Grade 12 Life Sciences Human Evolution Workbook
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The Partial Skeleton Stw 431 from Sterkfontein – Is It Time to Rethink the Plio-Pleistocene Hominin Diversity in South Africa?
doie-pub 10.4436/JASS.98020 ahead of print JASs Reports doi: 10.4436/jass.89003 Journal of Anthropological Sciences Vol. 98 (2020), pp. 73-88 The partial skeleton StW 431 from Sterkfontein – Is it time to rethink the Plio-Pleistocene hominin diversity in South Africa? Gabriele A. Macho1, Cinzia Fornai 2, Christine Tardieu3, Philip Hopley4, Martin Haeusler5 & Michel Toussaint6 1) Earth and Planetary Science, Birkbeck, University of London, London WC1E 7HX, England; School of Archaeology, University of Oxford, Oxford OX1 3QY, England email: [email protected]; [email protected] 2) Institute of Evolutionary Medicine, University of Zurich, Winterthurerstrasse 190, CH-8057 Zurich, Switzerland; Department of Anthropology, University of Vienna, Althanstraße 14, 1090 Vienna, Austria 3) Muséum National d’Histoire Naturelle, 55 rue Buffon, 75005 Paris, France 4) Earth and Planetary Science, Birkbeck, University of London, London WC1E 7HX; Department of Earth Sciences, University College London, London, WC1E 6BT, England 5) Institute of Evolutionary Medicine, University of Zurich, Winterthurerstrasse 190, CH-8057 Zurich, Switzerland 6) retired palaeoanthropologist, Belgium email: [email protected] Summary - The discovery of the nearly complete Plio-Pleistocene skeleton StW 573 Australopithecus prometheus from Sterkfontein Member 2, South Africa, has intensified debates as to whether Sterkfontein Member 4 contains a hominin species other than Australopithecus africanus. For example, it has recently been suggested that the partial skeleton StW 431 should be removed from the A. africanus hypodigm and be placed into A. prometheus. Here we re-evaluate this latter proposition, using published information and new comparative data. Although both StW 573 and StW 431 are apparently comparable in their arboreal (i.e., climbing) and bipedal adaptations, they also show significant morphological differences. -
The Taxonomy of Primates in the Laboratory Context
P0800261_01 7/14/05 8:00 AM Page 3 C HAPTER 1 The Taxonomy of Primates T HE T in the Laboratory Context AXONOMY OF P Colin Groves RIMATES School of Archaeology and Anthropology, Australian National University, Canberra, ACT 0200, Australia 3 What are species? D Taxonomy: EFINITION OF THE The biological Organizing nature species concept Taxonomy means classifying organisms. It is nowadays commonly used as a synonym for systematics, though Disagreement as to what precisely constitutes a species P strictly speaking systematics is a much broader sphere is to be expected, given that the concept serves so many RIMATE of interest – interrelationships, and biodiversity. At the functions (Vane-Wright, 1992). We may be interested basis of taxonomy lies that much-debated concept, the in classification as such, or in the evolutionary implica- species. tions of species; in the theory of species, or in simply M ODEL Because there is so much misunderstanding about how to recognize them; or in their reproductive, phys- what a species is, it is necessary to give some space to iological, or husbandry status. discussion of the concept. The importance of what we Most non-specialists probably have some vague mean by the word “species” goes way beyond taxonomy idea that species are defined by not interbreeding with as such: it affects such diverse fields as genetics, biogeog- each other; usually, that hybrids between different species raphy, population biology, ecology, ethology, and bio- are sterile, or that they are incapable of hybridizing at diversity; in an era in which threats to the natural all. Such an impression ultimately derives from the def- world and its biodiversity are accelerating, it affects inition by Mayr (1940), whereby species are “groups of conservation strategies (Rojas, 1992). -
Human Evolution: a Paleoanthropological Perspective - F.H
