Egypt. J. Prot. Res. Inst. (2021), 4 (1): 64 –83

Egyptian Journal of Plant Protection Research Institute www.ejppri.eg.net

Revision of the thysanopterous (: ) in Egypt Maaly, E. Wafy; Iman, I. El- Sebaey and Mahmoud, Y. H. Henaish Plant Protection Research Institute, Agricultural Research Center, Dokki, Giza, Egypt. ARTICLE INFO Abstract: Article History The updated checklist of Thysanoptera, Received: 5 / 1 /2021 Thripdiae are given. More than 70 and 37 genera are Accepted: 22 / 3 /2021 recorded in Egypt belonging to four subfamilies (Panchaetothripinae, , Dendrothripinae and Keywords Sericothripinae). Synonyms, geographical distribution and host are given. Thysanoptera, Thripdiae, checklist, host plants, geographical distribution and Egypt.

Introduction (Family and Subfamily (Thysanoptera: Idolothripinae). Other species are Thripidae) is widespread around the phytophagous inhabiting flowers and world especially in tropical region. leaves; they attack many parts of plant About 6246 species in 782 genera as buds, leaves, flowers, fruits and stem. were identified, derived from two They cause damage by sucking the suborders: the Terebrantia, whereas are contents of plant cell resulting leaf and classified into eight families: flower defoliation; fruit scarring in , , some fruit trees and often reduce its , , market value, also fruits become , Merothripidae, smaller than normal ones. Thripidae and Uzelothripidae and the Some species of Family tubulifera includes a single family, cause rolled leaf galls Phlaeothripidae. Thrips are small or in some oriental trees. Furthermore, minute slender-bodied (0.5- some species are known as tospoviruses 15mm). Antenna are short, 6-10 (Family Bunyaviridae) transmitted as segments. Body color varies from pale fusca Hinds, F. to dark black according weather occidentalis (Pergande), F. schultzei condition. Piercing mouthparts is (Trybom) Lindeman, T. asymmetrical (Right mandible absent) palmi Karny and dorsalis (Priesner, 1960; Richard and Davies, Hood. On the other hand, some species 1977 and Gullan and Cranston, 1994). are predators that feed on other thrips Most species are mycophagous species or other small as (More than 50%), feed on fungi existing scale insects and mites (Family in leaf litter or beneath the bark of trees Aeolothripidae; Scolothrips Family

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Thripidae and Karnyothrips Watson -Female ovipositor turned upwards Family Phlaeothripidae) (Priesner, toward body; antennae nine segments, 1960; Lewis, 1973 and Mound and segments III and IV usually with liner Kibby, 1998). sensoria Aeolothripidae (Figure2h, The aim of the present research 2i) work is to study the revision of the -Female ovipositor turned upwards thysanopterous in Egypt. toward body; antennal nine segments, Results and discussion clearly separate and with transverse Figure (1) showed that more sensoria. Melanthripidae (Figure 2j) than 150 species are described whereas The family Thripidae presently derived from five families in Egypt. includes about more than 2100 species, Key of Thysanoptera this is the second largest family of 1.Last abdominal segment usually Thysanoptera. Most genera of the conical; female with saw-like suborder Terebrantia are in this family. ovipositor; major anal setae arising In Egypt, this family includes 37genera from sub-apical region of last segment; and 77 species. It contains of four forewings with longitudinal or across subfamilies as follow: veins and setae. …………………….. Key to subfamilies of Egyptian Suborder Terebrantia(Figure 2a, 2b) Thripidae -Last abdominal segment tube-like; -Head and pronotum reticulated; female without external ovipositor; antennal segments III and IV without major anal setae arising from platelets sense cone; terminal segment long and attached to end of tube; forewings slender Panchaetothripinae (Figure without veins or setae . Suborder 3a,3b) Tubulifera (Figure 2c, 2d) -Head and pronotum not reticulated; Key to families of Egyptian antennal segments III and IV with sense Terebrantia cone; terminal segment rarely finely -Female with ovipositor weakly elongate…...Thripinae(Figure 3c,3d) developed; antennal eight-segments, -Metanotumfurca with lyre-shaped, segments III and IV with liner, forked extended to mesonotum; mesonotum or simple sensoria without spinula; fore wing curved at cone…Merothripidae(Figure 2c, 2d) apex join to posterior margin………… -Female ovipositor turned downwards Dendrothripinae(Figure 3e,3f) away from body; antennae usually -Metanotum without furca; mesonotum seven or eight segments, rarely six or with spinula; fore wing wide at apex nine segment, segments III and IV and not join to posterior developed into slender forked or simple margi…………….Sericothripinae sense cone .. Thripidae(Figure 2f, 2g) (Figure 3g,3h)

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Egypt. J. Plant Prot. Res. Inst. (2021), 4 (1): 64 –83

Order :Thysanoptera

Suborder: Terebrantia Suborder: Tubulifera Phlaeothripidae

Merothripidae (52 species) (1genus) Idolothripinae (3 species)

Melanthripidae (10 species) Phlaeothripinae (49 species) Aeolothripidae (13 species)

Thripidae

Panchaetothripinae (6 species)

Thripinae (65 species)

Dendrothripinae

(2 species)

Sericothripinae (4 species)

Figure (1): Families of Thysanoptera in Egypt.

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a b c

d e

f g h

i j

Figure (2): a, b Terebratia a (Last segmented), b (Forewing); c, d Tubulifera c (Last segmented), d (Forewing); e Merothripida (Last segmented); f, g Thripidae f (Ovipositor), g (Antennae); h, i Aeolothripidae h (Ovipositor), i (Antennae) and j Melanthripidae (Antennae).

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a b c d

e f

g h

Figure (3): a, b Panchaetothripinae a (Head and pronotum), b (Antennae); c, d Thripinae c (Head and pronotum), d (Antennae); e, f Dendrothripinae e (Meso and metanotum), f (Forewing); g, h Sericothripinae g (Meso and metanotum) and h (Forewing).

