THE GENUS TRAJANA (MOLLUSCA: GASTROPODA) IN THE NEW WORLD E:t\IILY II. VOKES TULANE UNIVERSITY
CONTENTS I. ABSTRACT __ - 75 II. INTRODUCTION 75 III. ACKNOWLEDGMENTS 77 IV. SYSTEMATIC DESCRIPTIONS 77 V. LOCALITY DATA _ 83 VI. LITERATURE CITED (8" )
ILLUSTRATIONS
TEXT FIGURE 1. Radula of Trajct1Ut ctcap11lcana ( Pilsbry and Lowe) 76 TEXT FIGUEE 2. N eoteron ariel Pilsbry and Lowe 76 PLATE 1 _ 81
I. ABSTRACT off the coast of western Mexico. All species The nassarioid gastropod genus T1'ajanct are treated systematically. s.s. includes those species with a closed siphonal canal and a circular aperture, sur II. INTRODUCTION rounded by a raised p eristome. There are Those gastropods possessing a short, but four species in the fossil record of the slightly recurved, closed siphonal canal, a New World, occurring in the upper Mio circular aperture surrounded by a raised cene of North Carolina, Florida, Mexico, peristome, and a single terminal varix have and Peru, and the Pliocene of Ecuador. One presented a problem to writers for many of these four is a new species: T. ve1'ctcru years. Dall ( 1910, p. 32-33) suggested that zana E. H . Vokes, from the upper Miocene they be referred to the genus Hindsict A. Agueguexquite Formation of Mexico, and Adams, 1851, which was a troubled group represents the first record of the genus in from the start. Dell ( 1967, p. 309-310) the "Tertiary Caribbean Province," linking has summarized the problem nicely, stating: the eastern United States and the western "The name Hindsia had an uncertain intro South American occurrences. Another new duction, and has had a chequered subsequent species, from the middle Miocene Gatun history in molluscan nomenclature. Almost Formation of Paname:l., is believed tO be before it had been properly proposed it had ancestral tO the Tt'ajana s.s. group as it been listed as a synonym of Nassa-ria and possesses an open siphonal canal and den its subsequent history has been linked with ticulations within the subcircular aperture this latter generic name." Dell presented a with a raised p eristome like that of Tra resume of the tangled history of the name, jana s.s. Becaus e of the fundamental differ which will not be repeated here, but con ences this new species is made the type of cluded by naming "Hindsia ni-vect Pfeiffer" a new subgenus with the name T1rajmzct as the type species of the genus, thus re ( N erva) woodringi E. H . Vokes. moving it from consideration for the species In addition to the fossil species, there are under discussion. two Recent species of T1rajana s.s. found These species are not left without a name,
EDITORIAL COMMITTEE FOR THIS PAPER: A. MYEA KEEN, Department of Geology, Stanford University, Stanford, California DRUID WILSON, United States Geological Survey, Washington, D. C. 75 T11l,nu: St/1{/ie.r m Geolog) cmd Pctleontology Vol. 7
to11" ctcmninatz;s by Reeve ( 1844, pl. 14, fig. 59), and is a true Hindsia. Another west coast species was named "Hindsict": "Hindsict" ( N eoteron) ariel Pils bry and Lowe, 1932, type of N eotetron ( text fig. 2). This species is clearly related to the Trajcma line, but is so co mpletely bi Tc'\t figure l. Hadnla of Tmimw acaJmlcmw zarre that it seems better tO separate it as ( Pihhr) ancl Lowe). \ Iagnification appro'\i a distinct, though monotypic, genus. N eo nwkl) X 150. (From photograph h) ::\1) ra teron ariel is distinguished from the typical Keen.) Trajct7Zct by the expanded wing-like terminal varix and by a conspicuous nassarioid fossa however, for Gardner ( 19'-±8, p. 221) pro circling the base of the body whorl. The posed the genus Trctjcmct for a species of the Trajmzct species have only a slight tendency group from the upper Miocene of North toward this fossa. It may be descended from Carolina, with the monotype Trttjct7Ztt pytct. the Miocene Trctjcma z;erctcntzana) n. sp. The She was of the opinion that this new genus two species share a peculiar channeled su was most closely related to the muricid sub ture (see pl. 1, fig. 1 b) not seen in any genus Hc~drimzitl, which is placed in the other species of Trctjana. The spiral orna subfamily Tritonaliinae (sec Vokes, 1964, mentation crossing the terminal varix is p. 20), the name Trctjcma being derived markedly similar in N eoteron and T1ra jcmct) from that of the Roman emperor Trajan, suggesting that N eoteron is a derivative who immediately preceded Hadrian. The form. However, there