PHYSICAL (BIOLOGICAL) ANTHROPOLOGY - Human Evolution: A Paleoanthropological Perspective - F.H. Smith HUMAN EVOLUTION: A PALEOANTHROPOLOGICAL PERSPECTIVE F.H. Smith Department of Anthropology, Loyola University Chicago, USA Keywords: Human evolution, Miocene apes, Sahelanthropus, australopithecines, Australopithecus afarensis, cladogenesis, robust australopithecines, early Homo, Homo erectus, Homo heidelbergensis, Australopithecus africanus/Australopithecus garhi, mitochondrial DNA, homology, Neandertals, modern human origins, African Transitional Group. Contents 1. Introduction 2. Reconstructing Biological History: The Relationship of Humans and Apes 3. The Human Fossil Record: Basal Hominins 4. The Earliest Definite Hominins: The Australopithecines 5. Early Australopithecines as Primitive Humans 6. The Australopithecine Radiation 7. Origin and Evolution of the Genus Homo 8. Explaining Early Hominin Evolution: Controversy and the Documentation- Explanation Controversy 9. Early Homo erectus in East Africa and the Initial Radiation of Homo 10. After Homo erectus: The Middle Range of the Evolution of the Genus Homo 11. Neandertals and Late Archaics from Africa and Asia: The Hominin World before Modernity 12. The Origin of Modern Humans 13. Closing Perspective Glossary Bibliography Biographical Sketch Summary UNESCO – EOLSS The basic course of human biological history is well represented by the existing fossil record, although there is considerable debate on the details of that history. This review details both what is firmly understood (first echelon issues) and what is contentious concerning humanSAMPLE evolution. Most of the coCHAPTERSntention actually concerns the details (second echelon issues) of human evolution rather than the fundamental issues. For example, both anatomical and molecular evidence on living (extant) hominoids (apes and humans) suggests the close relationship of African great apes and humans (hominins). That relationship is demonstrated by the existing hominoid fossil record, including that of early hominins. -
Title: Drimolen Crania Indicate Contemporaneity of Australopithecus, Paranthropus and Early Homo Erectus in S
Submitted Manuscript: Confidential Title: Drimolen crania indicate contemporaneity of Australopithecus, Paranthropus and early Homo erectus in S. Africa Authors: Andy I.R. Herries1,2*†, Jesse M. Martin1†, A.B. Leece1†, Justin W. Adams3,2†, Giovanni Boschian4,2†, Renaud Joannes-Boyau5,2, Tara R. Edwards1, Tom Mallett1, Jason Massey3,6, Ashleigh Murszewski1, Simon Neuebauer7, Robyn Pickering8.9, David Strait10,2, Brian J. Armstrong2, Stephanie Baker2, Matthew V. Caruana2, Tim Denham11, John Hellstrom12, Jacopo Moggi-Cecchi13, Simon Mokobane2, Paul Penzo-Kajewski1, Douglass S. Rovinsky3, Gary T. Schwartz14, Rhiannon C. Stammers1, Coen Wilson1, Jon Woodhead12, Colin Menter13 Affiliations: 1. Palaeoscience Labs, Dept. Archaeology and History, La Trobe University, Bundoora, 3086, VIC, Australia. 2. Palaeo-Research Institute, University of Johannesburg, Gauteng Province, South Africa. 3. Department of Anatomy and Developmental Biology, Biomedicine Discovery Institute, Monash University, VIC, Australia. 4. Department of Biology, University of Pisa, Italy 5. Geoarchaeology and Archaeometry Research Group (GARG), Southern Cross University, Military Rd, Lismore, 2480, NSW, Australia 6. Department of Integrative Biology and Physiology, University of Minnesota Medical School, USA 7. Department of Human Evolution, Max Planck Institute for Evolutionary Anthropology, Germany. 8. Department of Geological Sciences, University of Cape Town, Western Cape, South Africa 9. Human Evolution Research Institute, University of Cape Town, Western Cape, South Africa 10. Department of Anthropology, Washington University in St. Louis, St. Louis, USA 11. Geoarchaeology Research Group, School of Archaeology and Anthropology, Australian National University, Canberra, ACT, Australia 12. Earth Sciences, University of Melbourne, Australia 13. Department of Biology, University of Florence, Italy 14. Institute of Human Origins, School of Human Evolution and Social Change, Arizona State University, U.S.A. -
Unravelling the Positional Behaviour of Fossil Hominoids: Morphofunctional and Structural Analysis of the Primate Hindlimb
ADVERTIMENT. Lʼaccés als continguts dʼaquesta tesi queda condicionat a lʼacceptació de les condicions dʼús establertes per la següent llicència Creative Commons: http://cat.creativecommons.org/?page_id=184 ADVERTENCIA. El acceso a los contenidos de esta tesis queda condicionado a la aceptación de las condiciones de uso establecidas por la siguiente licencia Creative Commons: http://es.creativecommons.org/blog/licencias/ WARNING. The access to the contents of this doctoral thesis it is limited to the acceptance of the use conditions set by the following Creative Commons license: https://creativecommons.org/licenses/?lang=en Doctorado en Biodiversitat Facultad de Ciènces Tesis doctoral Unravelling the positional behaviour of fossil hominoids: Morphofunctional and structural analysis of the primate hindlimb Marta Pina Miguel 2016 Memoria presentada por Marta Pina Miguel para optar al grado de Doctor por la Universitat Autònoma de Barcelona, programa de doctorado en Biodiversitat del Departamento de Biologia Animal, de Biologia Vegetal i d’Ecologia (Facultad de Ciències). Este trabajo ha sido