Newsletter No. 28

INTERNATIONAL ORGANIZATION OF PLANT BIOSYSTEMATISTS •

NEWSLETTER NO. 28

ED ITED BY L. Borgen & B. Jonsell DJ. Cra wford & CA. St"( c

Oslo 1997 ...... _.. _. ..

.... ~ ~ . ~-. '" . , .."

IS ED FROM The Bergius Foundation, Royal Academy of Sciences, Stockholm Botanical Garden & Museum. University of Oslo

"/listrations: Kro ' & Almquist. Svensk Flora . ed, B, & I. lonsell. 27 th ed, 1994 Btidtke r. ,\<\" Norsk Flora, Illustrasjonsbind. 1970 Lindman. CA,M,. Bilder (Jr Nordens Flora, 1922 Curtis's Botanical Magazine, 1834

Graphic de it;n c", layout: Xtina Wootz

Printed by: GCS t\S, Oslo, Norway, t 997

ISSN 0254-8844

IOPB C ' The Authors 2 ~ OCT. 1997

IOPB NEWSLETTER NO. 28

I£!J. J; .RMN BOT !;m"'o c. S. I . C. lUBLJOTECA Contents ADQUmlOD EX

1 Editors' Column 3

2 News & Notes 4

3 Requests 5

4 The Profiles 6 Biosystematics in Gbteborg and Lund, Sweden today • 5 IOPB Chromosome Data 12 13 6 News from Molecular Biosystematists 8 20

7 Meetings - Past and Future 23 VII IOPB Symposium, Amsterdam, Netherlands Other meetings Valdivia, Chile Harare, Zimbabwe Paris, France Uppsala, Sweden 8 Points of View 26 , 9 IOPB Executive and Council 1995-98 28

10 Changed Addresses & New Members 30

Editors' Column When you read this the proceedings of the IOPB symposium in Troms0 should be out. Dear IOPB members, We regret that it has taken a long time -in Also this ewsletter has a spite of all technical and electronic facilities 1 pronounced Scandiavian fla- of today publishing times seem rather to go vour, not only through its the other way. You will find more about the editors which for the time Proceedings under "2. News and Notes" . being is inevitable, but more importantly The first circular for the IOPB symposium tluough the contributions this time found in Amsterdam August 1998 was sent out in under the standing heading "The Profile". In the spling. You find some additional infonna this issue biosystematic activities- in the tion under "7. Meetings". Before that meeting broad sense we would like to take it - are elections oflOPB executives and council will presented from ' nstitutions at the univer have to take place. With the next Newsletter, sities of Goteborg (Gothenburg) and Lund. probably in January 1998, voting fonns will Although we are sure that much more of this be sent out. Now already we would like to kind can be told from Scandinavia we would request you to be active in that matter - do very much like to see examples from other vote, and do before the 1st of January send in parts of the world as well. This holds true for nominations to the President. In this News the "News & I otes" too, where we have re letter you will also find Executives and Coun ported about some fresh Scandinavian the cil members - with updated addresses - and ses. Similar infonnation, particularly about you may make your opinion about the books and theses, would be welcome from all composition. In Amsterdam important de over the world . You may well e-mail con cisions about the future of IOPB have to be tributions to [email protected] - also as taken - see previous Newsletter under "8. attached documents. Points of View" and a few comments in this From all we have heard the new face and ~ one. We eagerly await more comments from layout of the IOPB ewsletter was very well members upon this important subject! received - contrary views have at least not With all best wishes and a hope that very reached us. We editors would like to express many of you are sending in the forms for taking oLir thanks to Xtina Wootz, Uppsala Univer part in the Amsterdam meeting. sity Library, who put such inspired eff0l1s into our Newsletter and continued to do it for the one you now have at hands. Liv Borgen & Bengl }onsell 3 News & Notes habitat selection. 39 pp. Lund 1997. Pub lished together with 5 papers printed or to be printed in international journals, three 2 Recently published of them with H. C. Prentice as co-author.

IOPB Proceedings Three doctoral theses with biosystematical The proceedings of the VI International themes have in June 1997 been defended and IOPB Symposium held in TromSI1 in 1995 published at Swedish universities, viz. have now appeared as Opera Botanica Vol. 132. The ordinary price is 750 Danis.h Crowns Black-Samuelsson, Sanna. 1997. Genetic (approximately USS 108); Individual IOPB variation and phenotypic plasti city in the members at good standing may buy it at a rare plant species Vicia pisiformis L. and V reduced price DKK 550. You may mail your dumetorum L. (Fabaceae). Acla Uni\'. Agri order to address "[email protected]". It s culturae Sueciae. Silvestria 30. 37 pp. ISBi\ title corresponds to the symposium theme, 91-576-5314-3. Summary published 10- "Variation and evolution in arctic and gether with 5 papers printed or to be print alpine plants". The contents are as follows. ed in international journals. Co-authors to une or more of those papers are S. Anders DIVERSITY AND SPECIATIO:'< son, Lund and G. Eriksson, L. Gustafsson, Murray. D.F. Regional and local vascular P. Gustafsson, M. Lascoux & K. Lundkvist, plant diversity in the arctic. Uppsala. Petrovsky, V.V. Areas of intensive plant speciation in the Beringian arctic shell. Lennartsson, Tommy. 1997. Demography, Yurlsev, B.A. Analysis of evolutionary dif reproductive biology and adaptive traits in ferentiation in some key arctic-alpine Genl,ianella campestris and G. amarella. taxa: Dryas, OX)'lropis sect. Arctobia. and Evaluating grassland management for con Tara :r;acum. sect. Arctica. servation by using indicator plant species. Stace, C.A., Cornall, RJ. & Ying, Shi. Acta Un iv. Agriculturae Sueciae, Agraria Cytological and molecular variation in 46.47 pp. ISBN 91-576-5253-8. Summary apomictic Hieracium sect. Alpina. published together with 6 papers p,;nted or Bayer, RJ. Antennaria rosea (Asteraceae) - to be printed in interllational journals. Co a model group for the study of the evolu authors to one or more of those papers are tion of polyploid agamic complexes C. Oostermeijer, J. Van Dijk and H. van der Nijs. Amsterdam, P. Nilsson, Lund, J. REPnODUCTIVE STRATEGIES Tuomi, Oulu & R. Svensson, Cppsala. Tikhmenev, E.A. The reproductive features of the northern angiosperms as a factor of Rlmyeon, Helena. 1997. Variation in Si plant diversity and community stability. lene vulgaris and S. unif/om (Caryophylla Nordal, I., Hestmark, C. & Solstad, H. ceae): genetic d iv-ersity, gene flow and Reproductive biology and demography of

4 Papaver radicatum - a key species in Nor . rbanska, K.M . Restoration ecology of alpine dic plant geography. allCI arctic areas: are the classical concepts Philipp, M. Genetic diversity, bTeeding sys of niche and succession directly applicable? tem and population structure in Silene Crawford, R.M. 1. & Smith, L.c. Responses acaulis (CaryophyUaceae) in West Green of some high arct.ic shore plants to variable land. lengths of growing season. Pickering, C.M. Reproductive strategies and Wookey, P.A. & Robinson, C.H. Respon const.raints of alpine plants as illust.rated siveness and resilience of high arctic eco by five species of Australian alpine Ranun systems to em·ironmental change. culus. Bergstrom, D., Selkirk, P.M., Keenan, H.M. & Wilson, M. E. Reproductive behaviour of Requests ten flowering plant species on subantarctic Macquarie Island. ... from Kudo, G. Sex expression and fruit set of an Dr. Jan Kirschner andromonoecious herb, Peucedanum Tnulti 3 viltatum ( mbelliferae) along a snowmelt gl·adient. Institute of Botany, Academy of Sciences. Khodachek, E.A. Seed reproduction in arctic CZ-25243 Pruhonice 1, Czech Republic. environments. Fax: +420 2 6775 003L E-mail: [email protected]