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1. Subfamily: Panchaetothripinae Common name: Cotton leaf thrips The species belonged to Distribution: Ethiopia, India, Kenya, Panchaetothripiae seem to be associated Nigeria, South Africa, Sudan, and with leaf; some species feed on grasses Zimbabwe (Moritz et al., 2013). and few causes damage to crop Host plant: Glycine max (Abd El- seedling. This subfamily consists of Wahab, 2016). forty-two genera and one-hundred 1.2. Heliothrips Haliday forty-five species around the world. In Heliothrips, Haliday, 1836, Ent. Mag., Egypt, it represents by four genera and 443. seven species. This attacks wide range 1.1. Genus: Caliothrips Daniel of hosts specially, the hard leaves one. Caliothrips Daniel, 1904, Ent. News, Five species belong to this genus, on the 296. other hand, one species only valid in The species belongs to this Egypt. genus are young leaf-feeder, several of 1.2.1. Heliothrips haemorrhoidalis them are pests of many crops, and few Bouche, 1833 species attacks grasses. This genus is Thrips haemorrhoidalis, Bouche, 1833, widespread in all countries, especially Naturg. Schadl. Garten-Ins., 206. tropical ones and many different plants. Heliothrips haemorrhoidalis, Priesner, It consists of twenty- three species in 1926, Thys. Eur., 126. the world, while in Egypt it represented Heliothrips adonidum Haliday, 1836. by three species. Ent. Mag., 3:443. 1.1.1. Caliothrips deserticola Priesner Common name: Greenhouse thrips, Caliothrips deserticola Priesner, 1960, Black greenhouse thrips and Black tea Publ. Inst. Desert Egypte 13: 226. thrips. Distribution: Sudan (Moritz et al., Distribution: Widely distributed 2013). (Mirab-balou, 2013). Host plants: Scrub on the rocky slopes Host plants: Acalypha Sp., Camellia (Priesner, 1960). Sp., Citrus Sp., Croton Sp., Laurus 1.1.2. Caliothrips graminicola Bagnall nobilis, Mangifera indica, Prunus and Cameron persica, Pyrus malus and Vitis vinifera Hercothrips graminicola Bagnall and (Priesner, 1960 and El-Wakkad, 2007). Cameron. 1932. Ann. Mag. Nat. Hist. 1.3. Hercinothrips Bagnall (10) 10: 415. Hercinothrips Bagnall, 1932. Ann. Caliothrips graminicola Priesner, 1960. Mag. nat. Hist., 10 (10): 506. Publ. Inst. Desert Egypte, 13: 227. Hercinothrips has been known Distribution: Australia, Iran, India, as a pest of many host plant. The origin Nepal, Sri Lanka, South Africa, Sudan, distribution is Africa. This genus Thailand, Yemen and Zimbabwe consists of nine species widespread. (Kudo, 1995 and Mirab-balou, 2013). Hercinothrips femoralis is the only Host plant: Glycine max (Abd El- species recorded in Egypt. Wahab, 2016). 1.3.1. Hercinothrips femoralis Reuter 1.1.3. Caliothrips sudanensis Heliothrips femoralis Reuter, 1891. (Bagnall) Medd. Soc. Fauna F1. Fenn., 17:166. Hercothrips sudanensis Bagnall and Heliothrips cestsi Pergande, 1895, Cameron, 1932, Ann. Mag. rlat. Hist. U.S.D.A., Life, 7 (5):390. (10) 10: 415. Hercinothrips femoralis, Bagnall. CaIiothrips sudanensis (Bagnall and 1932. Ann. Mag. Nat. Hist., Ser. 10, Cameron; Priesner, 1949, Bull. Soc. 10(59):506. Fouad I Ent. 33: 132.

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Common name: Banded greenhouse This subfamily includes sixteen genera thrips and sugar-beet thrips. and about one-hundred described Distribution: Widespread in the tropics species in the world. In Egypt it and subtropics; common in temperate represents by two genera and two areas in greenhouses, Denmark, species. Finland, Florida, Georgia, Iceland, 2.1. Dendrothrips Uzel Kenya, Nigeria, Norway, Sierra Leone, Dendrothrips, Uzel, 1895, Mon. Ord. Sweden, Tanzania and Uganda (Diffie Thys. 159. et al., 2008; Kobro, 2011 and Moritz et Several species of Dendrothrips al ., 2013). are particularly associated with the Host plant: Citrus sp., 2007). family Oleaceae and herbs. This genus 1.4. Retithrips Marchal now includes fifty-six species, and D. Retithrips, Marchal, 1019, Bull. Soc. R. eremicola is the only species that Ent. D`Egypte, 17. recorded in Egypt. One widespread species 2.1.1. Dendrothrips eremicola (Included Egypt) and the second one Priesner from Indonesia and Northern Australia. Dendrothrips eremicola Priesner, 1960, 1.4.1. Retithrips syriacus Mayet Publ. Inst. Desert Egypt 13: 281. Heliothrips syriacus Mayet, 1890, Host plants: Wild shrubs and Olea Insectes de la vigne, 451. europapea (Priesner, 1960 and Wafy, Retithrips aegytiacus Marchal, 1910, 2018). Bull. Soc. R. Ent. Egypte, 17-21. Recently, in 2018 this species Dictyothrips aegytiaca Zanon, 1917- occurred as an outbreak on olive trees at 18, L`Agric. Colon., Firenze, XI-XII. Ismailia Governorate (Wafy, 2018). Dictyothrips zanoniana Del Guercio, 2.2. Pseudodendrothrips Priesner Note Osservat., Ent. Agr. Firenze, 106- Pseudodendrothrips Priesner, 1960, 119. Publ. Inst. Desert Egypt 13:283. Common name: Castor thrips and Pseudodendrothrips is Black vine thrips commonly associated with Moraceae, Distribution: America, Brazil, Congo, and live on Poaceae, it lives on leaves Europe, Ghana, India, Iran, Kenya, of plants. There are now twenty-one Lebanon, Libya, Malawi, Morocco, species listed in this genus and Mozambique, Nigeria, Palestine, represents in Egypt by one species. Puerto Rico, Senegal, Somalia, South 2.2.1. Pseudodendrothrips aegyptiacus Africa, Sudan, Syria, Tanzania, Togo, Priesner Tunisia, Uganda and U.S.A . (Mirab- Pseudodendrothrips aegyptiacus balou, 2013 and Moritz et al., 2013). Priesner, 1960, Publ. Inst. Desert Host plants: Acalypha Sp., Gossypium Egypte 13:284. Sp., Quercus Sp., Eucalyptus indica; Distribution: Abu Dhabi, Canary Juglans regia, Lawsonia inermis, Islands, Israel and Southern Africa Mangifera indica, Myrtus communis, (Moritz et al., 2013). Psidium guajava, Pyrus cydonia, Host plants: Convolvulus arvensis, communis, Rosa centifolia, Euphorbia cuneata and Lycium Schinus molle, Sc. terebinthifolius, arabicum (Priesner, 1960). Terminalia arjuna and Vitis vinifera 3. Subfamily: Sericothripinae (Priesner, 1960 and El-Wakkad, 2007). Flower and leaf-feeding thrips. 2. Subfamily: Dendrothripinae The subfamily Sericothripinae is a This subfamily comprises a largely tropical group of about three group of leaf- feeding thrips (Live and genera and one-hundred, sixty-eight breed on green leaves not in flowers). species. On the other hand, there are