dirigido por el Dr. Salvador Moyà Solà (Institut Català de Paleontologia Miquel Crusafont) y el Dr. Sergio Almécija Martínez (The George Washington Univertisy). Director Co-director Dr. Salvador Moyà Solà Dr. Sergio Almécija Martínez A mis padres y hermana. Y a todas aquelas personas que un día decidieron perseguir un sueño Contents Acknowledgments [in Spanish] 13 Abstract 19 Resumen 21 Section I. Introduction 23 Hominoid positional behaviour The great apes of the Vallès-Penedès Basin: State-of-the-art Section II. Objectives 55 Section III. Material and Methods 59 Hindlimb fossil remains of the Vallès-Penedès hominoids Comparative sample Area of study: The Vallès-Penedès Basin Methodology: Generalities and principles Section IV. -
Little Foot, a South African Australopithecus As Old As
Press release Little Foot, a South African Australopithecus as 14 March 2014 old as Lucy? Laurent Bruxelles of Inrap has co-published an article with Ronald Clarke, Richard Maire, Richard Ortega and Dominic Stratford in the Journal of Human Evolution on the dating of Little Foot (StW 573) through stratigraphic analysis (Silberberg Grotto, Sterkfontein, South Africa). Sterkfontein, located to the north-west of Johannesburg in the province of Gauteng, is part of a zone classified by UNESCO in 1999 as a World Heritage Site called the “Cradle of Humankind”. Since its discovery in 1997 in Silberberg Grotto, a skeleton known as Little Foot has been meticulously excavated. Found under exceptional circumstances, this Australopithecine, which is the most complete ever identified, contributes unique elements to our knowledge of human origins. Although its age, somewhere between 2 and 4 million years, is still debated, a team of French researchers, in collaboration with the University of the Witwatersrand (Johannesburg), has contributed new elements to its dating. An exceptional discovery On September 6, 1994, in a box of animal bones, Ronald J. Clarke discovered four left foot bones belonging to a hominid found during an excavation of the lime miners’ rubble in the Sterkfontein cave system. This first discovery was dubbed “Little Foot” by P.V. Tobias, due to the small size of the foot that was identified. In May 1997, in a different box, he identified more bones from the same foot and a fragment of right shin bone.Being certain that these bones belonged to the same individual, Ron Clarke sent his assistants, Stephen Motsumi and Nkwane Molefe, to find the entire skeleton. -
Little Foot’ Fossil Chiselled out of Stone Yields Secrets Mysterious Ancient Hominin Retrieved After 20-Year Effort Might Be a Distinct Species
NEWS IN FOCUS HEALTH Mysterious polio- PUBLISHING China gets behind INFECTIOUS DISEASE Ebola NEUROSCIENCE Unlocking like disease probed with bold European open-access cases slip under how the brain makes machine learning p.170 plan p.171 the radar p.174 sense of faces p.176 PATRICK LANDMANN/SCIENCE PHOTO LIBRARY LANDMANN/SCIENCE PHOTO PATRICK The unique skeleton was embedded in rock deep inside a South African cave. ARCHAEOLOGY ‘Little Foot’ fossil chiselled out of stone yields secrets Mysterious ancient hominin retrieved after 20-year effort might be a distinct species. BY COLIN BARRAS a musculoskeletal biologist at the University He realized that a handful of small bones in the of Liverpool, UK, who collaborated with the collection belonged to a species of Australo- fter a tortuous 20-year-long excavation, research team that excavated the skeleton. pithecus — ape-like hominins in Africa between an ancient skeleton is starting to reveal The nickname Little Foot, echoing the mythi- about 4 million and 2 million years ago, before new information about early human cal ‘Bigfoot’, refers to the small foot bones that the human genus Homo rose to dominance1. Aevolution. The first of a raft of papers about ‘Lit- were among the first parts of the skeleton to Clarke and his colleagues then found more tle Foot’ suggests that the fossil is a female who be discovered. In 1994, Ronald Clarke, a pal- bones embedded in a matrix of solid rock deep showed some of the earliest signs of human-like, aeoanthropologist at the University of the Wit- in the caves. -
Senior Phase
