Death Defying Morphologies: Mass Extinction and Disparity in the Order Harpetida

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Death Defying Morphologies: Mass Extinction and Disparity in the Order Harpetida

EGU Galileo Conferences GC5-Mass-27, 2019 Mass extinctions, recovery and resilience © Author(s) 2019. CC Attribution 4.0 license. Death defying morphologies: mass extinction and disparity in the order Harpetida James Beech and James Lamsdell West Virginia University, Eberly College of Arts and Sciences, Geology, United States ([email protected]) This study seeks to better understand the phylogenetic relationships within the trilobite order Harpetida, with a view towards using this group to explore the relationship between extinction intensity and disparity. The harpetid response to the Late Ordovician mass extinction is of particular interest. A discrete morphological character matrix was created from the formal descriptions of harpetids in the published trilobite literature, and refined using first-hand observations of harpetid fossils. The final matrix consists of 76 discrete characters, including 69 cephalic characters, three thoracic characters, and four pygidial characters. This matrix is the first attempt of its kind to characterize the morphology of Harpetida as a whole, rather than focusing on individual harpetid genera. Exemplar species from a broad selection of harpetid genera, along with ptychopariid and redlichiid out groups, are included in the matrix, coded according to published figures and direct observation of specimens held in the collections of the Yale Peabody Museum of Natural History. The matrix was used to generate a hypothetical tree of harpetid phylogenetic relationships, the topology of which indicates support for harpetid monophyly but throws doubt onto the previous hypotheses of the internal relationships of the group. Disparity analysis of the group reveals a significant decline in morphological diversity across the Late Or- dovician mass extinction boundary, with slow recovery beginning in the Silurian and continuing into the Devonian..

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PUBLICACIONES DEL INSTITUTO GEOLÓGICO Y MINERO DE ESPAÑA Serie: CUADERNOS DEL MUSEO GEOMINERO. Nº 9 ADVANCES IN TRILOBITE RESEARCH ADVANCES IN TRILOBITE RESEARCH IN ADVANCES ADVANCES IN TRILOBITE RESEARCH IN ADVANCES planeta tierra Editors: I. Rábano, R. Gozalo and Ciencias de la Tierra para la Sociedad D. García-Bellido 9 788478 407590 MINISTERIO MINISTERIO DE CIENCIA DE CIENCIA E INNOVACIÓN E INNOVACIÓN ADVANCES IN TRILOBITE RESEARCH Editors: I. Rábano, R. Gozalo and D. García-Bellido Instituto Geológico y Minero de España Madrid, 2008 Serie: CUADERNOS DEL MUSEO GEOMINERO, Nº 9 INTERNATIONAL TRILOBITE CONFERENCE (4. 2008. Toledo) Advances in trilobite research: Fourth International Trilobite Conference, Toledo, June,16-24, 2008 / I. Rábano, R. Gozalo and D. García-Bellido, eds.- Madrid: Instituto Geológico y Minero de España, 2008. 448 pgs; ils; 24 cm .- (Cuadernos del Museo Geominero; 9) ISBN 978-84-7840-759-0 1. Fauna trilobites. 2. Congreso. I. Instituto Geológico y Minero de España, ed. II. Rábano,I., ed. III Gozalo, R., ed. IV. García-Bellido, D., ed. 562 All rights reserved. No part of this publication may be reproduced or transmitted in any form or by any means, electronic or mechanical, including photocopy, recording, or any information storage and retrieval system now known or to be invented, without permission in writing from the publisher. References to this volume: It is suggested that either of the following alternatives should be used for future bibliographic references to the whole or part of this volume: Rábano, I., Gozalo, R. and García-Bellido, D. (eds.) 2008. Advances in trilobite research. Cuadernos del Museo Geominero, 9.
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