Fo Ra Mini Fera from the Late Ju Ras Sic and Early Cre Ta Ceous Car Bon Ate Plat Form Fa Cies of the Southern Part of the Crimea Mountains; South Ern Ukraine

Annales Societatis Geologorum Poloniae (2007), vol. 77: 291–311.

FO RA MINI FERA FROM THE LATE JU RAS SIC AND EARLY CRE TA CEOUS CAR BON ATE PLAT FORM FA CIES OF THE SOUTHERN PART OF THE CRIMEA MOUNTAINS; SOUTH ERN UKRAINE

Mar cin KRA JEWSKI1 & Bar bara OLSZEWSKA2

1 AGH Uni ver sity of Science and Technol ogy, al. Mickiew icza 30, 30-059 Kraków, Poland, kra mar@ge olog.geol.agh.edu.pl 2 Polish Geologi cal Insti tute Car pathi an Branch, ul. Skrzatów 1, 31-560 Kraków, Poland, bar [email protected].

Krajewsk i, M. & Olszewska, B. 2007. Fora mini fera from the Late Juras sic and Early Cre taceous car bonate platform fa cies of the south ern part of the Cri mea Mountains; South ern Ukraine. An nales So cie ta tis Ge olo go rum Po lo niae, 77: 291–311.

Ab stract: Upper Ju ras sic and Lower Cre ta ceous de posits of the Cri mea Pen in sula are rich in micro fossils frequently used for stratigraph ic inter pre ta tions . In case of foramini fera the research has been car ried predom i- nantly on as semblages ob tained by washing the rock sam ples. The present pa per is based on in ves ti ga tions of thin sec tions from the more in du rated sedi ments that sel dom were objects of study. Its goal was to ob tain ad di tional in for mation on age and en vi ron ment of sedi ments studied. Over 250 thin sec tions from 16 surface out crops yielded abun dant fora minifera from which over forty are de scribed herein. Many fo rami nif eral spe cies (e.g. Labiryn thina mirabilis, Pa rur gonina caelin en sis, Neok ili an ina ra honen sis, Amijella amiji, An chis pi ro cyclina lu si tanica) are stra tigraphi cally signifi cant and known from the Kim meridgian–Ti tho nian of the Medi ter ra nean Tethys. The Early Cre taceous fauna is repre sented by Pro topen eroplis ul tra granu lata, Ever ti cy clam mina kel leri, Nau ti lo cu lina bron ni manni, Mon sale via saleven sis, Mayn cina bul garica. Gen er ally, the in ves ti gated fauna is typi cal for pa leoen vi ron ment of the car bon ate plat form. Older (Kim me ridgian–Ti tho nian) as sem blages rep re sent in ner part, younger (Ber ria sian) outer part of the plat form. Paleo geo graphic dis tri bu tion of many spe cies de scribed from the stud ied area in di cates its af filia tion with cos mopoli tan bi ota known from the north Te thyan shelf. Ad di tion ally few cal care ous cysts of di no flag el lata have been iden ti fied and de scribed.

Key words: fo ra mini fers, di no flag el lata, Up per Ju ras sic, Lower Cre ta ceous, Cri mea.

Manu cript re ceived 9 July 2007, ac cepted 17 De cem ber 2007

IN TRO DUC TION from thin sec tions (Fig. 1A). The exam ined sam ples yielded rich foram inif eral fauna, which can be used for stra tigraphi - The Cri mean mi cro fos sil stra tigra phy of the Up per Ju- cal and en vi ron mental in ves ti ga tions. Based on mi cro fau nal rassic –Lower Creta ceous depos it s has been based mainly on data, the paper pres ents the result s of new studies from bed- fora mini fera de scribed by Rus sian and Ukrain ian pa le on - ded and massive fa cies of the Up per Ju ras sic sedi ments tologist s (vide Kuznet sova & Gor batchik, 1985; Gorbatc hik which rep re sent cen tral part of the Cri mea Moun tains. & Kuznetsova, 1994). In major it y of cases, micro fos sil s were ex tracted from soft or mod er ately com pact rocks by wash ing sam ples with wa ter. Mi cro pa le on to logi cal stud ies GEO LOGI CAL SET TING of thin sec tions were rare (Volosh ina et al., 1965; Gorba- tchik & Moha m ad, 1997). Indu rat ed rocks, however, supply The Crim ea Mountai ns occupy the southern, mari time very im por tant pa le on to logi cal in for ma tion use ful in strati- part of the Cri mea Penin sula and form a narrow belt extend - graph i cal or pa leoen vi ron men tal in ter pre ta tions (Sliter, ing nearly W–E at a distanc e of more than 150 km (Fig. 1989, 1999). This encour aged present authors to com plete 1A). The basem ent of the Upper Juras sic rock shows a com - mi cro bi os tra tigra phy of the Up per Ju ras sic sedi ments of the pli cated structure , includ ing a number of intru sive bodies , SW segm ent of the Crim ea Mountai ns based on the data thrusts of cha otic com plexes, faults and tec tonic melange s 292 M. KRAJEWSKI & B. OLSZEWSKA

Fig. 1 Lo cal iza tion study area. A. Upper Ju ras sic–Lower Cre ta ceous de posits in the Cri mea Mountains; af ter Yudin 2001, simplified. B. Lo cal iza tion of sec tions be tween Iograf Ridge and At-Baœ Moun tain

(Nik ishin et al. 1998; Yudin, 1999, 2001; Mileev et al., formably mainly overli es folded flysch strata of the Tauride 2006). se ries (Up per Trias sic–Lower Ju ras sic; Fig. 2). Five main The main ridge of the Crim ea Mountai ns include s an al- se ries were dis tin guished within the Cri mea Moun tains: lochthonous com plex that is com posed of thrusts of Upper Eskiorda, Taurida, Kara dag, Sudak and Yaila (Fig. 2; Juras sic and Lower Cre taceous rocks. This com plex uncon - Mileev et al., 2006; cf. Leshukh et al., 1999). FORAMINIFERA FROM THE LATE JU RAS SIC AND EARLY CRE TA CEOUS, UKRAINE 293

Rocks building the main part of the Cri mea Mountai ns span a time inter val be tween Cal lovian and Ber ria sian (Su - dak and Yaila Serie s), al though strati graphic se quence is some times dis turbed due to com pli cated tec tonic de for mations (cf. Mileev & Bara bosh kin, 1999), and addi ti onall y in cer tain re gions strata of some stages do not oc cur at all. De- posi ti on in the Crim ea Mountai ns area procee ded in a back- arc ba sin, which was filled with shal low- to rela tively deep- wa ter marine sedim ents, close to land ar eas within marginal parts of an epicon ti nental basin that surrounded the Te thys from the north (Zonnen shai n & Le Pichon, 1986; Golonka, 2004). The Crim ea Mountai ns are subdi vided into several smaller massifs (called Yaila), up to 1.500 m a.s.l. Indi vid - ual massifs , al though situate d side by side, fre quently repre - sent tec toni cally iso lated frag ments that are char ac ter ised by dif fer ent mor phol ogy, li thol ogy and stra tigraphic po si tion of Up per Ju ras sic and Lower Cre taceous strata. The sub ject of in ter est: the Aj- Petri and Jalta Yaila massifs (Fig. 1B) are mainly com posed of Titho nian and Ber ria sian rocks be long- ing to the Yaila Se ries (Krajewsk i & Olszewska, 2006; Mileev et al., 2006). The gross part of Aj- Petri and Jalta Yai las is mainly com posed of thick com plexes of bedded limestone s, show- ing variable bed thicknesse s: from finely laminate d to thick-bedded ones. Thin-bedded marly limestone s are ubiq- ui tous. Mas sive lime stones fa cies of car bon ate build ups occur rarely within car bon ates of the area. The stud ies of bed - ded and massive fa cies in the west ern Cri mea Moun tains in - di cate that the Aj- Petri and Jalta Yailas mas sifs are mainly built up of limestone s repre sent ing mostly shallow wa ter fa- cies (pe loi dal, on coi dal, de tri tal, coral, stro ma to por oid, micro bial and marly) as well as sandy limestones and sand - Fig. 2. Gen eral stra tig ra phy of the Cri mea Moun tains af ter Mileev et al., 2006; mod i fied stones (e.g., Leshukh et al., 1999; Kra jewski & Olszewska, 2006; Mileev et al., 2006) Further more, due to com pli cated tecton ics in some ar- eas, the strata are disturbe d (Mileev & Ba raboskhin, 1999) METH ODS and it is dif fi cult to es timate thick ness of the de pos its and its stra tigraphy. Ac cording to some older papers, the total The presented mate rial was col lected from mas sive and thickness reaches a few thousand meter s, but it is proba bly a bedded limestone s and from marly limestone s. A few hun- tectonic ef fect (cf. Le shukh et al., 1999). Therefore more dred sam ples were collec ted, from which thin and polishe d probable thickness would be esti mated from hundreds to sec tions were made. The mate rial was col lected from sev en - thousand meter s for each sedim entar y unit. It is diffi cult to teen sec tions betwee n Iograf Ridge and At-Baœ Mountai n cre ate re al is tic gen eral lithos tra tigraphi cal sec tion for the (Fig. 1B). Over 250 thin sec tions with micro fos sil s have area. been exam ined under Nikon Eclipse LV100 Pol micro - scope. Photos of micro fos sil s have been taken with the aid of Nikon pho to mi cro graphic at tache ments Mi cro flex HFX- PALEONTOLOGICAL CHART DX and NIS- Elements Docu men ta tion al ter na tively. As a re sult of com plex fault tec tonics of this region, the Fo ra mini fera pre vail in all mi cro fos sils as sem blages stra tigraphic po si tion of the Aj- Petri and Jalta Yai las sedi - from the inves ti gated sedi ments. Benthi c forms are the main ments is un cer tain. Since only a few amm on ites were found com ponents . More than forty benthi c spe cies have been in the Jalta Yaila limeston es (Oviech kin, 1956; M. A. identi fied and descri bed, many for the first time from the re- Rogov, pers. comm.), this paper deals with data provided by gion (Figs 4–9). In one sam ple only, repre sen ta ti ve of fo rami nif eral stud ies. Al though stra tigra phy based on mi- plank tic Globu li gerina was spotted. Kim meridgi an as sem - crofos sil s is not as precis e as the or thostra tigraphic scheme blages are more di versi fie d and contai n large, im perfo rat e based on am mon ites, fo ra mini fers are ubiq ui tous in the forms with com plex inte rior typi cal for carbon ate plat forms stud ied sedi ments, un like am mon ites. (Pseudo cy clam mina, Ever ti cy clam mina, Rec to cy clam mina, 294 M. KRAJEWSKI & B. OLSZEWSKA