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Wafy et al. , 2021 two genera and five species belong to 4.1. Uzel this subfamily in Egypt. Anaphothrips, Uzel, 1895, Mon, Ord. 3.1. Sericothrips Haliday Thy. 142. Haliday, 1836, Ent. Mag., Many species are grass-living. 444. Eighty-six species are belonged to The genus Sericothrips includes Anaphothrips. Only seven species are about seven species worldwide. In recorded In Egypt. It considered as Egypt, this genus represents by three minor pests of cereals. species. 4.1.1. Anaphothrips alternans Bagnall 3.1.1. Sericothrips arenarius Priesner Anaphothrips alternans Bagnall, 1913, Sericothrips arenarius Priesner, 1960, Ann. Mag. Nat. Hist. (8), 12: 291. Publ. Inst. Desert Egypte 13: 244. Common name: Corn thrips and Host plant: herbs (Priesner, 1960). sugarcane thrips. 3.1.2. Sericothrips kassimianus Distribution: Central America and Priesner (Priesner, 1960). Sericothrips kassimianus Priesner, Host plant: Scirpus Sp.; Scirpus 1950, Bull. Soc. Fouad I Ent., 25. pungens; Panicum turgidum; Hordeum Hydatothrips kassimanus .Bhatti, 1973. vulgare; Saccharum officinarum; Oriental Insects. 7 (3): 403-449. Sorghum halepense; Cynodon Common name: Blood flower thrips. dactylon; Polypogon monspeliensis Distribution: India (Bhatti, 1973). (Priesner, 1960). Host plant: Cynanchum acutum 4.1.2. Anaphothrips crocatus Priesner (Priesner, 1960 and Wafy, 2018) Anaphothrips crocatus Priesner, 1938, 3.1.3. Sericothrips masrensis Priesner Bull. Soc. Fouad I Ent., 131. Sericothrips masrensis Priesner, 1960, Host plant: Zygophyllum simplex Publ. Inst. Desert Egypte 13: 245. (Priesner, 1960). Host plant: unknown (Priesner, 1960). 4.1.3. Anaphothrips hieroglyphicus 3.2. John Priesner Neohydatothrips John, 1929. Zootaxa. Anaphothrips hieroglyphicus Priesner, 1575: 47–68. 1939, Bull. Soc. Fouad I Ent., 352. This genus now includes almost Common name: Western desert thrips. one-hundred and eighteen species. Host plant: Desert shrubs (Priesner, 3.2.1. Neohydatothrips samayunkur 1960). Priesner 4.1.4. Anaphothrips obscurus Muller Hydatothrips (Neohydatothrips) Thrips obscurus Muller, 1776, O. F., samayunkur Kudo, 1995. Zootaxa. Zool. Dan Prodrom., 96. 1575: 62. Anaphothrips obscurus Uzel, 1895, Common name: Marigold thrips Mon. Ord. Thys. 279. Distribution: Africa, Asia, Australia, Anaphothrips striatus Hinds, 1902, Central and South America, China, Mon. Thys. N. Amer., 161. India, Kenya, New Zealand, North Common name: American grass thrips America and Uganda (Mirab-balou et and Grain thrips. al., 2011 and Rachana and Varatharajan Distribution: Widely distributed, 2017 ). Algeria, Australia, Canary Islands, Host plant: Tagetes erecta (Abd El- China, Denmark, Finland, Iceland, Iran, Wahab, 2015). Hawaii, North America, Norway, 4. Thripinae Morocco, Sweden, and Turkey (Kobro, Thripinae represents In Egypt 2011; Mirab-balou et al., 2011; Tunc et by twenty-nine genus and sixty-four al., 2012 and Mirab-balou, 2013). species as follow:

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Host plants: Gramineae, Citrus sp., This species has been known as Mangifera indica, Polypogon a pest of grasses, cereal and sugar cane monspeliensis, Psidium guajava, Pyrus and distributes in tropics regions. malus and Vitis vinifera (Priesner, 1960 4.2. Haliday and El-Wakkad, 2007). Aptinothrips Haliday, 1836, Ent. Mag., It feeds on the leaves of grasses 3: 445. and cereals rather than the Aptinothrips represented inflorescences. It is common in through the world by five species while temperate area. in Egypt, only one species valid. Europe 4.1.5. Anaphothrips parviceps being the center of distribution. They Priesner live in grasses and have some degree of Anaphothrips parviceps Priesner, 1960, host specificity. Publ. Inst. Desert Egypte 13: 265. 4.2.1. Aptinothrips rufus (Gmelin) Host plant: Composite (Priesner, Aptinothrips rufus Gmelin, 1788, Linne 1960). Syst. Nat. 2224. 4.1.6. Anaphothrips retamae Priesner Thrips rufa Gmelin, 1788. Carolia Anaphothrips retamae Priesner, 1934, Linn; Syst. Nat., 2224. Bull. Soc. R. Ent. Egypte, p.275. Common name: Grass thrips. Host plant: Retama raetam (Priesner, Distribution: Algeria, China, Costa 1960). Rica, Croatia, Denmark, Ethiopia, 4.1.7. Anaphothrips sudanensis Europe, Finland, Iceland, Iran, Italy, Trybom Norway, South North Africa, Sweden, Anaphothrips sudanensisTrybom, Tunisia and Turkey (Raspudici et al., 1911, Bull. Bri. Mus. Ent. 9, II: 21. 2009; Kobro, 2011; Tunc et al., 2012; Euthrips flavicinctus Karny, 1912, Bull. Mirab-balou, 2013 and Marullo and De Bri. Mus. Ent. 9, II: 21. Grazia, 2013). Euthrips (Anaphothrips) alternans Host plant: Cynodon dactylon Bagnall, 1913, Bull. Bri. Mus. Ent. 9, II: (Priesner, 1960) 21. 4.3. Ascirtothrips Priesner Euthripscitricinctus Bagnall, 1919, Ascirtothrips Priesner, 1960, Publ. Inst. Bull. Bri. Mus. Ent. 9, II: 21. Desert Egypte 13: 252. Euthrips bicolor Morgan, 1925, Bull. This genus comprises two Bri. Mus. Ent. 9, II: 21. species in Egypt. Euthrips transvaalensis Faure, 1925, 4.3.1. Ascirtothrips antilope Priesner Bull. Bri. Mus. Ent. 9, II: 21. Anaphothrips antilope Priesner,1923, Common name: Wheat thrips and Ent. Mitteil., XII: 63. Maize thrips. Scirtothrips antilope Priesner, 1932, Distribution: In subtropical and Bull. Soc. R. Ent. Egypte, 143, 152. tropical regions around the world, Ascirtothrips antilope Priesner, 1957, China, Ethiopia, India, Kenya, Libya, Zool. Anz., 159: 7-8. Mozambique, Morocco, Nigeria, Spain, Distribution: Cyprus, India, Israel and Senegal, Sri Lanka, South Africa, Morocco (Moritz et al., 2013). Sudan U. S. A. and Zimbabwe (Reitz et Host plants: Aerva tomentosa, Alhagi al., 2011; Tillekaratne et al., 2011; maurorum, Iphiona mucronata, Goldarazena et al., 2012; Moritz et al., Nitraria retusa, Pulicaria crispa and 2013 and Rachana and Varatharajan, Zilla Spinosa (Priesner, 1960). 2017). 4.3.2. Ascirtothrips efflatouni Host plant: Glycine max (Abd El- Priesner Wahab, 2016). Ascirtothrips efflatouni Priesner, 1960, Publ. Inst. Desert Egypte 13: 253.

