DOCSLIB.ORG

La Double Symbiose Mycorhizienne Chez Deux Espèces Forestières, Taxus Baccata L

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
La Double Symbiose Mycorhizienne Chez Deux Espèces Forestières, Taxus Baccata L République Algérienne Démocratique et Populaire Ministère de l’Enseignement supérieur et de la recherche Scientifique Université Mouloud Mammeri de Tizi-Ouzou Faculté des Sciences Biologiques et des Sciences Agronomiques THESE Présentée Par Bournine épouse Harchaoui Chafia En vue de l’obtention du titre de DOCTEUR EN SCIENCES BIOLOGIQUES THEME La double symbiose mycorhizienne chez deux espèces forestières, Taxus baccata L. et Populus nigra L., situées dans la région de Tizi-Ouzou (Tikjda, Akfadou et Ait zikki) Soutenue publiquement le 03/05/2017 Devant le jury composé de : Président : Pr Derridj A. Université de Tizi-Ouzou Rapporteur : Dr Adjoud- Sadadou Dj. Université de Tizi-Ouzou Examinateurs : Pr BOUTEKRABT A. Université de Blida Pr BEDDIAR A. Université MB d’Annaba Pr Belhadj S. Université de Djelfa Dr Boudiaf M. Université de Tizi-Ouzou Dédicaces A la mémoire de ma maman, A mon papa que Dieu lui prête longue vie, A mon mari Hakim, A ma raison de vivre mes enfants chéris Rimo, Lili et Dido. i REMERCIEMENTS Le travail présenté dans cette thèse a été réalisé au laboratoire Production, Amélioration et Protection des végétaux. Cette thèse est le résultat d’un travail de recherche de plusieurs années. En préambule, je veux adresser tous mes remerciements aux personnes qui m’ont aidé à la rédaction de cette thèse. En commençant par remercier tout d’abord Mme Adjoud-Sadadou Djamila, directrice de recherche du présent travail, pour m’avoir permis de continuer ce doctorataprès le départ de mon directeur de thèse. Merci pour votre aide précieuse et pour le temps que vous m’avez consacré malgré votre état de santé. Merci pour votre confiance, votre amitié et pour m’avoir transmis une passion inlassable pour les champignons mycorhiziens. Je ne peux oublier de remercier Mme Hargas-Halli Rosa professeur retraitée de l’université Mouloud Mammeri qui m’a encouragé et conseillé. Je garde d’elle l’image d’une grande dame. Je tiens à remercier les membres de jury qui ont accepté d’examiner ce travail : Au Professeur Derridj Arezki qui a accepté de présider le jury. Qu’il trouveici toute ma gratitude et mon sincère respect pour son esprit scientifique et son amabilité. Au Professeur Beddiar Aarifa de l’université d’Annaba qui a bien voulu accepter d’examinerce présent travail et à apporter ses remarques de professionnel dans le domaine des mycorhizesafin d’améliorer la qualité de ce travail. ii Au Professeur Boutekrabt Ammar de l’Université de Blida qui n’a pas hésité à répondrefavorablement à notre invitation à participer à l’examen de notre travail ; Au Professeur Belhadj Safia de l‘université de Djelfa qui n’a pas hésité à répondre favorablement à faire partie du juryde cette thèse. Merci d’enrichir ce travail par vos remarques constructives. Au Docteur Boudiaf Malika de l’UMMTO d’avoir accepté de rapporter et examiner mon travail de thèse et de faire partie de ce jury. Un vif merci en particulier à l’équipe du district d’Ait Zikki, de la circonscriptiond’Azzazga et à mon collègue Mr Bensidhoum qui nous ont accompagné lors de nos sorties sur le terrain. Merci également à mes amies lynda, Najat, Karima (l’ange du labo) et Fazia pour leurs présences, leur soutien moral, les discussions et leurs amitiés. Vous avez été au cœur de la réussite de ce projet. Trouvez là, messincères sentiments d’amitié. Je ne sais comment exprimer ma gratitude à deux personnes Mr et Mme KADI pour leur aide merci infiniment que Dieu vous récompense. J’envoie ensuite mes plus sincères remerciements au chef de département et à toute l’équipe dudépartement de pédologie de l’Ecole Nationale d’Agronomie d’ El Harrach pour m’avoiraccueilli et permis de réaliser les analyses physico- chimiques du sol des différentes placettesd’étude. Un grand merci pour Redouane qui a été très gentil et très serviable et pour son grandprofessionnalisme, qu’il trouve ici toute ma gratitude. Je tiens à m’excuser auprès de mon mari,mes enfants, mon papa et toute ma famille et mes amies pour mes absences aux moments où vous aviez besoin de moi.Merci de me pardonner et de me comprendre. Un grand merci aux étudiants qui sont passés par notre laboratoire pour leur sérieux et pour les beaux moments passer ensemble au labo et sur le terrain. iii Articles et communcations issus de la thése 1- Bournine- Harchaoui Chafia, Adjoud -Sadadou Djamila, Kadi-Bennane Saliha., Mezaour Najet (2016). Estimation of the ecto- and endomycorrhizal colonization of the black poplar - Populus nigra- of Ait zikki (Kabylia, Algeria). IMPACT: International Journal of Research in Applied, Natural and Social Sciences (IMPACT: IJRANSS) ISSN (P): 2347-4580; ISSN (E): 2321-8851.Vol. 4, Issue 9, 65-74. 