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5 NOMINA INSECTA NEARCTICA

TABLE OF CONTENTS

Introduction ------7 Structure of the check list ------8 Original Orthography ------11 Diphyllostomatidae Species and genus group names indices ------11 Structure of the data base ------12 Elateridae End date of the List ------12 Quality Control ------13 Endecatomidae Alternative family names ------14 Some Statistics ------19 Classification ------21 Coleoptera ------41 ------43 Eulichadidae ------43 Amphizoidae ------44 Anobiidae ------44 Glaresidae ------50 Gyrinidae ------53 Archeocrypticidae ------55 Artematopodidae ------55 ------55 ------57 ------57 ------57 Hydroscaphidae ------57 Ithyceridae ------59 ------59 Lampyridae Brachypteridae ------59 Languriidae Limnichidae Lucanidae Callirhipidae Lutrochidae Cantharidae Carabidae Cerambycidae Ceratocanthidae Meloidae Micromalthidae Chrysomelidae Microsporidae Monommatidae Mycteophagidae Colydiidae Corylophidae Nitidulidae Dascillidae

TABLE OF CONTENTS 6 NOMINA INSECTA NEARCTICA

Omethidae Pleocomidae Psephenidae Pyrochroidae Rhipiceridae Rhipiphoridae Scydmaenidae Sphaeritidae Staphylinidae Telegeusidae Tenebrionidae Throscidae Coleoptera Species Index Coleoptera Generic Index Strepsiptera Stylopodiae Strepsiptera Species Index Strepsiptera Generic Index

TABLE OF CONTENTS 7 NOMINA INSECTA NEARCTICA

INTRODUCTION

list was compiled. Many users of this list will A thing is defined as much by what it isn't as by find a politically based division more useful than what it is. Nomina Insecta Nearctica is not a the more scientific one. Finally the division catalog. Nor is it even a check list by between the Nearctic and Neotropical zones is conventional standards. Rather Nomina Insecta nebulous at best, both in the United States and Nearctica is a directory of the scientific names Mexico and the Antillies. Mexico is considered applied to the of North America. These to be entirely Neotropical for purely practical names include senior synonyms, junior reasons and will be so treated in any future synonyms, junior homonyms, unavailable names portion of the Nomina series covering the (in the sense of the Code of Zoological Neotropical fauna. Nomenclature), and in some cases misspellings and misidentifications. Junior synonyms, Nomina Insecta Nearctica has very specific and homonyms, and so forth, are arranged limited goals. The ultimate ideal of every chronologically under the valid species name. systematist is to create a network of printed All other levels of organization are alphabetical. publications and computer databases containing Species are arranged alphabetically under the all known information about every described current generic name. Genera are alphabetical species in the world. The first step in attaining within families, and families are arranged this Holy Grail of systematics is a listing of the alphabetically within orders. species of the world and of the names that have been applied to them. The Nomina series is such Nomina Insecta Nearctica is not an original a first step and Nomina Insecta Nearctica is the piece of scientific research. It is a compilation of part covering the insects of North America, the the current literature, a snapshot of the current dominant component (perhaps as much as three- published status of the classification. This check fourths) of the flora and fauna of the Nearctic list contains no new combinations or region. synonomies. No objective replacement names are proposed even for clearly identified junior The Nomina series has two guiding principles: homonyms. Homonyms without available Stability and utility. Complete stability, of replacement name are indicated only by brackets. course, is impossible in any developing science, Further clearly wrong arrangements of junior and such as systematics, and is not even completely senior synonyms in the literature are not desirable. On the other hand systematics must corrected. For example if a younger name is also consider the needs of the users of systematic listed in the literature as the valid name for a systems. Systematics does not happen in species in preference to an older name, the isolation nor are systematists the final users of synonomy is given as recorded, although the systems proposed. Therefore the Nomina indicated by a question mark. This happens series will approach classifications from a more commonly that you might think. conservative position and will not adopt major changes in the classification of a group unless or Although Nomina Insecta Nearctica contains the until it meets one or more of two criteria. Is the word Nearctic, a more proper definition of the change necessary to break up polyphyletic or region covered by this publication is North paraphyletic groupings? Has the change gained America north of the Mexican border. general acceptance within the systematic and Significant portions of Mexico are properly part user communities? Users of systematic systems of the Nearctic region. The check list has been tend to concentrate on primary levels of limited to north of the Mexican border for classification; order, family, genus and species. practical reasons. Adding the Nearctic portions An unfortunate tendency eixsts these days to of Mexico to the list would have made the break up already monophletic order, family, and compilation of the list considerably more generic concepts into smaller and smaller groups. difficult, if not nearly impossible. The Mexican The usual reason given for such subdivision of border has been used as the standard boundary in existing and utilized groupings is that the new the majority of publications from which this groupings more accurately reflect the evolutionary history of the group. It is the

