How Trees Modify Wood Development to Cope with Drought
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EVIDENCE for AUXIN REGULATION of BORDERED-PIT POSITIONING DURING TRACHEID DIFFERENTIATION in LARIX LARICINA By
IAWA Journal, Vol. 16 (3),1995: 289-297 EVIDENCE FOR AUXIN REGULATION OF BORDERED-PIT POSITIONING DURING TRACHEID DIFFERENTIATION IN LARIX LARICINA by Mathew Adam Leitch & Rodney Arthur Savidge 1 Faculty of Forestry and Environmental Management, University of New Brunswick, Fredericton, New Brunswick, E3B 6C2, Canada SUMMARY Chips containing cambium intact between xylem and phloem were cut from 8-year-old stem regions of 20-30-year-old dormant Larix laricina in late spring and cultured under controlled conditions for six weeks on a defined medium containing varied concentrations of I-naphthalene acetic acid (NAA, a synthetic auxin). Microscopy revealed that auxin was essential for cambial growth and tracheid differentiation. Low (0.1 mg/l) and high (10.0 mg/l) auxin concentrations were conducive to bordered pits forming in tangential walls, whereas an intermediate concentration (1.0 mg/l) of NAA favoured positioning of pits in radial walls. INTRODUCTION Two decades ago, Barnett and Harris (1975) pointed out that the processes involved in bordered-pit formation were incompletely understood, in particular the way in which the pit site is determined and how adjoining cells form a perfectly symmetrical bor dered-pit pair. Observing that radial walls of enlarging cambial derivatives were vari ably thick and thin, Barnett and Harris (1975) advanced the hypothesis that bordered pit development occurs at sites where the primary wall is thinned through the process of centrifugal displacement of microfibrils. Others, however, considered that a thick ening rather than a thinning of the primary wall was indicative of the site where pit development commenced (FengeI1972; Parham & Baird 1973). -
Eat Like an Mvp! Nj Devils Star Taylor Hall Tells How
A Publication of WINTER 2019 ROBERT WOOD JOHNSON UNIVERSITY HOSPITAL NEW BRUNSWICK healthytogether PROGRESS FOR PARKINSON’S EMERGENCY CARE FOR KIDS LIVING ORGAN DONORS EAT LIKE AN MVP! NJ DEVILS STAR TAYLOR HALL TELLS HOW RWJUHNB_Cover_Winter19_final.indd 1 12/18/18 2:41 PM A MESSAGE FROM LEADERSHIP HEALTH NEWS Joint Commission We’re Focused on Community Recognizes RWJUH oretta Scott King once said that the greatness of a community is best measured Surgical Programs by the compassionate actions of its members. At RWJBarnabas Health, we share King’s belief in the power of compassionate action. Each of our hospitals is Robert Wood Johnson University C Hospital New Brunswick has earned actively engaged in making a difference on critical community issues, including housing, the Joint Commission’s Gold Seal of Approval® for employment, food security and economic empowerment. That includes everything from Bariatric Surgery and Hip and Knee Joint Replacement hiring locally to finding ways to make fresh, affordable produce widely available. System- Programs. The certification award recognizes RWJUH wide, we’ve added RWJBarnabas Health TeleMed, a telemedicine service that improves New Brunswick’s dedication to continuous compliance access to care for people with transportation or scheduling challenges. with the Joint Commission’s state-of-the-art standards. To expand our community reach, we partner with other organizations, like the New Jersey Devils, to bring about positive change. One example: Collaborating with the staff of the Barnabas Health Hockey House at Newark’s Prudential Center, our RWJUH Cancer specialty physicians and nutrition experts have developed a youth hockey program Program Earns that promotes optimal performance and good health while building confidence, Prestigious sportsmanship and life skills. -
YELLOW-POPLAR (LIRIODENDRON TULIPIFERA L.)' George Lowerts E. A. Wheeler Robert C. Kellison