is a peculiar lamellar writer was originally of the same opinion due to the closed siphonal canal, but ad ditional work proved the group to be most closely related to the Nassariidae. The radula of the Recent species T. ctcctpttlcmzct ( Pilsbry and Lo\ve) is nassarioid (see text fig. 1 ) and the nature of the closure of the canal is different from that in the Tritonaliinae. In the latter group a thin plate extends from the columellar wall over the siphon, meeting the outer lip at the abaxial edge. But in Tra;mzct the outer lip folds over and joins the columellar lip at the adaxial edge, so these can truly be said to be analogous structures with no relationship other than function. Trajcmct s.s. is a small group, there being only four fossil species and two Recent spe cies known exclusively from the New World. The fossil species are upper Miocene and Pliocene in age, ranging from North Caro lina ro Peru. The two Recent forms occur on the west coast of Mexico. The shell figured by Reeve ( 1841, pl. 20, fig. 96) as "Triton 7l ctuoides Gray," from the Philippines, is not that species but probably is a specimen of one of the West Mexican species of Trct jcmcr. The shell originally figured as 11Triton" 7zaJsoides, in Griffith and Pidgeon ( 1834, Text figure 2. Neoteron ariel Pilsbry ancl pl. L·11, fig. 4), is not a Trajana but is prob Lowe. ANSP 155564 ( holotype); height 9.9 mm, diameter 5.6 mm; locality: Corinto, Nica ably the species subsequently named 11Tri- ragua. X 6. No.2 Trajana in the New tV orld 77 axial sculpture more akin to the Muricidae. Family N ASSARIJDAE This is particularly well developed on the Genus Trajanct Gardner, 1948 apertural face of the terminal varix, which together with the wing-like expansion of Subgenus Nervct E. H. Vokes, n. subgen. this varix combines to give a muricid aspect Type species: Trajmza (Nen)a) woodringi to the shell. E. H. Vokes, n. sp., Gatun Formation, The genus T1tajmza is almost certainly Panama, middle Miocene. descended from a N assarius-like ancestor. Etymology: Nerva, a Roman emperor; In the Gatun Formation of Panama there gender: masculine. is a species that strongly suggests T raj ana Description: Shell scalarifonn, high-spired; in having marked axial folds, a subcircular protoconch multiwhorled, smooth; axial costae aperture surrounded by a raised peristome, strong, separated by relatively smooth inter a high spire, and a short recurved canal. spaces; numerous spiral ribs; a single varix developed at aperture; aperture subcircular, However, the canal is open and there are almost enbre, surrounded by a slightly raised typical N assa1tius denticulations within the peristome; on tbe interior of the outer lip a few aperture, on both inner and outer lips. elongate dentides and on the columellar wall several weak denticulations, together with a The genus T1ta jana has a completely smooth larger tooth at the adaptical end; siphonal canal ap erture. Because of these very basic dif slightly extended, rccurved, open by a narrow ferences this ances tral form cannot in good slit. conscience be placed in Trajana s.s.; there TRAJANA (NERVA) WOODRING! fore, a new subgenus N Mva) named for the E. H. Vokes, n. sp. Roman emperor who preceded Trajan,* is Plate 1, figs. 2a, 2b here proposed. Description: Shell moderate m size for the genus, high-sph·ed; protoconch of two and III. ACKNOWLEDGMENTS one-half smooth whorls; ornamentation begin ning gradually with small axial costae, nine on The writer wishes to express her grati the first two post-nuclear whorls, then diminish tude to Druid Wilson, U . S. Geological Sur ing to seven or eight on each succeeding whorL vey; Myra K een, Stanford University; and spiral ornamentation of small ribs, three on early whorls gradually increasing to five on R . K . D ell, Dominion Museum, Wellington, penultimate and eight on body whorl, with two New Zealand, all of whom worked to un more on siphonal canal; where spiral ribs cross tangle the Hindsia problem. She would also axial costae small nodes are formed, spiral ribs almost completely obsolete between the axial like ro thank the fo llowing persons who costae; single varix formed at apcrtural lip, re loaned material necessary to the study: sorbed to form axial costae with each growth Copeland MacClintock, Peabody M useum stage; aperture almost circular, surrounded by a raised peristome; interior of outer lip beadng of Natural History, Yale