hadibadhangensis n. sp. (Plate II: 33–36) Holotype — BHU2001I 44 (Plate II: 35–36) Type locality — Hadibadhang, Khadir Island, Kachchh, western India. Type strata — Hadibadhang Shale Member of Khadir Formation, Bathonian. Species name — From the type locality of the type strata. Diagnosis — Axially ovate, gradate shells with rather short spire of cyrtoconical outline; feebly concave to flat, moderately wide subsutural ramp below suture; feebly concave outer face on whorls; convex base with narrow umbilicus; aperture pyriform, peristome simple. Material — Twenty-nine specimens in various state of preservation. Measurements H HL HP D WP AA AL BHU2001I 45 Plate II: 33–34 30.0* 22.5 17.0 19.0 16.5 80°* 68° BHU2001I 44 Plate II: 35–36 28.0 21.0 16.0 18.5 13.0 75° 62° Description — Ovate, gradate shells with whorls, shell is not ovate and peripheral angulation is lacking. having concave to flat, abaxially feebly sloping ramp, Protoconch of the available specimens are absent or delimited by rounded angulation. Wide, feebly concave poorly preserved but the juvenile part of the shells show outer face present between ramp angulation and suban- close morphological relation to the Ampullospira–Pictavia gular periphery. Base convex and narrowly umbilicate. species group. That is why Pseudamaura hadibadhangensis n. Columellar lip arched, slightly thickened and extending sp. has been regarded as member of Ampullinidae in this into umbilical region, sometimes furrow runs on col- paper. umellar lip parallel with its edge. Parietal region covered KOWALKE &BANDEL (1996) established Pseud- by thin callus. amauridae on a protoconch character of the type species. Growt lines orthocline (as main orientation), some- KOWALKE (1998) suggested Cerithioidea inc. sed. as sys- what opisthocyrt on outer face; slightly parasigmoidal on tematical position for Pseudamauridae on the studies of base. the protonch of another species. However, further investiga- Remarks — A comparable species from the similarly tions on the protoconchs of the other genera, traditionally aged ones is “Natica (Euspira) pyramidata” MORRIS & regarded as members of the Ampillinidae, are also needed LYCETT, 1851 (Cloughtonia in COX &ARKELL 1948–50) to support distinction of Pseudamauridae. because it has also concave belt below the subsutural Distribution in Kachchh — Hadibhadang Shale ramp. However, this species has much wider spire angle, Member, Khadir Island (Bathonian).

Superfamily ?Carinarioidea BLAINVILLE, 1818 Family ?Carinaridae BLAINVILLE, 1818

BANDEL &HEMLEBEN (1987) removed Coelodiscus linked it to Heteropoda LAMARCK, 1812 (Caenogastro- BRÖSAMLEN, 1909 from Euomphalidae and, based on poda/Mesogastropoda). Because homoeomorphism may similarities in protonch morphology to extant forms, exist also between protoconchs, further evidences are

Fragmenta Palaeontologica Hungarica 22, 2004 Jurassic gastropods from Kachchh 21 necessary to justify the caenogastropod nature of the tion, the planktonic mode of life can be excluded in case shells of Coelodiscus (and Kosmomphalus, FISCHER, 2001). of the Kachchh Coelodiscus and also the closely related Because of the size and the rather heavy shell construc- Kosmomphalus.