CHARACTERISTICS OF WOUND-ASSOCIATED WOOD OF YELLOW-POPLAR (LIRIODENDRON TULIPIFERA L.)' George Lowerts Forest Geneticist Woodlands Research, Union Camp Corp. Rincon, GA 31426 E. A. Wheeler Associate F'rofessor Department of Wood and Paper Science, North Carolina State University Raleigh, NC 27695-8005 and Robert C. Kellison Professor, Department of Forestry and Director, Hardwood Research Cooperative School of Forest Resoumes Raleigh, NC 27695-8002 (Received May 1985) ABSTRACT Selectedanatomical characteristicsand specificgravity ofydlow-poplar wood formed after wounding and adjacent to the wound were compared to similar characteristics of yellow-poplar wood formed before and after wounding and away from the wound. The wood formed immediately after wounding was similar anatomically to the bamer zones described for other species. Vessel volume, vessel diameter, percentage of vessel multiples, and vessel elcment length were significantly lower in wound- associated wood, while ray volume, ray density, and specific gravity were significantly greater. Such changes in the vessel system would result in a decrease in conductivity in the wounded area, while the increase in parenchyma would increase the potential for manufacture of fungitoxic compounds. With increasing radial distance from the wound area, the anatomical features of the wound-associated wood -~radualh . a~~roached .. those of normal wood, although. by. four years after wounding, the wood still had not returned to normal. The specific gravity stayed significantly greater. Keywords: Liriodendron lulipirpra L., yellow-poplar, barrier zones, wood anatomy, wounding, dis- coloration and decay. INTRODUCTION Wounds extending into the wood of branches, stems, or roots of a tree create an opportunity for the initiation of discoloration and decay. -
Yin-Yang, the Five Phases (Wu-Xing), and the Yijing 陰陽 / 五行 / 易經
Yin-yang, the Five Phases (wu-xing), and the Yijing 陰陽 / 五行 / 易經 In the Yijing, yang is represented by a solid line ( ) and yin by a broken line ( ); these are called the "Two Modes" (liang yi 兩義). The figure above depicts the yin-yang cycle mapped as a day. This can be divided into four stages, each corresponding to one of the "Four Images" (si xiang 四象) of the Yijing: 1. young yang (in this case midnight to 6 a.m.): unchanging yang 2. mature yang (6 a.m. to noon): changing yang 3. young yin (noon to 6 p.m.): unchanging yin 4. mature yin (6 p.m. to midnight): changing yin These four stages of changes in turn correspond to four of the Five Phases (wu xing), with the fifth one (earth) corresponding to the perfect balance of yin and yang: | yang | yin | | fire | water | Mature| |earth | | | wood | metal | Young | | | Combining the above two patterns yields the "generating cycle" (below left) of the Five Phases: Combining yin and yang in three-line diagrams yields the "Eight Trigrams" (ba gua 八卦) of the Yijing: Qian Dui Li Zhen Sun Kan Gen Kun (Heaven) (Lake) (Fire) (Thunder) (Wind) (Water) (Mountain) (Earth) 0 1 2 3 4 5 6 7 The Eight Trigrams can also be mapped against the yin-yang cycle, represented below as the famous Taiji (Supreme Polarity) Diagram (taijitu 太極圖): This also reflects a binary numbering system. If the solid (yang) line is assigned the value of 0 and the broken (yin) line is 1, the Eight Trigram can be arranged to represent the numbers 0 through 7. -
Intra-Organismal Variation in the Structure of Plant Vascular Transport Tissues in Poplar Trees
Trees (2018) 32:1335–1346 https://doi.org/10.1007/s00468-018-1714-z ORIGINAL ARTICLE Intra-organismal variation in the structure of plant vascular transport tissues in poplar trees Anna L. Jacobsen1 · Jessica Valdovinos‑Ayala1 · F. Daniela Rodriguez‑Zaccaro1 · M. Angela Hill‑Crim1 · Marta I. Percolla1 · Martin D. Venturas2 Received: 8 December 2017 / Accepted: 17 May 2018 / Published online: 24 May 2018 © Springer-Verlag GmbH Germany, part of Springer Nature 2018 Abstract Key message Phloem and xylem conduit structure vary greatly throughout the body of Populus trichocarpa trees, par- ticularly between roots and shoots. This has implications for understanding organ and whole plant vascular function. Abstract Woody plant vascular transport occurs predominantly within secondary xylem and phloem, which are both produced by the vascular cambium during secondary growth. We examined how vessel and sieve tube structure varied throughout the plant body of P. trichocarpa trees and whether xylem and phloem conduit structure was correlated across different positions within the