MOLECULAR APPROACHES Bachmann, K. uclear D A markers in plant There are two projects based on a worldwide biosystematic research. study of live plant material. They concern Crawford, D.]. Molecular markers for the Juncaceae (a taxonomic monograph within study of genetic variation within and be- t.he framework of IOPI - Species Plantarum tween populations of rare plants. project, A Flora of the World, 1997-1999) Warwick, S.I. & Black, L.D. Molecular phylo- and the genus Taraxacum (a cpDNA study genies from thoery to application in Brassica of tlle evoluti on \vithin the genus). Any mate and allies (tribe Brassiceae, Cruciferae) rial, even a ~ing l e specimen with viable seeds, or merely seeds from various parts of the world ECOLOGY - NATURAL CONSTRAINTS AJliD would mean a great help for the success of HUMAN IMPACT the projects. Please send the seeds or plants Razzhivin, V. Yu. Life form adaptations of • to the address given above. vascular plants to the most extreme en vironments of the Asian arctic Galen, c., Stanton, M. , Shore, J.5. & Sherry, R.A. Source-sink dynamics and the effect of an environmental gradient on gene flow and genetic substructure of the alpine buttercup, Ranunculus adoneus. The Profiles: Biosystematics in key). Her projects during later years have concerned evolution, systematics, karyology Goteborg and Lund, Sweden today (including C-banding), and reproductive Goteborg biology of bulbous and tuberous plants in K. Persson & B. ErikseT/ the East Mediterranean and SW Asia, above 4 all Colchicum, Tulipa and Fritillaria. Hy In Goteborg studies related to bridization, important as part of the specia t·raditional biosystematics tion processes in some of ,these genera. is in started in the mid 1970s with two projects vestigated in spontaneous as well as artificial focusing on the families Li iaceae s. lat. and hybrids by means of e.g. chromosomal me Papal'eraceae. In 1974 Karin Persson moved thods which will be complemented by gen to the University of GOteborg., Department of omic in situ hybridization techniques. Ira" Systematic botany, from Ihe Institute of Sys nian species of the genus Fritillaria are also tematic botany at Lund, a centre of biosys subject to studies of morphometry, cytology tematic research since the 19605. At the Uni (C-banding) and reproduction by a doctoral versity of Lund her main project concerned student coming from Iran, Gholarnreza Bakh the Artemisia maritima complex in Europe shi Khaniki. More recently cooperation with and included a number of classical biosystem two international projects has been initiated, atic methods such as chromosome studies, "Bulbous and tuberous plants of Greece" analyses of morphological vaJiation within (Botanical Institute and The Royal Veteri populations, crossing experiments, and repro nary and Agricultural University, Copen ductive studies (thesis for Ph.D. presented in hagen, in cooperation with staff from Lund 1974). In Goteborg she switched to monocots, and Patras, Greece) connected to the ongoing stal1ing a project on liliaceous plants in co Flora Hellenica project and intended to in operation with Per Wendelbo, then Director d ude morphological, karyological and mo of the Botanical Garden in Goteborg and an lecular aspects; furthermore "Generic bound expert on the systematics of SW Asian plant aries in Colchicaceae" with Joan Pedrola genera with a special interest in bulbous MonfOli and other researchers from Jardin plants. Their main scientific approach was Botanic Marimurtra, Blanes and Javier generic problems in the family Hyacintha fuertes-Aguilar, Real Jardin Botanico, Mad ceae. A lIumber of genera were studied with rid, Spain, which will include analyses of an emphasis on karyological aspects, resulting genetic variation by means of isoenzymes, in a series of publications, e. g. on Bellevalia, RAPDs, karyology and morphology. The lat HyacinclllLs, Alrawia (a genus described by ter project is moreover connected to a recent the two coworkers), and Hyacinthella. After ly developing project "Phylogenetic analysis Per \Vendelbo's untimely death in 1981, Ka of the family ColclIicaceae" at the Department rin Persson continued on her own or in co of Botany, University of Austin, Texas. operation with Jimmy Persson, with whom she Biosystematic research in the subfamily has travelled 011 a number of extensive re Fumarioideae of Papaveraceae was started by search expeditions above aU in the East Magnus Liden in the mid 19705 with work on Mediterranean area (mainly Greece and lnr- a synopsis of the subfamily and a monograph

6 of the tribe Fumarieae (thesis present ed in A biosystematic project to gain knowledge 1986), and continued by Aslog Dahl with a about the reproducti ve biology of the genus th esis (1990) on the genus HJpecoum. Both Campanuta was initiated by Yvonne Nyman made use of morphologic and cytologic studies in the mid 19805. The in ves tigation was based of Vel} large living materials, and the latter on studies of £lower morphology and £lower included extensive artificial crossing experi development. and a large number of crossing ments. Aslog Dahl has later turned to studies ability tests as well as pollen germinability on tbe evolution of breeding systems and tests. A detailed study on the fun ction of the reproductive strategies of wind-pollinated pollen collecting hairs was also made. From trees, e.g. Bel.u.la penaula, Corylus avellana the results obtained conclusions were drawn and Ulmus glahra. . Magnus Lid6n on the other about reproductive strategies within species. hand ha~ continued work in the Papaveraceae, The projected was completed with a Ph.D. concentrating on the large genus Corydalis (c. thesis in 1994. 4.30 species), at first the tuberous species Fanny Astholm is about to complete her whi ch have only recently been published in th esis on the South American, Andean genus the form of a book together with Henrik Zet A/ansaa (Scrophulariacae). The genus com terlund of the Botan.ical Garden, Gm eborg. prises about 16 species of perennial herbs and Most of the other groups are planned to be subshrubs. The herbaceous species. in par monographed during the years to come in ticular, are widespread and highl y variable cooperation with Su Zhi-Yun, Kunming. Mag morphologicall y, and bios),stemati c studies nus Lid6n and c workers have also devot ed have been used ext ensively to solve some of studies on diverse phyloge netic aspects of the taxonomi cal problems. Divergence in th e Fumarioideae in general and the genus position of the reproducti ve organs within the Corydalis in particular by mean.s of e.g. ITS fl ower and thereby reproducti ve isolation is and the plastid gene rps 16 intron. likely to be the reason for the ongoing spe Molecular studies also comprised the main ciati on within th e complex studied. By con bulk of Bengt Oxelman's studies in the genus ducting extensiv e greenhouse exp eriments Silene (thesis 1995). Phylogenetic reconstruc she also showed th at naturally occurring pink tions in the S. sedoides group were based on fl owered plants are hybrids between red- and RAPD patterns, sequences from the ITS sec- white-flowered species, and that the hybrid tions of nuclear rO NA, as well as on mor- has lowered fertility. phological data. Together with Magnus Lid6n, In 1994, Bente Eriksen was granted a Oxelman also used rON A sequences to con- post-doctoral fe llowship by the Swedis h struct a phylogeny for selected OT . s in the ~ I alural Science Research Council (NFR) and tribe Si1eneae, comparing the results with initiated studies on the genus Potelllilla structural data, with the aim to circumscribe (Rosaceae) in Fairbanks, Alaska , SA. Since natural genera and sections. These s tudies the spring of 1996 she has been employed as have recently been complement ed by a phy a research associate at the depal1ment. A logeny of the rps 16 intron in the same tribe. number of species of POlelltilla have been Bengt Oxelman has now moved to the l ni shown to be facultative apomi cts and, as such. versity of Stockholm. are obvious cases for biosystematic inves- 7 tigation. Presence or absence of apomixis is Dahl, A.E. 1989. Taxonomic and mor established for the various species under phological studies in Hypecoum sect.. study and, in case oCapomixis, the relation H)pecoum (papaveraceae). - PI. Sysl. Evol. between sexual and asexual reproduction is 163: 226--280. quantified within individuals by using ad Dahl, A.E. 1990. Infrageneric division of vanced microscopic techniques and molecu Hypecoum L. (Papavera.ceae). - Nord. J. Bot. lar markers for paternity analyses. Apomixis 10: 129-140. and ploidy level variation seem to interact Dahl, A.E. 1992. Artificial crossing experi and the mechanism is under investigation. ments in Hypecoum L. sect Hypecoum Autopolyploidization in diploid sexuals may (Papaveraceae). Nord . J. Bot. 12: 13-30. trigger both development of apomixis and Dahl, A.E. 1993. Hypecoum L. In: Flora ploidy level variation within populations. [n Europaea, vol. 1 (2nd ed.), pp. 302-303. situ hybridization techniques will reveal the Dahl, A.E. 1997. H)pecoum L. In: Jonsell, B. presence of autopolyploids in natural popu & al. (eds.), Flora ordica, vol. 1 (in press). lations. Allopolyploids resulting fTom hy Dahl, A.E. & Fredrikson, M. 1996. The time bridization are often postulated to be the main table for development of maternal tissues cause of taxonomical problems within Potent sets the stage for male genomic selection in ilia. The role of hybridization will be studied Betulapendllla. - Amer. 1. Bot. 83: 895-902. by means of artificial interspecific crossing Dahl, A.E. & Strandhede, S.O. 1996. programs and molecnlar leehniques. Predicting the intensity of t.he birch pollen season. - Aerobologia 12: 97-lCl6. SCtEi\.IFlC PUBLICATIONS MAli\LY FRmI THE Dahl, A.E., Wassgren, A.-B. & Bergstrom, G. 1990'S 1990. Floral scents of Hypecoum L. sect. Astholm, F. & Nyman Y. 1994. ?I'lorphomet H'ypecollnl (Papaveraceae): Chemical com ric variation in the Alonsoa meridionalis position and relevance to taxonomy and complex. - PI. Syst. Evol. 193: 53-68. mating system. - Biochem. Syst. Eco!. 18: Bakhshi Khaniki, G. 199.5. Meiotic studies 157-168. on some Iranian Cen.lal/rea (Compositae). - Eriksen, B., Molau, U. & Svensson, M. 1993. Cytologia 60: 341-346. Reproductive strategies ,in two arctic Pedi Bakhshi Khaniki, G. 1996. Karyologieal cularis species (Scrophulariaceae).-Eco studies in six taxa of the genus Cenlal/rea graph)' 16: 154-166. (Compositae). - Bot. Chmn. 12: 55-65. Eriksen, B. 1996. ;\1ating systems in two Bakhshi Khaniki, G. 1997. Fritillana alro species of Poten/,illa from Alaska. - Folia lineala (Liliaceae) a new species from Iran. Geobol. Phytotax. 31 : 333-344. - Edinb. J. Bot. (in press). Eriksen, B. 1997. Morphometric analysis Bakhshi Khaniki, K. 1997. Fritillaria chlo of Alaskan members of the genus POlentilla. rorhabdota (Liliaceae), a new species from L. sect. Nivea.e (Rosaceae). - Nord. J. Bol. Iran. - Herbertia (in press). 17 (in press). Bakhshi Khaniki & Persson, K. 1997. Nec Fukuhara, T. & Liden, ',1. 1995a. Pericarp tary morphology in South West Asian F rilil anatomy in Fumariaceae. - Bol. Jahrb. Sysl. Larin (Liliaceae). - Nord. 1. Bot. (submitted). 117: 499-530. Fukuhara, T. & Liden, M. 1995b. Seed coat B. & al. (eds.), Flora Nordica, vol. 1 (in anatomy and phylogeny in Fumariaceae. - press). Bot. 1. Linn. Soc. 119: 323-365. Liden, 1. . Fukuhara, T.. & Axberg, T. 1995. Liden, M. 1991a. I otes on Corydalis sect. Phylogeny of Corydalis, ITS and mor Corydalis in the Baltic area. - ! ord. 1. Bot phology. - PI. Syst. Evol. Suppl. 9: 183-188. II: 129-133. Liden, M. , Fukuhara, T., Rylander. 1.. & Liden M. 1991b. De svenska nunneortema. - Oxelman, B. 1997. Phylogeny and classi S". Bot. TIdskr. 85: 65--69. fication of Fumanaceae. with emphasis on Liden, M. 1991c. New tuberous species of Dicentra. s.l. based on th e plastid gene Corydali~ (Papaveraceae). - Willdenowia 21: rpsl6 intron. - PI. Syst. Evol. (in press). 175-79. Liden, M. & Staaf, R. 1995. Embryo growth Liden, M. 1991d. Phenetics and Fumaria - a in tuberous Corydalis species. - Bull. Torrey comment. - PI. Syst. Evol. 176: 221-225. Bot. Club 122: 312-313. Liden, M. 1991e. Rev ision of Corydalis sect. Liden, M. & Zetterlund. H. 1997. Corydalis, Flimarioides, part I. - Rheedea 1: 29-35. a gardener's guide and a monograph of the Liden M. 1992a. Evolution and systematics tuberous species. - AGS Publication Ltd. of seed plants. In: Behnke, H.-D. , Esser, K. , Friary Press, Dorset. 144 pp. Kubitzki, K. , Runge, M. & Ziegler, H., (eds.), Nyman, Y. 1991. Campanu[a occicientalis Progress in Botany 53: 282-317. - Springer (Campanulaceae), a new species from th e Verlag, Berlin, Heidelberg. Canary Islands. Willdenowia 20: 113-116. Liden, M. 1992ct- Fumario pelleri ssp. cal Nyman, Y. 1991. Crossing experiments within cara.ta Liden & Soler, first record (rom Al the CamparlUla dichotoma group (Campa meria. - Anal. Jard. Bot. Madrid. 50: 257. nulaceae). - PI. Syst. Evol. 177: 185-192. Liden, M. 1993a. Pteridophyllaceae. In: Nyman, Y. 1992. Pollination mechanisms in Kubitzki, K. (ed.), Families and genera of six Campanula species (Campanulaceae). - vascular plants, vol. 2. - Springer-Verlag. PI. Syst. Evol. 181: 97-108. Liden, M. 1993b. Fumariaceae. In : Kubitzki, . ·yman. Y. 1992. Reproduction in Camparwia K. (ed.), Families and genera of vascular afra (Campanulaceae): mating system and plants. vol. 2. Springer-Verlag. lhe rol e of the pollen- collecting hairs. - PI. Liden, M. 1993c. Corydalis cornula, a Hi Syst. Evol. 183: 33-41. malayan! Afro-montane disjunct species. - Nyman, Y. 1993. The pollen-collecting hair~ Opera Bot. 121: 4546. of Campanala. I. Morphological "anation Lid en, M. 1995a. A rev ision of Corydalis sect. and the retractive me(:hanism. - Amer. J. Fumarioides Lid en. Part U. - Rheedea 5: Bot. 80: 1427-1436. 1-36. . yman, Y. 1993. The pollen- col.lecting hair;; Liden, M. 1995b. Papaveraceae. In: Harling, of Campallula. II. Functi on and adaptive G. & Andersson, L. (eds.), Flora of Ecuador significance in relation to pollination. - 52. 13 pp. Amer. J. Bot. 80: 1437-1443. Liden, M. 1996. New taxa of tuberou s Cory Oxe\man, B. 1991. Si lene diversifolia Ollh dalis species. - Willdenowia 26: 22-34. and related species in Europe. In: Newton , Lid en, M. 1997f. Fumarioideae. In: Jonsell. 1. E. (ed .), Flora Europaea: Notulae sys- 9 -