INTRODUCTION 8 NOMINA INSECTA NEARCTICA opinion of the editors of the Nomina series that weaknesses. A computer database is readily and the concepts of superfamily, subfamily, tribe, and quickly searched for information. A series of subgenus exist for just this purpose. We suggest CD-ROMS or online databases occupies far less that these subordinate categories be used for space than a series of printed volumes (if one refinement of the classification and that order, ignores the space taken by the computer). A family, and generic concepts be kept at stable as printed publication, on the other hand, is far possible for the user community. The splitting of more portable and easier to use for checking primary groups will not be followed unless they specific names or curating collections. A printed satisfy one of the two criteria listed above. publication is easier on the eyes, is easier to learn to use, and just "feels" better. Some of the Nomina Insecta Nearctica employs only primary weaknesses of "searchability" of the printed groups; Class, Order, Family, Genus, and publication can be made up for with a thorough Species. Subordinate groupings such as indexing of the material such as we have Superfamily, Subfamily, Tribe, Subgenus, and attempted in the printed version. Subspecies are not listed in the main body of the directory, although subordinate categories to the This volume is divided into three main parts. subfamily level are given in the classification The first part contains introductory material and portion of this publication. Names originally an explanation of how to use the volume. This proposed, or currently treated, as subgenera or introductory material also contains a synopsis of subspecies are treated as simple synonyms. The the current classification of each order to the absence of subordinate categories in the body of subfamily level with genera listed alphabetically the check list, particularly subgenera and within each subfamily. Higher categories are subspecies, is not a statement, positive or listed in "phylogenetic order", the source or negative, about the utility of such categories. sources of the order given in the introduction to Rather subordinate categories are absent for this section of the check list. The second section practical reasons implicit in the nature of the composes the check list proper with an Nomina series. The lack of subordinate alphabetical arrangement of families with the categories is implicit because Nomina Insecta Coleoptera, genera within each family, and Nearctica is a dictionary of names. An accurate species within each genus. The final portion of representation of the evolutionary history of a the volume consists of two indices. The first group is not among the limited goals of the index comprises all of the species group names Nomina series nor should it be. Other and where to go to find them. The second index publications and databases exist, or should exist, contains the genus group names. Because of the for this purpose. In addition the compilers of alphabetical arrangement employed in the series, Nomina Insecta Nearctica are authorities only in no page numbers are employed in the index. their own limited groups and could never Rather for any species group name one finds the satisfactorily produce a phylogentically arranged family in the body of the list by searching the list. Secondly the addition of subordinate page footers, then finding the genus, and then categories would have materially lengthened finding the valid species. More details on the each volume in the series when the volumes are index are given later in the introductory material. already enormous. Although information about subordinate categories and phylogenetic arrangements is not given in either the printed STRUCTURE OF THE CHECK version or in the CD-ROM accompanying the series, references to these categories and LIST arrangements can be often be found in the source field of the database contained on the CD-ROM. The source field is discussed later in the The structure of the check list is very simple. introductory material. The list consists of the primary check list followed by two indexes. All names are arranged Both printed and computer versions of the alphabetically in the two indices. The body of Nomina Insecta Nearctica database have been the check list is arranged alphabetically by produced for practical reasons. Printed family. The genera are listed alphabetically publications and computer databases have within the family, and the valid species names different and complimentary strengths and alphabetically within genera. Junior synonyms,

ALTERNATIVE FAMILY NAMES 9 NOMINA INSECTA NEARCTICA junior homonyms, unavailable names, Nomenclature. Specific reasons for the misspellings, and so forth are listed unavailability of the name are contained chronologically under the valid species. A valid in the database and will be available on species is defined as the senior synonym for a the CD-ROM version of the check list. species. A typical portion of the check list, in A generic name may also be listed as this case from the Anthicidae, is: Unav. if the name has been suppressed by the ICZN in favor of a younger Anthicus Paykull 1798 name. Omonadus Mulsant and Rey 1866 Syn. Emend. - An emendation. Original orthography Nathicus Casey 1895 Syn. Hemantus Casey 1895 Syn. is utilized throughout this check list. By the standards of this check list Anthicus ancilla Casey 1895 (Anthicus) there is no such thing as a justifiable Anthicus parallelus Casey 1895 Homo. emendation. Emendations are Anthicus caseyi Pic 1896 Syn. Anthicus antiochensis Werner 1975 (Anthicus) available names. Specific details Anthicus barbatus Werner 1964 (Anthicus) about the emendation are contained in Anthicus bellulus Leconte 1851 (Anthicus) the database and will be available on Anthicus minusculus Champion 1890 Syn. the CD-ROM version of the check list. Anthicus helvinus Casey 1895 Syn. Anthicus biguttulus Leconte 1851 (Anthicus) Missp. - A misspelling. Misspellings are not Anthicus ovicollis Casey 1895 Syn. available names. Specific information Anthicus protectus Casey 1895 Syn. about misspellings are contained in the Anthicus cervinus Laferte 1848 (Anthicus) database and will be available on the Anthicus bifasciatus Say 1826 Homo. Anthicus bizonatus Laferte 1848 Syn. CD-ROM version of the check list. Anthicus terminalis Leconte 1850 Syn. Misid. - A misidentification. Misidentifications Anthicus gilensis Casey 1895 Syn. are almost never used in the generic Anthicus seminotatus Casey 1895 Syn. listing, but the possibility is kept for Anthicus comanche Werner 1964 (Anthicus) cases requiring them for clarity.