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Host plant: Halocnemon strobilaceum Yemen, and Uzbekistan (Mirab-balou, (Priesner, 1960). 2013). 4.4. Uzel Host plant: Gramineae; Cyperaceae; Baliothrips Uzel, 1895, Mon. Ord. Citrus Sp.; Cyperus Sp.; Cynodon Thys. 204. dactylon; Cladium mariscus; Imperata Three species have been cylindrical; Eragrostis bipinnata; recorded in this genus. On the other Panicum turgidum; Mangifera indica; hand, one species only is valid in Egypt. Psidium guajava; Pyrus malus and Vitis 4.4.1. Baliothrips vittipennis Bagnall vinifera (Priesner, 1960 and El- Baliothrips vittipennis Bagnall, 1927, Wakkad, 2007). Ann. Mag. Nat. Hist. (9), 20: 574. 4.6.2. meridionalis Common name: Mat sedge thrips. Bagnall Distribution: Australia, and Chirothrips meridionalis Bagnall, Poland (Priesner, 1960). 1927, Ann. Mag. Nat. Hist. (9), XIX: Host plant: Cyperus Sp (Priesner, 566. 1960). Common name: Broomcorn thrips. 4.5. Bolacidothrips Priesner Distribution: Algeria,Cyprus, France, Bolacidothrips Priesner, 1930, Bull. India, Iran, Italy, North and South Soc. R. Ent. Egypte, 6. Africa Palestine, Morocco Nigeria and Old World genus, it lives on Yemen (Minaei, 2013 and Moritz et al Poaceae. The genus consists of twelve ., 2013). species, the only species B. graminis Host plants: Andropogon halepensis, has been recorded in Egypt. Polypogon monspeliensis and Zea 4.5.1. Bolacidothrips graminis maiyz (Priesner, 1960). Priesner 4.6.3. Chirothrips mexicanus Bolacidothrips graminis Priesner, Crawford 1930, Bull. Soc. R. Ent. Egypte, 6. Chirothrips mexicanus Crawford, 1909, Common name: beard grass thrips. Pomona College Jour Ent., I, 114. Host plants: Scirpus sp., Triticum sp., Chirothrips floridensis Watson, 1920, Cynodon dactylon, Imperata Fla. Ent., IV (2) :21. cylindrical, Phragmites communis, Distribution: Hawaii, India, Mexico, Panicum colonum, Polypogon Philippines and South America (Moritz monspeliensis, Saccharum officinarum, et al., 2013) Sorghum vulgare and Zea maiyz Host plant: Glycine max (Abd El- (Priesner, 1960). Wahab, 2016). 4.6. Chirothrips Haliday 4.6.4. Chirothrips taxanus Andre Chirothrips, Haliday, 1836, Ent. Mag., Chirothrips taxanus Andre, 1939, Proc. 444. Ent. Soc. Wash., XLI (6): 200-202. Widespread genus across old Distribution: Brazil, Georgia, Mexico world, it lives in f lowers of Poaceae. and Texas (Diffie et al., 2008 and Forty-two species spread around the Monteiro, 2001). world. Four species recorded from Host plant: Cucumis sativus (Abd El- Egypt. It is common in the Holarctic, Wahab, 2012). Neotropical, and Ethiopian regions. 4.7. Dorcadothrips Priesner 4.6.1. Chirothrips africanus Priesner Dorcadothrips Priesner, 1932, Bull. Chirothrips africanus Priesner, 1932, Soc. R. Ent. Egypte, 49. Bull. Soc. R. Ent. Egypte, 46. 4.7.1. Dorcadothrips caespitis Common name: African thrips. Priesner Distribution: Algeria, China, Cyprus, Dorcadothrips caespitis Priesner, 1932, Ethiopia, Iran, India, Italy, Sudan, Bull. Soc. R. Ent. Egypte, 50.

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Common name: Cat`s tail thrips. Frankliniella sulphurea Karny, 1926, Host plants: Cyperus sp., Typha sp. Mem, Dept. Agric. Ind., Ent. Ser., 9: and Panicum colonum (Priesner, 1960). 195. Distribution: Sudan (Priesner, 1960). Common name: African borage thrips 4.8. Eremiothrips Priesner and Lantana thrips. Eremiothrips Priesner, 1950, Bull. Soc. Distribution: India, Italy, Palestine and Fouad I Ent., 29. Sudan (Priesner, 1960). This genus contains of twenty- Host plants: Alhagi maurorum, one species and represents in Egypt by Arnebia hispidissima, Inula one species. crithmoides, Iphiona mucronata, 4.8.1. Eremiothrips imitator Priesner Pulicaria crispa, Trichodesma Eremiothrips imitator Priesner, 1950, africanum, Triumfetta flavescens and Bull. Soc. Fouad I Ent., 29. Withania somnifera (Priesner, 1960). Host plant: Haloxylon schweinfurthii 4.10.2. Frankliniella fusca Karny (Priesner, 1960). Euthrips fuscus Hinds 1902. Zool. Stud. 4.9. Euphysothrips Bagnall 49 (6): 826 Euphysothrips Bagnall, 1926. Ann. Frankliniella fusca Karny 1912. Zool. Mag. Nat. Hist. (9), 18: 646. Stud. 49 (6): 826 Two species only have been Common name: Tobacco thrips. known in this genus. Distribution: Brazil, China, Florida, 4.9.1. Euphysothrips minozzii Bagnall Georgia, Guadeloupe, Martinique, Euphysothrips minozzii Bagnall, 1926. Spain and U. S. A (Diffie et al., 2008; Ann. Mag. Nat. Hist. (9), 18: 646. Monteiro, 2001; Goldarazena et al., Common name: Sea rush thrips. 2012; Reitz et al., 2011 and Etienne et Distribution: Austria, Canary Islands, al., 2015). India, Iran, Iseral, Mozambique, Host plant: Glycine max (Abd El- Southern Africa, Southern France, Wahab, 2016). Turkey, and Yemen (Zur Strassen and Kuslitzky, 2012). 4.10.3. Frankliniella occidentalis Host plants: Gramineae and Juncus Pergande acutus (Priesner, 1960). 4.10 . Frankliniella Karny Euthrips occidentalis Pergande. 1895. Frankliniella Karny, 1910, Mitt. Nat. U.S.D.A., Div. Ent. Insect Life, 7 Ver. Univ. Wien, 46. (5):392. Most species live in neotropics Euthrips tritici califomicus Moulton, area. Some species as F. fusca, F. 1911. U.S.D.A., Bur. Ent., Tech. Ser., occidentalis and F. schultzei have been 21: 28. known as virus transmitted. The Frankliniella occidentalis (Pergaade), members of this genus are flower Kamy, 1912. Zool. Ann., 4:335. feeding. Two-hundred and thirty-four Fmnkliniella tritici Moultoni, Hood. species are listed from this genus. On 1914. Proc. Ent. SOC. Wash., 16:38. the other hand, only five species Frankliniella californica Moulton, represent in Egypt. 1948. Rev. de Ent., 19(1-2):98. 4.10.1. Frankliniella dampfi Priesner Common name: , Frankliniella pallid Karny (nec Uzel), Alfalfa thrips. 1922, Denkschr. Akad. Wiss. Wien, 98: Distribution: Widespread around the 114. world (Mirab-balou, 2013). Frankliniella dampfi Priesner, 1923, Host plants: Citrus sp., Antirhinum Ent. Mitteil., 12: 64. majus, Begonia tuberhybrida, Capsicum annuum, Celosia argentea,

