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New Materials of Chalicotherium Brevirostris (Perissodactyla, Chalicotheriidae)
Geobios 45 (2012) 369–376 Available online at www.sciencedirect.com Original article New materials of Chalicotherium brevirostris (Perissodactyla, Chalicotheriidae) § from the Tunggur Formation, Inner Mongolia Yan Liu *, Zhaoqun Zhang Laboratory of Evolutionary Systematics of Vertebrate, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, 142, Xizhimenwai Street, PO Box 643, Beijing 100044, PR China A R T I C L E I N F O A B S T R A C T Article history: Chalicotherium brevirostris was named by Colbert based on a skull lacking mandibles from the late Middle Received 23 March 2011 Miocene Tunggur Formation, Tunggur, Inner Mongolia, China. Here we describe new mandibular Accepted 6 October 2011 materials collected from the same area. In contrast to previous expectations, the new mandibular Available online 14 July 2012 materials show a long snout, long diastema, a three lower incisors and a canine. C. brevirostris shows some sexual dimorphism and intraspecific variation in morphologic characters. The new materials differ Keywords: from previously described C. cf. brevirostris from Cixian County (Hebei Province) and the Tsaidam Basin, Tunggur which may represent a different, new species close to C. brevirostris. The diagnosis of C. brevirostris is Middle Miocene revised. Chalicotheriinae Chalicotherium ß 2012 Elsevier Masson SAS. All rights reserved. 1. Abbreviations progress in understanding the Tunggur geology and paleontology (Qiu et al., 1988). The most important result of this expedition so far is -
High-Resolution Magnetostratigraphy of the Neogene Huaitoutala Section
Earth and Planetary Science Letters 258 (2007) 293–306 www.elsevier.com/locate/epsl High-resolution magnetostratigraphy of the Neogene Huaitoutala section in the eastern Qaidam Basin on the NE Tibetan Plateau, Qinghai Province, China and its implication on tectonic uplift of the NE Tibetan Plateau ⁎ Xiaomin Fang a,b, , Weilin Zhang a, Qingquan Meng b, Junping Gao b, Xiaoming Wang c, John King d, Chunhui Song b, Shuang Dai b, Yunfa Miao b a Center for Basin Resource and Environment, Institute of Tibetan Plateau Research, Chinese Academy of Sciences, P. O. Box 2871, Beilin North Str. 18, Beijing 100085, China b Key Laboratory of Western China's Environmental Systems, Ministry of Education of China & College of Resources and Environment, Lanzhou University, Gansu 730000, China c Department of Vertebrate Paleontology, Natural History Museum of Los Angeles County, 900 Exposition Boulevard, Los Angeles, CA 90007, USA d Graduate School of Oceanography, University of Rhode Island, URI Bay Campus Box 52, South Ferry Road, Narragansett, RI 02882-1197, USA Received 31 December 2006; received in revised form 23 March 2007; accepted 23 March 2007 Available online 31 March 2007 Editor: R.D. van der Hilst Abstract The closed inland Qaidam Basin in the NE Tibetan Plateau contains possibly the world's thickest (∼12,000 m) continuous sequence of Cenozoic fluviolacustrine sedimentary rocks. This sequence contains considerable information on the history of Tibetan uplift and associated climatic change. However, work within Qaidam Basin has been held back by a paucity of precise time constraints on this sequence. Here we report on a detailed paleomagnetic study of the well exposed 4570 m Huaitoutala section along the Keluke anticline in the northeastern Qaidam Basin, where three distinct faunas were recovered and identified from the middle Miocene through Pliocene. -
Chapter 1 - Introduction
EURASIAN MIDDLE AND LATE MIOCENE HOMINOID PALEOBIOGEOGRAPHY AND THE GEOGRAPHIC ORIGINS OF THE HOMININAE by Mariam C. Nargolwalla A thesis submitted in conformity with the requirements for the degree of Doctor of Philosophy Graduate Department of Anthropology University of Toronto © Copyright by M. Nargolwalla (2009) Eurasian Middle and Late Miocene Hominoid Paleobiogeography and the Geographic Origins of the Homininae Mariam C. Nargolwalla Doctor of Philosophy Department of Anthropology University of Toronto 2009 Abstract The origin and diversification of great apes and humans is among the most researched and debated series of events in the evolutionary history of the Primates. A fundamental part of understanding these events involves reconstructing paleoenvironmental and paleogeographic