2- Bournine-Harchaoui Chafia., Adjoud-Sadadou Djamila., Halli-Hargas Rosa., Mezaour Nadjat., Kadi-Bennane Saliha (2014). Les mycorhizes : un moyen pour préserver Taxus baccata, espèce endémique menacée dans deux régions de la Kabylie (Algérie). International Congress On Mycorrhizae- October, 15-17 -Cadi Ayyad University, Marrakesh- Morocco. 3- Bournine-Harchaoui Chafia., Adjoud-Sadadou Djamila., Halli-Hargas Rosa., Mezaour Nadjat., Kadi-Bennane Saliha (2015). Le cortège mycorhizien de Taxus baccata, espèce endémique, dans le parc national du Djurjura : Tikjda (Kabylie). Séminaire International sur l’Environnement et le Développement Durable (CIE DD) Université de BATNA (3 et 4 juin). 4- Bournine-Harchaoui Chafia ; Adjoud-saddadou Djamila ; Kadi-Benane Saliha ; Mezaour Najet.( 2015). Diversité du cortège mycorhizien de deux espèces de peuplier dans la région de la Kabylie(Algérie), en rapport avec certains facteurs écologiques. Conférence Internationale sur la Biotechnologie et l’Agriculture, Tunis. 5- Bournine-Harchaoui Chafia., Adjoud-Sadadou Djamila., Kadi-Bennane Saliha, Mezaour Nadjat (2016).. Estimation de la mycorhization de quelques arbres de Taxus baccata dans la forêt d’Akfadou. Effet des caractères physicochimiques du sol sur cette mycorhization. Une Conférence Internationale sur les Symbioses Végétales et leurs Applications Biotechnologiques, Agronomiques et Environnementales. Khemis Miliana 19 – 20 Avril iv Liste des abréviations AM : Mycorhizes à Arbuscules. ECM : Ectomycorhize. F.P.A : Formol, Acide propionique et Ethanol. M: Morphotype. MPN: Most Probable Number N: Azote P: Phosphore PVLG : Polyvenyl-lacto-glycerine. Rh : Réseau de Hartig. T : Tannins. TC : Type exploratoire contact. TCD : Type exploratoire courte distance. TLMD : Type exploratoire longue et moyenne distance. v Dédicaces ..................................................................................................................................... i Remerciements ........................................................................................................................... ii Publications ............................................................................................................................... iii Abréviations ............................................................................................................................... iv Table des matières…………………………………………………………………………………………………………….…v Introduction .............................................................................................................................. 2 Chapitre I: Rappels bibliographiques sur la symbiose mycorhizienne .................................... 5 1- la symbiose mycorhizienne .................................................................................................. 6 2-Les différents types de mycorhizes ......................................................................................... 6 2-1-Les ectomycorhizes .............................................................................................................. 6 2-1-1-Classification typologique des ectomycorhizes ............................................................... 7 2-2-Les endomycorhizes. ............................................................................................................ 9 2-2-1-Les mycorhizes arbusculaires ......................................................................................... 10 2-2-2-Les mycorhizes éricoïdes ................................................................................................ 11 2-2-3-Les mycorhizes arbutoïdes et monotropoides ............................................................... 12 2-2-4-Les mycorhizes des orchidées ........................................................................................ 12 2-3-Les ectendomycorhizes ...................................................................................................... 13 2-4-Les pseudomycorhizes à endophytes bruns cloisonnés .................................................... 13 3- Processus de colonisation mycorhizienne ........................................................................... 13 3-1- La colonisation par les champignons les ectomycorhizes ................................................ 14 3-2- Processus de colonisation par les champignons mycorhiziens à arbuscules .................. 14 3-2-1-Potentiel mycorhizogène du sol ..................................................................................... 15 4- Rôles des mycorhizes ........................................................................................................... 16 4-1-Amélioration de la nutrition hydrique et minérale ........................................................... 16 4-2-Activité hormonales ..........................................................................................................