Fig. 3. The south ern es carp ment of the Jalta and Aj-Petri Yaila with lo cal iza tion of the sam ples pre sented in Figs 4–9 FORAMINIFERA FROM THE LATE JU RAS SIC AND EARLY CRE TA CEOUS, UKRAINE 295

Amijella, Labiryn thina). Ti tho nian–Ber ri as sian assembla- Holbourn & Kaminski, p. 46-47, pl. 16, figs 6-8; pl. 17, figs ges are rich in small forms, es pe cially miliolids and “tro cho - 1-4. li nas” as so ci ated with car bon ate build- ups. In both groups, Re marks. Lon gi tu di nal sec tions show an early, short biserial there are nu mer ous species use ful for stra tigraphical in ter - stage with bul bous chambers and a fol low ing uniserial stage with pre ta tion of in ves ti gated sedi ments (Fig. 9). Notewor thy is low cham bers sub di vided by hor i zon tal su tures. the pres ence of cal care ous cysts of di no flag el lata. Three Range. Tithonian–Barremian. characteristic spe cies of these groups have been also de - Oc cur rence. Sections: KE, KL, KO. scribed. Ge nus Haghimashella Neagu & Neagu, 1995 FORAMINIFERA Haghimashella arcuata Haeusler, 1890 Fora m ini feral taxon om y foll ows schemes elaborat ed by Fig. 4 D Kam inski (2004) for agglu t inat ed foram inifera as well as 1890. Bigenerina arcuata n.sp.: Haeusler, p. 73. (fide Ellis & Loeblich & Tappan (1988), Neagu (1984, 1994, 1995) and Messina, 1941-2007). Septfon ta ine (1988) for calcare ous foram inifera. 1968. Bigenerina arcuata Haeusler: Oesterle, p.742, text-fig. 37-39. 1995. Haghimashella arcuata (Haeusler): Neagu & Neagu, p. Class Foraminiferea d’Orbigny, 1926 216, pl. 2 figs 1-11. Order Lituolida Lankester, 1885 Re marks. Lon gi tu di nal sec tions show the early biserial stage fol - Suborder Lituolina Lankester, 1885 lowed by var i ously in clined adult, uniserial part. Commonly oc cur Family Lituolidae de Blainville, 1827 isolated biserial parts caused by break ing of fragile speci mens . Ge nus Ammobaculites Cushman, 1910 Range. Middle Oxfordian–Berriasian. Ammobaculites coprolithiformis Schwager, 1867 Oc cur rence. Sec tions: KB, KC, KJ, KN. Fig. 4A 1867. Haplophragmium coprolithiformis n.sp.: Schwager, p. 654, Suborder Verneuilina Mikhalevich & Kaminski, 2004 pl. 34, fig. 3 (fide Ellis & Messina, 1941-2007). Fam ily Verneuilinidae Cushman, 1911 1970. Haplophragmium coprolithiforme Schwager: Win ter, p. 8, Ge nus Paleogaudryina Said & Bakarat, 1958 pl. 1, figs 1-21, text-fig. 6. 1981. Ammobaculites coprolithiformis (Schwager): Barnard, Paleogaudryina magharaensis Said & Bakarat (1958) Cordey & Shipp, p. 389-391, pl. 1, fig. 9, text-fig 4. Fig. 4E Re marks. Lon gi tu di nal sec tion shows a tightly coiled, planispiral 1958. Paleogaudryina magharaensis n.sp.: Said & Bakarat, p. early part and a short rec ti lin ear, un coiled adult part. 243, pl.3, fig. 42; pl. 4, figs 33-36. Range. Oxfordian–Kimmeridgian. 2005. Paleogaudryina magharaensis Said & Bakarat: Olszewska, Oc cur rence. Sec tion KC. p. 4, fig. 12. Re marks. Common species, usu ally oc curs in sep a rate parts of the Ge nus Troglotella Wernli & Fookes, 1992 triserial and biserial stages. Differs from the Paleogaudryina var- Troglotella incrustans Wernli & Fookes, 1992 soviensis (Bielecka & Po¿aryski, 1954) in larger triserial stage and Fig. 4B flat tened chambers of the biseral stage giv ing al most rect an gu lar outline in the transversal sections. Simi lar in shape and strati- 1992. Troglotella incrustans Wernli & Fookes n. sp.: Wernli & graphic dis tri bu tion Gaudryina bukowiensis Cushman & Glazew- Fookes, p. 97-102, pl. 1 fig.15; pl. 2, figs 1-12. ski (1949) from the Nizhniow suite of Ukraine dif fers in be ing 1996. Troglotella incrustans Wernli & Fookes: Bucur, Senow- much larger. bari-Daryan & Abate, p. 69, pl. 2, fig. 3; pl. 5, figs 6, 9, 10. Range. Late Kimmeridgian–Middle Berriasian. 1999. Troglotella incrustans Wernli & Fookes: Schlagintweit & Oc cur rence. Sec tions: KA, KB, KF, KG, KK, KL, KN, KR. Ebli, p. 404, pl. 3, fig. 4; pl. 6, figs 7, 9, 10. Re marks. Longi tu di nal sections show typ ical set of slightly in - flated cham bers of variable shape. The early stage, uniserial, bor- Paleogaudryina varsoviensis ing, is fol lowed by an adult stage hor i zon tally at tached to the (Bielecka & Po¿aryski, 1954) sub strate. Fig. 4F Range. Kimmeridgian–Berriasian. Oc cur rence. Sec tions:KA, KB, KC, KG, KJ, KK, KR. 1954. Neobulimina varsoviensis n.sp.: Bielecka & Po¿aryski, p. 65, pl. 10, fig. 50. 1980. Paleogaudryina varsoviensis (Bielecka & Po¿aryski): Suborder Spiroplectamminina Mikhalevich, 1992 Bielecka, In: Malinowska (ed.), p. 303, pl. 82, fig. 10. Fam ily Textulariopsidae Loeblich & Tappan, 1982 Re marks. Mode of occur rence of the species resem bles that of Ge nus Aaptotoichus Loeblich & Tappan, 1982 Paleogaudryina magharaensis Said & Bakarat. It differs in be ing Aaptotoichus challengeri Holbourn & Kaminski, 1997 lon ger, much slen der, hav ing a shorter triserial stage and in more Fig. 4C inflated cham bers of the biserial part. Range. Late Oxfordian–Tithonian. 1997. Aaptotoichus challengeri Holbourn & Kaminski n. sp.: Oc cur rence. Sec tions: KA, KC, KD, KE, KG, KL, KO. 296 M. KRAJEWSKI & B. OLSZEWSKA

Fig. 4. A – Ammobaculites coprolithiformis (Schwager), (KC 30a); B – Troglotella incrustans Wernli & Fookes, (KA-6a); C – Aaptotoichus challengeri Holbourn & Kaminski, (KE 9a); D – Haghimashella arcuata (Haeusler), (KC 4a); E – Paleogaudryina magharaensis Said & Bakarat, (KL 5); F – Paleogaudryina varsoviensis (Bielecka & Po¿aryski), (KA 2a); G – Uvigerinammina uvigeriniformis (Seibold & Seibold), (KL 6); H – Nautiloculina bronnimanni Arnaud-Vanneau & Peybernès, (KJ 40a); I – Nautiloculina oolithica Mohler, (KF 4a); J, K – Mayncina bulgarica Laugh, PeybernÀs & Rey, (KJ 12a) FORAMINIFERA FROM THE LATE JU RAS SIC AND EARLY CRE TA CEOUS, UKRAINE 297