Genus Coelodiscus BRÖSAMLEN, 1909

Coelodiscus sadharaensis n. sp. (Plate I: 3–5) Holotype — BHU2001I 29 (Plate I: 3–5). Type locality — Pachchham Island, Kachchh, western India, Sadhara village. Type strata — Gora Doongar Yellow Flagstone Member, Bathonian. Species name — From Sadhara village. Diagnosis — Naticiform shell with feebly convex spire; protoconch smooth, planispiral and having narrow apical umbilicus; base strongly convex, narrowly phaneromphalous. Material — A specimen without last peristome. Measurements H HL HP D WP AA AL BHU2001I 29 Plate I: 3–5 9.0* 8.5* 8.5* 10.0* 4.2* 171° 164° Description — Small, naticiform species with almost Remarks — Disregarding the ornament, the whole plane but convex, feebly cyrtoconical spire. Protoconch shell morphology well correspond to that of the type (~one whorl) planispiral, smooth (? or ornament eroded) species [Coelodiscus minutus (ZIETEN, 1832)] but Coelodiscus and has also apical umbilicus; initial chamber globular. fluegeli BANDEL &HEMLEBEN, 1987 resembles also in the Whorls convex and rather rapidly increasing. Last whorl low spire and the dominantly spiral ornament (both bears wide, feebly convex ramp, abaxially delimited by species is Toarcian); the only significant difference is in strongly rounded angulation. Suture deeply impressed. the size; the Kachchh specimen is much larger (height Base strongly convex and narrowly phaneromphalous; and width roughly double). The more similarly aged umbilical rim rounded angular. Whorl cross-section suggests (Bajocian) C. brevispira CONTI &FISCHER, 1984 and axially elongate peristome of more than double height than Coelodiscus sp., (GRÜNDEL 2001) are discoidal and also width. Inner lip extends over umbilical region. Shell wall minute species of a few mm size. thick, especially in parts of subsutural ramp. In the Kachchh fauna, Kosmomphalus reticulatus FISCHER, Whorls and base ornamented by sparse, regularly 2001 is a comparable species but that is a discoidal, spaced spiral grooves of nearly equal depth and width. broadly phaneromphalous form. Growth lines prosocline, and tangential to former whorl; Distribution in Kachchh — Bathonian Gora shallow, wide, and opisthocyrt sinus visible below ramp. Doongar Yellow Flagstone Member, Sadhara, Pachchham (Peristome probably also tangential.) Island.

Genus Kosmomphalus FISCHER, 2001

Kosmomphalus reticulatus FISCHER, 2001 (Plate I: 6–10)

2001: Kosmomphalus nov. gen. reticulatus nov. sp. — FISCHER, p. 71, Pl. 1, Figs 1a–c. Material — One shelly specimen, peristome not preserved. Measurements H HL HP D WP AA AL BHU2001I 32 Plate I: 6–10 8. 8.5 8.0 15.5 168° 157° Description — Depressed, almost planispiral shell, shell near (last?, apparent?) peristome. consisting of few (~4) thick-walled whorls with nearly Remarks — The available specimen is somewhat circular cross-section. Suture deeply canaliculate, base different from the figures of the holotype. The Kachchh convex and broadly phaneromphalous. Equally strong, specimen has a more regularly rounded whorl cross-sec- marked spiral incisions/furrows cut whorl-surface into tion, the coiling is slightly trochospiral and the reticulate uniform, cord-like belts in interval between suture and nature of the ornament is hardly observable. outer third of base. On remaining part of base and in As FISCHER (2001) has already expressed, Coelodiscus umbilical wall, both spiral ornamental elements become BRÖSAMLEN, 1909 is morphologically closely related to wider. Growth-threads feebly prosocline; more markedly Kosmomphalus. The only notable difference is the presence visible in spiral furrows. No peristome part found pre- of a marked spiral ornament in Kosmomphalus, however, served but cross-section (see Plate I: 7) suggests thickened similar ornament exist also in some Coelodiscus species in

Fragmenta Palaeontologica Hungarica 22, 2004 22 SZABÓ &JAITLY weaker development (like in C. sadharaensis n. sp). Further needed to clarify the relation more accurately. differences seem to exist in the peristome structure but Kosmomphalus reticulatus is Callovian in Arabia. these characters are unsatisfactorily known in both Distribution in Kachchh — Upper part of Callovian genera. Additional material in good state of preservation to Oxfordian Chari Formation, Ler.

Superfamily Xenophoroidea TROSCHEL, 1852 Family Lamelliphoridae KOROBKOV, 1955 Genus Lamelliphorus COSSMANN, 1915

Lamelliphorus cf. ornatissimus (D’ORBIGNY, 1853) (Plate I: 50–51)