Load more

Recommended publications
  • Show Schedules 2012 Ver Finale
    119. 1 pan rock plant native to the Southern Hemisphere 120. 1 pan dwarf shurb THE SCOTTISH ROCK GARDEN CLUB 121. 1 pan rock plant raised from seed by the exhibitor. Date of sowing to be stated. Botanical notes permitted, AGS note 23(e) SECTION III Open to Amateur Members of AGS and SRGC who have not won an AGS Bronze Merit Medal or more than ten First Prizes at Shows run by either Society prior to 1st January 2011. Pan size not to exceed 19 cm outside diameter 130. 3 pans rock plants, distinct 131. 1 pan rock plant in flower 132. 1 pan Gentiana 133. 1 pan Cyclamen 134. 1 pan bulbous plant 135. 1 pan rock plant native to the Southern Hemisphere 136. 1 pan rock plant native to the Northern Hemisphere 137. 1 pan rock plant for foliage effect 138. 1 pan dwarf shrub or conifer 139. 1 pan rock plant. For exhibitors who have never won a first prize at an AGS or SRGC National show SHOW SCHEDULES 2012 DUNBLANE EARLY BULB DISPLAY 18th February* BLACKPOOL SHOW 17th March* STIRLING SHOW 24th March† New Location - Show this Year is in KINCARDINE NORTHUMBERLAND 40th ANNIVERSARY SHOW, HEXHAM 31st March EDINBURGH & THE LOTHIANS SHOW 14th April* PERTH SHOW 21st April HIGHLAND SHOW, NAIRN 28th April GLASGOW SHOW 5th May* ABERDEEN SHOW 19th May* GARDENING SCOTLAND (Joint Rock Only) 2nd June* LATE BULB DISPLAY, RBGE 8th September DISCUSSION WEEKEND, DUMFRIES 29th - 30th September NEWCASTLE SHOW 13th October* AGM 10th November† *Joint Rock Garden Plant Committee meetings 48 †Photographic/Art Competition SHOWS 2012 SHOW RULES 1.
    [Show full text]
  • The Rock Garden 136 the Ro
    January 2016 January 2016 THE ROCK GARDEN 136 THE ROCK GARDEN 136 January 2016 THE ROCK GARDEN Volume XXXIV Part 3 - 136 January 2016 THE ROCK GARDEN Volume XXXIV Part 3 - 136 PostalPostal Subscriptions Subscriptions from from 1st October, 1st October, 2015 2015 Postal subscriptionsPostal subscriptions are payable are payable annually annually by October by October and provide and provide membership membership of the of the SRGC untilSRGC 30 thuntil September 30th September of the following of the following year. year. SubscriptionSubscription Rates Rates UK UK OverseasOverseas Single annualSingle annual £18 £18 £23 £23 Junior Junior £3 £3 £7 £7 (under 18(under on 1 18st Oct) on 1st Oct) Family Family £21 £21 £25 £25 (Two adults(Two andadults up and to two up childrento two children under 18 under on 1 18st Oct) on 1st Oct) Three yearThree subscriptions year subscriptions are available are available at three at times three the times above the aboveannual annualrates. Renewals rates. Renewals for threefor year three subscriptions year subscriptions may only may be only made be atmade the end at the of endthe three of the year three period. year period. All subscriptionAll subscription payments payments to the club to the must club be must made be inmade GB Pounds in GB Pounds Sterling. Sterling. ChequesCheques should shouldbe made be payablemade payable to ‘The Scottishto ‘The Scottish Rock Garden Rock Garden Club’ and Club’ must and be must be drawn ondrawn a UK on bank. a UK bank. SubscriptionSubscription payments payments may be may made be throughmade through the post the by post Visa byor MastercardVisa or Mastercard providingproviding the following the following information information is sent: is sent: The longThe number long number on the cardon the card The nameThe ofname the cardholder of the cardholder as shown as onshown the cardon the card The cardThe expiry card date expiry date The cv2The 3 digit cv2 number3 digit number (from back (from of back the card) of the card) The cardholder’sThe cardholder’s signature.
