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Contributions to the Jurassic of Kachchh, Western India VIII. the Gastropod Fauna

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Contributions to the Jurassic of Kachchh, Western India VIII. the Gastropod Fauna FRAGMENTA PALAEONTOLOGICA HUNGARICA 22, BUDAPEST, 2004 Contributions to the Jurassic of Kachchh, western India VIII. The gastropod fauna. Part II: Discohelicidae, Neritomorpha, Caenogastropoda by János SZABÓ & Anand K. JAITLY Abstract — Twenty-seven species of gastropods have been identified and systematically described from the Middle to early Late Jurassic of Kachchh (western India). Most of them have been already known from the deposits of the epicontinental seas around the western part of the Tethys but a few of them are new: Neritoma kachchhensis n. sp., Globularia khadirensis n. sp., Coelodiscus sadharaensis n. sp. and Pseudamaura hadibadhang- ensis n. sp. Keywords — Jurassic, Gastropoda, systematics, Kachchh, India. SZABÓ,J.&JAITLY, A. K.: Contributions to the Jurassic of Kachchh, western India VIII. The gastropod fauna. Part II: Discohelicidae, Neritomorpha, Caenogastropoda. — Fragmenta Palaeontologica Hungarica, 22: 9–26. Introduction This contribution is the third one in a series of papers, gastropod genera (Bhujnerita, Neritidae), which seem to be which are based on the collections of Jurassic fossils from endemic in the “Ethiopian Faunal Province”. localities of Kachchh (State Gujarat, western India). The Paralelly, another team has been also working on the type specimens of this paper are deposited at the Banaras Jurassic gastropods of the same area; they published their Hindu University, Varanasi (India); complementary mater- results some months earlier (DAS et al. 1999) than JAITLY ial is kept in the collections of the Würzburg University. et al. (2000). Therefore some of the new species, found in A sketch of the geology and stratigraphy of the localities Kachchh, have already synonyms. In the following list, are added to the first part of the systematical results revised names of the species, published from Kachchh, is (JAITLY et al. 2000). Another paper (JAITLY &SZABÓ given (bold letters indicate valid names; junior synonyms 2002) wishes to draw attention to one of the few Jurassic immediately follow the senior ones in square brackets). 1. Pleurotomaria sp. 29. Biarmatoidella tagorei (DAS,BARDHAN &LAHIRI, 1999) 2. Anodomaria latigradata JAITLY,SZABÓ &FÜRSICH, 2000 [Riselloidea pileiformis JAITLY,SZABÓ &FÜRSICH, 2000] 3. Anodomaria depressa JAITLY,SZABÓ &FÜRSICH, 2000 30. Helicanthus chanda DAS,BARDHAN &LAHIRI, 1999 4. Anodomaria? sp. 31. Metriomphalus (Metriomphalus) cf. lyelli (D’ARCHIAC, 1843) 5. Obornella granulata (J. SOWERBY, 1818) 32. Amphitrochus angulatus (HUDLESTON, 1894) 6. Obornella aff. granulata (J. SOWERBY, 1818) [Colpomphalus jumarense DAS,BARDHAN &LAHIRI, 1999] 7. Obornella wuertembergensis (SIEBERER, 1908) 33. Amphitrochus acutispira JAITLY,SZABÓ &FÜRSICH, 2000 8. Leptomaria fraga (J. A. EUDES-DESLONGCHAMPS, 1849) 34. Amphitrochus orbis JAITLY,SZABÓ &FÜRSICH, 2000 9. Leptomaria pseudoumbilicata JAITLY,SZABÓ &FÜRSICH, 2000 35. Neritopsis (Neritopsis) benoisti COSSMANN, 1900 10. Leptomaria simplex JAITLY,SZABÓ &FÜRSICH, 2000 [Neritopsis (Neritopsis) patchamensis