patterns in the Eurasian Miocene; a time period and geographic expanse rich in evidence of lineage origins and dispersals of numerous mammalian lineages, including apes. Traditionally, the geographic origin of the African ape and human lineage is considered to have occurred in Africa, however, an alternative hypothesis favouring a Eurasian origin has been proposed. This hypothesis suggests that that after an initial dispersal from Africa to Eurasia at ~17Ma and subsequent radiation from Spain to China, fossil apes disperse back to Africa at least once and found the African ape and human lineage in the late Miocene. The purpose of this study is to test the Eurasian origin hypothesis through the analysis of spatial and temporal patterns of distribution, in situ evolution, interprovincial and intercontinental dispersals of Eurasian terrestrial mammals in response to environmental factors. Using the NOW and Paleobiology databases, together with data collected through survey and excavation of middle and late Miocene vertebrate localities in Hungary and Romania, taphonomic bias and sampling completeness of Eurasian faunas are assessed. -
Eomoropus, an American Eocene Chali- Cothere. by Henry Fairfield Osborn
56.9,725E (1181 :7) Article XIV.- EOMOROPUS, AN AMERICAN EOCENE CHALI- COTHERE. BY HENRY FAIRFIELD OSBORN. With Eleven Text Figures. 1. Triplopus amarorum Cope, a new chalicothere type. 2. Resemblances to Moropus. 3. Early European chalicotheres. 4. Description of Eomoropus, type skull and skeleton. 5. Relations to the Perissodactyla. 1. Triplopus amarorum COPE, A NEW CHALICOTHERE TYPE. The recognition of an upper Middle Eocene ancylopod or chalicothere in the type of Triplopus amarorum Cope is a matter of great interest: first, because it shows that a supposedly European family was established as early in America as in Europe and may have been of American origin; second, because geologically the family is carried back from the American Lower Miocene into the close of Middle Eocene times, the Washakie or Bartonian stage; third, the knowledge of considerable portions of the skeleton of this most primitive known chalicothere strengthens the relations of the Chalicotheres to the Perissodactyla. The species T. amarorum has always appeared to the present writer quite distinct from the genus Triplopu, the type species of which is T. cubitalis Cope. Dep4ret was the last author to examine the type of T. amarorum and he pointed out the resemblance of its superior molar teeth, with their interrupted anterior crests, to those of his genus Lophiaspis. But Lophi- aspis is a true lophiodont in the structure of the ectoloph of its superior molar teeth, which exhibits no mesostyle and a concave metacone. The type specimen of T. amarorum, as described by Cope in 1884 in the 'Tertiary Vertebrata', was not thoroughly worked out from the matrix so that his description included chiefly the skull and the pes. -
Eastern Georgia and Western Azerbaijan, South Caucasus)
Synopsis of the terrestrial vertebrate faunas from the Middle Kura Basin (Eastern Georgia and Western Azerbaijan, South Caucasus) MAIA BUKHSIANIDZE and KAKHABER KOIAVA Bukhsianidze, M. and Koiava, K. 2018. Synopsis of the terrestrial vertebrate faunas from the Middle Kura Basin (Eastern Georgia and Western Azerbaijan, South Caucasus). Acta Palaeontologica Polonica 63 (3): 441–461. This paper summarizes knowledge on the Neogene–Quaternary terrestrial fossil record from the Middle Kura Basin accumulated over a century and aims to its integration into the current research. This fossil evidence is essential in understanding the evolution of the Eurasian biome, since this territory is located at the border of Eastern Mediterranean and Central Asian regions. The general biostratigraphic framework suggests existence of two major intervals of the terrestrial fossil record in the area, spanning ca. 10–7 Ma and ca. 3–1 Ma, and points to an important hiatus between the late Miocene and late Pliocene. General aspects of the paleogeographic history and fossil record suggest that the biogeographic role of the Middle Kura Basin has been changing over geological time from a refugium (Khersonian) to a full-fledged part of the Greco-Iranian province (Meotian–Pontian). The dynamic environmental changes during the Quaternary do not depict this territory as a refugium in its general sense. The greatest value of this fossil record is the potential to understand a detailed history of terrestrial life during demise of late Miocene Hominoidea in Eurasia and early Homo dispersal out of Africa. Late Miocene record of the Middle Kura Basin captures the latest stage of the Eastern Paratethys regression, and among other fossils counts the latest and the easternmost occurence of dryopithecine, Udabnopithecus garedziensis, while the almost uninterrupted fossil record of the late Pliocene–Early Pleistocene covers the time interval of the early human occupation of Caucasus and Eurasia. -