University; H. G. four strong, elongate denticlcs; columellar wall Richards and R . Tucker Abbott, Academy with several small pustules and one larger of Natural Sciences, Philadelphia; and Mrs. tooth at adaptical end; siphonal canal slightly extended, recurved, open hy a narrow slit; R. C. Hoerle, W est Palm Beach, Florida. small siphonal fasciole developed. The financial assistance of National Science Dimensions of holotype: height 14.0 mm, Foundation Grant no. G .B. 6048 is grate diameter 8.0 mm. Holotype: USNJ\1 646229. fully acknowledged. Type locality: TU 757, roadcnt on south side of Boyd-Roosevelt Ilighway at junction of IV. SYSTEMATIC D ESCRIPTIONS road to "Refineria Panam
curved, open siphonal canal. The Trttjcmct the periphery and later either 1 or 2 lirae in front of the p eriphery, those nearest the sutures characteristics are the strong axial costae, the least conspicuous; primaries on the body the circular, almost entire aperture, sur whorl 8 or 9 including the 2 on the shoulder; rounded by a raised peristome, the terminal finer secondaries commonly introduced mid varix, and the very high spire. There are way between the primaries, that at the base four specimens in the type lot, all from the of the hody approximating in strength the primary behind it. Anterior siphonal canal same general area, which is approximately closely sculptured with 9 or 10 sharply ele locality 138b of Woodring (1959, p. 148) . vated lirae. the 1 or 2 posterior spirals set a The geographic position of this ancestral little apart from those in front of them. Si phonal canal cut off from the base of the body form, in the Panamanian area, is ideal from whorl by a convex area that is strongly rip the standpoint of subsequent distribution pled by the axial costae but not spirally both north to Mexico and the United States th readed; delicate filamentary incrementals and south ro Peru and Ecuador. visible under magnification in the interspiral areas. Suture line distinct, impressed, crenu latecl by the axials of the preceding whorl. Subgenus TRAJANA s.s. A narrow shoulder in front of the suture Trajana CAHD'\EH, 1948, U. S. Ccol. Surv. Prof. strongly defined, persistent almost to the apex; Paper 199, p . 221. concave near the aperture, sloping and feebly undulated by the axials on the earlier volu Type species: Trajana pyta Gardner, by tions and occasionally threaded with a single original designation. fortuitous spiral near the anterior margin. Aper ture elliptical, smoothly and evenly glazed Description: "Shell more or less scalarifonn withi11 , with no trace of any angularity either in outline. Spire elevated. Protoconch small, un anteriorly or posterioly nor of any dcnticula sculptured. A\.ial costae prominent and some tion or rugosity on either the outer or the what varicose hut not foli ated or spinose. Spiral inner lip. Labrum strongly varicose, the varix sculpture dense. Anal fasciole strongly arched . obtuse and prominent, encroaching posteriorly Aperture regularly elliptical in outline; inner on the preceding whorl. An terior canal closed , lining continuous. Outer lip varicose, nonlirate short, and rather feebly recurved. Umbilical within. Inner lip evenly concave, nonplicate. chink not visible." (Gardner, 1948.) Anterior canal closed, short, and rather feebly Dimensions of holotype: height 17.3 mm, recun eel. Umbilical chink closed." (Gardner, diameter 10.8 mm. 1948) Ilolotype : USN1\f 497151. Type locality: Natural Well, two miles south TRA JAN A ( TRA J AN A) PYTA Gardner west of l\1agnoli a. D uplin County, North Caro Plate 1, fi gs . 3a, 3 b lina ( = TU 376 ). Occ urrence : Duplin 1\ Iarl, North Carolina; Tmjana puta CAHD);"EH, 1948. U . S. C eol. Surv. Jackson Bluff Formation and Pinecrest Beds, Prof. Papcr 199, p . 221, pl. 29, fi gs. 10, 11. F lorida; upper l\ h ocene. II indsia puta (Gardner ) . 