The genus begins with the valid generic name The species group names follow a similar followed by a chronological listing in italics of structure. The valid species names are listed the junior synonyms, junior homonyms, alphabetically under the genus name. Junior emendations, and so forth, for the genus. The synonyms, homonyms, emendations, and so genus, in this case, Anthicus, is followed by the forth, are listed chronologically under the valid junior synonyms Omonadus Mulsant and Rey species name. All species group names are listed 1866, Nathicus Casey 1895, and Hemantus as binomials even if the species group name was Casey 1895. All genus group names are listed originally proposed as a subspecific or without indication of whether or not the genus infrasubspecific category such as subspecies, group name was originally or subsequently form, variety, or aberration. The original status treated as a subgenus. If a genus group name of these subspecific and infrasubspecific names was originally proposed as a subgenus, this is listed in the source field of the database and information is available in the CD-Rom version will be available in the CD-ROM version of the of the check list if this information was readily check list. The listing for the valid species available to the compilers of the database. The Anthicus cervinus is: abbreviation at the end of each synonomy gives Anthicus cervinus Laferte 1848 (Anthicus) the current status of the name. Anthicus bifasciatus Say 1826 Homo. Anthicus bizonatus Laferte 1848 Syn. Syn. - A junior synonym Anthicus terminalis Leconte 1850 Syn. Anthicus gilensis Casey 1895 Syn. Homo. - A junior homonym. Specific details Anthicus seminotatus Casey 1895 Syn. about the nature of the homonomy are The valid name is listed first and in plain text. contained in the database and will be Junior synonyms, homonyms, at so forth, are in available on the CD-ROM version of italics, indented, and listed chronologically the check list. below the valid species group name. Unav. - An unavailable name, either because the name is a nomen nudum or in The valid species group name consists of the some other way fails to satisfy the current generic name, the species group name, requirements of the Code of Zoological the author, the date of publication, and followed

INTRODUCTION 10 NOMINA INSECTA NEARCTICA by the original generic name in parentheses. The name may be a misidentification. Any field in a name not in the database at the The systematic literature, and the time this check list was published is indicated by Coleoptera literature in particular, is the notation [no entry]. Although an effort has plagued with misidentifications listed been made to eliminate all of these [no entry] but not identified as misidentifications. fields, a few remain because of either time The compilers have tried to eliminate as constraints or the unavailability of the literature many of these as possible, but some still needed to find the information. remain unidentified. The name may have been treated as a nomen oblitum The species group name for both valid names by the author or authors of the and subordinate names uses original source used in compiling the name, but orthography, i.e. the original spelling as used not specifically stated as such. Finally by the author in the original description. The sometimes the authors of the sources names have not been changed to agree in used just didn't want to use the older number and gender with the current generic name. We emphasize that this list is a assignment. For example if the original compilation. Therefore we have not description listed the name as Anthicus albus tried to correct these synonomies and Jones 1912 and albus was later transfered to the simply use the (?) convention to denote genus Striata, the name would be still be listed as the problem. Striata albus Jones 1912 (Anthicus). Original Homo. - The name is a junior homonym. A orthography is also used for all subordinate junior homonym may be a junior names. More will be said about the reasons for primary homonym, a junior secondary using original orthography in a later section of homonym, and a former junior the introduction. The check list, however, secondary homonym. These three types supplies all of the information necessary to make of homonomy cannot be distinguised in the changes in endings if the user so wishes. the printed check list. However, the database usually contains this Junior synonyms, homonyms, unavailable names, information and it will be available on and so forth, are listed chronologically under the the CD-ROM version of the check list. valid species name and consist of the the original Emend. - A emendation of a species group generic name, species group name, authors, and name. Original orthography is date. The date given is the actual date of employed throughout this list. publication. The convention of listing both the Therefore for the purposes of this puported and actual date of publication, if check list, there is no such thing as a different, is not used in this check list. The name justified emendation even if the is followed by an abbreviation indicating the original spelling is patently incorrect. status of the name. These abbreviations are Emendations are available names. essentially the same as those employed for genus Missp. - A misspelling of a species group name. group names. Misspellings are listed in this check list when they were readily available and Syn. - The name is a junior synonym of the identifiable in the sources used during valid name. In some cases the compilation. No special effort, abbreviation Syn. may be followed by a however, was made to find question mark in parentheses. The misspellings. question mark signifies an older species Unav. - The name is unavailable in the sense of group name than the listed valid name the Code of Zoological Nomenclature. and for which no reason was found in Several reasons can exist for a name to the sources used in compiling this list be unavailable. Most commonly the for its synonomy. Although an effort name is a nomen nudum or was made to resolve these problems, infrasubspecific. some remain and can probably be Misid. - The name is a misidentification. This traced to one of several causes. The category is rarely used in this list and name may be a junior homonym but only where absolutely necessary to was not listed as such in any of the clarify a particular situation. Listing all sources examined by the compilers. misidentifications would be an exercise