Eppelsheim 2000 - New Discoveries at a Classic Locality
Jens Lorenz Franzen1, Oldrich Fejfar2, Gerhard Storch1 & Volker Wilde1 1 Forschungsinstitut Senckenberg, Frankfurt am Main 2 Charles University, Prague Eppelsheim 2000 - new discoveries at a classic locality Franzen, J. L., Fejfar, O., Storch, G. & Wilde, V., 2003- Eppelsheim 2000 – new discoveries at a classic locality - in: Reumer, J.W.F. & Wessels, W. (eds.) - DISTRIBUTION AND MIGRATION OF TERTIARY MAMMALS IN EURASIA. A VOLUME IN HONOUR OF HANS DE BRUIJN - DEINSEA 10: 217- 234 [ISSN 0923-9308] Published 1 December 2003 First scientific excavations at the classic mammal locality of Eppelsheim carried out by the Senckenberg Research Institute during the years 1996 - 2000 are described (Senckenberg research project no. 150: Franzen & Storch). They resulted in the discovery of the first flora and determi- nable micromammals (Talpa vallesensis, Plesiosorex aff. schaffneri, Crusafontina sp.) as well as a phalange of a large hominoid (cf. Dryopithecus sp.) from that locality. The excavations also deli- vered new aspects for understanding the early formation of the Rhine river system. Erste wissenschaftliche Ausgrabungen an der klassischen Säugetierfundstelle Eppelsheim durch das Forschungsinstitut Senckenberg (Senckenberg-Forschungsprojekt Nr. 150: Franzen & Storch) führten in den Jahren 1996 - 2000 zur Entdeckung der ersten Flora und bestimmbaren Kleinsäuger (Talpa vallesensis, Plesiosorex, Crusafontina sp.) sowie der Phalanx eines großen Hominoiden (cf. Dryopithecus sp.) an dieser Lokalität. Darüber hinaus ergaben die Grabungen neue Aspekte hin- sichtlich der Entstehung des rheinischen Flußsystems. Correspondence: J. L. Franzen, Jakobistr. 10, D-79822 Titisee-Neustadt, Germany, e-mail: jlfran- [email protected]; O. Fejfar, Department of Palaeontology, Charles University, Albertov 6, CZ-128 43 Praha 2, Czech Republic, e-mail: [email protected]; G. -
Hyarna and Probably Many Other Phyla, the Middle Siwalik Stage Has
19291 Matthew, Critical Observations upon Siwalik Mammals 457 Hyarna and probably many other phyla, the Middle Siwalik stage has a somewhat more primitive representative type in the Pikermi fauna, but a directly ancestral type (so far as appears) in the Chinji. This does not prove that India was the center of dispersal of these types, but that it was accessible to them both in Chinji and Dhok Pathan, more so, one would judge, than Western Europe, and less so than the present Egaean region. III. PRELIMINARY REVIEW OF SIWALIK COLLECTIONS IN INDIAN MUSEUM, CALCUTTA PROBOSCIDEA.-Dinotherium, skull. Palmer, 1924, Pal. Ind., N.S., VII, No. 4. Partial skull coll. Pilgrim, 1912, Lower Chinji. All that is saved is the basicranial region and set of teeth. Basicranial agrees so far as stated with Eppelsheim skull. No condyloid foramen. An alisphenoid fora- men. Postglenoid and posttympanic processes unite, enclosing a "false meatus" better developed than in Elephas, D. indicum and D. pentapotami.e. Type of D. indicum is part of a molar identified by Lydekker as hinder part of m'1. It is, according to Palmer, front part of m21. Probably is from Lower Siwalik beds of Dera Ghazi Khan. Type of D. pentapotamize, p3, probably Lower Siwalik, near Attock. Agrees rather closely with corresponding tooth of Palmer's skull, which is intermediate in size between Lydekker's type and D. giganteum. Such differences as there are may be regarded as indicating one rather variable species. Specimens of p4, m1 and m' from the Chinji beds also show much variability in the characters used by Lydekker to distinguish pentapotamixe from giganteum. -
Order PERISSODACTYLA – Equids, Rhinoceroses, Tapirs