1853: Trochus ornatissimus — D’ORBIGNY, p. 272, Pl. 312, Figs 5–8. 1894: ornatissimus D’ORBIGNY — HUDLESTON, p. 330, Pl. XXVII, Figs 8–b. 1915: Lamelliphorus ornatissimus D’ORBIGNY — COSSMANN, p. 191, Pl. 7, Figs 27–32. 1997: Lamelliphorus ornatissimus (D’ORBIGNY, 1853) — FISCHER &WEBER, p. 106, Pl. 22, Figs 1a–b, 2. 2003: Lamelliphorus ornatissimus (D’ORBIGNY, 1853) — GRÜNDEL, p. 85, Pl. 11, Figs 9–11. Material — Single specimen with eroded apex and base not cleanable from matrix. Measurements H HL HP D WP AA AL BHU2001I 43 Plate I: 50–51 19.0 73° 85° Description — Medium sized, moderately wide base, be cleaned, however, the available characters support its conical species with feebly concave shell outline and identification with Lamelliphorus ornatissimus (D’ORBIGNY, blunt apex. Whorls low and flat; thin incision like suture 1853). “Onustus Heberti” LAUBE, 1868 bears a similar separates them. Periphery undulating frill-like, sharp and number of ribs but the measurements of the whorls are carinated. different, and the shell is not coeloconical. Strong, slightly opisthocline, suture-to-suture ribs on Lamelliphorus ornatissimus (D’ORBIGNY, 1853) is late all whorls, just perpendicular to extremely prosocline Bajocian in France, England and Germany. growth-lines. Number of ribs on last whorl: 30. Distribution in Kachchh — Sponge Limestone Remarks — The base and the peristome could not member, Patcham Formation (Bathonian), Jumara Dome. * * * Acknowledgements — The authors are thankful to F. T. Fürsich, Würzburg, J. H. Callomon, London, I. B. Singh, Lucknow and D. K. Pandey, Jaipur, who were associated in the collection of gastropods within a project, supported by the National Geographic society of America (grant Ȳ 3597-87). The authors are indebted to F. T. FÜRSICH (Würzburg), providing also complementary information to the studies. The field works and the consultations of the authors were supported in the frame of the Hungarian–Indian cultural exchange program in 2000–2001 (IND–4/99) and the Ȳ T 031873 project of the Hungarian Scientific Research Fund. We are thankful for D. SZABÓ for his help to the preparation of the photo plates. Ÿ Explanation to Plate I

1–2 Discohelix cf. sapho (D’ORBIGNY, 1853) — u2; Upper part of Callovian to Oxfordian Chari Formation, Ler. 3–5 Coelodiscus sadharaensis n. sp. — 3: u1; 4–5: u 2.2; Upper part of Callovian to Oxfordian Chari Formation, Ler. 6–10 Kosmomphalus reticulatus FISCHER, 2001 — 6: u1; 7–10: u2; Upper part of Callovian to Oxfordian Chari Formation, Ler. 11–12 Neritopsis (Neritopsis) benoisti COSSMANN, 1900 — u2; Coral Bed of Jhurio Formation (Bathonian), Jumara Dome. 13–16 Neritopsis sp. — 13: u1; 14–16: u2; Coral Bed of Jhurio Formation (Bathonian), Jumara Dome. 17–19 Neritopsis (Neritopsis) benoisti COSSMANN, 1900 — 17: u1; 18–19: u2; Coral Bed of Jhurio Formation (Bathonian), Jumara Dome. 20–21 Neritopsis (Neritopsis) benoisti COSSMANN, 1900 — u2; Coral Bed of Jhurio Formation (Bathonian), Jumara Dome. 22–25 Hayamia bajocensis (D’ORBIGNY, 1852) — u1; Lower Callovian part of Chari Formation, Jumara Dome. 26 Hayamia subsulcosa (D’ORBIGNY, 1850) — u1; Lower Callovian Chari Formation of Jumara Dome. 27–29 Neridomus involuta (LYCETT, 1863) — u1; Lower Yellow Flagstone Member (Bathonian), Pachchhmai Pir, Pachchham Island 30–32 Neridomus sp. — u1; Hadibhadang Shale Member, Khadir Island (Bathonian) and ?Bajocian/Bathonian, Kuar Bet. 33-37 Neritoma kachchhensis n. sp. — 33–35: u1; 36–37: u2; Hadibhadang Shale Member, Khadir Island (Bathonian). 38–39 Ampullospira (Pictavia) aff. tanganyicensis (COX, 1965) — u1; ?Bajocian/Bathonian, Kuar Bet and Coral Bed, Jhurio Formation, Jumara Dome. 40–43 Ampullospira (Pictavia) bajocensis (D’ORBIGNY, 1852) — u1; 40–41: Lower Callovian Chari Formation of Jumara Dome; 42–43: Gora Doongar Yellow Flagstone Member (Bathonian), Sadhara village, Pachchham Island. 44–45 Ampullospira (Pictavia) sp. — u1; Gadhada Formation (Lower Callovian), Umrapur section, Khadir Island. 46–47 Pseudomelania sp. — u1; Gadhada Formation (Lower Callovian), N of Mouwana village, Mouwana Dome. 48–49 Ampullospira (Pictavia) tanganyicensis (COX, 1965) — u1; Gadhada Formation (Lower Callovian), north of Mouwana village, Mouwana Dome. 50–51 Lamelliphorus ornatissimus (D’ORBIGNY, 1853) — u1; Sponge Limestone member, Patcham Formation, Jumara Dome.