    [Show full text]
  • PDF Document
    Cyclamen Notes by Wilhelm (Bill) Bischoff Flowers of Atlantis? Page 2 Cyclamen Blooming Times Page 4 Cyclamen Species, Subspecies, Page 5 Forma, & Varieties in Alphabetical Order Cyclamen Descriptions Page 6 (photos referenced are not included) Wilhelm (Bill) Bischoff is available for lectures & garden tours for Cyclamen & Hardy Orchids 604-589-6134 wbischoff @ shaw.ca The Flowers of Atlantis? By Wilhelm (Bill) Bischoff / member BC Council of Garden Clubs If you can accept that the island called Santorini in the central Mediterranean, also known as Thira / Tera, is the original Island of Atlantis; if you also can agree that this Island had a terrific volcanic explosion more than 3,000 years ago, than I can share with you an equally fantastic botanical story with you. That today’s Thira is the remnant of an exploded volcano is quite evident when one looks at a map of this region of the Mediterranean. Located as part of the Aegean Islands, just north of Crete, it shows the unmistakable shape of a water filled volcanic caldera with a center-cone island. Scientists have identified volcanic ash taken from the bottom of the Mediterranean Sea, close to the Lebanese coast, as originating from Thira. The time frame of some 3300 years ago also coincides with the beginning of a rather tumultuous time in this part of the ancient world, the end of the “Bronze Age”. The possible cause of that could well have been a natural disaster, in the very heart of the ancient world as we know it. Now that I have your attention and possibly have whetted your curiosity, let me introduce you to one of the small wonders of this very ancient world, the beautiful Cyclamen, all 22 species of them.
    [Show full text]
  • WINTER 2005 VOL. 63 Aniei-K-Im Primrose Noddy WINTHR 2005 Primroses the Quarterly of the American Primrose Society
    Primroses E QUARTERLY OF THE AMERICAN PRIMROSE SOCIETY WINTER 2005 VOL. 63 Aniei-k-im Primrose Noddy WINTHR 2005 Primroses The Quarterly of the American Primrose Society www.americanprimrosesoc.org Volume 63 No \VINTI-R200S The purpose of this society is to bring the people interested in Primula together in an organization to increase the general knowledge of and interest in the collecting, growing, breeding, showing and using in the landscape and garden the genus Primula in all its forms and to serve as a clearing house for collecting and disseminating information about Primula. President's Message by Ed Buyarski 5 From the Editor by Matt Mattus 7 Androsace cylindrica Photo: Matt Mattus Ncpalcsc Androsace of the Upper Marsyandi Valley by Josef Lcmmcns s This Issue, which focuses on the greater family of Primulaceae, will Discovering and Cultivating Hardy Cyclamen by John Lonsdale 12 take you from Nepal to Canada, and from Belgium to Nevada. Dodecatheon Variants by James L. Reveal 27 Some Hardy Alpine Primula by Margaret Brown 30 PRIMROSES • The Quarterly of the American Primrose Society Androsace 101: An Introduction by Jozef Lemmens 35 Editor Editorial Committee Matt Mattus Robert Tonkin 26 SpofTord Road Judy Sellers Worcester, MA 01607 Pam Evclcigh [email protected] Ed Buyarski About the Cover EDITORIAL Manuscripts for publication in the ADVERTISING Advertising rates per issue: full quarterly arc invited from members and other page. $100; half page, $50: quarter page, $25; A vast population of Androsace rvbusta xsp. purpurea blooming in the Sabche LMrdeiicrs. although there is no payment. Please cigth page and minimum.