Order PERISSODACTYLA Order PERISSODACTYLA – Equids, Rhinoceroses, Tapirs Perissodactyla Owen, 1848. Quarterly Journal of the Geological Society of London 4: 103–141. Upper toothrows in altungulate Radinskya (late Paleocene) and Hyracotherium (Eocene). Tentative phylogenetic tree of Perissodactyla after Beninda-Emonds, 2007. Equidae (1 genus, 4 species) Asses, Zebras p. xx Rhinocerotidae (2 genera, 2 Rhinoceroses p. xx for true horses. North America became the centre of evolution of species) true horses, which occasionally migrated to other continents. The The perissodactyls are the order of herbivorous ‘odd-toed’ hoofed descendants of Protorohippus (once called Hyracotherium; Froehlich mammals that includes the living horses, zebras, asses, tapirs, 2002) evolved into many different lineages living side by side. The rhinoceroses and their extinct relatives. They were originally named collie-sized three-toed horses Mesohippus and Miohippus (from beds by Richard Owen (1848) as a group including horses, rhinos, tapirs dated about 30–37 mya) were once believed to be sequential segments and hyraxes, although no recent authors have accepted the inclusion on the unbranched trunk of the horse evolutionary tree. However, of hyraxes in Perissodactyla. Perissodactyls are recognized by a number they coexisted for millions of years, with five different species of two of unique specializations (Hooker 2005), but their single most diagnostic genera living at the same time and place. From Miohippus-like ancestors, feature is the structure of their feet. Most perissodactyls have either horses diversified into many different ecological niches. One major one or three toes on each foot, and the axis of symmetry of the foot lineage, the anchitherines, retained low-crowned teeth, presumably runs through the middle digit. -
The Late Miocene Mammal Faunas of the Mytilinii Basin, Samos Island, Greece: New Collection - 10
ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Beiträge zur Paläontologie Jahr/Year: 2009 Band/Volume: 31 Autor(en)/Author(s): Giaourtsakis Ioannis X., Koufos George D. Artikel/Article: The Late Miocene Mammal Faunas of the Mytilinii Basin, Samos Island, Greece: New Collection - 10. Chalicotheriidae 189-205 ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien Beitr. Paläont., 31:189-205, Wien 2009 The Late Miocene Mammal Faunas of the Mytilinii Basin, Samos Island, Greece: New Collection 10. Chalicotheriidae by Ioannis X. Giaourtsakis1* & George D. Koufos2* G iaourtsakis, I.X. &K oufos, G.D., 2009. The Late Miocene Mammal Faunas of the Mytilinii Basin, Samos Island, Greece: New Collection. 10. Chalicotheriidae. — Beitr. Palaont., 31:189-205, Wien. Abstract (Griechenland). Einige der postcranialen Elemente wer den zum ersten Mal aus dieser Fundstelle beschrieben. Die In the present study, we describe several postcranial neuen Funde werden mit dem bisher bekannten Material elements ofAncylotherium pentelicum (Chalicotheriidae, dieser A rt, sowie mit dem anderer Chalicotherien-Arten Schizotheriinae) from the island of Samos (Greece), some verglichen. Die miozänen Fundstellen mit Chalicothe of which are recorded for the first time from this site. riinae- und Schizotheriinae-Funden aus dem östlichen The material is compared to the known hypodigm of the Mittelmeerraum und benachbarter Regionen werden species, as well as with other chalicotheres. The known zusammengefasst und ausgewertet. Potenzielle paläo- Miocene record of the two chalicothere subfamilies, ökologische Präferenzen beider Unterfamilien werden Chalicotheriinae and Schizotheriinae, in the Eastern kurz besprochen. Die Anwesenheit des Schizotheriinen Mediterranean and adjacent regions is briefly reviewed A. -
(Perissodactyla, Mammalia) from the Late Miocene of Bulgaria. Denis Geraads, Nikolaï Spassov, Dimitar Kovachev
New Chalicotheriidae (Perissodactyla, Mammalia) from the late Miocene of Bulgaria. Denis Geraads, Nikolaï Spassov, Dimitar Kovachev To cite this version: Denis Geraads, Nikolaï Spassov, Dimitar Kovachev. New Chalicotheriidae (Perissodactyla, Mam- malia) from the late Miocene of Bulgaria.. Journal of Vertebrate Paleontology, Society of Vertebrate Paleontology, 2001, 21 (3), pp.596-606. halshs-00067712 HAL Id: halshs-00067712 https://halshs.archives-ouvertes.fr/halshs-00067712 Submitted on 10 May 2006 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. NEW CHALICOTHERIIDAE (PERISSODACTYLA, MAMMALIA) FROM THE LATE MIOCENE OF BULGARIA 1 2 3 DENIS GERAADS , NIKOLAÏ SPASSOV and DIMITAR KOVACHEV 1 UPR 2147 CNRS, 44 rue de l'Amiral Mouchez, 75014 PARIS, France; 2 National Museum of Natural History, 1, Tsar Osvoboditel blvd, 1000 SOFIA, Bulgaria; 3 Museum of Paleontology (Branch of the National Museum of Natural History, Sofia), ASSENOVGRAD, Bulgaria Running title: Geraads et al.