tematicae ad Floram Europeam spectan and evolution in Colchicum. - Proc. 5th OP tes, series 2. no. 4. - Bot. 1. Linn. Soc. 106: TIMA Meeting, pp. 394-414. Istanbul. 115-1l7. Persson, K. 1993. Colchicum feinbruniae Oxelman, B. 1991. A revision of the Silene sp. n. and allied species in the Middle East. sedoides-group. - Willdenowia 25: 143-169. - Israel 1. Bot. 41: 75-86. Oxelman, B. 1996. RAPD pattems, nrDNA Persson, K. 1997. The genus Colchicum in ITS sequences and morphological patterns Turkey. 1. New species. - Edinb. 1. Bot. (in in Silene section Sedoideae (CaryophyJla press). ceae). - PI. Syst. Evol. 201: 93-116. Persson, K. 1997. The genus Colchicum in Oxelman, B. & Liden, M. 1995a. Generic Turkey. 2. Revision of the large-leaved boundaries in the tribe Sileneae (Caryo autumnal species. Edinb. 1. Bot. (in press). phyllaceae) as inferred by nuclear rONA Persson, K. & Persson, J. 1992. A new spe sequences. - Taxon 44: 525-542. cies and additional chromosome counts of OxeLnan, B. & Liden, M. 1995b. The posi Hyacinlhella in Tllrkey. - Nord. 1. Bot. 12: tion of Circaeasler - evidence from nuclear 615--620. ribosomal DNA. - PI. Syst. Evol. Suppl. 9: Su Z.- Y. & Liden, M. 1997. Corydalis in China 189-193. I: Some new species. - Edinb. 1. Bot. 54: Oxelman, B., Liden, M., & Berglund, D. 55-84. 1997c. Chloroplast rpsl6 intron phylo geny of the tTibe SileTleae (Caryophyllaceae). - PI. Syst. Evol. (in press). Lund Persson, 1. & Persson, K. A new species of Mikael Hedren Fritillaria fTom Turkey. The New Plantsman (submitted). At the Uni\'ersity of Lund there is a long Persson, K. 1988. New species of Colchu'um standing tradition of experimental plant (Colchicaceae) from the Greek mountains. biology. Below, I will describe some on-going - Willdenowia 18: 29-46. projects at the department of systematic bo Persson, K. 1989. The genus Tulipa in Greece: tany. However, many significant projects have taxonomy karyology and hybridization. - been carried out at other departments as well, Abstr. 6th OPTIMA Meeting Delphi, p. 32. notably the department of genetics where Persson, K. 1991. Colchicum 1., in A. Strid much of biosystematic studies is still done & Kit Tan (ed.), Mountain Flora of Greece today. Yet, because the undersigned has 2: 650-662. Edinburgh Univ. Press. moved to Lund recently and does not have Persson, K. 1991. Tulipa 1., in A. St.rid & the full view of projects being performed at Kit Tan (ed.), Mountain Flora of Greece 2: other departments at the University I will give 667-672. Edinburgh Univ. Press. a sholi review of the projects that are perfor Persson, K. 1992. Liliaceae subfam. Wurm med at "my" department, that of systematic baeoideae Buxbaum, in K. H. Rechinger botany. (ed.), Flora Iranica 170: 1-40, tt. 1-14. - The full professor at the department is Dr. Graz. Honor Prentice, who came down to Lund from Persson, K. 1993. Reproductive strategies Uppsala University about four years ago. Most

10 of Honor's work is related to how genetic gArd station of the Swedish Agricultural Uni diversity is stl1Jctured within species and how versity. this structuring correlates to other parameters Dr. Bjorn Widen, now professor at the de such as microhabitat, geography and immi partment, has a number of long-teml studies gration history to Scandinavia after the last with which he is still working actively. The ice age. Her field work has mainly been per first of these is centered around the Helian formed on the Baltic Island of Oland where themum oelandiwm complex which is en there is a well-equipped field station run by demic on Oland. After describing the popu the University of Uppsala. Most taxa studied lation biology, flowering biology and geo by Honor are in the CaryophyUaceae, among graphic distribution of different flowel;ng others the disjunct Gypsophilafastigiata,