1 SENIOR PHASE • Social Sciences 2 • Mathematics 9 • Life Orientation 14 • Natural Sciences 21 • Languages 29 • Technology 37 • EMS 40 2 SENIOR PHASE Social Sciences Senior Phase: Educator Page 3 Learning Area: Social Sciences Theme: World Heritage Sites and Sustainability Grade 8 Pages 3 - 8 Settlement and land use Urban settlements • Land use within urban settlements – including the central business district, zones for light and heavy industry, residential areas (high-, middle- and low-income), shopping centres, services and recreation Rural settlements • Types of rural settlement – including farming, mining, forestry, fishing Land use on aerial photographs and large-scale maps • What aerial photographs look like (oblique and vertical) Information from aerial photos – natural and constructed features • Identifying land uses in urban settlements (aerial photographs and large-scale maps *) Investigation of a settlement (project) • An independent study of a settlement known to the individual learner • Describe the settlement and the different types of land use. • Identify specific features or landmarks (natural and/or human-made). • Suggest reasons for the location of this settlement • Discuss decline and/or growth of population of the settlement and suggest reasons. Background knowledge Africa is the birthplace of humankind. This is where our collective umbilical cord lies buried. Hominids – the ancestors of modern humans – first emerged about 7-million years ago, in Africa. Many significant fossil finds have been made in the Cradle of Humankind World Heritage Site, including the famous fossils “Mrs Ples” and “Little Foot”. The first stone tools were made and used in Africa, at least 2.6-million years ago. Let us go and visit. -
Part 7 Humans
How do we recognise th 6 Extinction a human? • Walking on two legs (bipedal) We humans have destroyed over 80% of the habitat All non-human great apes are in the 34 continental biodiversity hotspots on Earth critically endangered. The biggest • Large brain relative to body size Biodiversity (see Part 2 of this series). threat is the loss of their habitat • Language, Symbolism, Happiness due to commercial logging, and the conversion of rainforest into palm oil plantations and biofuel Croplands of the Grazing agriculture. This in turn has led to lands take Earth occupy an an increase in human populations School activity & Extinction area about the up an area in forest areas and has led to an about the Part 7 size of South increase in bush meat trade. Why not set up a weekly Compiled by Dr John Anderson, Hominins America size of Africa debate on Democracy. Prof Francis Thackeray (ESI, Wits, anthropology) We humans, Homo sapiens, have arrived very late in the history of life. We’ve been Commercial Logging – The Why should we see the & Katherine Visser (Jhb Zoo, curator primates); cutting down of forest trees to layout by Ditshego Madopi & Anmari Hanekom around on Earth for only some 200,000 years. Our subtribe, the Hominina, have 7 bil world as ours alone? Bring all the cities sell wood and make room for Wouldn’t it be a good been around for some 7 million years; and our subfamily, the Homininae including Homo sapiens population Chimps agriculture is making less and less and towns and explosion population plan to give our close the chimps and gorillas, for some 10 million years. -
A Unique Middle Miocene European Hominoid and the Origins of the Great Ape and Human Clade Salvador Moya` -Sola` A,1, David M
A unique Middle Miocene European hominoid and the origins of the great ape and human clade Salvador Moya` -Sola` a,1, David M. Albab,c, Sergio Alme´ cijac, Isaac Casanovas-Vilarc, Meike Ko¨ hlera, Soledad De Esteban-Trivignoc, Josep M. Roblesc,d, Jordi Galindoc, and Josep Fortunyc aInstitucio´Catalana de Recerca i Estudis Avanc¸ats at Institut Catala`de Paleontologia (ICP) and Unitat d’Antropologia Biolo`gica (Dipartimento de Biologia Animal, Biologia Vegetal, i Ecologia), Universitat Auto`noma de Barcelona, Edifici ICP, Campus de Bellaterra s/n, 08193 Cerdanyola del Valle`s, Barcelona, Spain; bDipartimento di Scienze della Terra, Universita`degli Studi di Firenze, Via G. La Pira 4, 50121 Florence, Italy; cInstitut Catala`de Paleontologia, Universitat Auto`noma de Barcelona, Edifici ICP, Campus de Bellaterra s/n, 08193 Cerdanyola del Valle`s, Barcelona, Spain; and dFOSSILIA Serveis Paleontolo`gics i Geolo`gics, S.L. c/ Jaume I nu´m 87, 1er 5a, 08470 Sant Celoni, Barcelona, Spain Edited by David Pilbeam, Harvard University, Cambridge, MA, and approved March 4, 2009 (received for review November 20, 2008) The great ape and human clade (Primates: Hominidae) currently sediments by the diggers and bulldozers. After 6 years of includes orangutans, gorillas, chimpanzees, bonobos, and humans. fieldwork, 150 fossiliferous localities have been sampled from the When, where, and from which taxon hominids evolved are among 300-m-thick local stratigraphic series of ACM, which spans an the most exciting questions yet to be resolved. Within the Afro- interval of 1 million years (Ϸ12.5–11.3 Ma, Late Aragonian, pithecidae, the Kenyapithecinae (Kenyapithecini ؉ Equatorini) Middle Miocene). -
What Is the Viewpoint of Hemoglobin, and Does It Matter?