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Chenopodium amaranticolor, Thrips tritici Fitch 1948. Bull. Bri. Chrysanthemum morifolium, Coleus Mus. Ent. 9, II: 39. hybridus, Convolvulus arvensis, Common name: Eastern flower thrips. Crinum moorei, Cucumis sativus, Host plant: Glycine max (Abd El- Euphorbia seguieriana, Gazania Wahab, 2016). rigens, grandiflora, Glycine 4.11. Isoneurothrips Bagnall max, Gomphrena globosa, Isoneurothrips Bagnall 1915, Ann. Hippeastrum hybrid, H. vittatum, Mag. Nat. Hist. (8), 15: 592. Ipomoea tricolor, Kalanchoe 4.11.1. Isoneurothrips australis blossfeldiana, Lantana camara , Malus Bagnall pumila, Mangifera indica, Olea Isoneurothrips australis Bagnall 1915, europeana, Pelargonium hortorum, Ann. Mag. Nat. Hist. (8), 15: 592. Petunia hybrid, Psidium guajava, Thrips mediolinus Girault 1926, Ins. Ranunculus ficaria, Rosa hybrid, Insc. Menstr., 18. Solanum nigrum, Tropaeolum majus, Anomalothrips amygdali Morgan, Vinca minor and Vitis vinifera (El- 1929, Proc. Ent. Soc. Wash, 31: 5. Wakkad, 2007; Abd El-Wahab et al., Common name: Oat thrips. 2011; Shalaby, 2015; Abd El-Wahab, Distribution: Cyprus, Europe and 2016 and Wafy, 2018). Turkey (Tunc et al., 2012; Marullo and 4.10.4. De Grazia, 2013 and Mirab-balou, (Trybom) 2013). Frankliniella schultzei (Trybom, 1910), Host plant: Avena sativa (Priesner, Raff. Bull. Zool. 42(2):220. 1960). Physopus schultzei Trybom, 1910, 4.12. Limothrips Haliday Raff. Bull. Zool. 42(2):220. Limothrips Haliday, 1836, Ent. Mag., Frankliniella insularis Moulton, 1936, 445. Raff. Bull. Zool. 42(2):220. These species all bread in grass Common name: Common blossom and can cause reduction of cereal yield. thrips, cotton bud thrips, Limothrips recently includes eight Tomato thrips and Yellow flower species spread around the world. Of thrips. these, two species recorded in Egypt. Distribution: Tropical to subtropical 4.12.1. Limothrips angulicornis and southern Mediterranean, America, Jablonowski Angola, Asia, Cape Verde, Benin, Limothrips angulicornis Jablonowski, Botswana, Burkina Faso, Cameroon, 1894, Term. Fuzetek, XVII: 45. Central African Republic, Chad, Distribution: Europe, Iran, Italy, North Congo, Ethiopia, Gabon, Gambia, America, Sardinia, Turkey and Western Ghana, Kenya, Libya, Madagascar, Asia (Moritz et al., 2013). Maurititius, Morocco, Namibia, New Host plants: Hordeum Sp. and Wild Zealand, Niger, Nigeria, Senegal, grasses (Priesner, 1960). Somalia, South Africa, Sudan, 4.12.2. Limothrips cerealium Haliday Tanzania, Togo, Tunisia, Uganda, Limothrips cerealium Haliday, 1836, Zambia and Zimbabwe (Moritz et al ., Ent. Mag., 445. 2013). Limotbrips auenae Hinds, 1902. Proc. Host plant: Glycine max (Abd El- U.S. Nat. Mus., 26(1310):139-141. Wahab, 2016). Common name: Barley thrips, Cereal 4.10.5. Frankliniella tritici (Fitch) thrips, Grain thrips and thunder thrips. Frankliniella varicorne Bagnall, 1919. Distribution: Worldwide (Moritz et al., Bull. Bri. Mus. Ent. 9, II: 39 2013).

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Host plants: Citrus sp., Agrostis 4.14.1. Microcephalothrips verticillata, Alopecurus agrestis, abdominalis (D. L. Crawford) Avena fatua, Cynodon dactylon, Thrips abdominalis D. L. Crawford, Imperata cylindrical, Hordeum 1910, Pomona, Coll. Journ., 157. vulgare, H. maritimum, Mangifera Thrips microcephalus Priesner, 1923, indica, Panicum turgidum, Phalaris Ent. Mitteil., 12:160. paradoxa; Phragmites communis, Microcephalothrips abdominalis Polypogon monspeliensis, Psidium Priesner, 1937, Bull. Soc. R. Ent. guajava, Pyrus malus, Saccharum Egypte, 209. officinarum, Sorghum halepense, Common name: Composite thrips and Triticum vulgare and Vitis vinifera Sunflower thrips. (Priesner, 1960 and El-Wakkad, 2007). Distribution: Widespread (Moritz et 4.13. Bagnall al., 2013). Megalurothrips Bagnall, 1915. Ann. Host plants: Compositae, Mag. nat. Hist., (8) 15: 589. Chenopodium sp, Chrysanthemum sp, All species seem to associate Tagetes sp, Ageratum conyzoides, with Legumin. Fourteen species are Erigeron crispus, Mangifera indica, listed in this genus have been recoded. Pulicaria crispa and Vitis vinifera 4.13.1. Megalurothrips sjostedti (Priesner, 1960 and El-Wakkad, 2007). (Trybom) 4.15. Mycterothrips Karny Physopus sjostedti Trybom. 1908, Mycterothrips Karny, 1926, Mem. Wisr. Ergeb. Schwed. Zool. Exped. Dept. Agric. Ind., Ent., IX, 199. KiIimandjaro. Meru 1905-06, 3 (16): 4- Widely spread in Asia, Europe 6. and America in flowers and leaf litter. Physopus variabilis Bagnall, 1913, All species concentrate in old world. Ann. Mag. nat. Hist., (8) 12: 294. Thirty-sex species have been variabilis Steinweden, identifying in this genus. 1933, Trans. Amer. Ent. Soc., 59: 273- 4.15.1. Mycterothrips acacia Priesner 274. Mycterothrips acacia Priesner, 1932, Taeniothrips sjöstedti Steinweden, Bull. Soc. R. Ent. Egypte, 5, 142. 1933, Trans. Amer. Ent. Soc., 59: 275. Common name: Acacia flower thrips. Common name: Bean flower thrips Distribution: India (Rachana and and African bean thrips. Varatharajan, 2017). Distribution: Afrotropical Region, Host plant: Acacia nilotica (Priesner, Angola, Burundi, Cameroon, Central 1960). African Republic, Cape Verde Islands, 4.15.2. Mycterothrips echii Priesner Chad, Congo, Gabon, Ghana, Gambia, Mycterothrips echii Priesner , 1931, Ivory Coast, Kenya, Mozambique, Bull. Soc. R. Ent. Egypte, 129. Niger, Nigeria, Senegal, Seychelles, Host plant: Echium sericeum (Priesner, South Africa, Tanzania, Togo, Uganda, 1960). Yemen and Zimbabwe (Moritz et al ., 4.16. Serville 2013). Odontothrips Serville, 1843, Ins. Host plant: Glycine max (Abd El- Hemipt., 643. Wahab, 2016). This genus is flower-feeding 4.14. Microcephalothrips Bagnall thrips, especially Legume plant. It Microcephalothrips Bagnall 1926, consists of thirty-five species spread Ann. Mag. Nat. Hist. (9), 18: 113. around the world. It lives in flowers of . 4.16.1. Odontothrips elbaensis This genus presents by only one species Priesner widespread.