12 700013. 700014). e. buxba/lmii Wahlenb. 2n = 106 (39 J] + 7 IV; 33 II + 10 1\ ; 31 II + 11 IV; 29 II + 12 IV; 2 I + 28 II + 12 IV: 25 II + 14 I ). Qu ebec: comte de Gaspe, rivi ere Dart mouth, fos se "Snake", platiere de gravier et de sable, JC J85-138-1 (DAD 700006, IOPB 700007, 700008). Chromosome Data 12 e. lenticularis Michx. n = 44. Quebec: re gion du lac Chavigny, lac all sud du lac edited by Clive A. Stace Chavigny, rive nord , en bordure d'un es Department of Botany ker (58°02'30"N 75°04'00"W), rives sab 5 University of Leicester Ie uses d'un lac asseche. JC J82-297-1 (DAD Leicester LEI 7RH 700012). England. e. limosa L. 2n = 64. Quehec: comte de E-mail: ca .. [email protected]. Please send contri- Saguenay. lIets-Uremi e, baie des lIets, butions to Professor Stace at the above address vers la pointe E et la sortie de la baie (preferably bye-mail, but failing that on a 3.5 (48°53'50". 68°45'45"W), zone de contact inl'll mi crodisc with text in ASCII-[jle and a entre u.n marecage sale et la decharge d'un printed ropy) using the exacl la.yout of the marecage d'eau douce, JC J79-178-1 (QFA present list and ating whether or not you are :393181). a member of 10PB. Neither proofs nor reprints C. roslrala Stokes x e. saxatilis L. 2n = 80 will be lIIade available, but the editor will (30 II + 4 rn + 2 IV; 2 [ + 37 II + 1 rv; 38 II acknowledge rereipt of contributions and + 1 Ill). No uveau-Quehec: environs du raise queries with authors if necessary. lac Wasatimis, dans la partie nord-ouest du lac Bienville (55°15'N 72°12'W; UTM Reports by: 18UXS768266), sur nanc de petite dune • Jacques Cayouette, Centre de recherches Ie long du rivage du lac, aux endroits saison de l'Est sur les cereales et oleagineux, Agri nierement inondes, avec les deux parents culture et agroalimentaire Canada, edifice presumes, P. MOl'isset, J. Deshayes, R. Wm Saunders, F.E.C.. Ollawa, Onl., Canada Pelletier & 1. Murdock 84-74-1 (DAD KIA OC6. Les specimens te moins sont 700004.700010). deposes a DAD ou A QFA. All localities in e. UlrU:llloio Booll. n = 41. Quebec: comte c Canada. de Saguenay, Pointe-au Boisvert (48 34 "25N 69°1O'05"W), tourbiere lillorale a CYPERACEAE MenJonthes, Carex paleacea. JC 179-174-1 Carex ade/osloma Krecz. 2n = ca 106. (QFA 393180); 2n =8] (39 II + 1 Ill. 11 + Quebec: region du lac Chavigny, lac au sud 40 1I). same locality, JC ]79-175-2 (QFA). du lac Chavigny, extremite sud , vallee de la C. viridula Michx. subsp. viridllia. D = 36. charge (57°59'N 75°05'W), rives rocheuse Quebec: comte de Saguenay, Pointe-des du ruisseau, JC J82-230-1 (DAD 700009, Monts (49°19"20N 67°22'20"W), au fon d 13 d'une pelile anse. avec Myrica gale el Scirpus (QUE. Ste-Foy. Quebec, Canada. and 1.1. rufus, JC r9-15-1 (QfA 19360. DAO Oldham, Ontario lalural Heritao-e Centre, 700002). Peterborough, Ontario. Canada. \ouchers in QfA. Numbers in brackets corrrespond • Jacques Cayouette & Marcel Blondeau. to greenhouse accession numbers and iden Cenlre de recherches de l'Esl sur les cere tify some of th e vou cher specimens. AU ales el oleagineux Agriculture el agro localities in Canada. alimenlaire Canada, edifice Wm Saunders. EE.c., 0llaw8, Onl.. Canada KIA OC6. and J Ul\CAGI.i'iACEAE 2400 Chemin Sie-foy. te-foy. QU libec. Trigloclzin debile (Jones Love & Love. Canada GIV IT2. Les specimens temoins 2n=c. 96. Ontario: ='Io rth Dorchester so nt deposes a DAO. AJI localities in Ca Middlesex. saline highway interchange. nada. ~ I.J. Oldham. 17665.28-0--1995. (95-13). 2n=96. Ontario: Sturgeon River, 0.9 km west CYPERACEA.E of eine River (Rainy River. roadside eLtch. Carex rufina Drej er. 2n = 88. TelTitoires du M.1. Oldhanl & W. Bakowsky 1784-, 06- ! ord-Ouest: district de Keewalin, lles 08-1995, (95-46). Belcher (ile Flaherty). Sanikiluaq (56°32'N Triglochin ela/u/1l NutL 2n= '.1 42. Quebec: 79°14'W; UTM 17VPN 109 608), rivage Het Canuel, rive sud (Rimouski). C. Gen'ais. d'un lac, entre les cailloux MB SK89295a 1--08-199.5, (95-19O)(from ;;eeds) . DAO -00015}. 2n=<:.144. Quebec: Bridge\'ille (Gaspe-Est. POACEAE larune de Barachoi s. marais saM, :\1. Gar Arctophiia fulua Trin.) Rupr. 2n = 42. Ter neau & 1. Labrecque. 92-102M, 06-08- ritoi res du i\ord-Ouest: district de Keewatin. 1992. (92-163 (from seeds of a herbarium lies Belcher (ile flah "rty), eJl\'irons de Sani specimen): Havre-Aubert tl es-de-l.a-Made kiluaq (56°32'N 79°14'W; UTM 17\ PN leine) lagune du Bassin, marais sale, C. Roy 06.5-23), au bord d'un petit etang a 3.50 m & G. Lavoie, 92-2182-C 18-08-1992 (in du rivage, MB SK89363a (DAO 700011). situ fixation); Havre-Aubert (tie -de-la-Ma del ein e). lagune du Bassin, marais sale,1- • C. Gervais, J. Ga",UJIon & R. Trall8I1. Gagnon & S. Pereira, 95-120, 29-08-1995, Direction de la conservation et du patri (95-95); Havre-Aubert (iles-de-Ia-Made moine ecologique. Ministere de l'Environ leine). tourbiere minerotrophe du Bassin, 1- nement et de la Faune du Quebec. 2360. Gagnon & S. Pereira. 95-126. 29-08-1995. ch. Ste-foy. Ste-f0)". Quebec. Canada. G 1 V (95-112): aint-Andre-de-Kamouraska 4H2. Live material was collected by the {Kamourasla . rive de l'estuaire maritime. first two authors and by: 1. Labrecqu and C Gen'ais. 05-09-1993. (93-31 )(from G. Lavoie above address: S. Lamoureux. seeds: Brador, (Saguenay). M. Gameau & fleurbec, St-Henri-de-L.evis, Quebec, Ca P. Mori s -t , 94-818M. 19-08-1994. (94-20); nada; M. Garneau and C. Roy, Louis-Marie Bl anc-Sablon. (Saguenay), M. Garneau & P. Herbarium (QFA) , Ste-Foy, Quebec, Ca MOl'issel, 94-786M, 15-08-1994, (94-26); nada: N. Dignard Quebec Herbarium Blanc-Sablon. (Saguenay). M. Garneau & P. Morisset, 94-783M, 15-08-1994, (94-30). 2n=c.96. Quebec: Havre-Aubert (iles-de-la 2n=c.148. Quijbec: Riviere-OueUe (Kamou Madeleine), lagwle du Havre aux Basques, raska), rochers en bordure du fleuve, C. marais salij, M. Garneau, J. Gagnon, G. la Gervais, 04-08-1995, (95-12). voie & C. Roy, 92-2001\1, 23-08-1992, (92- 2n=c.150. Quebec: Riviere-Ouelle (Kamou 133A); Havre-Aubert (iles-de-la-Made raska), rochers en bordure du fleuve, C. leine), lagune du Havre aux Basques, marais Gervais, 04-08-1995, (95-17); Blanc sale, M. Garneau, J. Gagnon. G. Lavoie & Sablon, (Saguenay), M. Garneau & P. Moris C. Roy, 92-200M, 23-08-1992, (92-147) set, 94-784M, 15-08-1994, (94-33); Brador. (from seeds); Cacouna-Sud (Riviere-du (Saguenay), M. Garneau & P. Mori sset, 94- Loup), marais sale a rest du Gros-Cacouna, 819M, 19-08-1994, (94-21). S. Lamoureux & J. Labrecque, TG-5-1, 04- 2n=e.156. Quijbec: Blanc-Sablon, (Sague 08-1994, (94-12); Saint-Simeon-Est (Bona nay), M. Garneau & P. Morisset, 94-785M, venture), battures maritimes, S. Lamoureux 15-08-1994, (94-35). & J. Labrecque, TG-25-1, 20-08-1994. (94- Trigloehin etatum Nutl. x T. gaspense Lieth & 15). D. Love (back crosses with other hybrids?). Trigloehin gaspense Lieth & D. Love x T. 2n=e.126. Quebec: 11et Canue\., rive sud etalum Nutl. (backcrosses with T. gaspense). (Rimouski), C. Gervais & M. Garneau, 17- 2n=e.99. Quebec: Havre-Aube.rt (tles-de-la 08-1995, (95-52). Madeleine), lagune. du Bassin, marais sale, Trigloehin ela/um . utI. x T. gaspense Lieth & C. Roy & G. Lavoie, 92-2181-C, 18-08- D. Love (bac crosses with T. elaturn). 1992 (in situ fixation); Havre-Saint-PielTe. 2n=e.138. Nouveau-Brwlswick: Lam~que, coulee a Paul (Saguenay), rivage maritime, C. Gervais, 08-08-1992, (92-158)(from S. Lamoureux & J. Labrecque, s.n., 10-08- seeds). 1995, (95-22A); Baie-Saint-Paul, (Charle Trigloehin elatum Nutl. x 1: palu.s/re L. voix-Ouest), rivage de l'estuaire maritime, (unreduced poUen?). 2n=c.96. Quebec: baie au sud-ouest du quai, S. Lamoureux & J. de Rupert (Nouveau-Quebec), batture mari Labrecque, s.n., 07-08-1995, (95-18); Pe lime au nord-ouest de la pointe LeJa.ivre, J. tit-Pabos, (Gaspe-Est), marais sale de la Gagnon, 2634, 06-08-1991. (95-8)(from baie, S. Lamoureux & J. Labrecque, TG-18- seeds). 1, 18-08-1994, (94-09). Trigloehingaspense Lieth & D. Love. n=c.48. 2n=e.102. Quebec: tlet Callue!. rive sud, QU ij bec: Parc national de Forillon (Gaspe (Rimouski), C. Gervais & C. Roy, 17-08- Est), marais sale de la baie de PenouiHe, S. 1995, (95-37,\, 95-51); Douglastown, LamourelL': & J. Labrecque, TG-14-1, 06- (Gaspe-Est), marais sale a l'embouchure 08-1994, (94-11). de la riviere Saint-Jean, S. Lamoureux & 2n=96. Quebec: Riviere-Pentecote (Sague J. Labrecque, TG-16-1, 07-08-1994 (94- nay), embouchure de la riviere Riverill, N. 13); Rimouski (Rimouski), marais sale 11 Dignard, 95-644, 25-08-1995, (95-74); l'embouchure de la riviere Rimouski , S. Etang-du-Xord (iles-de-la-Madeleine) lag LamourelLx & J. Labrecque, TG-1O-1 , 05- une du Havre aux Basques, marais sale, J. 08-1994, (94-08); Saint-Majorique (Gaspe Gagnon, 94-77, 08-09-1994, (94-42). Est), marais sale du bassin nord-ouest de la 15 baie de Gaspe, S, Lamoureux & J, La-brecque, 38,94-41); Havre-Aubert (lles-de-Ia-Made TG-15-1 , 06-08-1994, (94-14); leine), lagune du Bassin, marais sale, J. 2n=c,105. Quebec: Baie de Moisie (Sa Gagnon & S. Pereira, 95-117, 29-08-1995, guenay), embouchure d'un ruisseau au nord (95-113). de l'anse Le Cormoran, S. Lamoureux & J. Triglochin rn.ariJimum L. 5.S. 2n=48. Quebec: Labrecque s.n., 12-08-1995, (95-20). Reserve faunique des Laurentides (Charle Triglochin gaspense Lieih & D. Love x (T. voix-Ouest), tourbiere minerotrophe de la gaspense x T. elatum). 2n=c.l08. Quebec: Jiviere Pikauba, J. Gagnon, 95-204, 23-09- Havre-Aubert (Iles-de-la-Madeleine), baie 1995, (95-178); Languedoc (Abitibi), du Havre aux Basques, marais sale, M. Gar tourbiere minerotrophe de la riviere Au neau & G. Lavoie, 92-202M, 23-08-1992, thier, J. Gagnon, 95-34, 27-07-1995, (95- (92-134). 23). Triglochin gaspense Lieth & D. Love x T. elatum Nutl. 2n=120. Quebec: Havre-Au • C. Gervais, M. Parent, R. Trahan & S. bert (Hes-de-Ia-Madeleille), lagune du Plante, Direction de la conservation et du Havre aux Basques, marais sale, M. Garn patrimoine ecologique, Ministere de I'En eau & G. Lavoie, 92-218M, 25-08-1992, vironnement et de la Faune du Quebec, (92-135). and HerbieI' Louis-Matie, Universite ulval, 2n=c.120. Quebec: Havre-Aubert (Iles-de pavilion C.-E. Marchand, Sainte-Foy, la-Madeleine), lagune du Bassin, marais Quebec, Canada, GIK 7P4. Vouchers are sale, M. Garneau, 92-202M, 23-08-1992, deposited at QFA. Some specimens collect (92-146)(from seeds). ed by F. Coursol (MT) may be considered n=61, 2n=122. Quebec: Grosse-lIe, anse au as substitute vouchers of those collected Sable, (Iles-de-la-Madeleine), marais sale, by Plante and Coursol or Plante, Coursol, J. Gagnon, H. Chevrier & A. Richard, 95- Bouchard and Labrecque. The numbers in 151,02-09-1995, (95-111). brackets correspond to greenhouse acces 2n=c.123. Quebec: archipel de Mingan.1:le sion numbers at QFA. All localities in Ca Saint-Charles, (Saguenay), lac Sale, S. nada: Quebec except Festl/.CC/vivipara (q.v.). Lamoureux & J. Labrecque, s.n., 1l-08- 1995, (95-21). BRASSICACEAE 2n=c.124. Quebec: .Iet Canuel, rive sud Carda mine belli.difolia L. 2n=16. CornIe de (Rimouski), M. Garneau, 95-141M, 1l-08- Matane: Mont Logan, Bassin de Pease, 1995, (95-43). 900m, 07.1992, Trahan (92-43). Triglochinpalllslre L. 2n=24. Quebec: Lauzon, Draba aurea M. Vahl. 2n=c.72. Comte de anse au Sauvage (Levis), rive de I'estuaire Rimouski: !Jet Canuel, rive Slid, rochers fluvial, J. Gagnon, 95-113, 17-08-1995, (95- marilimes, 15.08.1995, Gervais (95-183B); 47); Ilet Canuel, pointe ouest (Rimouski), rive nord, 16.08.1995, Gervais & Roy (95- C. Gervais & C. Roy, 16-08-1995, (95-50). 188A). 2n=37. Quebec: Etang-du-Nord, (lles-de-Ia Draba clivicola Fern. 2n=c.48. Comte de Madeleine), lagune du Havre aux Basques, Matane: Mont Griscom, ravin humide, marais sale, J. Gagnon, 08-09-1994, (94- 900m, 28.07.1992. Trahan (92-92A).