Ge nus Verneuilinoides Loeblich & Tappan, 1949 1967. Nautiloculina oolithica Mohler: Brönimann, p. 54-61, p;. 1, figs 1-6; pl. 2, figs 1-9; pl. 3, figs 1-9; text-figs 1-4. Verneuilinoides polonicus (Cushman & Glazewski, 1949) 1971. Nautiloculina oolithica Mohler: Ramalho, p. 143, pl. 13, Fig. 8 B figs 12, 13. 1984. Nautiloculina oolithica Mohler: Bernier, p. 514, pl. 16, figs 1949. Verneuilina polonica n.sp.: Cushman & Glazewski, p. 7, pl. 7-9. 1, figs 14, 15. 1989. Verneuilina cf. polonica Cushman & Glazewski: Arnaud- Re marks. The species dif fers from Nautiloculina bronnimanni in Vanneau & Masse, p. 264-265. smaller size, larger num ber of cham bers and in much broader pe - 1997. Verneuilinoides polonicus (Cushman & Glazewski): riph ery. It has also lon ger strati graphi cal dis tri bu tion. Neagu, p. 313, Fig. 4 (13-19); Fig. 5 (39-49). Range. Late Oxfordian–Berriasian. Oc cur rence. Sec tions: KA, KC, KD, KF, KG, KK, KO, KR. Re marks. The subaxial sec tions show dis tinct triserial arrange - ment of the slowly en larg ing weakly in flated chambers with char - ac ter is tic thick walls. Fam ily Mayncinidae Loeblich & Tappan, 1985 Range. Tithonian–Early Valanginian. Ge nus Mayncina Neumann, 1965 Oc cur rence. Sec tions: KD, KF. Mayncina bulgarica Laugh, PeybernÀs & Rey, 1968 Fig. 4 J, K Fam ily Reophacellidae Mikhalevich & Kaminski, 2004 Ge nus Uvigerinammina Majzon, 1943 1968. Mayncina bulgarica n.sp.: Laug, PeybernÀs & Rey, p. 68-76; fig. 3, 1-16. Uvigerinammina uvigeriniformis 1986. Mayncina? aff. bulgarica Laug, PeybernÀs & Rey: Luperto (Seibold & Seibold, 1960) Sinni & Masse, pl. 7, figs 1-3. Fig. 4G 1988. Mayncina cf. bulgarica Laug, PeybernÀs & Rey: Bucur, pl. 1, fig. 14. 1960. Gaudryina uvigeriniformis n.sp.: Seibold & Seibold, p. 334, 1991. Mayncina? sp.: Altiner, pl. 12, figs 1, 2. 335; text-fig. 8b, pl. 7, fig. 4. 2004. Mayncina bulgarica Laug, PeybernÀs & Rey: Ivanova & 1995. Uvigerinammina uvigeriniformis (Seibold & Seibold): Koleva-Rekalova, pl. 3, fig. 10. Neagu & Neagu, p. 218, pl. l2, figs 28-43; pl. 6, figs 11-14. 2005. Uvigerinammina uvigeriniformis Seibold & Seibold): Re marks. Subequatorial sec tions of the macrospheric spec i mens Olszewska, p 34, p. 5, fig. 1. show two whorls composed of slowly en larg ing, rect an gu lar cham bers, finely ag glu ti nated walls. Sec tions of the microspheri- Re marks. Axial sections show typ ical for the species sphaerical cal spec i mens show more nu merous and nar row chambers and ten - ini tial cham ber, al ter nat ing at tach ment of cham bers and their dency to un coil ing. The subaxial sec tions show succes sive open - sack-like shape. ings be tween cham bers and acute pe riph ery. Range. Middle Oxfordian–Early Valanginian. Range. Tithonian–Barremian. Oc cur rence. Sections: KL, KN. Oc cur rence. Sec tions: KA, KC, KJ, KL, KN, KO, KR.

Fam ily Nautiloculinidae Loeblich & Tappan, 1985 Order Loftusiida Kaminski & Mikhalevich, 2004 Ge nus Nautiloculina Mohler, 1938 Suborder Loftusiina Kaminski & Mikhalevich, 2004 Nautiloculina bronnimanni Arnaud Vanneau Fam ily Mesoendothyridae Voloshinova, 1958 & PeybernÀs, 1978 Ge nus Mesoendothyra Dain, 1958 Fig. 4H Mesoendothyra izjumiana Dain, 1958 1978. Nautiloculina bronnimanni n.sp.: Arnaud Vanneau & B. Fig. 5A PeybernÀs, p. 70, pl.1, figs 6-8; pl. 2, figs 4-11. 1958. Mesoendothyra izjumiana n.sp.: Dain, In: Bykova et al., p. 1998. Nautiloculina bronnimanni Arnaud-Vanneau & PeybernÀs: 20, 21 pl. 4, figs 7-9. Ebli & Schlagintweit, p. 13, pl. 2, figs 5, 6. 1991. Mesoendothyra izjumiana Dain: Altiner, pl. 4, figs 1-3. 2003. Nautiloculina broennimanni Arnaud-Vanneau & 2004. Mesoendothyra izjumiana Dain: Ivanova & Koleva-Reka- PeybernÀs: Dragastan & Rich ter, p. 93, pl. 1, fig. 2; pl. 9, lova, p. 219, pl. 1, figs 6-9. figs 10, 11, 16 n. 2005. Mesoendothyra izjumiana Dain: Olszewska, p. 35, pl. IV, 2004. Nautiloculina bronnimanni Arnaud-Vanneau & PeybernÀs: figs 5, 6. Ivanova & Koleva-Rekalova, p. 220, pl. 1, fig. 5. Re marks. Axial and subaxial sec tions show typ i cal, early Re marks. Axial sections show, typ ical for the spe cies, slightly streptospiral part followed by planispiral late whorl, small number acute periph ery, 6 whorls of semicir cu lar cham bers and charac ter - of cham bers and a broad ex ter nal mar gins. istic pro jec tions (septa) over ap er tural part of the pre ced ing cham - Range. Late Oxfordian–Tithonian. ber. Oc cur rence. Sec tions: KB, KD, KG. Range. Berriasian–Hauterivian. Oc cur rence. Sec tions: KA, KB, KC,KD, KF, KJ. Ge nus Labirynthina Weynschenk, 1951 Nautiloculina oolithica Mohler, 1938 Labirynthina mirabilis Weynschenk, 1951 Fig. 4I Fig. 5B 1938. Nautiloculina oolithica n.sp.: Mohler, p. 19, pl. 4, figs 1-3 1951. Labirynthina mirabilis n.sp.: Weynschenk, p. 793. (fide Ellis & Messina 1941-2007). 1984. Labirynthina mirabilis Weynschenk: Bernier, p. 515-517, 298 M. KRAJEWSKI & B. OLSZEWSKA

Fig. 5. A – Mesoendothyra izjumiana Dain, (KD 5); B – Labirynthina mirabilis Weynschenk, (KA 8a); C, D – Everticyclammina praekelleri Redmond (KP 4a); E – Everticyclammina kelleri Henson (KL 10); F – Rectocyclammina chouberti Hottinger (KN 1a); G, H – Charentia evoluta Gorbatchik, (KA 1a); I – Melathrokerion spirialis Gorbatchik (KC 16a); J – Scythiolina camposaurii (Sartoni & Crescenti) (KF 3a); K – Montsalevia salevensis (Charollais, Brönnomann & Zaninetti) (KC 36a) FORAMINIFERA FROM THE LATE JU RAS SIC AND EARLY CRE TA CEOUS, UKRAINE 299