DAS,BARDHAN &LAHIRI, 1999] 11. Bathrotomaria kachchhensis JAITLY,SZABÓ &FÜRSICH, 2000 36. Neritopsis (Hayamiella) sankhamata DAS,BARDHAN &LAHIRI, 1999 12. Bathrotomaria siebereri JAITLY,SZABÓ &FÜRSICH, 2000 37. Neritopsis sp. 13. Bathrotomaria millepunctata (J. A. EUDES-DESLONGCHAMPS, 1849) 38. Hayamia subsulcosa (D’ORBIGNY, 1850) 14. Bathrotomaria sp. [Hayamia mitra DAS,BARDHAN &LAHIRI, 1999)] 15. Pyrgotrochus? sp. 39. Hayamia bajocensis (D’ORBIGNY, 1852) 16. Emarginula karuna DAS,BARDHAN &LAHIRI, 1999 40. Neridomus involuta (LYCETT, 1863) [Emarginula (Tauschia) jumarensis JAITLY,SZABÓ &FÜRSICH, 2000] 41. Neridomus sp. 17. Discohelix cf. sapho (D’ORBIGNY, 1853) 42. Neritoma kachchhensis n. sp. 18. Proconulus jadavpuriensis DAS,BARDHAN &LAHIRI, 1999 43. Bhujnerita bhattii JAITLY &SZABÓ, 2002 [Proconulus (Proconulus) bhujensis JAITLY,SZABÓ &FÜRSICH, 2000] 44. Pseudomelania remtsaensis COX, 1969 19. Proconulus (Proconulus) haboensis JAITLY,SZABÓ &FÜRSICH, 2000 45. Pseudomelania calloviensis (HEBERT &DESLONGCHAMPS,1860) 20. Proconulus (Proconulus) wagadensis JAITLY,SZABÓ &FÜRSICH, 2000 46. Pseudomelania ? sp. 21. Proconulus (Laeviconulus?) sp. 47. Bourgetia saemanni (OPPEL, 1856) 22. Calliostoma? sp. 48. Ampullospira (Ampullospira) brevispira J.–C. FISCHER,2001 23. Ataphrus (Ataphrus) kaladongarensis JAITLY,SZABÓ &FÜRSICH, 2000 49. Ampullospira (Pictavia) lorierei (D’ORBIGNY, 1852) 24. Adeorbisina? sp. 50. Ampullospira (Pictavia) bajocensis (D’ORBIGNY, 1852) 25. Eucyclus laevijugatus (QUENSTEDT, 1884) 51. Ampullospira (Pictavia) tanganyicensis COX, 1965 26. Onkospira? kutchensis DAS,BARDHAN &LAHIRI, 1999 52. Ampullospira (Pictavia)cf. pyramidata (MORRIS &LYCETT, 1851) [Onkospira? praecursor JAITLY,SZABÓ &FÜRSICH, 2000] 53. Ampullospira (Pictavia) aff. tanganyicensis Cox, 1965 27. Onkospira? aff. kutchensis DAS,BARDHAN &LAHIRI, 1999 54. Ampullospira (Pictavia) sp. [Onkospira? aff. praecursor JAITLY,SZABÓ &FÜRSICH, 2000] 55. Globularia spathi DAS,BARDHAN &LAHIRI, 1999 28. Biarmatoidella biarmata (MÜNSTER, 1844) 56. Globularia khadirensis n. sp. [Riselloidea elongata DAS,BARDHAN &LAHIRI, 1999] 57. Globularia michelini (D’ARCHIAC, 1843) Fragmenta Palaeontologica Hungarica 22, 2004 10 SZABÓ &JAITLY 58. Globularia ? aff. michelini (D’ARCHIAC, 1843) 62. Lamelliphorus cf. ornatissimus (D’ORBIGNY, 1853) 59. Naricopsina cornelia (LAUBE, 1868) 63. Coelodiscus sadharaensis n. sp. 60. Megatylotus sp. 64. Kosmomphalus reticulatus FISCHER, 2001 61. Pseudamaura hadibadhangensis n. sp. Abbreviations to measurements — H = total height; HL = height of last whorl; HP = height of peristome; D = diameter of last whorl; WP = width of peristome; AA = apical angle; AL = coiling angle of last whorl (if different from apical angle). Asterisks indicate reconstructed measurements taken on more (**) or less (*) damaged shell parts. Class Gastropoda CUVIER, 1797 Subclass Prosobranchia MILNE EDWARDS, 1848 Order ?Vetigastropoda SALVINI-PLÄWEN, 1980 Superfamily ?Cirroidea COSSMANN, 1916 Family Discohelicidae SCHRÖDER, 1995 Genus Discohelix DUNKER, 1848 Discohelix cf. sapho (D’ORBIGNY, 1853) (Plate I: 1–2) 1853: Straparollus Sappho — D’ORBIGNY, p. 315, Pl. 323, Figs 9–13. 