ALTERNATIVE FAMILY NAMES 11 NOMINA INSECTA NEARCTICA

in futility in any case. Indeed one of esthetic value for those brought up speaking a the major exercises in compiling this romance language. However, one of the two list was removing all of the compilers of the database sees no scientific value misidentifications listed in synonomies in such changes. In addition professional as if they were available names. systematists often disagree about the correct Nomob. - Nomen oblitum. In those cases where endings of species group names. This the oldest name for a species is listed complication can be avoided by simply not as a junior synonym and has been worrying about it. Never the less, if one wishes treated by the author of the source to conform to the current number and gender used by the compilers as a nomen provisions of the Code of Zoological oblitum, and if he or she clearly Nomenclature, the check list contains all of the indicated this, the abbreviation information necessary to make such changes. Nomob. is used. Changes in specific endings to reflect number and gender are not emendations in the sense of the Code of Zoological Nomenclature. ORIGINAL ORTHOGRAPHY

Original orthography has been employed SPECIES AND GENUS GROUP throughout this list whenever possible. Two NAMES INDICES reasons are advanced for using original orthography. A typical sample from the index of species group names is as follows: 1. In the opinion of the compilers of the Nomina series, computers will come to play an ever abdominalis White Tricorynus (Anobiidae) Tricorynus increasing role is keeping track of biosystematic abducens Casey Gnypeta (Staphylinidae) Gnypeta information. The changing of specific endings to abducens Casey Hippodamia (Coccinellidae) Hippodamia reflect the number and gender of the current glacialis abducens Casey Hister (Histeridae) Atholus nubilus generic assignment is destabilizing and difficult abducens Leconte Hormops (Curculionidae) Hormops to keep track of in a computer database such as abductus Horn Agrilus (Buprestidae) Agrilus the one employed in compiling the check lists aberrans White Tricorynus (Anobiidae) Tricorynus and databases of the Nomina series. In short aberti Hatch Bembidion (Carabidae) Bembidion dejectum aberti Hatch Philonthus (Staphylinidae) Philonthus computers do no speak Latin nor is there any simple way to make them. A critical element in A valid species name is indicated by a regular designing a relational database or distributed font and a junior synonym, homonym, database is the choice of a primary field for emendation, and so forth by italics. Use joining tables in the relationship. None of the Tricorynus abdominalis White as an example. standard categories such as genus or species is The entry for a valid name begins with the sufficient. However the combination of the species group name (abdominalis) followed by original generic and the species name as the author or authors (White), the original originally spelled is invariant over time. For generic name (Tricorynus), the current family example a field such as ogenus_species where assignment in parentheses (Anobiidae), ending ogenus is the original generic name, species is with the current generic assignment of the the species group name, and the character " _ " species (Tricorynus). To find this entry in the merely serves to join the two names into a single check list, go to the family in the alphabetical name does not change with time. Moreover any arrangement using the page footers, and then find valid species has one and only one the current genus in the alphabetical arrangement ogenus_species because of the laws of of genera. homonomy. The name is invariant with time and will not change regardless of later shifts in A typical synonym is generic assignment. 2. Changes in the endings of species group abducens Casey Hister (Histeridae) Atholus nubilus names to reflect the number and gender of the current generic assignment of the species has an

INTRODUCTION 12 NOMINA INSECTA NEARCTICA

The entry begins with the species group name and are not listed here. The final field in the (abducens) followed by the authors (Casey), the database is a general listing of information called original generic name (Hister), the family "source". The information in this field is not (Histeridae), the current generic assignment available in the printed publication, but will be (Atholus) and the current valid name for this given in the CD-ROM version of Nomina Insecta species (nubilus). To find this synonymous Nearctica released after completion of the name, simply find the current valid name, published version. The source field contains Atholus nubilus, using the same alphabetical specific information about the name in question. procedure given above. If the the homonymy, e.g. if the name is a junior secondary homonym, what name is it a junior The genus group name index works in the same secondary homonym of. Secondly the source way as the species group list except that for field contains a listing of the publication or synonymous names only the valid genus name publications used during the compilation of the needs to be listed. database. This "source" might be a printed check list, revision, or any publication including the STRUCTURE OF THE paper or book in which the name was originally described. The source field can often be used to DATABASE track down further informaiton about a name or species including its original description, Each species group and genus group name subsequent papers about its biology or consists of a record in a computer database from distribution, and the like. However the database which the information in this printed publication is a check list and a compilation and is not is drawn. A species group record contains the intended to take the place of catalogs or species name as originally spelled by the author databases specifically created to provide this type of the name (original orthography), the author or of information. authors of the name, the date of publication, the original generic name used by the describer of the species, and the current status of the name. If a name is the not valid name for a species, the ENDING DATE FOR THE valid name for the species is also listed. In this CHECK LIST printed publication the subordinate status of a name is indicated by listing it in italics This check list claims to cover all names chronologically beneath the valid name for the published prior to June of 1994. This cut-off species. date corresponds to volume 130 of the Zoological Record. The database also includes a breakdown of the distribution of a species by biogeographical region. Biogeographical breakdown is not applicable to Nomina Insecta Nearctica because all species occur in the Nearctic Region. Nor will further information on biogeographical QUALITY CONTROL regions be available in the accompanying CD- ROM because any further volumes in the Nomina series on insects will be compiled by Every effort has been made to make the data in biogeographical region. Any volume in the this directory has accurate and complete as Nomina series covering a group small enough to possible. The compilers of the Nomina Insecta be done in a single volume will contain Nearctica series, however, are limited by two information about occurrence by biogeographical important contstraints. region, the regions depending on whether the organism is terrestrial or marine. 1. Nomina Insecta Nearctica is a compilation from the literature, not an original piece of Some fields of the database are internal to the scientific research. Although thousands of hours functioning of the database (such as fields for of were spent checking original descriptions in creating relations between tables and a field order to compile correct spellings and original denoting the current status of the editing process)