Fragmenta Palaeontologica Hungarica 22, 2004 Jurassic gastropods from Kachchh 23 Plate I

Fragmenta Palaeontologica Hungarica 22, 2004 24 SZABÓ &JAITLY

Ÿ Explanation to Plate II

1–5 Pseudomelania remtsaensis COX, 1969 — 1: u3; 2–3: u1; 4–5: u1: Chari Formation (Amphiastrea Bed), Dhrang section, northern Habo Dome. 6–8 Bourgetia saemanni (OPPEL, 1856) — u1; 6–7: Chari Formation (?Callovian), Jumara Dome; 8: Upper Chari Formation (?Oxfordian), Fakirwari. 9–10 Pseudomelania calloviensis (HEBERT &DESLONGCHAMPS, 1860) — u1; Kaladongar, Pachchham Island, Goradongar Formation, Bathonian 11–13 Globularia michelini (D’ARCHIAC, 1843) — u1; Chari Formation (Amphiastrea Bed), Dhrang Section, Northern Habo Dome. 14–16 Globularia ? aff. michelini (D’ARCHIAC, 1843) — u1; ?Bajocian/Bathonian, Kuar Bet. 17–19 Globularia khadirensis n. sp. — u1; Hadibhadang Shale Member, Khadir Island (Bathonian). 20–21 Ampullospira (Ampullospira) brevispira J.–C. FISCHER, 2001 — u1; Gora Doongar Yellow Flagstone Member (Bathonian), Sadhara village, Pachchham Island. 22–23 Globularia khadirensis n. sp. — u1; Kala Doongar Sandstone member (Bajocian), Pachchmaipir section, Pachchham Island. 24–25 Ampullospira (Pictavia) lorierei (D’ORBIGNY, 1852) — u1; Gora Doongar Yellow Flagstone Member (Bathonian), Sadhara village, Pachchham Island. 26–28 Ampullospira (Pictavia)cf.pyramidata (MORRIS &LYCETT, 1851) — 26: u2; 27–28: u1; Gadhada Formation (Lower Callovian), N of Mouwana village, Mouwana Dome. 29–30 Megatylotus sp. — u1; Hadibhadang Shale Member, Khadir Island (Bathonian). 31–32 Ampullospira (Pictavia) lorierei (D’ORBIGNY, 1852) — u1; Amphiastrea Coral Bed, Chari Formation, Habo Dome. 33–36 Pseudamaura sp. — u1; Hadibhadang Shale Member, Khadir Island (Bathonian). 37–39 Naricopsina texata (LYCETT, 1863) — u1; Khadir Island, Khadir Formation (Bajocian to Oxfordian) 40–41 Ampullospira (Pictavia)cf.pyramidata (MORRIS &LYCETT, 1851) — u2; Coral Bed Jhurio Formation, Jumara Dome (?Bajocian).

Fragmenta Palaeontologica Hungarica 22, 2004 Jurassic gastropods from Kachchh 25 Plate II

Fragmenta Palaeontologica Hungarica 22, 2004 26 SZABÓ &JAITLY

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Authors’ addresses: Dr. János SZABÓ Dr. Anand K. JAITLY Geological and Palaeontological Department Department of Geology Hungarian Natural History Museum Banaras Hindu University Map: Budapest, VIII, Ludovika tér 2 Varanasi 221005 Mail: 1431 Budapest, pf. 137 India Hungary E-mail: [email protected] E-mail: [email protected]

Fragmenta Palaeontologica Hungarica 22, 2004