    [Show full text]
  • For Peer Review
    Pollen sterols are associated with phylogenetics and environment but not with pollinators Journal: New Phytologist ManuscriptFor ID NPH-MS-2020-34747.R1 Peer Review Manuscript Type: MS - Regular Manuscript Date Submitted by the n/a Author: Complete List of Authors: Zu, Pengjuan; University of Zurich, Geology; Royal Botanic Gardens Kew, Natural Capital and Plant Health Koch, Hauke; Royal Botanic Gardens Kew, Natural Capital and Plant Health Schwery, Orlando; New Mexico Consortium, Plant biology Pironon, Samuel; Royal Botanic Gardens Kew, Biodiversity Informatics and Spatial Analysis Phillips, Charlotte; Royal Botanic Gardens Kew, Natural Capital and Plant Health Ondo, Ian; Royal Botanic Gardens Kew, Biodiversity Informatics and Spatial Analysis Farrell, Iain; Royal Botanic Gardens Kew, Natural Capital and Plant Health Nes, David; Texas Tech University, Dept of Chemistry & Biochemistry Moore, Elynor; Oxford University, Department of Zoology Wright, Geraldine; Oxford University, Department of Zoology Farman, Dudley; University of Greenwich, Natural Resources Institute Stevenson, Phillip; Royal Botanic Gardens, , Jodrell Laboratory,; University of Greenwich, Natural Resources Institute Phytosterol diversity, Pollen nutrient, Pollinator assemblages, Key Words: Environmental factors, Phylogenetic, Plant-insect interactions, Chemical ecology, Chemotaxonomy Manuscript submitted to New Phytologist for review Page 1 of 44 1 Pollen sterols are associated with phylogenetics and environment but not with 2 pollinators 3 4 Pengjuan Zu1,2*, Hauke Koch1, Orlando Schwery3, Samuel Pironon1, Charlotte 5 Phillips1, Ian Ondo1, Iain W. Farrell1, W. David Nes4, Elynor Moore5, Geraldine A. 6 Wright5, Dudley I. Farman6, Philip C. Stevenson1,6 7 8 1 Royal Botanic Gardens, Kew, Richmond, Surrey, TW9 3AB, UK 9 2 Swiss Federal InstituteFor of Aquatic Peer Science Review and Technology, Seestrasse 79, CH- 10 6047, Kastanienbaum, Switzerland 11 3 New Mexico Consortium, 4200 W.
    [Show full text]
  • Cytotoxic Activity of Cyclamen Persicum Ethanolic Extract on MCF-7, PC-3 and Lncap Cancer Cell Lines
    Journal of Natural Sciences Research www.iiste.org ISSN 2224-3186 (Paper) ISSN 2225-0921 (Online) Vol.10, No.2, 2020 Cytotoxic Activity of Cyclamen Persicum Ethanolic Extract on MCF-7, PC-3 and LNCaP Cancer Cell Lines Walid M. Khalilia Forensic Science Department, Al- Istiqlal University, PO. Box 10, Jericho, Palestine Abstract It is important to develop new approaches to increase the efficacy of cancer treatments. Nowadays, the uses of natural products to treat cancer are very common. In addition, working with plants that are endemic to Palestine and determining the biological activities of these plant extracts, is extremely important due to the potential for new drug development . Cyclamen persicum is used in traditional medicinal to treat anti-rheumatic, diarrhea, abdominal pains, edema, abscesses, eczema, cancer and other ailments. In this study the cytotoxic effect of C. persicum tubers and leaves ethanolic extracts were studied against MCF-7, PC-3 and LNCaP cancer cell lines, using mitochondrial dehydrogenase enzyme method . Results showed the remarkable cytotoxic activity of C. persicum extracts, against breast and prostate adenocarcinoma. For tubers extract the IC 50 value was found to be 0.05 mg/ml for the three cell lines. Although the leaves extract the IC 50 value was found to be 0.25 mg/ml for PC-3 and MCF-7 cell lines, while LNCaP cell inhibition were less than 30% at all tested leaves extract concentrations. MCF-7 cells exhibited the highest sensitivity to the C. persicum extracts, compared to PC-3 and LNCaP cell lines evaluated. In contrast, LNCaP cells generally exhibited the lowest sensitivity to extracts.