pl. 19, fig. 3. 2004. Rectocyclammina chouberti Hottinger: Ivanova & Koleva- 1991. Labirynthina mirabilis Weynschenk: Altiner, pl. 3, figs 17, Rekalova, pl. 1, fig. 1. 18, 20-22, 24, 19, 23. Re marks. Axial sec tions show the early short planispiral whorl 1997. Labirynthina mirabilis Weynschenk: Bassoulet, p. 301- fol lowed by uniserial, rec ti lin ear later part com posed of the slowly 302. in creas ing, over lap ping cham bers with thick septa. In some sec - 2004. Labirynthina mirabilis Weynschenk: Ivanova & Koleva- tions, charac ter is tic alveoles in cham ber walls may be observed . Rekalova, p. 218, pl. 2-4. Range. Late Kimmeridgian–Tithonian (?Valanginian). 2006. Labirynthina mirabilis Weynschenk: Krajewski & Olszew- Oc cur rence. Sec tions: KB, KE, KF, KL, KN. ska, pl.1, fig. 5b. Re marks. Lon gi tu di nal sec tions show an early in vo lute stage (with charac teris tic large initial chamber) followed by an evolute, recti lin ear late stage. Inter nal pillars within chambers and their Suborder Biokovinina Kaminski, 2004 microgranular walls are also vis ible. Fam ily Charentiidae Loeblich & Tappan, 1985 Range. Lat est Oxfordian–Early Tithonian. Ge nus Charentia Neumann, 1965 Oc cur rence. Sec tions: KA, KB, KC, KG. Charentia evoluta (Gorbatchik, 1968) Fig. 5G, H Family Everticyclamminidae Septfontaine, 1988 1968. Tonasia evoluta n.sp.: Gorbatchik, p. 8, 9; pl. 2, figs 1-5. Ge nus Everticyclammina Redmond, 1964 1975. Charentia evoluta (Gorbatchik): Kuznetsova & Gorbat- chik, p. 82, 83; pl. 3, figs 5, 6. Everticyclammina kelleri (Henson, 1948). 1999. Charentia evoluta (Gorbatchik): Neagu, p. 292, pl. 3, figs Fig. 5E 24-29; pl. 9, figs 25, 26. 2005. Charentia evoluta (Gorbatchik): Olszewska, p. 35, pl. IV, 1948. Pseudocyclammina kelleri n.sp. Henson, p. 16, 17; pl. 9, figs 7, 8. figs 4, 5, 7 (fide Ellis & Messina, 1941-2007). 1964. Everitcyclammina eccentrica n.sp.: Redmond, p. 408, pl. 1, Re marks. Hori zon tal sec tions of the early, planispiral part show figs 16-18; pl. 2, figs 12, 13. rect an gular cham bers subdi vided by thin septa. In ax ial sec tions 1964. Everticyclammina elegans n.sp.: Redmond, p. 408-409, pl. (un like in the ge nus Nautiloculina) the base of cham bers lack in- 1, figs 19-21; pl. 2, figs 14-16. ter nal pro jec tions. Sec tions of spec i mens with uncoiled late part 1990. Everticyclammina kelleri (Henson): Ban ner & High ton, p. oc cur rarely. 6, pl. 1, figs 2-6; pl. 2, figs 1-4; pl. 3, figs 1, 2. Range. Late Kimmeridgian–Valanginian. Oc cur rence. Sec tions: KA, KC, KD, KF, KJ, KK, KN, KR. Re marks. In the ma te rial studied usually occur, planispirally coiled, early stages of the spe cies fol lowed by one cham ber of the uncoiled part. To char ac ter istic fea tures be long two whorls and the Genus Melathrokerion Brönnimann & Conrad, 1967 non al ve olar walls of chambers in the coiled stage. Melathrokerion spirialis (Gorbatchik, 1968) Range. Berriasian–Valanginian. Oc cur rence. Sec tions: KD, KE, KK. Fig. 5I 1968. Melathrokerion spirialis n.sp.: Gorbatchik, p. 6, 7; pl. 1 figs Everticyclammina praekelleri Banner & Highton, 1990 1-6. Fig. 5C, D 1985. Melathrokerion spirialis Gorbatchik: Kuznetsova & Gorba- tchik, p. 81, pl. 3, fig. 4. 1990. Everticyclammina praekelleri n.sp.: Ban ner & High ton, p. Re marks. Ax ial sec tions show typ i cal sub acute pe riph ery, 8-10, pl. 1, fig. 1; pl. 3, fig. 5; pl. 4, figs 1-11. streptospiral early whorl, thick septa be tween cham bers (un like in Re marks. The species is charac ter ized by thick cham ber walls of the ge nus Charentia) and coarse alve o lar canaliculi. ir reg u lar thick ness, ir reg u lar shape of cham bers and dis tinct Range. Tithonian–Valanginian (pre domi nantly on the Carpathian- alveoles even in the early part. It differs from Evericyclammina Cri mea area). virguliana (Koechlin) in having thicker walls, ir reg ular shape of Oc cur rence. Sec tions: KB, KC, KE, KF, KL, KR. cham bers and a smaller num ber of cham bers per whorl. Range. Kimmeridgian–Tithonian. Oc cur rence. Sec tions: KA, KB, KC, KD, KE, KF, KG, KJ, KL, KN, KO, KP, KR. Family Montsaleviidae Zaninetti, Salvini-Bonnard, Charollais & Decrouez, 1987 Ge nus Rectocyclammina, Hottinger, 1967 Ge nus Montsalevia Zaninetti, Salvini Bonnard, Charollais & Decrouez, 1987 Rectocyclammina chouberti Hottinger, 1967 Fig. 5F Montsalevia salevensis (Charollais, Brönnimann & Zaninetti, 1966) 1967. Rectocyclammina chouberti n.sp.: Hottinger, p. 55, 56, pl. Fig. 5K 9, figs 19-21; text-figs 26, 27. 1971. Rectocyclammina chouberti Hottinger: Ramalho, p. 144, 1966. Pseudotextulariella salevensis n.sp.: Charollais, Brönni- 145, pl. 14, figs 1-4. mann & Zaninetti, p. 28-34, pl. 1, figs 1-5; pl. 2, figs 2, 6; 1984. Rectocyclammina chouberti Hottinger: Bernier, p. 513-514, text-fig. 1. pl. 20, fig. 3. 1987. “Montsalevia” salevensis (Charollais, Brönnimann & Zani- 1997. Rectocyclammina chouberti Hottinger: Bassoulet, p. 303. netti): Zaninetti, Charollais & Decrouez, p. 168. 300 M. KRAJEWSKI & B. OLSZEWSKA

Fig. 6. A – Siphovalvulina varaibilis Septfontaine (KE 11a); B – Dobrogelina ovidi Neagu (KC 44a); C – Amijella amiji (Henson) (KG 5a); D, E – Anchispirocyclina lusitanica (Egger) (KL 11); F – Pseudocyclammina lituus (Yokoyama) (KJ 30a); G – Parurgonina caelinensis Cuvillier, Foury & Pignatti Morano (KD 9); H – Neokilianina rahonensis (Foury & Vin cent) (KD 10); I – Bigenerina erecta Dain (KA 14a); J – Andersenolina alpina (Leupold) (KA 1a); K – Andersenolina elongata (Leupold) (KF 2a); L – Ichnusella burlini (Gorbatchik) (KB 3a) FORAMINIFERA FROM THE LATE JU RAS SIC AND EARLY CRE TA CEOUS, UKRAINE 301