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Odontothrips elbaensis Priesner, 1933, Phlehothrips aegyptiacus Priesner, Bull. Soc. R. Ent. Egypte, 6. 1960, Publ. Inst. Desert Egypte 13: 284. Common name: Gebel elba thrips. Common name: Egyptian thrips. Host plant: Weeds (Priesner, 1960). Distribution: South Africa (Moritz et 16.2- Odontothrips karnyi Priesner al., 2013) Odontothrips karnyi Priesner, 1924, Host plants: Convolvulus arvensis, Konowia, 3: 1. Euphorbia cuneata, Lycium arabicum Common name: Broad bean thrips. (Priesner, 1960). Distribution: Canary Islands, Cyprus, 4.20. Psilothrips Hood Italy, Palestine and Sudan (Moritz et Psilothrips, Hood 1927, Proc. Biol. al., 2013). Soc. Washington, 40: 198. Host plants: Leguminosae, Lypinus sp., Thamnothrips Priesner, 1932, Bull. Lathyrus odoratus, Medicago hispsida, Soc. R. Ent. Egypte, 2. Melilotus messanensis,Retama raetam This genus represents by five and Vicia faba (Priesner, 1960). species through the world. 4.17. Uzel 4.20.1. Psilothrips bimaculatus Oxythrips Uzel, 1895, Mon. Ord. Thys. Priesner 133. Thamnothrips bimaculatus Oxythrips includes thirty-five Priesner,1932, Bull. Soc. R. Ent. species. Egypte, 3. 4.17.1. Oxythrips tamaricis (Bagnall) Common name: Bimaculated thrips. Anaphothrips tamaricis Bagnall, 1926, Distribution: Iran (Mirab-balou, Ann. Mag. Nat. Hist. (9), 18: 645. 2013). Oxythrips narasi Bagnall, 1926 Ann. Host plant: Lycium arabicum Mag. Nat. Hist. (9), 18: 648. (Priesner, 1960). Oxythrips tamaricis Priesner, 1938, 4.21. Prosopothrips Uzel Bull. Soc. Fouad I Ent., 123. Prosopothrips Uzel, 1895, Mon. Odr. Common name: Tamarisk thrips. Thy. 165. Host plants: Juncus sp., Suaeda sp. and Widespread genus associated Tamarix nilotica (Priesner, 1960). with Poaceae. Nine species have been 4.18. Parexothrips Priesner recorded around the world. Parexothrips Priesner 1960, Publ. Inst. 4.21.1. Prosopothrips matrouhenisis Desert Egypte 13:285. Priesner Old World genus, from Prosopothrips matrouhenisis Priesner, Poaceae. Nineteen species have been 1960, Publ. Inst. Desert Egypte 13: 277. recoded of this genus. Common name: Matrouh thrips. 4.18.1. Parexothrips tenellus Priesner Host plant: Low vegetation (Priesner, Exothrips tenellus Priesner, 1950, Bull. 1960). Soc. Fouad I Ent., 30. 4.21.2. Prosopothrips nigriceps Parexothrips tenellus, Pricsner, 1960, Bagnall Publ. Inst. Desert Egypte, 13: 286. Prosopothrips nigriceps Bagnall, 1927, Distribution: China and India (Moritz Bull. Bri. Mus. Ent. 9, II: 49. et al., 2013). Distribution: Southern France and Host plant: Imperata cylindrical Turkey (Priesner, 1937 and Tunc et al., (Priesner, 1960). 2012). 4.19. Phlebothrips Priesner Host plant: Grasses (Priesner, 1937). Phlehothrips aegyptiacus Priesner, 4.21.3. Prosopothrips salloumensis 1960, Publ. Inst. Desert Egypte 13: 283. Priesner 4.19.1. Phlebothrips aegyptiacus Prosopothrips nigriceps Priesner, 1938, Priesner Bull. Soc. R. Ent. Egypte, 210.