plant. We excavated entire juvenile P. trichocarpa trees and measured vessel and sieve tube structural traits of current-year growth in 1 m increments along the main root:shoot axis. Trees were > 4 m tall and had roots that extended 4–5 m at their longest length. We found that both sieve tube and vessel diameters greatly varied throughout the plant body and with organ diameter. Roots had wider diameter conduits than shoots. Sieve tube diameter was strongly correlated with vessel diameter, which may be related to their common developmental origin. Other structural traits, such as pit membrane area and pit density for xylem, and sieve plate area and number of sieve areas per plate for phloem, also varied and were correlated with changes in conduit diameter. -
Redalyc.Stem and Root Anatomy of Two Species of Echinopsis
Revista Mexicana de Biodiversidad ISSN: 1870-3453 [email protected] Universidad Nacional Autónoma de México México dos Santos Garcia, Joelma; Scremin-Dias, Edna; Soffiatti, Patricia Stem and root anatomy of two species of Echinopsis (Trichocereeae: Cactaceae) Revista Mexicana de Biodiversidad, vol. 83, núm. 4, diciembre, 2012, pp. 1036-1044 Universidad Nacional Autónoma de México Distrito Federal, México Available in: http://www.redalyc.org/articulo.oa?id=42525092001 How to cite Complete issue Scientific Information System More information about this article Network of Scientific Journals from Latin America, the Caribbean, Spain and Portugal Journal's homepage in redalyc.org Non-profit academic project, developed under the open access initiative Revista Mexicana de Biodiversidad 83: 1036-1044, 2012 DOI: 10.7550/rmb.28124 Stem and root anatomy of two species of Echinopsis (Trichocereeae: Cactaceae) Anatomía de la raíz y del tallo de dos especies de Echinopsis (Trichocereeae: Cactaceae) Joelma dos Santos Garcia1, Edna Scremin-Dias1 and Patricia Soffiatti2 1Universidade Federal de Mato Grosso do Sul, CCBS, Departamento de Biologia, Programa de Pós Graduação em Biologia Vegetal Cidade Universitária, S/N, Caixa Postal 549, CEP 79.070.900 Campo Grande, MS, Brasil. 2Universidade Federal do Paraná, SCB, Departamento de Botânica, Programa de Pós-Graduação em Botânica, Caixa Postal 19031, CEP 81.531.990 Curitiba, PR, Brasil. [email protected] Abstract. This study characterizes and compares the stem and root anatomy of Echinopsis calochlora and E. rhodotricha (Cactaceae) occurring in the Central-Western Region of Brazil, in Mato Grosso do Sul State. Three individuals of each species were collected, fixed, stored and prepared following usual anatomy techniques, for subsequent observation in light and scanning electronic microscopy. -
Five Elements Note
Five Elements Note: This is NOT our work and we are only providing this because we want to share the link with you, but links often get deleted. If the link has been removed, the info is below. Otherwise, find it at http://blog.aoma.edu/blog/bid/307306/Chinese-Medicine-School-Basic- Five-Element-Theory The theory of the natural elements is an enduring philosophy across cultures, appearing in separate countries in vastly different eras around the world. The ancient Greeks used the five elements of earth, water, air, fire, and “aether” (quintessence/spirit) as a guiding principal to better understand the universe. Both ancient Egyptians and Buddhists understood the elements as fire, water, air, and earth. Hinduism utilizes the five elements (earth, water, fire, wind, and “aether”) as well. In fact, the seven chakras pair with Hindu and Buddhist five element theory. Western astrology also makes use of the four classical elements in astrological charting. In Traditional Chinese Medicine (TCM), Five Element theory (also called Wu Xing) is a powerful, foundational lens through which medicine, our bodies, and the world at large can be viewed. Fire, Earth, Metal, Water, and Wood are understood to be the Five Elements in TCM. Each element is awarded a number of characteristics and correspondences. They all have their separate natures, movements, directions, sounds, times of the day, and much more. Similar to Yin Yang theory, many specific aspects of life and the world can be attributed to a certain element. In addition to these basic qualities, the elements also correspond with certain internal organs, tastes, emotions, and sense organs in Traditional Chinese Medicine—a very important feature of the theory with great implications to the medicinal practice. -