1988. Pseudotextulariella salevensis Charollais, Brönnimann & Ge nus Amijella Loeblich & Tappan, 1985 Zaninetti: Bucur, pl. 2, figs 11, 12. 2004. Montsalevia salevensis (Charollais, Brönnimann & Zani- Amijella amiji (Henson, 1948) netti): Ivanova & Koleva-rekalova: p. 220, pl. 2, figs 4-6. Fig. 6C 2004. Montsalevia salevensis (Charollais, Brönnimann & Zani- 1948. Haurania amiji n. sp.: Henson, p. 12; pl. 15, figs 5-10. netti): Ivanova & Ko³odziej, pl. 1, fig. K. 1967. Haurania amiji Henson: Hottinger, p. 52, pl. 8, figs 1-6, Re marks. Oblique sec tion shows succes sion of low cham bers and 20-21, text-fig. 2. traces of verti cal parti tions in the biserial part. 1991. Amijella amiji (Henson): Schlaginweit, p. 248-250, pl. I, Range. Late Berriasian–Hauterivian. figs 1-10. Oc cur rence. Sec tion KC. 1991. Amijella amiji (Henson): Darga & Schlaginweit, p. 212, pl. 4, figs 9, 10, 12. Fam ily Cuneolinidae Saidova, 1981 1997. Bramkampella arabica Redmond: Gorbatchik & Moha- mad, pl. 1, figs 8, 9, 11. Ge nus Scythiolina Neagu, 2000 Re marks. The subaxial sec tion of typi cal club-like spec i men Scythiolina camposaurii (Sartoni & Crescenti, 1964) show a glob u lar ini tial chamber and slowly en larg ing suc ces sive Fig. 5J cham bers with in tense subepidermal net work of beams and hori - zon tal raf ters. Schlaginweit (1991) af ter the thor ough in ves ti ga tion 1964. Cuneolina camposaurii n.sp.: Sartoni & Crescenti, p. of gen era Amijella Loeblich & Tappan (1985) and Bramkampella 275-277, pl. 24, fig. 1; pl. 48, figs 1-6. Redmond (1964) came to con clu sion that they have iden ti cal 1984. Cuneolina camposaurii Sartoni & Crescenti: Luperto Sinni struc ture thus are synonimous. & Masse, pl. 41, figs 4, 5. Range. Tithonian–Berriasian. 1988. Cuneolina camposaurii Sartoni & Crescenti: Bucur, pl.1, Oc cur rence. Sec tions: KB, KC, KD, KF, KG, KJ, KK, KL, KN, figs 13, 14. KP, KR. 2000. Scythiolina camposauri (Sartoni & Crescenti): Neagu, p. 369, pl.1, figs 41-44; pl. 2, figs 18-29, pl. 4, figs 50, 54; pl. 7, figs 7-10. Ge nus Anchispirocyclina Jordan & Applin, 1952 Re marks. Sections paralell with the plane of biseriality show typ i - Anchispirocyclina lusitanica (Egger, 1902) cal for the species flabelliform shape of the test, short early planispiral stage and verti cal ra dial parti tions within late cham- Fig. 6D, E bers. 1902. Dicyclina lusitanica n.sp.: Egger, p. 585-586, pl. 6, fig. 3-5 Range. Latest Berriasian–Barremian. (fide Ellis & Messina 1941-2007). Oc cur rence. Section KF. 1971. Anchispirocyclina lusitanica (Egger): Ramalho, p. 148- 149, pl. 8, fig. 2; pl. 10, fig. 1; pl. 15, figs 4-9; pl. 16, figs 1, 2. 1991. Anchispirocyclina lusitanica (Egger); Darga & Schlagin- Suborder Orbitolinina Kaminski 2004 weit, p. 213, pl. 2, fig. 2; pl. 4, figs 2, 3. Fam ily Pfenderinidae Smout & Sugden, 1962 1997. Anchispirocyclina lusitanica (Egger): Bassoulet, p. 303. Ge nus Siphovalvulina Septfontaine, 1988 1998. Anchispirocyclina lusitanica (Egger): Ebli & Schlagint- Siphovalvulina variabilis Septfontaine, 1988 weit, p. 12, pl. 1, fig. 6. 1999. Anchispirocyclina lusitanica (Egger): Schlagintweit & Fig. 6A Ebli: p. 398, pl. 5, fig. 9. 1988. Siphovalvulina variabilis n.sp.: Septfontaine, p. 245. Re marks. The axial section (D) shows a slightly asymetrically 1991. Siphovalvulina variabilis Septfontaine: Darga & Schlagin- coiled early part fol lowed by planispiral later part with many weit, p. 214, pl. 4, fig. 14. cham bers (E) ir reg u larly subdi vided by beams and raf ters. 2004. Siphovalvulina variabilis Septfontaine: Ivanova & Koleva- Range. Tithonian–ear li est Berriasian. Rekalova, pl. 3, fig. 8. Oc cur rence. Sec tions: KB, KJ, KK, KL, KN, KO. Re marks. Lon gi tu di nal sec tions show in ter nal ca nal par al lel to the axis of coil ing and the sack-like shape of chambers. Range. Mid dle Ju ras sic–Tithonian Ge nus Pseudocyclammina Yabe & Hanzawa, 1926 Oc cur rence. Sec tions: KA, KE, KL, KR. Pseudocyclammina lituus (Yokoyama, 1890) Fig. 6F Ge nus Dobrogelina Neagu, 1979 1890. Cyclammina lituus n.sp.: Yokoyama, p. 26, pl. 5, fig. 7. Dobrogelina ovidi Neagu, 1979 1984. Pseudocyclammina lituus (Yokoyama): Bernier, p. 513, pl. Fig. 6B 19, figs 5, 6. 1991. Pseudocyclammina lituus (Yokoyama): Altiner, pl. 7, fig. 9. 1979. Dobrogelina ovidi n. sp.: Neagu, p. 494, pl. 1, figs 1-7; pl. 4, 1997. Pseudocyclammina littus (Yokoyama): Bassoulet, p. 303. figs 17, 18. 2004. Pseudocyclammina lituus (Yokoyama): Ivanova & Koleva- 2004. Dobrogelina ovidi Neagu: Ivanova & Koleva-Rekalova, pl. Rekalova, p. 219, pl. 1, fig. 10. 3, figs 9, 11. 2005. Pseudocyclammina lituus (Yokoyama): Olszewska, p. 35, Re marks. Axial and oblique sec tions show convex spi ral side, in - p. IV, fig. 10. flated cham bers and char ac ter is tic deep um bi li cus. 2006. Pseudocyclammina lituus (Yokoyama): Kobayashi & Range. Berriasian–Valanginian. Vuks, fig. 5 (7-14). Oc cur rence. Sec tions: KC, KJ, KL, KN, KR. Re marks. Axial sections show a planispiral early stage, coarsely 302 M. KRAJEWSKI & B. OLSZEWSKA

ag glu ti nated walls and typ i cal coarse subepidermal net work. Un- let, p. 303. coiled speci mens rarely oc cur. Re marks. Ac cord ing to Septfontaine (1988), gen era Neokilianina Range. Oxfordian–Berriasian. and Parurgonina are mor pho log i cally re lated, the for mer be ing an Oc cur rence. Sec tions: KA, KC, KF, KJ, KK, KL, KR. older homeomorph. Lon gi tu di nal-oblique sec tion of the poorly pre served spec i mens shows coni cal shape of the test with vis i ble Fam ily Parurgoninidae Septfontaine, 1988 cham bers of the rec ti lin ear part al ter nat ing in po si tion and sub di- Ge nus Parurgonina Cuvillier, vided into chamberlets. Range. Kimmeridgian–ear li est Tithonian. Foury & Pignatti Morano, 1968 Oc cur rence. Sec tions: KA, KD. Fig. 6G

1968. Urgonina (Parurgonina) caelinensis n.sp.: Cuvillier, Foury Suborder Involutinina Hohenegger & Piller, 1977 & Pignatti Morano, p. 150-154, pl. II, figs 1-12; pl. III, figs Fam ily Involutinidae Bütschli, 1880 1-9. 1975. Parurgonina caelinensis Cuvillier, Foury & Pignatti Ge nus Andersenolina Neagu, 1994 Morano: Schroeder, Guellal & Villa, p. 319-326, pl. 1, figs Andersenolina alpina (Leupold, 1936) 1-4; p. 2, figs 3-5. Fig. 6J 1984. Parurgonina caelinensis Cuvillier, Foury & Pignatti Morano: Bernier, p. 522-523, pl. 20, figs 4, 5, 7, 8. 1936. Coscinodiscus alpinus n.sp.: Leupold, p. 610, pl. 18, figs 1988. Parurgonina caelinensis Cuvillier, Foury & Pignatti 1-8 (fide Ellis & Messina, 1941-2007). Morano: Septfontaine, p. 248-249. 1991. Trocholina alpina (Leupold): Darga & Schlagintweit: p. 1993. Paururgonina caelinensis Cuvillier, Foury & Vicent: 214, pl. 4, fig. 1. Bucur, pl. 4, figs 1-8. 1994. Andersenolina alpina (Leupold): Neagu, p. 133, text-fig. 4, 1997. Parurgonina caelinensis Cuvillier, Foury & Pignatti figs 3, 4; pl. 7, figs. 8, 9; pl. 8, figs 1-10; pl. 12, figs 1-5. Morano: Bassoulet, p. 302. 2003. Andersenolina alpina (Leupold): Dragastan & Rich ter, p. Re marks. Subaxial sec tion shows a large glob u lar ini tial chamber 89; pl. 10, figs 1-4. with suc ces sive cham bers added in trochospiral coil what re sults Re marks. Lon gi tu di nal sec tions show a small cone with the api cal in a cone-like shape of the test. In transversal section ver tical pil- an gle of 80–95° and 4 to 5 whorls of low, cres cen tic chambers typ - lars in the central part of the test are also vis ible. ical for the species. Range. Lat est Oxfordian–Early Tithonian, mostly Kimmeridgian. Range. Tithonian–Early Valanginian. Oc cur rence. Sec tions: KA, KC, KR. Oc cur rence. Sec tions: KA, KB, KC, KD, KE, KF, KJ, KK, KL, KN, KR. Order Textulariidae Delage & Herouard, 1896 Suborder Textulariina Delage & Herouard, 1896 Andersenolina elongata (Leupold, 1936) Fam ily Textulariidae Ehrenberg, 1838 Fig. 6K Ge nus Bigenerina d’Orbigny, 1826 1936. Coscinodiscus elongatus n.sp.: Leupold, p. 617, pl. 8, figs Bigenerina erecta Dain, 1976 12-14 (fide Ellis & Messina 1941-2007). Fig. 6I 1988. Trocholina elongata (Leupold): Arnaud-Vanneau, Boi- sseau & Darsac, p. 356-357, pl. 1, fig. 4; pl. 2, figs 1-8. 1976. Bigenerina erecta n.sp.: Dain in Dain & Kuznetsova, p. 1991. Trocholina elongata (Leupold): Darge & Schlagintweit: p. 54-55, pl. 7, fig. 4. 214, pl. 4, fig. 4. Re marks. Lon gi tu di nal sec tions show the early, wedge-shaped 1994. Andersenolina elongata (Leupold): Neagu, p. 130, text-fig. biserial part and the directly adja cent uniserial, rectilineal late 3, fig. 7; pl. 4, figs 1-22; pl. 6, figs 12-14; pl. 12, figs 13-17. stage. 2003. Andersenolina elongata (Leupold): Dragastan & Richer, p. Range. Tithonian. 89, 90; pl.10, fig. 7. Oc cur rence. Section KA. Re marks. Lon gi tu di nal sec tions show a long, slen der shape of the spe cies com posed of over 7 whorls of low cham bers and a sharp api cal cone of 22°–30°. Family Paravalvulinidae Ban ner, Range. Tithonian–Early Valanginan. Simmons & Whittaker, 1991 Oc cur rence. Sec tions: KB, KF, KK, KN. Ge nus Neokilianina Septfontaine, 1988 Neokilianina rahonensis (Foury & Vincent , 1967) Ge nus Ichnusella Dieni & Massari, 1966 Fig. 6H Ichnusella burlini (Gorbatchik, 1959) 1967. Kilianina rahonensis n. sp.: Foury & Vin cent, p. 39-44, pl. Fig. 6L 2, figs 1-14. 1984. Kilianina rahonensis Foury & Vincent: Bernier, p. 520, pl. 1959. Trocholina burlini n.sp.: Gorbatchik, p. 81, pl. 4, figs 3-5. 20, fig. 6. 1995. Ichnusella burlini (Gorbatchik): Neagu, p. 271, 272; pl. 2, 1988. Neokilianina rahonensis (Foury & Vin cent); Septfontaine, figs 45-48; pl. 3, figs 13-36, 45-48; pl. 13, fig. 10. p. 249. Re marks. Char ac ter istic for the spe cies is a low cone of 100–115° 1993. Kilianina rahonensis Foury & Vincent: Bucur, pl. 3, figs 4, and 4–5 whorls of the low cham bers. In the lon gi tu di nal or trans - 5, 8, 9. verse sections of the well preserved speci mens close to the um bili - 1997. Neokilianina rahonensis (Foury & Vin cent), J.-P. Bassou- cal side the calcite crys tals are visi ble. FORAMINIFERA FROM THE LATE JU RAS SIC AND EARLY CRE TA CEOUS, UKRAINE 303