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Prosopothrips salloumensis Priesner, Scirtothrips dorsallis Hood, 1919, Bull. 1960, Publ. Inst. Desert Egypte 13:277. Bri. Mus. Ent. 9, II: 51. Common name: Salloum thrips. Heliothrips minutissimus Bagnall, Host plant: Grasses (Priesner, 1960). 1919, Bull. Bri. Mus. Ent. 9, II: 51. 4.22. Scirtothrips Shull Common names: Castor thrips, Chilli Scirtothrips, Shull, 1909, Ent. News, thrips, thrips and Yellow tea XX: 222. thrips. Species of the genus Distribution: Oriental and Pacific Scirtothrips are small, active thrips. Region, China, Israel, Ivory Coast, Sri They breed on the young leaves and Lanka, South Africa and U. S. A. flowers of plants. Several species are (Tillekaratne et al., 2011; Reitz et al., recognized as pests on citrus trees, 2011; Zur Strassen and Kuslitzky, 2012 coffee, and tea. The total number of and Moritz et al., 2013). described Scirtothrips is about one Host plant: Glycine max (Abd El- hundred and five species. In Egypt, the Wahab, 2016). genus consists of three species. 4.22.4. Scirtothrips mangiferae 4.22.1. Faure Priesner Scirtothrips aurantii Faure , 1929, Scirtothrips mangiferae Priesner, 1932, Transv. Univ. Coll., Bull. 18: 3. Bull. Soc. R. Ent. Egypte, 141-155. Common name: South African citrus Common name: Mango thrips. thrips. Distribution: Afrotropical and Eastern Distribution: Subtropics and tropics of Mediterranean, Gabon, India, Iran, Africa, introduced to Queensland Libya, Sudan and Yemen (Priesner, (Australia), Angola, China, Ethiopia, 1960; Mirab-balou, 2013 and Moritz et Ghana, Ivory Coast, Kenya, Malawi, al., 2013). Nigeria, South Africa, Sudan, Tanzania, Host plants: Citrus sp., Ficus carica, Uganda, U. S. A. and Zimbabwe (Reitz Mangifera indica, Parkinsonia et al., 2011 and Moritz et al ., 2013). aculeate, Pyrus malus and Vitis vinifera Host plants: Cassia sp., Citrus sp., (Priesner, 1932 and El-Wakkad, 2007). Cynodon sp., Panicum sp., Salix sp., 4.22.5. Scirtothrips nubicus Priesner Acacia nilotica, Convolvulus arvensis, Scirtothrips nubicus Priesner, 1936, Mangifera indica, Malus pumila, Olea Bull. Soc. R. Ent. Egypte, 84. europeana, Psidium guajava and Vitis Common name: common fig thrips. vinifera (Priesner, 1960; El- Wakkad, Distribution: Israel and Sudan (Zur 2007 and Wafy , 2018) Strassen and Kuslitzky, 2012). 4.22.2. Scirtothrips citri Moulton Host plants: Citrus sp., Ficus carica, Eutbrips Citri Moulton, 1909. Mangifera indica and Parkinsonia U.S.D.A., Bur. Ent., Tech. Ser. 12, pt. aculeate (Priesner, 1960). VII: 121-122. 4.23. Scolothrips Hinds Physothrips citri, Karny, 1912. 2001. Scolothrips Hinds 1902, Mon. Thys. N. Ann., 4:339. Amer. 157. Scirtothrips citri, Hood, 1914. Proc. Worldwide thrips. Originally, it Ent. Soc. Wash., 16(1):40. appears in old world country. It is Common name: Citrus thrips. predator of Tetranychid mites. Sixteen Distribution: China, Florida, Georgia, species have been described through the Iran and U. S. A. (Diffie et al., 2008 and world. Reitz et al., 2011) 4.23.1. Scolothrips latipennis Priesner Host plant: Citrus sp. (Salem et al., Scolothrips sexmaculatus uzeli 2017) Priesner, 1923, Entom. Mitteil., 12: 66, 4.22.3. Scirtothrips dorsallis Hood 115.

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Scolothrips uzeli Priesner, 1926, Thys. One species have been Eur., 241. identifying, it breads on Poaceae. Scolothrips latipennis Priesner, 1950, 4.25.1. Stenothrips graminum Uzel Bull. Soc. Fouad I Ent., 54. Stenothrips graminum Uzel, 1895, Distribution: Canary Isalnds Crimea, Mon. Ord. Thys. 210. Europe, Iran, Morocco and Spain Common name: Oats thrips. (Mirab-balou, 2013). Distribution: Austria, Croatia, Host plant: Predator of Tetranychidae Denmark, Hungary, Finland, Iceland, (Priesner, 1960). Iran, Norway, Poland, Sweden and 4.23.2. Scolothrips longicornis Turkey (Raspudici et al., 2009; Kobro, Priesner 2011; Tunc et al., 2012 and Mirab- Scolothrips longicornis Priesner , 1926, balou, 2013). Thys. Eur., 239. Host plant: Gramineae (Priesner, Common name: Tetranychid Predator 1937). thrips. 4.26. Taeniothrips Amyot and Distribution: Austria, China, Serville Denmark, Finland, Iran, Iceland, Taeniothrips Amyot and Serville, Norway, Sweden, Russia and Turkey 1843, Ins. Hemipt., 644. (Kobro, 2011; Mirab-balou et al., 2011; They live in flowers and young Tunc et al., 2012 and Mirab-balou, leaves. It includes thirty-one species. 2013). Six species only are valid in Egypt. Host plant: Predator of Tetranychidae 4.26.1. Taeniothrips dianthi Priesner (Priesner, 1960). Taeniothrips dianthi Priesner, 1921, 4.23.3. Scolothrips rhagebianus Wiener Ent. Zeitg., 38: 116, 117. Priesner Common name: Clover pink thrips and Scolothrips rhagebianus Priesner , carnation thrips. 1950, Bull. Soc. Fouad I Ent., 46. Distribution: Europe, Morocco, North Common name: Okra thrips. and South America (Priesner, 1960). Distribution: China, India, Iran, South Host plant: Syzygium aromaticum Africa and Sudan (Mirab-balou et al., (Priesner, 1960). 2011 and Mirab-balou, 2013). 4.26.2.Taeniothrips discolor Karny Host plant: Predator of Tetranychidae Euthrips discolor Karny, 1907, Berliner (Priesner, 1960). Ent. Zeitschr., 52:46. 4.24. Sitothrips Priesner Taeniothrips discolor Priesner, 1926, Sitothrips Priesner 1931, Bull. Soc. R. Thys. Eur., 292. Ent. Egypte, 127. Common name: Artemisia thrips. Five species belonged to this Distribution: Cyprus, Sibiria, South genus spread from Europe, Europe, and Turkey (Priesner, 1960 and Mediterranean region and Africa. Tunc et al., 2012). 4.24.1. Sitothrips arabicus Priesner Host plants: Artemisia sp., Iphiona Sitothrips arabicus Priesner 1931, Bull. mucronata, Linaria aegyptiaca and Soc. R. Ent. Egypte, 127. Odontospermum graveolens (Priesner, Distribution: Cyprus, Iran and Georgia 1960). (Priesner, 1960 and Mirab-balou, 4.26.3. Taeniothrips niloticus Priesner 2013). Taeniothrips niloticus Priesner 1930, Host plants: Grasses and Hordeum Bull. Soc. R. Ent. Egypte, 11. vulgare (Priesner, 1960). Host plants: Acacia nilotica, 4.25. Stenothrips Uzel Caesalpinia sepiaria and Punica Stenothrips Uzel, 1895, Mon. Ord. granatum (Priesner, 1960). Thys. 209.