ALTERNATIVE FAMILY NAMES 13 NOMINA INSECTA NEARCTICA generic names, this checking of original sources, could not be done for all, or even most, or the names listed. Ultimately the quality and accuracy of the list depends upon the sources used in compiling this list. Many of the errors encountered in these sources have been found and corrected during the compilation process, but many still remain. We also recognize that despite both direct and computer assisted proof reading, we will have committed our own mistakes. Entomological Information Services promises to correct such mistakes in the database whenever such mistakes are found or brought to our attention.

2. The Nomina series and its publisher, Entomological Information Services, is a commercial operation. The company receives no outside funds either public or private. EIS depends upon a rigid and rapid schedule of publication for its continued financial existence. Food must be bought and the mortgage paid. A leisurely search for perfection is not possible.

The compilers of Nomina Insecta Nearctica and the Nomina series will be extremely grateful to anyone sending us corrections of any errors in the check lists. These corrections will be immediately added to the database and incorporated in any future revisions. We also strongly encourage everyone to publish lists of corrections and additions to the list consistent with the goals of the series.

It is the intention of Entomological Information Services to continuously update the databases used in creating the volumes of the Nomina series and to put out updated versions of the checklists when they are needed. It would significantly assist us, and we would be extremely grateful, if authors could send us copies of their publications as they appear in print.

INTRODUCTION 14 NOMINA INSECTA NEARCTICA

ALTERNATIVE FAMILY NAMES

Blapsidae [see Tenebrionidae] The purpose of this section is to list some alternative ( not NA) family names the user might encounter in the Boridae Coleoptera. This list is not an exhaustive liste of family Bostrichidae Bostrychidae [see Bostrichidae] group names, but is included for the convenience of the Bothrideridae readers of the check list. The abbreviation (NA) stands Brachinidae [see Carabidae] for not North American, i.e. the family is not found in Brachyderidae [see Curculionidae] the Nearctic region. Brachypsectridae Brachypteridae Brachyrhinidae [see Curculionidae] COLEOPTERA FAMILIES Brachyrrhinidae [see Curculionidae] Brathinidae [see Staphylinidae] Brenthidae [see Brentidae] Acanthoceridae [see Ceratocanthidae] Brentidae (Acanthocnemidae not NA) Brontidae [see Silvanidae] Acanthoscelidae [see Chrysomelidae] Broscidae [see Carabidae] Aclopidae [see Scarabaeidae] Bruchelidae [see Anthribidae] Aculagnathidae [see Cerylonidae] Bruchidae [see Chrysomelidae] Aderidae Buprestidae Adimeridae [see Colydiidae] Byrrhidae Aegialiidae [see Scarabaeidae] Byrsopidae [see Curculionidae] Aegialitidae [see Salpingidae] Byturidae Agathidiidae [see Leiodidae] Aglycyderidae [see Belidae] Calendridae [see Curculionidae] Agnathidiidae [see Leiodidae] Callirhipidae Agridae [see Carabidae] Callistidae [see Carabidae] Agrypnidae [see Elateridae] Calophaenidae [see Carabidae] Agyrtidae Calopidae [see Oedemeridae] ( not NA) Calyptomeridae [see Clambidae] Alleculidae [see Tenebrionidae] Camiaridae [see Leiodidae] Amphizoidae Campylidae [see Elateridae] Anaspidae [see Scraptiidae] Cantharidae Anaspididae [see Scraptiidae] Carabidae Anchomenidae [see Carabidae] ( not NA) Anisosphaeridae [see Scydmaenidae] Carphuridae [see Melyridae] Anistomidae [see Leiodidae] Cassididae [see Chrysomelidae] Anobiidae Cateretidae [see Brachypteridae] Anommatidae [see Bothrideridae] Catogenidae [see Passandridae] Anoplodermatidae [see Cerambycidae] Catopidae [see Leiodidae] Anoplognathidae [see Scarabaeidae] Catopochrotidae [see Cryptophagidae] Anthicidae Cavicoxidae [see Elateridae] Anthiidae [see Carabidae] Cavicoxumidae [see Elateridae] Anthonomidae [see Curculionidae] (Cavognathidae not NA) Anthribidae Cebrionidae [see Elateridae] Apatidae [see Bostrichidae] Cephaloidae [see Stenotrachelidae] Aphodiidae [see Scarabaeidae] Cephaloplectidae [see Ptiliidae] Apionidae [see Brentidae] Cerambycidae Apoderidae [see Attelabidae] Cerasommatidiidae [see Endomychidae] Apotomidae [see Carabidae] Ceratocanthidae Archeocrypticidae Cerophytidae Artematopidae [see Artematopodidae] Cerylidae [see Cerylonidae] Artematopodidae Cerylonidae Ascleridae [see Oedemeridae] Cetoniidae [see Scarabaeidae] Aspidiphoridae [see Sphindidae] (Chaetosomatidae not NA) Atratoceridae [see Lymexylidae] (Chalcodryidae not NA) Attelabidae Chauliognathidae [see Cantharidae] Aculognathidae [see Latridiidae] Chelonariidae