16 CALLITRICHACE.U LAIIIlACEAE Calli1riche s1agnalis Scop. 2n=1O. Comte de Lycopus laurenlianus Rolland-Germain. Montmorency: Beaupre, rivage du 5t-Lau 2n=22. Comte de Portneuf: 10 km a I'ouest rent, 09.09.1995, Gervais (95-114): lie de NeuviUe, ri vage du St-Laurent, 20.09. d'Orleans, Village-des-Anglais, rivage du 1995, Plante, Coursol, Bouchard & La 5t-Laurent 21.09.1995, Plante, Coursol & brecque (95-122, 95-130). Comte de Bouchard (95-165B). Montmorency: lie d'Orleans. Village-des CARYOPIIYLLACEAE Anglais, rivage du St-Laurent, 21.09.1995, SperglLlaria. marina. (L.) Gri seb. 2n=36. Comt ~ Plante, Coursol, Bouchard & Labrecque (95- de Rimouski: lIet Canuel, rive sud, rivage 162). du 5t-Laurent, 17.08.1995, Gervais (95-70). Physoslegia vi.rginiana (L.) Benth. var. granul CYPERACEAE osa (Fassett) Fern. 2n=38. Comte de Port Eleochuris obtusa (Willd.) Schultes val". peasei neuf: 10 km a I'ouest de ieuville. 20.09. Svenson. 2n=10. Comte de Montmorency: 1995, Plante, Coursol, Bouchard & La lie d'Orieans, Battures des lIets, 21.09. brecque (95-121 , 95-123). 1995, Plante, Coursol & Bouchard (95-164, LILlACE.AE 95-166). Smilacinu sl.ellala (L.) Desf. 2n=:36. Comte Scirpus smithii Gray. 2n=40. Comte de de Rimouski: lIet Canuel. haut rivage mari Quebe : Sainte-Foy, plage Jacques Cartier, time. 16.08.1995, Gen'ais & Roy (95-40A). 07.09.1995, Plante & Coursol (95-101). 01\AGRACEAE Comte de Monvnorency: IIe d'Orieans, Vil Epilobillm ciliatum Raf. n=18, 2n=36. Cornte lage-des-Anglai:;, rivage du St-Laurent, de Rimouski: lIet Canuel, rive nord , lisiere 21.09.1995, Plante, Coursol & Boue-hard de foret, haut de greve rocail.l euse, 16.08. (95-161). 1995, Gervais & Roy (95-66). Comte de FABACEAE Portneuf: 10 km a I'ouest de euville, O.'t)"uopis borealis l\utl. var. hudsonicu 20.09.1995, Plante, Coursol, Bouchard & (Greene) Welsh. 2n=16. Territoire-du-Nou Labrecque (95-125). veau-Quebec: miujaq, terrasse de gravier, Epilobillm ecomosum (Fassett) Fern. n=18,

.56°32' I 76°33'0, 22.08.1995, Blondeau 2n=36. Cornt e de Quebec: Sainte-Foy, UM 95450, UM 95451 (95-196, 95-197). plage Jacques Cartier. 07.09.1995, Plame GEi'\TIA1\ACEAE & Coursol (95-81, 95-83); Cornte de P0l1- Gen1.i-a.nclla amarella (L.) Boerner ssp. acula neuf: 10 km a rouest de Neuville. 20.09. (Michx) Gillett. n=9. Comte de Rimouski: 1995, Plante, Coursol, Bouchard & La lIet Canuel, rochers maritimes. 15.08.1995, brecque (95-132, 95-140). Cornte de Mont Gervais (95-68). morency: Ile d'Orle8Jls, ViUage-des-Anglais, JNCAGINAC EAE rivage du St-Laurent, 21.09.1995, Plante. TriglochiTl gaspeTlse Lieth & D. Love. 11=48. Coursol, Bouchard & Labrecque (95-159). Comte du Saguenay: Grandes-B e.rgeronnes, PLUMBAGfriACE.U rivage maritime, 08.1995, Labrecque & Limonium rwshii Small. 2n=36. Comte de Lamoureux (95-19A). Karnouraska: Riviere-Ouelle, rivage mari time. 4.08.1995, Gervais (95-14). 17 POACEAE morency: lie d'Orleans, Village-des-Anglais, Festuca vivipara (L.) Sm. 2n=42. Canada: rivage du St-Laurent, 21.09.1995, Plante, Labrador: Anse-au-Clair, sommet expose de Coursol, Bouchard & Labrecque (95-171. falaise maritime, 3.08.1994, Morissel, Garn 95-174,95-175). eau & Fortin, PM 94-380 (95-86). VERBENACEAE PORTt:LACACEAE Verbena urticifolia L. n=7,8. CornIe de 1'lIe Clay/.Onia caroliniana Michx. 2n=16 + 0-4B Jesus: St-Martin, boise ouvert, 30.07.1995, (51 individuals studied). Comte de Megan Gervais, s.n. tic: St-Ferdinand, erabliere, 03.05.1991, Trahan & Gervais. 2n=16 + 0-IB (27 • B, Turk, DepaJiment of Biology. Biotech individuals studied). COlllte de Megantic: nical Faculty, Univ. of Ljubljana, Vecna pot Ste-Anne-du-Lac, erabuere, 05.05.1993, Ill, SI -1000 Ljubljana, Slovenija. Vouchers Trahan. in W . Author's name abbreviated to BI. ROSACEAE Pu/en/Wa palus/ris (L.) Scop. 2n=42. Comte VAlERIAl\ACEAE de Montmorency: Beaupre, rivage du St Valeriana officillalis L. 2n=14. Slovenia: Laurent, 09.09.1995, Gervais (95-106A, 95- Viniee near Ribnica, 520 m. 14.4.1990. 8T; I06B). Dolnje Jezero ne.ar Cerknica, 553 m, 14.4. Rosa blanda Ail. 2n=14. Comte de Cha.rle 1990, BT; on road between Crnuce and Trzin voix-Est: St-lrenee, haut de greve, 10.09. near Ljubljana, 2.5 km away from crossing 1995, Gervais (95-108B). in Cmuce. 300 m, 15.4.1990, B1'; Cerklje Rosa rugosa Thunb. 2n=14. Comte de na Gorenjskem near castle Strmol, 400 m, Rimouski: Bie, haut de greve a l'embou 15.4.1990, BT; between Medvode and chure de la riviere Hiitee, 18.08.1995, Medno at Ljubljana, near rivulet Mavel Gervais & Roy (95-65); Comte de Charle scica, 315 m, 15.4.1990, BT; on road Lesee voix-Est: St-Irenee, haut de greve. 10.09. to Bled, where river Reeica runs into river 1995, Genais (95-105). Sava Dolinka, 460 01,17.4.1990, 8T; Kozje, Rosa 'wUliamsii Fern. 2n=14. Comte de along river Bislrica, 280 m, 17.4.1990, BT; Rimouski: Bie, schistes a l'embouchure on road Ljubljana to Ig at crossing for Babna de la riviere Hiitee, 18.08.1995, Gervais & Gorica near bridge over river Izica, 290 m, Roy (95-58). 18.4.1990, BT; I 01'0 Mesto on road Novo Sanguisorba canadensis L. 2n=28. Comte de Mesto to Crmosnjiee, 200 m, 19.4.1990, BT; Rimouski: Ilet Canuel, rochers maritimes, Vahta Pass on road Novo Meslo to Metlika, 16.08.1995, Gervais & Roy (95-44A). 615 m, 19.4.1990, BT; right bank of river SCROPH LARIACEAE Lahinja in Primostek near Metlika, 138 m, Agalinis purpurea (L.) Pennell var. paroijlora 19.4.1990, BT; left bank of river Lahillja on (Benth.) Boivin. 2n=28. Comte de Belle road Gradae to Cmomelj at Okljuka, 145 c-hasse: Beaumont, greve du St-Laurent, m, 19.4.1990, 8T; on road Slovens ka 10.08.1994, Gervais, S. n. Bistrica to Pragersko, under the highway Lindern.ia dubia (L.) Pennell var. illulula./.a crossing, 280 m, 23.4.1990, BT; Zane near Pennell. n=9, 2n='18. Comte de Mont- Ormoz at crossing for Turski vrh, 200 m,