Range. Tithonian–Valanginian. recrystallised) hyaline walls of the test. Oc cur rence. Sec tions: KB, KF, KK, KO, KR. Range. Middle Late Tithonian–Valanginian. Oc cur rence. Sec tions: KB, KC, KD, KE, KJ, KK, KL, KN, KR. Ge nus Neotrocholina Reichel, 1956 emended Neagu, 1995 Neotrocholina molesta (Gorbatchik, 1959) Fig. 7A Suborder Miliolina Delage & Herouard, 1875 Fam ily Cornuspiridae Schulze, 1854 1959. Trocholina molesta n.sp.: Gorbatchik, pl 4 figs 1, 2. 1988. Trocholina molesta Gorbatchik: Arnaud-Vanneau, Bois- Ge nus Meandrospira Loeblich & Tappan, 1946 seau & Darsac, p. 359, pl. 6, figs 11-21. Meandrospira favrei Charollais, 1995. Neotrocholina burgeri molesta (Grobatchik): Neagu, p.16- Brönnimann & Zaninetti, 1966 19; pl. 1, figs 13-16, 21, 22, 25, 26; pl. 7, fig. 62-67, 70, 71; Fig. 7B pl. 9, figs 1-9; pl. 13, fig. 13, 25, 26. 1998. Trocholina molesta (Gorbatchik): Ebli & Schlagintweit, p. 1966. Citaella? favrei n.sp.: Charollais, Brönnimann & Zaninetti, 15, pl. 2, fig. 3. p. 37-47, pl. 2, figs 3, 4; pl. 3, figs 1-5; pl. 5, figs 1, 2; 2005. Neotrocholina molesta (Gorbatchik): Olszewska, p. 36, 37; text-figs 4-6. pl. V, fig. 12. 1988. Meandrospira favrei (Charollais, Brönnimann & Zani- Re marks. Test moder ately con ical with an apical angle of 90– netti): Bucur, pl. 2, figs 1-3. 120° and 4 to 6 whorls of the low, cres cen tic chambers. 1991. Meandrospira favrei (Charollais, Brönnimann & Zani- Range. Tithonian–Barremian. netti): Altiner, pl.13, figs 1-5. Oc cur rence. Sec tions: KC, KE, KN, KO. 1999. Meandrospira favrei (Charollais, Brönnimann & Zani- netti): Schlagintweit & Ebli, p. 399-400, pl. 4, figs 8,11. 2004. Meandrospira favrei (Charollais, Brönnimann & Zani- netti): Ivanova & Ko³odziej, pl.1, figs L, M. Fam ily Ventrolaminidae Weynschenk, 1950 Ge nus Protopeneroplis Weynschenk, 1950 Re marks. Loeblich & Tappan (1988) included ge nus Citaella into the ge nus Meandrospira. Ex am ined spec i mens in var i ous sec tions Protopeneroplis striata Weynschenk, 1950 reveal subsphaerical small initial cham ber and typi cally strepto- Fig. 8D, E spi ral un di vided tu bu lar, microgranular, sec ond cham ber. Range. Lat est Berriasian–Hauterivian. 1950. Protopeneroplis striata n.sp.: Weynschenk, p. 13, pl.2, figs Oc cur rence. Sec tions: KD, KE, KF, KG, KO. 12-14. 1991. Protopeneroplis striata Weynschenk: Altiner, pl. 3, figs 1-7. 1999. Protopeneroplis striata Weynschenk: Schlagintweit & Family Hauerinidae Schwager, 1876 Ebli, p. 402, pl. 6, figs 3, 4. Ge nus Decussoloculina Neagu, 1984 2005. Protopeneroplis striata Weynschenk: Olszewska, p. 37, pl. V, fig. 13. Decussoloculina barbui Neagu, 1984 Re marks. The ax ial sec tions show fully planispiral mode of coil - Fig. 7D ing of he spe cies. Ax ial, subaxial or transversal sec tions show 1984. Decussoloculina barbui n.sp.: Neagu, p. 81, 82; pl. 2, figs char ac ter is tic two lay ered cham ber walls (“striae”). The in ter nal 8-12. layer is built of calcite cristals (light in trans mitted light) while the 2003. Decussoloculina barbui Neagu: Dragastan & Rich ter, p. 93, exter nal layer is built of microgranular calcite (dark in trans mitted pl. 9, fig. 15. light). 2005. Decussoloculina barbui Neagu: Olszewska, p. 37, pl. VI, Range. Middle-Late Juras sic (up to Tithonian). figs 4, 5. Oc cur rence. Sec tions. KA, KB, KC, KF. Re marks. Transversal sec tions show “X” shaped arrange ment of four cham bers in one whorl what re sults in somewhat ir reg u lar Protopeneroplis ultragranulata (Gorbatchik, 1971) outline of the test. Fig. 7C Range. Middle Tithonian–Valanginian. 1971. Hoeglundina (?) ultragranulata n.sp.: Gorbatchik, p. 135, Oc cur rence. Sec tions: KA, KC, KD, KL, KO. pl. 26, fig. 2. 1991. Protopeneroplis trochangulata Septfontaine: Altiner, pl. 7, figs 1-5. Ge nus Quinqueloculina d’Orbigny, 1826 1996. Protopeneroplis ultragranulata (Gorbatchik): Bucur, Quinqueloculina semisphaeroidalis Danitch, 1971 Senowbari-Daryan & Abate, p. 69-70, pl. 3, figs 14-17. Fig. 7H 1999. Protopeneroplis ultragranulata (Gorbatchik): Schlagint- weit & Ebli, p. 420-423, pl., 6, figs 5, 6, 9. 1971. Quinqueloculina semisphaeroidalis n.sp.: Danitch, In: 2004. Protopeneroplis ultragranulata (Gorbatchik): Ivanova & Romanov & Danitch, p. 144-145, pl. 39, figs 1-4. Ko³odziej, pl. 1, fig. C. Re marks. Transversal sec tions show al most circu lar out line of the 2005. Protopeneroplis ultragranulata (Gorbatchik): Olszewska, test and a “Y” mode arrange ment of cham bers and rela tively thick p. 37, pl. V, figs 15, 16. walls. Re marks. Charac ter is tic for the species is trochospiral mode of Range. Late Oxfordian–Tithonian. coil ing, lack of the microgranular “striae” and the thick ened (of ten Oc cur rence. Sec tions: KA, KE, KG, KK, KL, KO. 304 M. KRAJEWSKI & B. OLSZEWSKA

Fig. 7. A – Neotrocholina molesta (Gorbatchik) (KC 25a); B – Meandrospira favrei (Charollais, Brönnimann & Zaninetti) (KO 3a); C – Protopeneroplis utragranulata (Gorbatchik) (KC 4a); D – Decussoloculina barbui Neagu (KC 41a); E – Rumanoloculina mitchurini (Dain) (KA 5a); F – Quinqueloculina stellata Matsieva & Temirbekova (KR 13a); G – Scythiloculina confusa Neagu (KB 17a); H – Quinqueloculina semisphaeroidalis Danitsch (KB 28a); I, J – Rumanoloculina verbizhiensis (Dulub) (KB 30a)