Chiloeidae [see Dryopidae] Barididae [see Curculionidae] Chlaeniidae [see Carabidae] Belidae Chlamydidae [see Chrysomelidae] (Belohinidae not NA) Chloevidae [see Leiodidae] Bembidiidae [see Carabidae] Cholevidae [see Leiodidae] Biphyllidae Chrysomelidae Cicindelidae [see Carabidae]

ALTERNATIVE FAMILY NAMES 15 NOMINA INSECTA NEARCTICA

Ciidae Elmididae [see Elmidae] Cioidae [see Ciidae] Elminthidae [see Elmidae] Cisidae [see Ciidae] Elodidae [see Scirtidae] Cistelidae [see Tenebrionidae] Empelidae [see Staphylinidae] Clambidae Endecatomidae Clavigeridae [see Staphylinidae] Endomychidae Cleonidae [see Curculionidae] Engidae [see Erotylidae] Cleridae Engididae [see Erotylidae] Clytridae [see Chrysomelidae] Epilachnidae [see Coccinellidae] Cnemacanthidae [see Carabidae] Erirrhinidae [see Curculionidae] (Cneoglossidae not NA) Erotylidae Coccinellidae Eubriidae [see Psephenidae] Collyridae [see Carabidae] Euchiridae [see Scarabaeidae] Colonidae [see Leiodidae] Eucinetidae Colydiidae Eucnemidae Cononotidae [see Pyrochroidae] Eucnemididae [see Eucnemidae] Copridae [see Scarabaeidae] Euglenidae [see Aderidae] Corylophidae Euglenesidae [see Aderidae] Corynetidae [see Cleridae] Eulichadidae Cossonidae [see Curculionidae] Eumoplidae [see Chrysomelidae] Cossyphodidae [see Tenebrionidae] Eurhynchidae [see Brentidae] Crioceridae [see Chrysomelidae] Eurypalpidae [see Psephenidae] (Crowsoniellidae not NA) Eurypogonidae [see Artematopodidae] Cryptocephalidae [see Chrysomelidae] Eurystethidae [see Salpingidae] Cryptomeridae [see Eucinetidae] Euxestidae [see Cerylonidae] Cryptophagidae Cryptophilidae [see Languriidae] Feronidae [see Carabidae] Ctenodactylidae [see Carabidae] Fulcidacidae [see Chrysomelidae] Ctenostomidae [see Carabidae] Cucujidae Galerucidae [see Chrysomelidae] Cupedidae Ganglbaueriidae [see Oedemeridae] Cupesidae [see Cupedidae] Gehringiidae [see Carabidae] Curculionidae Georissidae [see Hydrophilidae] Cyathoceridae [see Lepiceridae] Georyssidae [see Hydrophilidae] Cybocephalidae [see Nitidulidae] Geotrupidae Cyladidae [see Brentidae] Glaphyridae Cymbionotidae [see Carabidae] Glaresidae Cyphonidae [see Scirtidae] Glyptidae [see Carabidae] Gnostidae [see Anobiidae] Dacnidae [see Erotylidae] Gyrinidae Dacoderidae [see Salpingidae] Dascillidae Haliplidae Dasytidae [see Melyridae] Halticidae [see Chrysomelidae] (Decliniidae not NA) Harpalidae [see Carabidae] Dermestidae Helmidae [see Elmidae] Derodontidae Helminthidae [see Elmidae] Diaperidae [see Tenebrionidae] Helodidae [see Scirtidae] Dicronychidae [see Elateridae] Helopheridae [see Hydrophilidae] Diphyllidae [see Biphyllidae] Helophoridae [see Hydrophilidae] Diphyllostomatidae Helopidae [see Tenebrionidae] ( not NA) ( not NA) Discolomidae [see Discolomatidae] Hemipeplidae [see Mycteridae] [see Cerambycidae] Heteroceridae Ditylidae [see Oedemeridae] Hiletidae [see Carabidae] Dolosidae [see Cerylonidae] Hispidae [see Chrysomelidae] Donaciidae [see Chrysomelidae] Histeridae Doydirhynchidae [see Nemonychidae] (Hobartiidae not NA) (Drylidae not NA) Homalisidae [see Omalisidae] Dryopidae Hybosoridae Dryptidae [see Carabidae] Hydraenidae Dynastidae [see Scarabaeidae] Hydrochidae [see Hydrophilidae] Dytiscidae Hydrophilidae Hydroporidae [see Dytiscidae] Ectrephidae [see Anobiidae] Hydroscaphidae Elacatidae [see Salpingidae] (Hygrobiidae not NA) Elaphridae [see Carabidae] Hylophilidae [see Aderidae] Elateridae Hylurgidae [see Curculionidae] Elmidae Hyphydridae [see Dytiscidae]