0Lssox, 1967, T er Figured .sp ec im en: U SN1I 646230; h eight tiary mollusks south F lorida and Caribbean, 16.4 mm, diameter 9.2 rnm; locality TU 730. p. 3.5, pl. 6, fig. 8. Other occurrences TU locality nos. 60, 523, Description: "Shell scalarifonn, spire ele 729, 932, 933. ,·atecl, apex acute, body h road and relatively low, abruptly constricted at the base. W horls Discttssion : Although none of the species strongly convex, tabulated posteriorly, rapidly of Trctjana is common1 T. pyta is the rnost increasing in size, approximately 7. Proto nearly common and widespread of the conch of J ~~ to 2 small, unsculptmecl coils, group. The species was described from the earlier largely immersed . Axial sculp ture N atural Well, North Carolina, and was initiated at the beginnin g of first turn of conch with obtuse vertical rihlets that rapidly said to be based on two specimens. No speci develop into hroad, rounded, strongly ele mens have been collected by the writer at vated, undulatory costals ; 7 or 8 on the this locality ( = TU 3 7 6 ) . A single speci whorls of the spire, equal in size and spacin g men, in the collection of Mr. Paul Huddle C'\<.:ept on the fin al half turn, most prominent ston, of New Orleans, has been found at on the periphery of the whorl and tending to alternate in arrangement. Spiral sculpture of Jackson Bluff, Ochlochonee River, Florida sharply elevated lirations that override the ( = TU 60 ) , type locality of the Jackson costals and are equally promin ent on the Bluff Formation. Another single specimen costal and intercostal areas; spirals initiated was collected by the writer at TU 523, simultaneously with the axial sculpture by the appearance of the 2 lirae that crown the which represents the youngest known oc periphery of the whorl; first one, and then currence of the genus in the United States, a second finer thrcadlct introduced b ehind for this locality is in the uppermost level No. 2 T1rajctna in the New World 79
of the Pin~crest Beds, immediately below Discussion: At the tlme Spieker re the overlymg Caloosahatchee Formation. described the Nelson Collection of fossils This is approximately the same locality from the vicinity of Zorritas northern Peru where Olsson ( 1967, p. 35) noted that the exact locality for these ' specimens was' specimens had been found by Hughes. Only not known and Spieker noted ( 1922, p. along the Kissimmee River (or Canal) in 12-13): "The matrix in which most of the the vicinity of Fort Basinger, Florida (TU specimens so far collected [by J. T. Singe 7 ~0 & 932 ) , is the species really abundant, wald, Jr.] occur is far too resistant to per wlth over 200 specimens collected. A few mit the extraction by any known means of specimens have been found at the northern specimens at all susceptible to injury through n1ost end of TU 729, in beds correlative to moderate violence; . . . Loose matrix on those farther north at Fort Basinger, and a some of the specimens in the Nelson col few have been taken at TU 933, which is lection indicates that there are probably south of Lake Okeechobee, on "Alligator some localities at which the sands have not Alley." All of these occurrences are in the been cemented." Olsson subsequently re older strata of the Pinecrest Beds. located the Nelson locality ( 1932, p. 7 & 30) and observed that it was actually a TRAJANA (TRAJANA) LAQUEORATA down-faulted block of upper Miocene (Spieker) Tumbes Formation that carried Nelson's Plate 1, fig. 5 fossil fauna and not part of the Zorritas M urex laqueomtus SPIEKER, 1922, Johns Hop Group as supposed by Spieker. Originally kins Stud. Geol. , no. 3, p. 51, pl. 2, fig. 4 spelled "Tmnbez" by Olsson, the correct ( spelled laq uoratus in plate expl.). spelling is now "Tumbes" according to the M urex laqueoratus Spieker. HANNA AND IsRAEL Lexique Stratigraphique International ( 1956, SKY, 1925, California Acad. Sci., Proc., ( Ser. 4 ) v. 14, no. 2, p. 53. v. 5, fasc. 5b, p. 110). M urex laqueoratus Spieker. OLssoN, 1932, Although the siphonal canal on the unique Bulls. Am er. Paleontology, v. 19, no. 68, type specimen of T. laqueorata is open, this p. 31. seems to be the result of breakage and there D escription: "Sh ell small, turreted, fairly is little doubt that in life the canal would stout. Whorls ab out six in number, the first two smooth, the remainder delicately sculp have been sealed. The alternating large and tured and convex. Sculpture of regular longi small spiral ribs and the smooth aperture tudinal folds, eight to the whorl, which are indicates a closer alliance with the typical negligible at the suture, increasing rapidly to full strength at the mid-line of the whorl, Tttajana group than with the ancestral Nerva and decreasing again. On the body whorl all form. except the last two continue over the base as narrow rounded folds; the last two die TRAJANA (TRAJANA) VERACRUZANA out just b elow the shoulder. Spiral sculpture of fine threads, about six to the whorl, with