18 23.4.1990, BT· on road Verzej to Doklezovje crossing for Babna Gorica near bridge over on left bank of river Mura near bridge over river Izica, 290 m, 18.4.1990. BT. Mura, 182 m, 23.4.1990, BT; Frankolovo, 2n=28 + 3B. Slovenia: Ivanjkovci near 315 m 24.4.1990, BT; Hrusevje near Ormoz at rivulet Pado\'ski potok. 220 m. Postojna, 567 m, 27.4.1990, BT; Gornji 23.4.1990, BT. Dolic at Mislinja, 560 m, 8.5.1990, BT. Valeriano nemorensis Turk. 211=28. Slovenia: Valeriano collin a WaHr. 2n=28. Slovenia: Dolnje lezero nea.r Cerknica. on right side Gornji Dolic at Mislinja, 580 m. 8.5.1990, of road Dolnje l ezero to Gorica, 553 m, 14.4. BT 1990, BT; on road Ozeljan to ova Gorica Valeriana pratensis Di erbach ex Walther. at bridge over rivulet Lijak, 80 m, 28.4. 2n=28. Slovenia: :--iotranje Gorice near 1990. BT; Stan grad near Kobarid, 300 m, Ljubljana on road Notranje Gori ce to Pod 29.4.1990, BT. pec near rivulet Ku slj anov graben, 292 m, Va leriano sambucifolia Mikan f. 2n=56. 18.4.1990, BT; village lezero near Podpec Slovenia: Poborje, on road Lovrenc ua Po on S. E. s ide of lake l ezero , 290 m, horju to Pesek near collage on Pesek, 1:300 18.4.1990, BT; on road Ljubljana to Ig at m, l.5.1990, BT.

\ .UDt l"~ ." OfllCl.v,IJS L 19 SA, Mexico (Yucatan, Morelos, Puebla, Oaxaca, Chihuahua and Sonora) as well as in Chile and Peru. Collaboration with other cactus systematists and acquisition of plant samples from botanical gardens has made it possible to be fully representative of the phylogenetic and geographic diversity within News from Molecular the Cactaceae. Future field research eff0l1S Biosystematists 8 are planned for Brazil, Bolivia and Argentina, as well as more intensive studies of Mexican Editor: Dan J. Crawford (address: see p. 28) cactus diversification, particularly in Tribe Cacteae. Robert S. Wallace Department of Botany laboratory Research 6 Iowa State University, Ames, Much of the recent research in my lab has Iowa 50011-1020 USA utilized comparative D iA sequencing, aJ e-mail:[email protected] though we continue to examine restriction site variation when appropriate. The cacti present For the last several years the research efforts a significant methodological challenge for the in my laboratory have foeused on the eluci molecular systematist due to their production dation of the phylogenetic relationships with of extensive amounts of complex poiysaceha in the morphologically complex family Cacta rides in the form of mucilage w'ithin their ceae. The cacti have a long and complicated tissues. Specialized DNA isolation techniques taxonomic history. Many examples of paral have been developod in my laboratory which lel character changes are found within this enable the isolation of clean, genomic DNA family, which exacerbate the problems of the samples from normally recalcitrant tissues interpretation of its evolution. and also add (now close to 850 taxa sampled from ca. 18 to the taxonomic confusion brought about by plant families; 92% cacti) which provide ex overdescription of its ca 1800 species. M,y in cellent quality templates for PCR ampli volvement as part of a working group for the fication, or work extremely well for restriction Cactaeeae of the International Organization site/hybridization studies. We have con for Succulent Plant Study (IOS) has proven centrated on the plastid genome for our valuable in directing these research efforts, primary source of information due to its in particular to address the many previously relative consen'atism, ease of use, and free unanswered questions for which morpho dom from effects of polyploidy, concerted logical studies could oot provide adequate in evolution, and other factors involved with ,the formation. use of nuclear DNA. Initial comparative se quencing studies were done using rbcL and Field Research ndhF gene sequenees, although low level Our studies of cactus phylogeny have in phylogenetic resolution was lacking for many cluded field research' in various areas of the cactus lineages, due perhaps to effects of long

20 generation time. [n an attempt to obtain bet letic; others are polyphyleti c (e.g. Echinoce teT phylogenetic resolution non-coding se reeae) and thei r recognition is not warranted. quence- from the ImL-tmF intergenic spacer These studies have also enabled the more and from th e intron of rpLI6 have been used confident placement of morphologically ano successfully to provide resolution at the inter malous genera (e.g. Uebelmanni.n) into ap generic, and in several cases interspecific propriate tribes, and forced a more detailed le\·els. examination of morphology for many of these problematic groups. Previous Projects A major effort to determine the evolu We ha\'e been able to examine both struc tionary relationships among the columnar tural (inversions, insertions, deletions) .and cadi in both North and South America was sequence yariation in the plastid genome of supported by the National Geographic Society. the Cactaceae and have identified synapo The resulting 0 A and morphological data morphic stmctural changes which support the support the hypotllesis that the columnar cacti monophyly of the famil y (6 kb cpDNA in found in No rth Am eri ca are derived from version), monoph)'l )' of Subfamily Cacto South American ancestors. and that the cen ideae (loss of rpoCl intran), and monophyly tral Andes is the likely center of origin for of Subfamily Opuntioideae (deletion in dumo the Cactaceae. Further studies of the very plast gene aeeDfORF512). Sequence analy closely-related South American tribes Brown ses of chloroplast genes rbeL ndhF, rp1l6 ingieae, Cereeae, and Trichocereeae are re intron, and tm L-vnF intergeni c spacer all cor quired to better resolve the pattern of ph ylo roborate monophyly for these gmups, and genetic and biogeographic divergences in disclose that the most basal divergences of these lineages. Genus level studies have also the cacti resulted in four discrete lineages, been conducted (e.g. Ferocac/us by recent not three, as had been previously hypothes Ph.D. graduate, Dr. 1. Hugo Cota; Copiapoa, ized {these groups were recognized at the Mila, Hamsi.n , Echinocereus and others by subfamiliallevell· various coUaborators and students); such in Studies of the general phylogenetic stmct terspecific level studies will continue as ure using comparative DNA sequencing for further investigations of tribal/generi c rela each of these subfamilial Lineages within the tionships identify other systemati caUy-im Cactaceae have proven to be very useful. The portant questions. Master's thesis research of Mr. Steven Di ckie has shown that for the subfamily Opuntioi Current Projects deae as presently circumscribed (particularly. BASAL D1VERGE],(CES r1'O THE OPUiliTlOIDEAE with respect to the genus Opunlia and its ex Mr. Steven Di cki e is continuing his doctoral tensive paraphyly), the enti.re subfamily is in research on the subfamiJy Opuntioideae by need of a complete reevaluation of the generic examining the phylogenetic and biogeo concept for its members. Within th e largest graphic patterns of divergence in the South suhfamily Cactoideae, the molecular system American lineages of this subfamily, whi ch atic studies are continuing, however se\reral represent the extant plesiomorphic groups tribes have been demonstrated as monophy- within th e subfamily. 21 PHYLOGEi'iY AND SYSTE~L-\TICS OF TIfE GEN 'S Phytolaccaceae. Specifically, I have begun

-'L-\'~I"mL-\RU basic molecular systematic studies of the Ph.D. dissertation research by Mr. Charlie Aizoaceae to examine its phylogeny and as Butlenl'orth is commencing on the largest sess its relative levels of diversification in genus of the Cactaceas Mammillan:a. He will comparison with the cacti; the families are evaluate overall morphological evolution of comparable in size but have a considerably the genus in the context of a molecular phylo different biogeographic histol),. Future re geny, and then will concentrate on one of the search is planned to assess the pattern of better defined lineages within the genus to diversification of the order Cru'yophyllales, determjne its patterns of evolution. particularly with respect to the Phyto laccaceae. and its relationships to the other PHYLOGENY OT EPIPHYfIC CACTUS LINEAGES major clades of the order. To accomplish this, The epiphytic habit has evolved twice within continued study of DNA variation in both the Cactaceae, and comparisons between the nuclear and organellar genomes is planned. tribes Rhipsalideae ('Christmas Cactus' tribe) Collaboration with interested systematists is and the Hylocereeae ('Orchid Cactus' tribe) encouraged! show that they are vastly different in their rates and panerns of divergence. The role of hybridization (including intergeneric hyb ridization) in the Hylocereeae as a significant evolutionary mechanism is palticularly in teresting.