ALTERNATIVE FAMILY NAMES 16 NOMINA INSECTA NEARCTICA

Hypocephalidae [see Cerambycidae] Melyridae Hypocopridae [see Cryptophagidae] Merophysiidae [see Endomychidae] Merycidae [see Zopheridae in part; see in part] Inopeplidae [see Salpingidae] Metriidae [see Carabidae] Ipidae [see Curculionidae] Micromalthidae Ithyceridae Micropeplidae [see Staphylinidae] Microsporidae ( not NA) Migadopidae [see Carabidae] Monocrepidiidae [see Elateridae] Karumiidae [see Dascillidae] Monoedidae [see Colydiidae] [see Brachypteridae] Monommatidae Korynetidae [see Cleridae] Monommidae [see Monommatidae] Monotomatidae [see Monotomidae] Laemophloeidae Monotomidae Lagriidae [see Tenebrionidae] Mordellidae Lamiidae [see Cerambycidae] Murmidiidae [see Cerylonidae] (Lamingtoniidae not NA) Mycetaeidae [see Endomychidae] Lamprosomatidae [see Chrysomelidae] Lampyridae Mycteridae Languriidae Mylabridae [see Chrysomelidae] Laricobiidae [see Derodontidae] Myloechinae [see Leiodidae] Lariidae [see Chrysomelidae] Lathridiidae [see Latridiidae] Nacerdidae [see Oedemeridae] Latridiidae Nebriidae [see Carabidae] Lebiidae [see Carabidae] Necrophoridae [see Silphidae] Leiodidae Nemenychidae [see Nemonychidae] (Lepiceridae not NA) Nemonychidae Leptinidae [see Leiodidae] Nemosomidae [see Trogossitidae] Leptodiridae [see Leiodidae] Nilionidae [see Tenebrionidae] Lepturidae [see Cerambycidae] Niponidae [see Histeridae] Lichadidae [see Eulichadidae] Niponiidae [see Histeridae] Lichnidae [see Glaphyridae] Nitidulidae Licinidae [see Carabidae] Nosodendridae Limnebiidae [see Hydraenidae] Noteridae Limnichidae Notiophygidae [see Discolomatidae] Limniidae [see Elmidae] Notoxidae [see Anthicidae] Limulodidae [see Ptiliidae] Liodesidae [see Leiodidae] Ochodaeidae Liodidae [see Leiodidae] Odacanthidae [see Carabidae] Lissomidae [see Elateridae] Oedemeridae Lophocateridae [see Trogossitidae] Omalidae [see Staphylinidae] Loriceridae [see Carabidae] (Omalisidae not NA) Loroceridae [see Carabidae] Omalysidae [see Omalisidae] Lucanidae Omethidae Ludiidae [see Elateridae] Ommadidae [see Ommatidae] Lutrochidae Ommatidae Lycidae [see Lycidae] Omophronidae [see Carabidae] Lyctidae [see Bostrichidae] Orphnidae [see Scarabaeidae] Lymexylidae Orsodacnidae Lymexylonidae [see Lymexylidae] Orthoceridae [see Colydiidae] Lyttidae [see Meloidae] Orthogoniidae [see Carabidae] Orthoperidae [see Corylophidae] Malachiidae [see Melyridae] Ostomatidae [see Trogossitidae] Malacodermidae [see Lampyridae] Ostomidae [see Trogossitidae] Malalopidae [see Chrysomelidae] Othniidae [see Salpingidae] Mantichoridae [see Carabidae] Otiorhynchidae [see Curculionidae] Masoreidae [see Carabidae] Oxycorynidae [see Belidae] Megacephalidae [see Carabidae] [see Cerambycidae] Megalopidae [see Megalopodidae] Oxytelidae [see Staphylinidae] Megalopodidae Ozaenidae [see Carabidae] Megascelidae [see Chrysomelidae] Megatomidae [see Dermestidae] Pahypodidae [see Scarabaeidae] Melandryidae Panagaeidae [see Carabidae] Melanodidae [see Carabidae] Parandridae [see Cerambycidae] Melanophthalmidae [see Latridiidae] Parnidae [see Dryopidae] Melasidae [see Eucnemidae] Passalidae Meloidae Passandridae Melolonthidae [see Scarabaeidae] Patrobidae [see Carabidae]