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4.26.4. Taeniothrips Sinaiticus Achaetothrips loboptera Karny and Priesner Bagnallia, Amer. Publ. Co., New Delhi, Taeniothrips Sinaiticus Priesner 1935, Indica. 240. Bull. Soc. R. Ent. Egypte, 318. Distribution: Asia, Austria, Canary Host plants: Alcanna orientalis Islands, Czech Republic, Denmark, (Priesner, 1960). England, Europe, Iceland, Iran, Italy, 4.26.5. Taeniothrips traegardhi Finland, Morocco, Norway, Palestine, (Trybom) Sibiria, Sweden and Turkey (Kobro, Physopus traegardhi Trybom, 1911, 2011; Mirab-balou, 2013 and Moritz et Res. Swed. Zool. Exped. Egypt and c., al ., 2013). 4, Uppsala, 4. Host plants: Triticum Sp., Cistanche Taeniothrips niloticus Priesner, 1930, lutea, Chrysanthemum coronarium, Bull. Soc. R. Ent. Egypte, 11. Convolvulus arvensis, Hordeum Taeniothrips traegardhi Priesner , vulgare and tingitana 1938, Bull. Soc. R. Ent. Egypte, 210. (Priesner, 1960). Distribution: India and Sudan 4.28.2 . Thrips mareoticus Priesner (Priesner, 1960). Stenothrips mareoticus Priesner, 1932, Host plants: Grasses, Caesalpinia Bull. Soc. R. Ent. Egypte, 10. sepiaria, Lycium arabicum, Polypogon Thrips quadrisetosus Hood, 1932, Bull. monspeliensis, Pulica crispa, Punica Soc. R. Ent. Egypte, 124. granatum, Scirpus badius Sorghum Thrips tenuisetosus Ruxgs, 1935, (nec vulgare; Triticum vulgare and Zea Knechtel), Bull. Soc. Sci. Nat. Maroc, mays (Priesner, 1960). 15: 4. 4.26.6. Taeniothrips zillarum Priesner Thrips mareoticus Priesner, 1937, Taeniothrips zillarum Priesner, 1950, Bull. Soc. R. Ent. Egypte, 210. Bull. Soc. Fouad I Ent., 32. Common name: Lavender cotton Common name: Zilla thrips. thrips. Host plant: Zilla spionsa (Priesner, Distribution: Asia, Cyprus, Iran, 1960). Germany, Morocco, Palestine and 4.27. Thamnothrips Priesner Turkey (Mirab-balou, 2013 and Moritz Thamnothrips Priesner, 1932. Bull. et al., 2013). Soc. Fouad I Ent., 2. Host plants: Compositae, Achillea 4.27.1. Thamnothrips bimaculatus santolina, Chrysanthemum coronarium Priesner and Reichardia tingitana (Priesner, Thamnothrips bimaculatus Priesner, 1960). 1932. Bull. Soc. Fouad I Ent., 3. 4.28.3. Thrips mediterraneus Priesner Host plant: Desert bushes (Priesner, Thrips mediterraneus Priesner, 1934, 1932). Bull. Soc. R. Ent. Egypte, 276. 4.28. Thrips Linne Distribution: Armenia, Cyprus, Thrips Linne, 1776, Fauna Suecica, Ed. Europe, Iran, Iraq, Israel, Georgia, I, 220. Northern India, Lebanon, South It is considered as the largest Ukraine and Turkey (Mirab-balou, genus of Thysanoptera, it widespread of 2013 and Moritz et al., 2013). the world. Species are leaf and flower Host plant: Phlomis floccose (Priesner, feeding, most of them are pest of many 1960). plants. Two-hundred and ninety sex 4.28.4 . Thrips microchaetus Karny species have been described. Thrips flavus var. microchatus Karny, 4.28.1. Uzel 1920. Akad. Anzeig, 2. Thrips angusticeps Uzel, 1895, Mon. Thrips microchatus Hood, 1923, Ent, Ord. Thys. 191. Mitteil., 13: 116.

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Common name: Grapevine leaf thrips. Host plant: Gladiolus antakiensis Distribution: Sudan (Priesner, 1960). (Priesner, 1960). Host plants: Aerva tomentosa, Olea 4.28.8. Thrips tabaci Lindeman europeana, Pulicaria crispa, Thrips tabaci Lindeman, 1888. Schadl. Triumfetta flavescens, and Vitis vinifera Ins. d. Tabaks. i. Bessarabien, 15: 61- (Priesner, 1960 and Wafy, 2018). 65. 4.28.5. Thrips nigropilosus Uzel T. communis Uzel, 1895. Mon. Ord. Thrips nigropilosa Uzel, 1895. Mon. Thys., 176. Ord. Thys., 198. T. bremnenii Moulton, 1907. U.S.D. A., Thrips nigmpilosus, Priesner. 1927. Bull. Ent., Tech.Ser. 12: 59-60. Thys. Eur., 409-414. Common name: Onion thrips, cotton Common name: Chrysanthemum thrips. thrips. Distribution: Widespread (across all Distribution: Australia, Canada, the countries). China, Croatia, Denmark, Ethiopia, Host plant: Different kind of plants. Europe, Fiji, Finland, Hawaii, Iceland, 4.29. Trichromothrips Priesner Iran, Japan, Kenya, New Zealand, Trichromothrips Priesner, 1930, Bull. Norway, Russia, Sweden, Tanzania, Soc. R. Ent. Egypte, 9. Turkey and U.S.A. (Raspudici et al., Thirty-eight species have been 2009; Kobro, 2011; Mirab-balou, 2013 recorded. and Moritz et al., 2013). 4.29.1. Trichromothrips bellus Host plants: Althaea Sp. and Priesner Pyrethrum cinerariaefolium (Priesner, Trichromothrips bellus Priesner,1930, 1937). Bull. Soc. R. Ent. Egypte, 9. 4.28.6. Karny Common name: Papyrus thrips and Thrips palmi Karny, 1925, Raff. Bull. seedling barely thrips, wheat thrips. Zool. 42(2): 285. Distribution: India (Rachana and Distribution: Brazil, China, Denmark, Varatharajan, 2017). Finland, Guadeloupe, Iceland, India, Host plant: Cyperaceae (Priesner, Martinique, Norway, Sweden, Sri 1960). Lanka, Spain and U. S. A. (Monteiro, References 2001; Kobro, 2011; Reitz et al., 2011; Abd EL-Wahab, A.S. (2015): Tillekaratne et al., 2011; Etienne et al., Identification of a new species, 2015 and Rachana and Varatharajan, Marigold thrips 2017). Neohydatothrips samayunkur Host plants: Cucumis sativus and Kudô (Thysanoptera, Glycine max (Abd EL-Wahab et al., Thripidae) on marigold in 2012 and Abd El-Wahab, 2016). Egypt. Journal of Entomology 4.28.7. Morison and Zoology Studies, 3(5): 01- Physothrips simplex Morison, 1930, 06. Raff. Bull. Zool. 42(2):288. Abd EL-Wahab, A.S. (2016): Survey, Thrips simplex Bhatti, 1969, Raff. Bull. seasonal abundance of thrips Zool. 42(2):288. species and first record of two Common name: Gladiolus thrips. thrips species associated with Distribution: Australia, Canada, soybean and weed plants in Europe, Hawaii, India, Spain, South Egypt. Egypt. Acad. J. Biolog. Africa, Turkey and U. S. A. Sci., 9(3): 49–68. (Tillekaratne et al., 2011; Tunc et al., Abd EL-Wahab, A.S.; El-Sheikh, M. 2012 and Rachana and Varatharajan, A. K. and Elnagar, S. (2011): 2017) First record of Frankliniella

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