1997: Discohelix sapho (D’ORBIGNY, 1853) — FISCHER &WEBER, p. 124, Pl. 22, Figs 6a–c. Material — A single, badly damaged specimen. Measurements H HL HP D WP AA AL BHU2001I 30 Plate I: 1–2 4.2* 4.2* 4.2* 9* 2.2* 228° Description — Biconcave, sinistral, discoidal shell of permits only an open identification because the morpho- small size. Whorl cross-section symmetrically trapezoidal logy of the nodules are hardly visible. They appear as undu- with feebly convex spiral and umbilical sides and more lation of the inner mould and the shell cross section of the convex outer side on spire whorls. Last whorl cross-section outer face. Beside this, the spiral angle (concavity of spire) tends to be circular. Both outer angulations bear weak seems somewhat greater than in the holotype of D. sapho carina. Ornament consists of riblet like growth-lines on (D’ORBIGNY); name corrected by FISCHER &WEBER (1997). outer face, and tubercles and nodes on outer angulations Discohelix sapho (D’ORBIGNY, 1853) has been recorded (delimiting outer face). Tubercles gradually strengthen and from the Oxfordian in France. nodes develop on last whorl. Distribution in Kachchh — Upper part of Callovian Remarks — The available, poorly preserved specimen to Oxfordian Chari Formation, Ler. Order Neritopsina COX &KNIGHT, 1960 Superfamily Neritopsioidea RAFINESQUE, 1815 Family Neritopsidae GRAY, 1847 Genus Neritopsis GRATELOUP, 1832 Subgenus Neritopsis GRATELOUP, 1832 Neritopsis (Neritopsis) benoisti COSSMANN, 1900 (Plate I: 11–12, 17–21) 1900: Neritopsis Benoisti nov. sp. — COSSMANN, p. 567, pl. XVII, figs. 18, 23. 1918: Neritopsis Benoisti COSSMANN —ROLLIER, p. 23. 1964: Neritopsis benoisti COSSMANN — J.–C. FISCHER, p. 62, pl. II. figs 12a–b. 1999: Neritopsis (Neritopsis) patchamensis sp. nov. — DAS,BARDHAN &LAHIRI, p. 278, Figure 7: 3–5. Material — Seven, more or less incomplete specimens. Measurements H HL HP D WP AA AL BHU2001I 8 Plate I: 17–19 11.5 11.0 9.5 12.0 9.0 120° BHU2001I 9 Plate I: 20–21 11.5** 10.0** 122° BHU2001I 10 Plate I: 11–12 13.5 13.0 11.0 18.5 13.5* 133°* Description — Rather small species of the genus with whorl surface evenly arched. Sutural part and columellar lip low-spire shell, consisting of rapidly expanding whorls, of peristome partly preserved on one specimen. Frag- separated by widely canaliculate suture. Narrow, feebly ments, as well as sculpture, indicate nearly circular shape of convex ramp below sutural furrow; remaining part of peristome almost in one plane, being strongly prosocline Fragmenta Palaeontologica Hungarica 22, 2004 Jurassic gastropods from Kachchh 11 and almost tangential to base. Peristome fragments have has sparser collabral ribs than N. benoisti, moreover, sharp outer edge; columellar lip looks like keel. Adapically alternating stronger and weaker spiral cords cross the ribs. narrowing furrow forms false umbilicus along columellar The density of the stronger ones is similar to that of the lip. Ornament consists of marked, cancellate elements of collabral ribs. The Callovian Neritopsis (Neritopsis) guerrei dense spiral cords and sparser, somewhat stronger collabral HEBERT &DESLONGCHAMPS, 1860 is another species of ribs. Number of cords between suture and inner lip falls in high resemblance but the three figured specimens (Pl. I, range of 14 to 16 on latest segment of shells. Ribs run
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