Quinqueloculina stellata Matsieva & Temirbekova, 1989 Ge nus Rumanoloculina Neagu, 1986 Fig. 7F Rumanoloculina mitchurini (Dain, 1971) 1989. Quinqueloculina stellata n.sp.: Matsieva & Temirbekova, Fig. 7E p. 115, pl. 1, figs d, z, e. Re marks. Transversal sec tions show “Y” mode of chamber ar - 1971. Quinqueloculina mitchurini n.sp.: Dain, In: Dain & Kuzne- range ment and dou ble pro jec tions at outer walls of cham bers of tsova p. 114-115, pl. 1, figs 9, 10. the last whorl that mark ribs run ning along the test. 1989. Quinqueloculina mitchurini Dain: Matsieva & Temirbe- Range. Tithonian–Early Berriasian. kova, p. 115, pl. 1, figs a-g. Oc cur rence. Sec tions: KB, KC, KE, KF, KR. Re marks. Transversal sec tion shows “Y” mode of chamber ar - FORAMINIFERA FROM THE LATE JU RAS SIC AND EARLY CRE TA CEOUS, UKRAINE 305

range ment and tri an gu lar but rounded out line of the test. Sim i lar trochospiral mode of coil ing and a two lay ered wals: in ner-dark features of the transversal section dis play Quinqueloculina and microgranular, outer-clear, hyaline. jurassica Bielecka & Styk from the Late Oxfordian–Early Range. Oxfordian–Valanginian. Kimmeridgian of Poland and Quinqueloculina podlubiensis Oc cur rence. Sec tions: KA, KB, KC, KD, KE, KJ, KK, KL, KN, Tereshuk from the Kimmeridgian–Tithonian sed i ments of the KR. Western Ukraine. The au thors of both above men tioned spe cies re - late them to the Quinqueloculina sp. A and Quinqueloculina sp. B Suborder Globigerinina Delage & Herouard, 1896 re ported by Cushman & Glazewski (1949) from the Tithonian Fam ily Globuligerinidae Loeblich & Tappan, 1884 Nizhniov lime stone of the West ern Ukraine. More de tailed in ves - Ge nus Globuligerina Bignot & Guyader, 1971 ti ga tions are nec es sary to solve the prob lem. Range. Tithonian–Berriasian. Globuligerina terquemi (Iovcheva & Trifonova 1961) Oc cur rence. Sec tions: KA, KB, KC, KD, KG, KJ, KK, KN, KO, Fig. 8C KR. 1961. Globigerina terquemi n.sp.: Iovcheva & Trifonova, p. Rumanoloculina verbizhizhiensis (Dulub, 1964) 344-345, pl. II, figs 9-14. Fig. 7I, J Re marks. Hori zon tal sections of this small species show char ac - ter istic loose ar range ment of cham bers of the last whorl, while the 1964. Quinqueloculina verbizhiensis n.sp.: Dulub, p. 108, pl. 1, ax ial sec tions re veal two whorls of cham bers ar ranged in a low figs 3, 4. spire. Forms mentioned by Kuznetsova (In: Kuznetsova & 1989. Quinqueloculina verbizhiensis Dulub: Matsieva & Temir- Uspenskaya, 1980) as Globuligerina exgrterquemi (Iovcheva & bekova, p. 115, 117. pl. 1, figs z, c, k. Trifonova) and later de scribed as Globuligerina parva n.sp. (In: Re marks. Transversal sec tion shows a quinqueloculine cham ber Kuznetsova & Gorbatchik, 1985) from the Early Kimmeridgian of ar range ment and oval out line of the test. Ax ial sec tions show three Cri mea prob a bly be long to the spe cies. sets of cham bers making the whole test. Range. Kimmeridgian–Tithonian Range. Kimmeridgian–Tithonian. Oc cur rence. Section KP. Oc cur rence. Sec tions: KA, KB, KE, KF, KG. CALCAR E O US DINOCYSTS Ge nus Scythiloculina Neagu, 1984 (sys tem atics af ter Øehánek & Cecca, 1993) Scythiloculina confusa Neagu, 1984. Fig. 7G Or der Peridiniales Haeckel, 1894 1984a. Scythiloculina confusa n.sp.: Neagu, pl. 1, figs 1-8, 16. Family Calciodinellaceae Deflandre, 1947 emend. Bujak & 1984b. Scythiloculina confusa Neagu: Neagu, p. 205, 206 pl. 4, figs Davies, 1983 10-37, text-fig.1. Ge nus Comittosphaera Øehánek, 1985 2005. Scythiloculina confusa Neagu: Olszewska, p. 38, pl. VI, figs 9, 10. Comittosphaera sublapidosa (Vogler, 1941) Fig. 8F Re marks. Transversal section show “Y” type of cham ber arrange - ment in nu merous whorls what makes outline of the test al most 1941. Cadosina sublapidosa n.sp.: Vogler, p. 280, pl. 2, fig. 5 cir cu lar. 1994. Comittosphaera sublapidosa (Vogler): Ivanova, p.99, 100, Range. Late Berriasian–Valanginian. pl 2, figs 9,10. Oc cur rence. Sec tions. KB, KN, KR. 2005. Comittosphaera sublapidosa (Vogler): Olszewska, p. 31, pl. 3, fig. 7 Suborder Rotaliina Delage & Herouard, 1896 Re marks. Spher ical cyst with a two lay ered wall. The inner layer Fam ily Discorbidae Ehrenberg, 1838 of variable thickness is com posed of the microcrystalline calcite. Ge nus Mohlerina Bucur, Senowbari-Daryan & Abate, The outer layer, vit re ous in trans mitted light is composed of the ir - 1996 reg u lar, fine cal cite crys tals. Range. Tithonian–Hauterivian. Mohlerina basiliensis (Mohler, 1938) Oc cur rence. Section KP. Fig. 8A 1938. Conicospirillina basiliensis n.sp.: Mohler, p. 27, pl. 27, 28; Ge nus Cadosina Wan ner, 1940 pl. 4, fig. 5. Cadosina parvula Nagy, 1966 1984. “Conicospirillina “ basiliensis Mohler: Bernier, p. 525-526, Fig. 8G pl. 21, fig. 3. 1991. “Conicospirillina” basiliensis Mohler: Altiner, pl. 3, figs 8, 1966. Cadosina parvula n.sp.: Nagy, p. 93, pl. 5, fig. 17 9. 1993. Cadosina parvula Nagy: Øehánek & Cecca, p. 155, pl. 1, 1996. Mohlerina basiliensis (Mohler): Bucur, Senowbari-Daryan fig. 12, text-fig. 6A. & Abate, p. 70-74, pl. 3, figs 3-6; pl. 4, figs 2, 3, 5-9. Re marks. Sphaerical cyst with a one lay ered wall composed of 1999. Mohlerina basiliensis (Mohler): Schlagintweit & Ebli, p. microcrystalline calcite. Differs from Cadosina fusca Wanner in 400, pl. 6, figs 1-2. smaller size and opti mal dis tribu tion in the Late Oxfordian– 2005. Mohlerina basiliensis (Mohler): Olszewska, p. 38, pl. 6, fig. Kimmeridgian. 1. Range. Late Oxfordian–Tithonian. Re marks. Diversely ori ented sec tions show typi cal for the spe cies Oc cur rence. Section KB 306 M. KRAJEWSKI & B. OLSZEWSKA

Fig. 8. A – Mohlerina basiliensis (Mohler) (KC 14a); B – Verneuilinoides polonicus (Cushman & Glazewski) (KD 3); C – Globuligerina terquemi (Iovcheva & Trifonova) (KP 5a); D, E – Protopeneroplis striata Weynschenk (KB 2a); F – Comittosphaera sublapidosa (Vogler) (KP 4a); G – Cadosina parvula Nagy (KB 14a); H – Crustocadosina semiradiata (Wanner) (KL 1) FORAMINIFERA FROM THE LATE JU RAS SIC AND EARLY CRE TA CEOUS, UKRAINE 307

Fig. 9. Strati graphic ranges of foraminifers from inves ti gated area pre sented in Figs 4–8. For lo cal iza tion see Fig. 3