Hist. Phil. Life Sci., 31 (2009), 241-262 What is the Viewpoint of Hemoglobin, and Does It Matter? Jonathan Marks University of North Carolina at Charlotte Department of Anthropology Charlotte, NC 28223, USA ABSTRACT - In this paper I discuss reductive trends in evolutionary anthropology. The first involved the reduction of human ancestry to genetic relationships (in the 1960s) and the second involved a parallel reduction of classification to phylogenetic retrieval (in the 1980s). Neither of these affords greater accuracy than their alternatives; that is to say, their novelty is epistemic, not empirical. As a result, there has been a revolution in classification in evolutionary anthropology, which arguably clouds the biological relationships of the relevant species, rather than clarifying them. Just below the species level, another taxonomic issue is raised by the reinscription of race as a natural category of the human species. This, too, is driven by the convergent interests of cultural forces including conservative political ideologies, the creation of pharmaceutical niche markets, free-market genomics, and old- fashioned scientific racism. KEYWORDS – Molecular anthropology, Systematics, Classification, Human evolution Introduction In this paper I explore the intersection of genetics, taxonomy, and evolutionary anthropology. All three fields are scientific areas saturated with cultural meanings and associations. First, genetics is the scientific study of heredity, but has historically capitalized on non-scientific prejudices about heredity to curry support: hence James Watson’s epigrammatic proclamation in support of the Human Genome Project, “We used to think our fate was in the stars. Now we know, in large measure, our fate is in our genes” (Jaroff 1989; Duster 1990; Nelkin and Lindee 1995). -
New Findings – New Problems in Classification of Hominids+
Volume 46(1-2):57-60, 2002 Acta Biologica Szegediensis http://www.sci.u-szeged.hu/ABS SYMPOSIUM New findings – new problems in classification of hominids+ Gyula Gyenis Department of Biological Anthropology, Eötvös Loránd University, Budapest, Hungary ABSTRACT The criteria for the inclusion of species within the genus Homo have changed KEY WORDS over the years. There has been a stepwise relaxation of these criteria, therefore the gradistic classification classification and the evolutionary place of hominid fossils have never been free of controversy. cladistic classification It is the main reason that the discoveries of new hominid fossils have not helped in solving phylogenetic classification the generally accepted classification of hominids. Acta Biol Szeged 46(1-2):57-60 (2002) primates hominids “...neither the paleontological, nor the genetical, nor the second one was the Homo sapiens, the species to which all archeological records as they now stand can tell us exactly modern human populations belong. Since its introduction when, where, or how...” (Howells 1967) almost 250 years ago, our understanding of Homo has been Although almost 35 years passed after Howells (1967) changed by the addition of fossil species. This has resulted wrote the above cited statement concerning the evolution of in the step-by-step relaxation of the criteria for the inclusion the hominids, it remained as true as it was earlier. of species into the genus Homo. The species of the hominids that have been recognised Until the middle of the 1960s, all the classification of the since the late Pliocene fossils in the continents of the Old primates were based on Simpson’s classification (1945, World have always been in the center of a never ending 1961), which used only morphological characteristics, and debate: when, where and how they evolved into our species, a genus may be monopyletic or paraphyletic, too.