A Physiologically Explicit Morphospace for Tracheid-Based Water Transport in Modern and Extinct Seed Plants
A Physiologically Explicit Morphospace for Tracheid-based Water Transport in Modern and Extinct Seed Plants The Harvard community has made this article openly available. Please share how this access benefits you. Your story matters Citation Wilson, Jonathan P., and Andrew H. Knoll. 2010. A physiologically explicit morphospace for tracheid-based water transport in modern and extinct seed plants. Paleobiology 36(2): 335-355. Published Version doi:10.1666/08071.1 Citable link http://nrs.harvard.edu/urn-3:HUL.InstRepos:4795216 Terms of Use This article was downloaded from Harvard University’s DASH repository, and is made available under the terms and conditions applicable to Open Access Policy Articles, as set forth at http:// nrs.harvard.edu/urn-3:HUL.InstRepos:dash.current.terms-of- use#OAP Wilson - 1 A Physiologically Explicit Morphospace for Tracheid-Based Water Transport in Modern and Extinct Seed Plants Jonathan P. Wilson* Andrew H. Knoll September 7, 2009 RRH: PHYSIOLOGICALLY EXPLICIT MORPHOSPACE LRH: JONATHAN P. WILSON AND ANDREW H. KNOLL Wilson - 2 Abstract We present a morphometric analysis of water transport cells within a physiologically explicit three-dimensional space. Previous work has shown that cell length, diameter, and pit resistance govern the hydraulic resistance of individual conducting cells; thus, we use these three parameters as axes for our morphospace. We compare living and extinct plants within this space to investigate how patterns of plant conductivity have changed over evolutionary time. Extinct coniferophytes fall within the range of living conifers, despite differences in tracheid-level anatomy. Living cycads, Ginkgo biloba, the Miocene fossil Ginkgo beckii, and extinct cycadeoids overlap with both conifers and vesselless angiosperms. -
University of Michigan University Library
CONTRIBUTIONS FROM THE MUSEUM OF PALEONTOLOGY THE UNIVERSITY OF MICHIGAN VOL. XIX, NO.6, pp. 65-88 (7 pls., 1 fig.) SEFTEMBER25, 1964 A FOSSIL DENNSTAEDTIOID FERN FROM THE EOCENE CLARNO FORMATION OF OREGON BY CHESTER A. ARNOLD and LYMAN H. DAUGHERTY MUSEUM OF PALEONTOLOGY THE UNIVERSITY OF MICHIGAN ANN ARBOR CONTRIBUTIONS FROM THE MUSEUM OF PALEONTOLOGY Director: LEWISB. KELLUM The series of contributions from the Museum of Paleontology is a medium for the publication of papers based chiefly upon the collections in the Museum. When the number of pages issued is sufficient to make a volume, a title page and a table of contents will be sent to libraries on the mailing list, and to individuals upon request. A list of the separate papers may also be obtained. Correspondence should be directed to the Museum of Paleontology, The University of Michigan, Ann Arbor, Michigan. VOLUMEXIX 1. Silicified Trilobites from the Devonian Jeffersonville Limestone at the Falls of the Ohio, by Erwin C. Stumm. Pages 1-14, with 3 plates. 2. Two Gastropods from the Lower Cretaceous (Albian) of Coahuila, Mexico, by Lewis B. Kellum and Kenneth E. Appelt. Pages 15-22, with 2 figures. 3. Corals of the Traverse Group of Michigan, Part XII, The Small-celled Species of Favosites and Emmonsia, by Erwin C. Stumm and John H. Tyler. Pages 23-36, with 7 plates. 4. Redescription of Syntypes of the Bryozoan Species Rhombotrypa quadrata (Rominger), by Roger J. Cuffey and T. G. Perry. Pages 37-45, with 2 plates. 5. Rare Crustaceans from the Upper Devonian Chagrin Shale in Northern Ohio, by Myron T. -
An Analytical and Comparative Study of Indian and Chinese Cosmologies