E. H. Vokes, n. sp. occasional interstitial threads above or below Plate 1, figs. 1a, 1b, 1c the p eriphery of the whorls; there the two central threads are prominent, forming a band Description: Shell small for the group, high which h eightens the prominence of the nodes spired; protoconch of two and one-half, smooth, on the longitudinal ridges. On the body whorl bulbous whorls, seven post-nuclear whorls in there are twelve primary threads between the adult; cancellate ornamentation beginning grad b and and the base, with occasional interstitial ually and consisting of axial costae and spiral threads. Aperture almost circular, smooth ribs; twelve axial costae on the first post within, with a raised lip and a fan-like flare nuclear whorl, decreasing to about eight on bordering the outer lip over which the spiral medial whorls then increasing to about twelve sculpture continues to the raised peristome. again on the penultimate but becoming almost The canal is very narrow, open and short. completely obsolete on body whorl; three spiral Columella almost straight." (Spieker, 1922) ribs on first whorl gradually increasing to Dimension of holotype: height 17.8 mm, twelve major ribs with several smaller inter diameter 11.3 mm. calated riblets on the body whorl and an ad Holotype : Yale Peabody Museum no. 502. ditional eight riblets on the siphonal canal; Type locality : Near mouth of Quebrada on the medial whorls the two ribs at the Tucillal, just north of Zorritas schoolhouse, periphery larger than those above and be Zorritas, Peru ( Olsson, 1932, p. 7). low, becoming approximately the same on the Occurrence: Tumbes Formation, Peru; upper body whorl; the ril) at the shoulder greatly Miocene. raised causing suture to be deeply set into a Figw·ecl specimen: YPM 502 ( holotype). narrow channel formed by the rib at the 80 Tulane Studies in Geology ctnd Paleontology Vol. 7
shoulder and the peripheral rib of the preced represent the uppermost Pinecrest strata. ing whorl; one terminal varix at aperture with The appearance of an Agueguexquite fossil a deep excavation behind it; aperture circular, entire, surrounded by a raised peristome, in this relative position would suggest that smooth on the interior; siphonal canal short, perhaps the Mexican formation is even recurvccl, closed. younger than previously believed. On the Dimensions of holotype: height 12.0 mm, basis of the n1uricine species the writer has diameter 7.2 mm. IIolotype: USNM 646231. considered the Agueguexquite to be early Type locality: TU 638, roadcut and quarry upper Miocene in age. Correlation with the on J\Iexico Highway 180, 14 mHes east of Brighton beds would indicate a late upper junction with side road into Coatzacoalcos, Miocene age for the Agueguexquite Forn1a Veracruz, J\Iexico. Occurrence: Agueguexquite Formation, Mex tion. According to Mr. Wilson (personal ico; Pinecrest Beds, Florida; upper Miocene. communication) there are also other species Figured specimen: USNM 6462.31 ( halo in common between the Adine beds and type). the Agueguexquite. Discttssion: The type lot of this small In a paper on the Panama land bridge as species consists of two and one-half speci a sea barrier, Woodring ( 1966) noted that mens from the Agueguexquite Formation of there are a number of genera originally Mexico. The form is unique in having a found in the western Atlantic region now deeply channeled suture (see pl. 1, fig. 1 b ), extinct there and surviving only in the more pronounced than that in any other eastern Pacific today. The geologic record species of the group. A tendency toward of n1igration of these "paciphiles," as he this type of suture is seen in the other spe termed them, is often lacking and as an cies. only immediately behind the terminal example he stated ( 1966, p. 429) : "Miss vanx. ing records are illustrated by the occurrence In addition to the type lot, in the U. S. in Florida and North Carolina of a paciphile Geological Survey collections made by Druid ( T1rajana) that is so far unknown in the Wilson at Adine, Charlotte County, Florida western Atlantic part of the Tertiary ( = TU 200) , there are two specimens of province, although the only rational explana T. 11eracrztzana. The fauna at Adine is tion is that it migrated through that part." equivalent to that of the so-called "Brighton The writer is pleased to be able to fill in Beds" near Lake Okeechobee, and seems to one gap in the record.