PORTUUCACEAE ANn ALUED FA.\IIUES The dissertation research of Ms. Wendy Applequist will examine the diversification and specialization of members of families Portulacaceae, Basellaceae, and Didiereaceae (together with general studies of the Cacla ceae) to provide evidence regarding their relatively close relationships as a group within the Caryophyllales, as well as their implied Gondwanan affinities.

FUTURE RESEARCH In addition to examining the evolution31'y relationships within the Cactaceae, our studies also include the investigation of inter and intrafamilial relationships in other Caryophyllalean families such as the Aizoa ceae, Portulacaceae, Didiereaceae and OPll STlA BRE\1 fOlH.

2:2 Meetings - 'Past & Future To give VOll Ih e latest available infomla lion on the li st of speakers and chairmen. the programme of the different symposia are lisled Amsterdam '98 undemealh. By the time you will be reading 7 Ilus, I guess there \\;Il be maIlY more be Listed.

VII INTERNATJON,u. IOpn SDfPOSIUM I:\TERi'iATJONAL ORGAN IZATIOi'i Of PLA:"T BIO

A~JSTERDMI, NETKERLANDS 10-15 AUGUST ' 98 Sr SfEM.m srs - E\'Ol.lITION IN MA N- M~\DE HABrrAT5

snlP051 ~I 1998. A~tSITRD .HI The 1998 symposium is well on its way of Provis ional Time Table (chairmen and organization now. You all will have received speakers confirmed as per luly 1, 1997) the first circular with the basic information, and many of you sent the form that guarantees Tuesday, 11 August 1998 you the receipt of the second circular. In this, General Plenary Lecture the final registration fomls wilJ be included. Bengt JonseU, President lOPB The t.heme chosen obviously attracts interes t Bergius Botanical Garden. Stockholm, Sweden not only from IOPB members but also from other disciplines, particularly from plant Symposium 1: EYOLUTION IN DISfCRBED HABIT.U'S breeding and biotechnology. We may expect Chairman: Herbert Hurka interesting discussions on the para LI eUi sm niversity of Osnabrtlck. Gemlany between domesti cation and natural evolution. Speakers: There doubtless will also develop a debate Barhara Neuffer about the past inlrogressions of crops into wild 'niversit} ofOsnabriick. Germany relatives, and of course al so on the possible Krystyna Urhanska future consequences of such introgressions ETH Zurich, Swilzerland from transgenic races of crops if th ese will have been taken in full agricultural practice Symposium 2: EVOL 'lION OF CROrs. DO~15"TICA - an extremely important matter with respect TIOi'i: SI.\IUL\TING EVOL 'flO ' to conservation of genetic resources and bio Chairman & speaker: Paul Gepts diversit), in general. Universiry of Caljfornia. Davis, c.o\.. USA Up to now (the beginning of luly), some 90 people made th eir provisional registration. Symposium 3: E\'OLUTlo;, Of CROPS. MAPPIJ\G

Given th e lime table this is very promising. OF SPECL\L TR.~ ITS However, we would like to weicorne quite . Chainnan: Andreas Boerner more coUaegues, so don't hesitate to send in IPK Gatersleben, Gennany the form or to send me an E-mail message Speakers: ([email protected]). As indicated in the first • John Doehley cir 'ular you also could visit our website where Universil}' of Wisconsin. 1adisoll \\'1. USA regularly updated information is available: "Evolution of Maize" htlp:/Iwww/bio.uva.nl/conferences/iopb 98. • Andreas Boenler him "Mapping in Triticeae" 23 • Renate Sclmudt Royal Ontario Museum, Toronto, Canada Max-DelbIiick-Laboratorium Koln. Gemlany "Species concepts in agamic complexes" "Comparative Mapping in Brassicaceae" • Ron van der Hulst University of Amsterdam, The Netherlands Symposium 4: [VOLU 'ION or 1~· V.. \ SIVE PL~!';'r SPE "Evolution of agamic complexes in European Ta CIF.': ADAPTATtON .. \ ND .LIFE CYCLES raxacum (Asteraeene)" Chainnan: Joachim Kadereil University of Jlainz, Germany We all, the members of the organizing com Speakers: mittee, hope to welcome large numbers of • Suzanne I. Wandck you next year in Amsterdam. Biosystematics Research Centre, Ottawa On behalf of these, Hans C.M, den Nijs, "In\'asi,'e plant species: a peropeClive" secretary organizing committee Vnt.h sym • Shoichi Kawano posium: Institute of Systematjcs & Popula Universily of Kvoto. Japan tion Biology, Unjversity of Amsterdam, Kruis " Life history traits in naturalized populati()ns of laan 318, 1098 SM Amsterdam, The Nether Arabidopsis thaliana in Japan" lands, phone • 31 20 5257660. fax • 31 20 • Joachim Kadereil 5257662 "The e"olution of invasive species in Senecio"

Symposium 5: EVOLlJTtON OF CROP-WILD RELATIVE Other meetings COMPLEXES VALDIVIA, CHILE Chairman & speaker: KJaus Anllllrum II Southern Connection symposium Valdjvia, Botanical Garden Berne. Switzerland Chile 6-11 january 1997. "Hybrid speciation in sunflowers" "Southern Connection" is an informal as sociation for all scientists interested in the Symposium 6: EVOLUTION OF IWASiVE PLA NT natural history of the southern temperate areas SPECIES: APmnXtS: CLO:'I,\L VS. SEXUALSPECJATtON and includes botanists as well as e.g. zoo Chairman: Christopher S. Campbell logists and quaternary geologists, both basic niversity of Maine, Orono ME, USA research and applications in natural con Speakers: servancy in particular. Excursions to the • Christopher S. Campbell magnificent but threatened temperate rain " Hybridization and the evolution of agamic com forests close by were extremely instructive. plexes" Meetings are held every three years, the • Michael D. Hayward second one took place in Valdivia, temperate IGER, Aberyslwyth Chjle this January. The purpose is to promote "The Genetics of Apomixis" contacts both between the disciplines and • Randall J. Bayer between the world's south temperate areas. CSIRO, Canberra, Australia The Valdivia meeting counted over 300 "Polyploidy and the evolution of agamic com delegates among whom Argentine, Chile, plexes" Australia and New Zealand were particularly • Timothy Dickinson well represented, but also South Africa, USA,

24 4

and other S. American countries. In fact the PARIS, FRAI'I'CE Europeans were a small minority. The main LX OPTIMA Meeting in Paris, ~I a y 11-17 theme this time, for whic h a numbe r of 1998. speakers were invited, was the south tempe The th emes of the coming OPT IMA rate forests, fo cusing problems about mana meeting will be (I) French acti vity in botany. gement a nd conservancy. Numerous (2) Plants and serpentine formations in the conLI;buted papers and pos ters covered a great MeditelTanean (3) Molecular phylogeny of array of topics and gave a broad survey of Mediterranean groups, (4) Demonstrations of acti vities in south temperate taxa and biota. Mediterranean databases, (5) Plant limits of The nex t "Southem con-nection" meeLing will the Mediterranean region, (6) The uses of take place in New Zealand in 2000. Mediterranean plants. One excursion during General Secretary and primary organizer the conference and two post meeting ex of the symposium was Professor Mary Kalin cursions are planned. Arroyo, member of IOPB and previously in Professor J. Morat is General Secretary for our council. An yone with interests in the th e arrangement s. south temperate areas is welcome to join the Southem Connection and will receive a news UPPSALA , SWEDEN letter and symposium circular. You should Plant.a Europa - second European conference contact a board member, e.g. Mary AlToyo on th e conservation of wild plants in Uppsala. Lab. De Sistematica y Bi ologia Vegetal, Depto Sweden, 9-13 June 1998. Bi ologia, Uni v. De Chile, Casilla 653. San The first European conference on wild tiago, Chile. plant conservation in Hyeres, France created the Planta Europa network. whi ch now in HAItARE, 2mBABWE vites you to join th em in sharing experience, XV AETFAT Symposium in Harare, Zim developing the network and working out how babwe 2-9 February 1997. together we can better conserve Europe's " L'A ssociatjon pour l'etudes de la Oore de plants. For more infonnation - and if you l'Afrique tropicale" holds since nearly 50 would like to contribute a paper, pl ease years regular meetings at tlu-ee )·ear inten'als, contact the Conference Secretariat through nowadays every six years in Afri ca and in Dr Johan Samuelsson e-mail: PlantaEuropa between in Europe or some other temperate 98@dha. slu.se part of the world. In Ha rare about 200 participants appeared and a great number of Bengl. }onsell African countries were represented with the ~ large t delegations from South Africa and the host country itself. Taxonomy and phylogeny, biodiversit y, phytogeography, plant protection and ,·egetat.ion consen'ancy were among th e subjects treated. TIle next meeting. the 50th anniversary of th e organzalion. will be held in Brussels in 2000. 25

a Points of view From V;iell1er Greuter, Berlin, the following comment has arrived: We had only a few reactions Concerning the term Bios),stematics, my upon the views about the zoological colleagues tell me that they now 8 futuJ'e perspectives of IOPB widel)' use it in the geneml sense oj biolo expressed by the editors in gical systematics (i.e. (.0 include alpha ta the previous Newsletter. Should we take this xon{)my, et.c.), e.g. in Systematics Agenda as if members in general share our views - if 2000 - along with similar neologisms in so, we are of course happy, but we cannot fashion as biodiversity, bionomenclat1lre, help thinking that there are opinions we etc. In other words the lenll risks to become ought to know about thai llIay be hidden ambiguolls (no/ault oj IOPB, oJcollrse, but among you. Please let us hear from you! there we are).