AN ANALYTICAL AND COMPARATIVE STUDY OF INDIAN AND CHINESE COSMOLOGIES S. MAHDIHASSAN* ABSTRACT To understand Chinese Cosmology in terms of the Indian system we have to look upon Cninese elements as symbols of the qualities they incorporate. The elements rearranged with the corresponding qualities would be : As elements: Fire. Water, Metal, Earth and Wood. As quali- ties: Hot, Cold, Dry, Moist, and Wind. Nothing can be drier than a metal hence dryness is symbolized as Metal. Subsoil is invariabry moist, hence moisture is symbolized as Earth. Wood is fresh - wood, like a cutting which transplants another life- form. It is potential life, like an egg, or better still here cosmic egg, the source of all creation. Its content is wind, like pneuma in Greek philosophy. Life-breath, the source of cosmic move- ment, cosmic existence, in fact cosmic soul. Life-energy is creative energy and Akasha as container would have cosmic soul as content. This makes wind (as cosmic) = Akasha, 1. A problem in comparative cosmology: By cosmology is understood a system of interpreting the universe III terms of few irreducible factors called Cosmic elements. Now \\ hatever exists in the universe can be either a form of matter, like star, stone and plant, or a form of energy, like heat and light. Further we must recognize entities as being independent of others, like stone as matter and heat as energy. Then what is merely relative, like heat and cold, these would be one and same entity, which concentrated would be called heat and reduced would be felt as cold. Thirdly there would be hypothetical entities logically justifiable but not knowable. -
Ecological Wood Adaptation and Horizontal Variations of Vessel Element and Fibre Length of Calligonum Mongolicum
Electronic Electronic Journal of Biology, 2006, Vol. 2(2): 19-23 Ecological Wood Adaptation and Horizontal Variations of Vessel Element and Fibre Length of Calligonum mongolicum Shumin Yang Research Institute of Wood Industry, Chinese Academy of Forestry, Beijing 100095, China E-mail: [email protected] Abstract 2. Materials and Methods Wood anatomy of Calligonum mongolicum Turcz. Five healthy trees were felled and two discs (2-3cm was described from an ecological perspective. This thick) from each tree were taken at the height of 20- species showed similar wood structure to that 30cm above the ground. Some of the discs were species published in the same genus [1]. This immediately fixed in formalin-acetic-alcohol (5:5:90 species has distinct growth ring boundaries, ring- v/v). porosity, distinct helical thickenings, simple Wood samples were softened in 5% glycerin perforation plate, nonseptate fibre, axial solution, subsequently sectioned with a sliding parenchyma, uni- or 2-5-seriate heterogeneous rays, microtome moving on transverse, radial and and alternate intervessel pitting. It was observed tangential surfaces of the disks. Thin sections were that there is a slightly increase of fibre length as the stained with safranine, dehydrated in a graded diameter from the pith increases. However, the alcohol series and mounted in Canada balsam for vessel element length remained more or less light microscope examination. Small blocks constant from pith to bark. Furthermore, the exposing transverse, radial and tangential surfaces relationships between anatomical features and were respectively prepared according to Exley et adaptability to desert environments were discussed. al’s methods [3] for scanning electron microscope (X-650, Hitachi Ltd Tokyo, Japan) observations. -
Zoroaster and the Theory of Four Elements
Bull. Hist. Chem., VOLUME 25, Number 2 (2000) 109 ZOROASTER AND THE THEORY OF FOUR ELEMENTS th bh, v Unvrt Intrdtn trl, nd n thr tp. vr, h dvtd nl t p t th rlr rn phlphr h npt f fr lnt: r, tr, rth, nd fr, rtr nd h rln. t rprnl, prhp, th thht t hv t rn th th Gr phlphr d f fr "rd" lnt bnt. Epdl bt 440 .C., hld fr n nt A tpl rnt r. Arttl (8422 .C. prntd fll (. ddd t th npt tht th Ardn t Arttl, th prprt f btn r IE b f th trl rld th rlt f th ltn prn f rtn fndn pr ttr, hh hd nl ptntl xt tl prprt. h Arttln t r n ntl prd b dtrn thrfr n "fr." fr h dd nt rnd nt th ht drn n hp nl, bt ll ht ll lnt bt AI EA tht nfrrd pn bd th n btrt nptn f t pf prprt. In rtn ntrr prprt r Cld t plt nfttn, "lt," pll ld Mt fr v r t th "fr n, htn, drn, nd lnt," r, tr, rth, tn, hh b nd fr hh r dtn ntd n fr bntn: WAE hd fr n nthr drn nd ht (fr, ht b thr "lt." In nd tr (r, tr Figure 1. h fr lnt rprntd n lt h lnt, n lt nd ld (tr, nd ld nd ll htr nd htr f htr b. prdnt vr th drn (rth (. thr: n rth, drn n Arttl nd h fllr blvd tht ll btn tr, ld n r, fldt, nd n fr, ht.