PLATE 1 (all figures X 4) Figures Page 1. T rajana ( T rajana) veracruzana E. H. Vokes, n. sp. ______------7 9 USNM 646231 (holotype); height 12.0 mm, diameter 7.2 111m. Locality: TU 638. Agueguexquite Fm., upper Miocene. 2. Trajana ( N erva ) wood1ringi E. H. Vokes, n. sp. ______77 USNM 646229 ( holotype) ; height 14.0 mm, diameter 8.0 111m. locality: TU 757. Gatun Fm., middle Miocene. 3. Trajana ( Trajana) pyta Gardner______. ______7 8 USNM 646230; height 16.4 mm, diameter 9.2 mm. locality: TU 730. Pinecrest Beds, upper Miocene. 4. Trajana (Trajana) wheeleri (Pilsbry and Olsson ) ______82 ANSP 14483 ( holotype ); height 16.5 mm, diameter 9.5 111m. locality: Punta Blanca, Ecuador. Canoa Fm., Pliocene. 5. Trajana ( Trajana) laqtte orata (Spieker) ______79 Yale Peabody Mus. no. 502 (holotype); height 17.8 mm, diameter 11.3 111m. Locality: Zorritas, Peru. Tumbes Fm., upper Miocene. No.2 Trctjcma zn the N en· World 81 1a
1 c
2a 3a
2b
4a
PLATE 1 82 T11lrt11e St;;d/es In Geology cmd Paleontolog)' Vol. 7
TRAJANA (TRAJANA) WHEELER! six slightly protractive axial ribs, strong and rounded, following each other up the spire ( Pilsbry and Olsson) in a gently receding spiral, with much wider Plate 1, figs. 4a, 4b interspaces, most prominent at the p eriphery of the whorls, but traceable to the canal; these II iJl(lsia u;heeleri PILSHRY A'\D 0Lsso~, 1941, are crossed between the sutures by six rounded, Acacl. 1\at. Sci. Phila., Proc., v. 93, p. 29, subequally spaced spiral threads with much pl. 5, fig. 10. wider interspaces ( the space b etween the Description: "Shell small, solid; whorls about posterior thread and the su ture widest of all) , H of which the first 2 are nuclear. smooth, a little swollen where they override the ribs; except the last quarter turn which has weak on the last whorl finer intercalary threads ap spirals~ the post-nuclear whorls are strongly pear, and on the canal are seven or eight sculptured by ribs and spirals; the sculpture stronger, spirally striated cords; ap erture sub of the spire-whorls consists principally of 2 circular, not lirate within, with a produced, strong spiral cords which form the middle of simple peritreme, and behind it a wide, strong, the whorls, these spirals are crossed, and rounded varix over-ridden by the spiral sculp rendered strongly nodose, by axial ribs; on ture; canal short, closed when adult, pointed the shoulder area, there are 2 smaller spirals and strongly recurved. The color of the shell \Vith 2 others on the area between the lower is mostly yellowish-white, with a basal and suture and the lower of the 2 primary middle sometimes a sutural hrownish spiral band, spiral cords; the sculpture is similar on the which shows more or less in the interior of body-whorl with 4 spirals on the base and the aperture." ( D all, 1910 ) about 8 strong ones on the beak; there are Dimensions of holotype: h eight 28.5 mm, 9 ribs on the last whorl beside the enlarged, diameter 15.0 mm. thickened lip; on the last whorl the sutural Holotype: U SNM 96658. zone becomes a deep, smooth channel which Type locality: off La Paz, Baja California, unites with another channel behind the outer ~1ex i co, in 21 fms. lip; aperture is small, ovate, continuous with Occurrence: Recent only, Gulf of California a raised, inner