Krystyna Urbanska, Ziirich, has sent us the From David Hawks worlh, London , the following points of view on the actual issue: following observation was received. 1. If and when IOPB merges with another Th e word "bios),stematics" (as "bios),stem organization, a balanced intem<1tional char at.ists") was evidentlyjirst. used in 1937 in ucter should be considered. This condition the San Fransisco area oj the USA for an. is indispellSable to many decisi.ons, elections in terdisciplinary group in/omwlly carrying oj Executives, etc. on the work oj the then declining Society Jor 2. If and when IOPB merges wit.h another the Study oj Speciation.. This inJorn!<1tion organization, a stmct.ured membership Jee comes Jrom V.B . Smocovilus (J. HiS/aT)' should be considered. In the presenl time both Biology 27: 241-309, 1994), who also wrot.e individu<1ls and libran:es all Olier the world a book I h.ave not seen that. may inclu.de fight with increasingly restricted budgets, Jurther injormation ("Botany a.nel the and the situation in post-communistic Evolutionary Synthesis"). The often cited coU/uries or those in the Third World is usage by Camp & Gill'X (Briuonul 4: 323- particul<1rlydifficult. For thaI reason, IOPB 385, 1943), as "biosystemat.y", is t.hus later. members should be given a free choice be t.rueen (a) a Jee including a Newsletter onl); and (b) a considembly higher fee covering also a subscription to one journal. This S)'S tem has been adopted in many societies of UK and USA , and I consider it Jair to the people who may nal be able to pay outmgeous anl!ualJees but wish to participate in society activities all the same. For m<1ny it is t.he only "lifeline" to Joreign colleagues. I Jear that a ':rorced" subscriptiOll to any - how ever good - journal, may cost IOPB 1/l<1n.y members.

26 Q:) V

27 IOPB phone. +1 6142928952 Executive and Council 1995-98 fax· +16142926345 e-mail. [email protected] Co-Editor Newsletter "Chromosome Data". Clive A. Stace 9 Dept. of Botany niversity of Leicester Executive LE[CESTER LEI 7RH, .. K. President, Bengl E. Jonsell phone. +44 116 252 3381 Bergiu5 Bot. Garden fax. +44 116252 2791 P.0.B.50017 e-mail.(secr.)[email protected]/ S-I0405 STOCKHOLM, Sweden (prof. Stace)[email protected] phone. +46 8 156 896 Secretaryffreasurer, Peter C. Hoch fax • +4686129005 Missouri Bot. Garden e-mail [email protected] p. O. B. 299 Vice-President - President Elect. ST. LO '[S, 1\'10 63166-0299, USA Konrad Bachmann phone. +13145775175 Dept. of Taxonomy fax· +13145779589 [PK Gatersleben e-mail. [email protected] CorrensslTaBe 3 Regional Treasurer (European currency) 0-06466 GATERSLEBEN , Germany Leo W.O. van Raamsdonk phone. +49 39482 5465 Centre for Plant Breeding & fax· +4939482 5155 Reproduction Research, CPRO-DLO e-mail [email protected] P.0.B.16 Past President. Peter H. Raven NL-6700 AA WAGE, [NGE Missouri Bot. Garden The Netherlands P.0.B.229 phone. +31 317477279 ST. LOUTS, MO 63166-0299, USA fax. +31 317416513 phone. +1314577 .5111 [email protected] fax· +1314577 9595 Member ex officio for 1998 Symposium, e-mail. [email protected] Hans C. 1\1. den Nijs Ed.itor Newsletter, Liv Borgen Hugo de Vries Lab. Bot. Garden & Museum University of Amsterdam University of Oslo Kruislaan 318 Trondheimsveien 23 B NL-1098 SM AMSTERDA)1 ~-0562 OSLO, Norway The Nelherlands phone. +47 22 85 17 78 phone. +31 205257660 fax. +47 22 85 18 35 fax· +31 20 525 7662 [email protected] [email protected] Co-Editor lewsletter "Molecular I ews" Danie[ J. Crawford Dept. of Botany Ohio State 'niversity 1735 Neil Ave COL MBUS. 0043210-1293. USA

28 • Council e-mail. [email protected] Randall J. Bayer u.ac.jp CSIRO - Plant Industry Jan Kirschner Molecular Systematics Lab institute of Botany Australian National Herbarium Academy of Sciences GPO Box 1600 CZ-25243 PRUHONICE 1 CA l BERRA, ACT, 2601, Australia Czech Republic phone. +61 6 246 5514 fax. +420267750031 fax • +61 6246 5249 e-mail [email protected] e-mail [email protected] Tatsuyoshi Morita [email protected] Biological Lab. Jorge V. Crisci Faculty of Education (Museo de la Plata. 1900 La Ptala. Il.A.. Argentina) University of Niigata until Feb. 1998: 8050 Ikarashi-ninocho Dept. of Botany NIIGATA-SHI 950-21. Japan niv. of Wisconsin-Madison fax • +81 25263 1277 132 Birge Hall e-mail [email protected] 430 Lincoln Drive David F. Murray MADISON, WIS 53706-1381, SA niv. of Alaska Museum phone. + 1 608 262 8644 907 Yu.kon Drive fax· + 1 608262 7509 FAIRBANKS, AK 99775-6960, USA e-mail. [email protected] fax. +1 9074745469 Philip Garnock-J nes [email protected]. edu School of Biological Sciences JUrg Stock lin Victoria (Sniv. of Wellington Botanical Institute P.O. Box 600 University of Basel Schonbeinstrasse 6 WELLlNGTO I , New Zealand CH4056 BASEL, Switzerland phone. +64 4 472 1000 phone. +41 61 267 3501 fax· +64 4 4715331 fax. +41 61 2673504 e-mail .phi.\[email protected] e-mail·[email protected] Herbert Hurka Suzanne L Wamick Institut fill Biologie Centre for Land & Biological Resources Lni\'.Osnabriick Research Agriculture and Agli-food Barbarastr. 11 Canada, K. W. eatby Bldg 0-49076 OSNABRCCK, Gennan), C.E.F., OTTAWA, Ontario fax. +49541 9692870 Canada KIA OC6 e-mail. [email protected]. un i phone. + 1 613 759 1829 osnabrueck.de fax· +1 6137591924 Shoichi Kawano e-mai.l • [email protected] Dept. of Biology Faculty of Science Kyoto niversity KYOTO 606, Japan phone. +81 75 753 4131 fax· +81 75 7534145

29 Changed addresses & Bente Eriksen Dept. of Systematic Botany New members Carl Skottsbergs Gata 22 10 S-413 19 GOTEBORG, Sweden Aslaug Hagen Karl Peter Buttler Botanical Garden and Museum Orber Strasse 38 Trondheimsveien 23 B D-60386 FRANKFURT AM MAIN N-0562 OSLO, Non,'a), Germany Mikael Hedren Hann-Chung Lo, Dr Dept. of Systematic Botany Department of Forestry O. Vallgatan 18-20 National Taiwan University S-223 61 L I D, Sweden P.O. Box 13-387 TAIPEI 100, Taiwan. Republic of China Hilde ! ybom e-mail: [email protected] Balsgllrd Dept. for Horticultural Plant Breeding Donna I. Ford-Wernitz Swedish Univ. of Agricultural Sciences Department of Biology Fjalkestadsvagen 123-1 P.O. Box 6057 S-291 94 KRISTIA NSTAD, Sweden MORGANTOWN, WV 26506-6057, USA Karin Persson Henrik Aerenlund Pedersen Botanical Garden Botanical fuserun Carl Skottsbergs Gala 22 Gothersgade 130 S-41319 GOTEBORG, Sweden DK-1123 COPENHAGEN, Denlllark

Your address! Roland von Bothmer Please send changes and additions to the editor Dept. of Plant Breeding Research Bengt Joosell - preferably bye-mail! Your Swedish niv. Agricultural Sciences changed address will be published in the next S-268 31 SVALOV, Sweden issue, which will also be correctly distrihuted. For GeT1lUJny in general should be obseroed that Anne Brysting 4-digit =ips are obsolete and misleading. German Trondheimsveien 23 B colleagues wilh such zips in the directory oj the previous Newsleuer are encouraged to inJorm. us Botanical Garden and Museum uDOIL/. the flew nrunber. The address oj Werner N-0562 OSLO, I on\'ay Grewer, lI.ho inJormed US on thi.! point, should IIOW run.: Bol. Garten & Museum Berlin-Dahlem An'e Elvebakk Ktlnigin Luise Strasse (Hl IBG, niv. ofTrom~o D-14191 BERLIN, Germany N-9000 TROMSO, orway

30 .. International Organization of Plant Biosystematists current research, requests for ma The International Organizalion of terial and information, announce Plant Biosystematists, founded in ments of meetings, etc. Two pernm 1960, acts on several levels £rom co nent features in the Newsletter are ordinating

I rimental and Molecular Approaches At present, IOPB Membership is to Plant Biosystematics) give exten for the three-year period between si\'e insight in the field lOPB deals the Symposia.The next Symposium with. will be held in The Netherlands in The lOPB Newsletter is pub 1998. Detailed information on the lished twice a year and maiJed to payment system will be pro\'ided by all Members. It includes reports on the TreasuTer upon request.

An)' inquiries about joining IOPB, IOPB ME~mERsmp APPLICATIOI'i membership fees, Newsletter sub FOR.'t (Please print!) scription. etc., as well as the applicat· ion form , should be mailed to the Mr J Ms. I..a;;t name Secrelaryffreasurer Peter C. Hoch Missouri Botanical Garden P.O.B.299 First name, middle initial ST LOUIS, Missouri 63166·0299 USA Addres. phone. +1 314577 5175 fax· +1 314 .~77 9589 e-mail· hochimobol.mobol.org

ResidenlS ill the Nordic cOUIUries may also apply to one of the News feller editors. D,te Signalure

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