-Chalicotheriidae from Bulgaria 1 ABSTRACT-Few fossils have been described from the very rich localities of Kalimantsi in southwestern Bulgaria, and none from Hadjidimovo, in the same area. These deposits have produced one of the best collections of Chalicotheriidae from the late Miocene of Europe, increasing the diversity of the family, and providing the first definite evidence of the co- occurrence of both subfamilies, Chalicotheriinae and Schizotheriinae. -
Eocene-Oligocene 1 - Paradise Lost
Eocene-Oligocene 1 - paradise lost - Jarðsaga 2 - Saga Lífs og Jarðar - Ólafur Ingólfsson Háskóli Íslands Climate started cooling towards the end of Eocene Global climate during the Late Eocene was still warm, but on a cooling trend. Glaciers begun to form in Antarctica. Oligocene (“almost recent”) • The Oligocene is the third epoch of the Cenozoic, 34 -23 MY. • Global cooling marked the start of the Oligocene, an environmental shift that led to changes in the flora and fauna. Global temperature dropped as much as 10° C, shrinking forests and introducing the grassy plains that would come to dominate in the Miocene. •Herbivorous species adapted to eating grass and carnivores adapted to hunting out in the open. Both groups evolved better eyesight to see predators and prey as well as elongated limbs to increase speed over open terrain. • During this epoch archaic species of the early Cenozoic were replaced by modern herbivores, such as horses, camels and deer, and carnivores such as cats and dogs. Oligocene World Oligocene Ocean Currents When Antarctica became isolated in a South Pole position, the coastal seas of Antarctica began to freeze, changing the ocean currents dramatically. Cold water and icebergs flowed north. Warm currents circulated around the equatorial continents. Oligocene climate During the Oligocene, glacial ice covered the South Pole but not the North Pole. Seasonal sea ice may have formed there. Increased pole-equator temperature gradients. Oxygen isotope records show cooling and growth of glaciers Mammals go marine Whales evolved and diversified during the Eocene and Oligocene, and the two groups of Mysticetes (“skíð- hvalir”) and Odonto- cetes (“tannhvalir”) originated between 40-35 MY ago Mesonychide – ancestors of whales and hippos Whales evolved from wolf-like creatures which 60-55 MY ago began searching for food in the water. -
Wednesday Morning, November 3, 2004
WEDNESDAY MORNING, NOVEMBER 3, 2004 ROMER PRIZE SESSION PLAZA BALLROOM A/B MODERATORS: RYOSUKE MOTANI AND RAYMOND ROGERS 8:00 Welcome 8:15 Beck, A.: THE ORIGINS OF MAMMALIAN LOCOMOTION: NEW METHODS FOR RECONSTRUCTING POSURE IN EXTINCT NON-MAMMALIAN SYNAPSIDS BECK, Allison, Univ. of Chicago, Chicago, IL The Synapsida, composed of living mammals and their extinct ancestors, are colloquially known as the ‘mammal-like reptiles.’ The extensive fossil record captures numerous transitional forms recording the transition from Permian, reptile-like pelycosaurs to primitive therians of the Triassic. A major part of this transition involved a change from a sprawling posture to one similar to the crouched posture of living small mammals such as the opossum. Despite our understanding of the postural endpoints, the question remains: What was the locomotory posture of taxa that are phylogenetically intermediate between pelycosaurs and modern mammals? Two major notions of postural change have been proposed, both supported by functional morphologic analyses and comparison to living mammals and reptiles. One suggests that intermediate taxa were capable of a dual-gait, much like modern crocodilians. The other outlines a series of increasingly upright intermediates. Neither hypothesis has been quantitatively evaluated. Here I set up a framework for interpreting function in extinct vertebrates, and apply it to reconstructing posture in extinct non-mammalian synapsids. Linear and angular measurements were taken on the limb and girdle bones of extant iguanian and varanid lizards, crocodilians, therian mammals and monotremes, and again on fossil synapsids. Multivariate and bivariate analyses were used to correlate suites of morphologic features with posture in the living forms.