Ge nus Crustocadosina Øehánek, 1985 mirabilis Weynschenk, Parurgonina caelinensis Cuvillier, Foury & Pignatti Morano and Neokilianina rahonensis Crustocadosina semiradiata (Wan ner, 1940) (Foury & Vincent ) are know predom inantl y from the Kim- Fig. 8H meridgian of the Medi ter ranean Tethys (Bassoulet, 1997). 1940. Cadosina semiradiata n.sp.: Wan ner, p. 81, figs 36, 37. In the same area spe cies Anchispirocyclina lusitanica 1994. Crustocadosina semiradiata (Wanner): Ivanova, p. 89, 90 (Egger) charac ter ises the Tithonian strata (Bassoulet, 1997; pl. I, figs 8, 9. Darga & Schlagintweit, 1991). In the Centra l and NW Cri- 2005. Crustocadosina semiradiata (Wanner): Olszewska, p. 33, mea Anchispirocyclina lusitanica (Egger) is pres ent in both pl. 2, fig. 1. Tithonian and Berriasian strata (Voloshina, 1977; Gorba- Re marks. Spheri cal to oval cyst with two lay ered walls. The in ner tchik & Mohamad, 1997; Zhabina, 1989). Inter est ing is the dark, microgranular layer has thick ness equal to larger than the per sis tent pres ence in the mate rial stud ied the long last ing thick ness of the outer, white, ra dial layer. Range. Late Oxfordian–Early Albian. (Li assic–Berriasian) Amijella amiji (Henson) com mon in Oc cur rence. Section KL. Tithonian strata of the Alpino-Cri mean segm ent of the Tethys (Voloshina, 1977; Schlagintweit, 1991; authors ob- serva ti ons). The specie s also consti tutes an index taxon for the lower Berriasian “beds with Bramkampella” repor ted by RE MARKS ON STRA TIGRA PHY Gorbatchik & Mohamad (1997) from the Crim ea. In the upper part of Tithonian makes its first appear ance Foraminiferal as semblages from the Aj-Petri and Jalta spe cies Protopeneroplis ultragranulata (Gorbatchik) fre- Yaila con tain many spe cies of small and large foraminifera quently used as an index taxon for the Early Berriasian of of the recog nise d strati graphi cal value for Juras sic carbon - the northern margin of the Tethys (Azema et al., 1977; ate sedi m ents (Fig. 9). Among the large forms Labirynthina Bassoulet & Fourcade, 1979; Kuznetsova & Gorbatchik, 308 M. KRAJEWSKI & B. OLSZEWSKA

1985; Sotak in Vašièek et al., 1994; Gorbatchik & Moha- tute of RAS, Mos cow), V. V. Yu din (Ukr. Min. Re sources State mad, 1997). The Early Cre taceous age of the topm ost part of Inst. Simferopol) and J. Ma tysz kiewicz (AGH, Kraków) for help the inves ti gated profil es is also suggest ed by the apearance in litera ture collec tion and discus sions . This research was financed of such spe cies as Everticyclammina kelleri (Henson), by the AGH grants no. 10.10.140.463 and 11.11.140.560. Nautiloculina bronnimanni Arnaud-Vanneau & PeybernÀs, Montsalevia salevensis (Charollais, Brönnimann & Zani- netti) or Scythiolina camposaurii (Sartoni & Crescenti), and REF ER ENCES Mayncina bulgarica Laug, PeybernÀs & Rey. Pa leoen vi ron metal, rather than stra tigraphic, sig nifi- Al tiner, D., 1991. Mi cro fos sil bi os tra tigra phy (Mainly Fo ra mini- cance have the oc cur rence of abun dant “tro cho linas” and fera) of the Ju ras sic–Lower Cre ta ceous car bon ate suc ces sions miliolids in Ti tho nian part of the Aj- Petri car bon ates and in North- Western Ana to lia (Tur key). Ge olo gia Ro mana, 27: Jalta Yaila. Devel op m ent of both groups (known also from 167–213. the Alpino-Carpa thian realm and Moe sian Platform ) may be Arnaud-Vanneau, A., 1986. Vari a tions dans la com po sition et attri buted to sea sonal variati ons of sea level dur ing the dans la diversité des faunes de ForaminifÀres benthiques du stage. Crétacé inférieur sur quelques plate-formes carbonatées To sum up the rem arks one may conclude that stra - téthysiennes de l’Europe et du moyen-Ori ent. Bul le tin de la tigraphic ranges of char ac ter is tic spe cies of fora mini fera Société Géologique de France (8), II: 245–253. (vide Fig. 9) identi fied in the inves ti gated sam ples suggest Arnaud-Vanneau, A., Boisseau, T. & Darsac, C., 1988. Le genre Trocholina Paalzow, 1922 et ses principales espÀces au the Kimme rid gian to Ber ria sian age for the Aj- Petri and Crétacé. Re vue de Paléobiologie, Vol ume Spécial 2: 353– Jalta Yaila car bon ates. 377. Cor re la tion of the thin- plate as sem blages ob tained from Arnaud-Vanneau, A. & Masse, J. P., 1989. Les ForaminfÀres the indu rat ed carbon ate s with those from the water proc- benthiques des For mations carbonatées de l’Hauterivien– essed soft sedim ents of the same region (vide Kuznetsova & Barrémien pro parte du Jura Vaudois et Neuchâtelois Gorbatc hik, 1985) is somewhat diffi cult . The latter do not (Suisse). MÀmoires de la Société neuchâteloise des Sciences re flect nei ther spa tial nor tem po ral origi nal dis tri bu tion of naturelles, 11: 257–276. taxons in the rock. They also refle ct differ ent sedim entar y Arnaud-Vanneau, A. & PeybernÀs, B., 1978. Les représentants re gime. éocretacés du genre Nautiloculina Mohler, 1938 (Forami- nifera, Fam. Lituolidae ?) dans les chaines subalpines septen- trionales (Vercors) et les Pyrénées franco-espgnoles. Géo- bios, 11: 67–81. REMARKS ON PALEOENVIRONMENT Azéma, J., Chab rier, G., Fourcade, E. & Ja frezzo, M., 1977. Nou - AND PALEOBIOGEOGRAPHY velles données mi cro paléon tolo giques, stra tigraphiques et OF FORAMINIFERA paléogéo graphiques sur le Port lan dien et le Néo comien de Sar daigne. Re vue de Mi cro paléon tolo gie, 20: 125–139. Flügel in his funda m ental work (Flügel, 2004, p. 660) Ban ner, F. T. & High ton, J., 1990. On Ever ti cy clam mina Red - states that “car bon ate platfor ms are dy namic system s that mond (Fo ra mini fera), es pe cially E. kel leri (Hen son). Jour nal change thorough time and space”. The right ness of the state- of Mi cro pa le on tol ogy, 9: 1–14. ment is confirm ed also by changes in foraminiferal assem - Bar nard, T., Cor dey, W. G. & Shipp, D. J., 1981. Fo ra minifera blages of the inves ti gated area. The Kimmeridgian– Titho- from the Oxford Clay (Cal lovian–Ox for dian of Eng land). Re - vista Es pañola de Mi cro palen tología, 13: 383–462. nian assem blages are predom inantl y made of the inter nal Barwicz- Piskorz, W., 1995. Ze spo³y ot wornic i straty grafia gór nej plat form genera such as Pseudocyclammina, Everticycla- jury obr ze¿enia Gór Œwiêtokrzyskich. (Eng lish summary ). mmina, Rectocyclammina, Parurgonina, Anchispirocy- AGH Rozprawy-Monografie , 21: 1–120. clina, Amijella or Neokilianina and Miliolidae (Septfon- Bassoulet, J. P., 1997. Les Grandes ForaminifÀres. In: Cariou, E. taine, 1980; Pélissié, PeybernÀs & Rey, 1984). The Early & Hantzpergue, P. (eds.), Biostratigraphie du Jurassique Cre ta ceous as sem blages con tain more outer plat form el e- Ouest-Européen et Mediterranén. ELF Ep Mémoire 17: 293– ments such as “trocholinas”, and genera: Mohlerina, Proto- 304. peneroplis, Charentia, Montsalevia (Chioccini et al., 1988). Bassoulet, J. P. & Fourcade, E., 1979. Essai de synthÀse de Known paleo geo graphi c occur rence s of many of Aj- répartition de ForaminifÀres benthiques du Jurassique Petri and Jalta Yaila foraminifera in di cates that they be long carbonaté mésogéen. Compte Rendu Sommaire des Séances to cos mo poli tan forms con nected pre domi natly with the de la Société Géologique de France, 2: 69–71. north Te thyan shelves during the end of Juras sic and the Bassoulet, J. P., Fourcade, E. & PeybernÀs, B., 1985. Paléo- biogéographie des grands ForaminifÀres benthiques des early Cre ta ceous (Pélis sié et al., 1982; Bassou let et al., marges néo-téthysiennes au Jurassique et au Crétacé infé- 1985; Arnaud-Vannea u, 1986). rieur. Bul le tin de la Société Géologique de France (8), I: 699–713. Ac knowl edge ments Ber nier, P., 1984. Les for mations car bonatées du Kim méridg ien et du Portlan dien dans le Jura Mé ridi onal. Docu ments des Labo- The authors thank the re view ers J. Smoleñ (PIG Warsaw) and ratoires de Géolo gie Lyon, 92: 445–803. J. Tyszka (ING PAN Kraków) for theirs sug ges tions; B. M. Rom - Bielecka, W., 1980. Rz¹d Fo ra miniferida Eichwald. (In Pol ish). anyuk (Lviv Na tional Uni ver sity), M.A. Ro gov (Geo logi cal In sti - In: Ma li nowska, L. (ed.), Bu dowa geo logic zna Pol ski. At las FORAMINIFERA FROM THE LATE JU RAS SIC AND EARLY CRE TA CEOUS, UKRAINE 309

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