ALTERNATIVE FAMILY NAMES 17 NOMINA INSECTA NEARCTICA

Paussidae [see Carabidae] Rhizophagidae [see Monotomidae] Pedilidae [see Pyrochroidae] Rhynchitidae [see Attelabidae] Peleciidae [see Carabidae] Rhynchophoridae [see Curculionidae] Pelobiidae [see Hygrobiidae] Rhysodidae Peltasticidae [see Derodontidae] Rhysopaussidae [see Tenebrionidae] Peltidae [see Trogossitidae] Rutelidae [see Scarabaeidae] Pentagonicidae [see Carabidae] Pericalidae [see Carabidae] Sagridae [see Chrysomelidae] Perigonidae [see Carabidae] Salpingidae (Perimylopidae not NA) Sandalidae [see Rhipiceridae] Perothopidae [see Eucnemidae] Sarothriidae [see Jacobsoniidae] Petriidae [see Tenebrionidae] Scalidiidae [see Passandridae] Phaenocephalidae [see Phalacridae] Scaphidiidae [see Staphylinidae] Phaenomeridae [see Scarabaeidae] Scarabaeidae Phaenomerididae [see Scarabaeidae] Scaritidae [see Carabidae] Phalacridae Schizopidae [see Buprestidae] Phengodidae Scirtidae (Phloiophilidae not NA) Scolytidae [see Curculionidae] Phloeophilidae [see Phloiophilidae] Scraptiidae (Phloeostichidae not NA) Scydmaenidae Phloephilidae [see Melyridae] Serropalpidae [see Melandryidae] Phreatodytidae [see Noteridae] Siagonidae [see Carabidae] (Phycosecidae not NA) Sibynidae [see Curculionidae] Phylloceridae [see Eucnemidae] Silphidae Pilipalpidae [see Pyrochroidae] Silvanidae Pimeliidae [see Tenebrionidae] Smicripidae (Plastoceridae not NA) Sogdiidae [see Leiodidae] Platostomatidae [see Anthribidae] Sparedridae [see Oedemeridae] Platypidae [see Curculionidae] Spercheidae [see Hydrophilidae] Platypodidae [see Curculionidae] Spermophagidae [see Chrysomelidae] Platypsyllidae [see Leiodidae] Sphaeridiidae [see Hydrophilidae] Platystomidae [see Anthribidae] Sphaeriidae [see Microsporidae] Platyrhinidae [see Anthribidae] Sphaeritidae Pleocomidae Sphaerosomatidae [see Alexiidae] (Podabrocephalidae not NA) Sphaerosomidae [see Alexiidae] Prionidae [see Cerambycidae] Sphindidae ( not NA) Spondylidae [see Cerambycidae] (Propalticidae not NA) Spondylididae [see Cerambycidae] Prostomidae Staphylinidae Proterhinidae [see Belidae] Stenidae [see Staphylinidae] Proterrhinidae [see Belidae] Stenostomatidae [see Oedemeridae] ( not NA) Stenotrachelidae Protorhinidae [see Curculionidae] Syncalyptidae [see Byrrhidae] Psallidiidae [see Curculionidae] Synchroidae Pselaphidae [see Staphylinidae] (Synteliidae not NA) Psephenidae Psephenoididae [see Psephenidae] Tachyporidae [see Staphylinidae] Pseudomorphidae [see Carabidae] Taurocerastidae [see Geotrupidae] Psoidae [see Bostrichidae] Telegeusidae Psydridae [see Carabidae] Telephoridae [see Cantharidae] Pterocolidae [see Attelabidae] Temnochilidae [see Trogossitidae] (Pterogeniidae not NA) Tenebrionidae Pterostichidae [see Carabidae] Tentyriidae [see Tenebrionidae] Ptiliidae Tetraonycidae [see Meloidae] Ptilodactylidae Tetraonychidae [see Meloidae] Ptinidae [see Anobiidae] Tetraphateridae [see Ommatidae] Pyrochroidae Tetratomidae Pyrophoridae [see Elateridae] Tetropiidae [see Cerambycidae] Pythidae Thorictidae [see Dermestidae] Thrachypachyidae [see Trachypachidae] Rhadalidae [see Melyridae] Threopteridae [see Carabidae] Rhagophtalmidae [see Phengodidae] Throscidae Rhinomaceridae [see Nemonychidae] Thylodriidae [see Dermestidae] (Rhinorhipidae not NA) Tillidae [see Cleridae] Rhipiceridae Tomicidae [see Curculionidae] Rhipidoceridae [see Rhipiceridae] ( not NA) Rhipiphoridae (Trachelostenidae not NA) Rhissodidae [see Rhysodidae] Trachypachidae

ALTERNATIVE FAMILY NAMES 18 NOMINA INSECTA NEARCTICA

Trechidae [see Carabidae] Tretothoracidae [see Salpingidae] Trichopterygidae [see Ptiliidae] ( not NA) Trixagidae [see Throscidae] Trogidae Trogossitidae Trogostitidae [see Trogossitidae]

(Ulodidae not NA) Urodontidae [see Anthribidae]

Vesperidae [see Cerambycidae]

Xylophilidae [see Aderidae]

Zarudniolidae [see Dascillidae] Zopheridae Zuphiidae [see Carabidae]

STREPSIPTERA FAMILIES

Callipharixenidae [see Stylopidae] Dioxoceridae [see Stylopidae] Elenchidae [see Stylopidae] Halictophagidae [see Stylopidae] Hylechthridae [see Stylopidae] Mengeidae [see Stylopidae] Myrmecolacidae [see Stylopidae] Stichotrematidae [see Stylopidae] Stylopidae Xenidae [see Stylopidae]

ALTERNATIVE FAMILY NAMES