rim; outer lip doubling back from La Paz to Guaymas, 20-30 fms. on itself, forming a wide varix, more or less hollow within; beak long, recurved at the Discussion: The two Recent species of tip and canying the anterior canal." ( Pilshry Trajana found on the western coast of Mex and Olsson, 1941) Dimensions of holotype: height 16.5 mm, ico are much alike. T. pe1rideris is a lighter diameter 9.5 mm. color than T. acapulcana and has fewer axial Holotype: ANSP 1448.3. ribs. The spiral rib at the periphery is Type locality: Punta Blanca, Ecuador. stronger, giving the shell a marked angu Occurrence: Canoa Formation, Ecuador; Pliocene. lation almost exactly at the mid-point. In Figured .specimen: ANSP 1448.3 ( holotype). some specimens there is a white spiral color band that corresponds to this peripheral Discussion: As noted by Pilsbry and Ols angulation. son this species is closely related to T . acapulcana (Pilsbry and lowe) from the TRAJANA (TRAJANA) ACAPULCANA Recent fauna of western Mexico. The fos sil ( Pilsbry and lowe) form differs in having a shorter spire and H indsia acapulcana PrLSBHY A ND LowE, 1932, more inflated body whorl. The other dif Acad. Nat. Sci. Phila., Proc., v. 84, p. 66, ferences cited by the original authors, such pl. 7, fig. 11. as the heavier lip, entire peristome, and Hindsia ( Hindsia ) acapulcana Pilsbry and closed canal, are due to the fact that the L owe. K EEN, 1958, Sea sl1 ells of tropical type of T. ctcapttlcana is a juvenile specimen West America, p. 404, fig. 553. with an incomplete aperture. Description: "Shell of about 91h strongly convex whorls, of which two are nuclear, TRAJANA ( TRAJANA) PERIDERIS smooth and white. Color chestnut brown, with a darker subsutural band, the intervals of ribs (Dall) and much of base cartridge buff. Axial ribs II indsia perideris DALL, 1910, Kautilus, v. 24, rounded, crossed on the penultimate whorl by no . .3, p . .32. six spiral cords, the two on the p eriphery b e II indsia ( H inclsia) ]Jerideris Dall. KEE~, 1958, in g stronger, swollen a little where they cross Sea shells of tropical West America, p . 404, the axial ribs; on the body whorl may b e fig. 554 ( holotype). counted about twenty spiral cords of varying strength, eight of which arc on the canal, Description: "Shell of about ten whorls, the which is recurvcd and almost closed. Outer nuclens smooth white of about two whorls, the lip thin, doubling back on itself forming a subsequent whorls (except the last ) slowly wide hollow varix, crossed by the spiral sculp enlarging. forming a slender acute spire, the ture, which is here emphasized." ( Pilsbry and last whorl suddenly larger; sculpture of about Lowe, 1932 ) No.2 T raj ana MZ the N etu U7 orld 83
Dimensions of holotype : height 17.5 n1n1, Basinger ( SE 111 Sec. 20, T.36S, R.33E), c1 iamet er 9 .7 mm. Okeechobee Co., Florida. Ilolotype: ANSP 155.334. 9.3.3. Pinecrest Beds, material exposed during . Type locality: off Acapulco, Guerrero, Mex construction of "Alligator Alley," 21 ..5 miles lCO, in 20 fms. east. of Florida Highway 29, Collier Co., Occ urrence: Recent only. off southwestern Flonda. Ivfexico, from Acapulco to Gulf of Tehuantepec, 958. Gatun Fm., hillslope on east side of road 20 to 40 fms. from Boyd-Roosevelt Highway to "Refineria ~anan:
July 16, 1969