    [Show full text]
  • Saxifraga and I Hope My Short Article Will Encourage More Gardeners to Grow These Superb Plants
    Alpine Garden Society Dublin Group www.alpinegardensociety.ie NEWSLETTER NO. 75 – WINTER 2021 CONTENTS A message from our Chairman--------------------------------------------- 3 Alpine Miscellany---------------------------------------------------------------5 Trillium rivale, Michael J. Campbell------------------------------------------10 Jimi Blake’s plantsperson’s course, Triona Corcoran---------------------14 Some of the plants that helped me through the Lockdown, H.C.A. McBride----------------------------------------------------------------17 Five gardeners and their five favourite plants, Various------------------ 21 Saxifrages, Billy Moore------------------------------------------------------- 35 Fixtures--------------------------------------------------------------------------41 Officers and Committee------------------------------------------------------43 Cover: Trillium rivale, - see page 10. (Photo: Michael J. Campbell). This newsletter is edited by Billy Moore who can be contacted at 32 Braemor Park, Dublin D14 YX39. Tel. 086 8183140. Printed by Snap Printing, Sandyford, Tel. (01) 2958860. 2 From our Chairman Greetings Alpinists! We’ve reached the end of a very difficult year, one which has brought changes and sadness to us all. But though I’m afraid things won’t get any easier for a while, at least we now have some hope for a return to normality. When I last wrote, in the Summer Newsletter, we still expected that this might happen soon, and optimistically planned for a carefully managed Plant Sale. Alas, the second wave came upon us and this wasn’t to be: I’m sorry to have disappointed the many of you that got in touch to offer plants, and to participate. But we still have the plants from Gerd Stopp, apart from the annuals that is, and have tentatively pencilled in March next year for the sale. Given the uncertainty of the times we think it would not be sensible to set a date just yet (you see I’ve learned my lesson on this at least!), but we will get in touch to confirm the arrangements as soon as it seems possible.
    [Show full text]
  • Biodiversity of Bulbous and Tuberous Geophytes from the El Kala National Park (North-Eastern Algeria): Checklist, Vulnerability and Conservation
    Anales de Biología 41: 25-38, 2019 ARTICLE DOI: http://dx.doi.org/10.6018/analesbio.41.05 Biodiversity of bulbous and tuberous geophytes from the El Kala National Park (North-Eastern Algeria): checklist, vulnerability and conservation Besma Dechir1, Atef Chouikh2, Tarek Hamel3, Nawel Nadia Azizi4, Nawel Ganaoui1, AbdSlem Grira5, Ahmed Abdiouene6, Mohamed Cherif Maazi1,7 & Azzedine Chefrour1,8 1 Department of Biology, Fac. Sciences of Nature and Life, Mohamed-Cherif Messaadia University-Souk Ahras, 41000, Algeria. 2 Biology Department, Faculty of Natural Science and Life, El Oued University, Algeria. 3 Department of Biology, Faculty of Science, Badji Mokhtar University, Annaba-Algeria. 4 Department of Biology, Faculty of Science, Chadli Bendjedid University, El Tarf, Algeria. 5 El Kala National Park, El Tarf-Algeria. 6 National Institute of Forest Research of Souarekh, El Tarf, Algeria. 7 Laboratory of Aquatic and Terrestrial Ecosystems, Mohamed Cherif Messaadia University, Souk Ahras, 41000, Algeria. 8 Lab. Development and Control of Hospital Pharmaceutical Preparations, Department of Pharmacy, Faculty of Medicine, Badji Mokhtar University, Annaba, 23000, Algeria. Resumen Correspondence Biodiversidad de geófitos bulbosos y tuberosos del parque A. Chouikh nacional El Kala (noreste de Argelia): lista de especies, E-mail: [email protected] vulnerabilidad y conservación Received: 18 November 2018 Treinta estaciones en el Parque Nacional El Kala (Noreste de Accepted: 13 March 2019 Argelia) fueron objeto de un inventario florístico que se centró en el Published on-line: 8 May 2019 estudio de la ecología de geófitos bulbosos y tuberosos. La flora se caracteriza por una alta proporción de taxones raros y/o endé- micos, entre ellos, 6 especies son endemismos algero-tunecinos pertenecientes a la familia Orchidaceae.
    [Show full text]
  • Nieuwe Oogst
    NIEUWE OOGST LEGENDA NIJSSEN BULBS Verklaring van de gebruikte symbolen: (Her)introductie in 2018. Nijssen Bulbs Keurmerk Duurzaam geteeld, met respect voor het milieu. | | NIEUWE OOGST NIEUWE OOGST Drachtplanten. o.a. honingbijen, wilde bijen en hommels verzamelen volop nectar en stuifmeel uit de bloemen van dit bolgewas. ! Gemiddelde hoogte in cm. De bloei valt in deze periode, @ opgave in maanden. Geadviseerde plantafstand in cm. 2018 # Gemiddelde plantdiepte in cm. $ Waarde geeft de diepte aan van de onderkant van de bol. | | BOL- EN KNOLGEWASSEN BOL- EN KNOLGEWASSEN BOL- EN KNOLGEWASSEN Herfstbloeier. Bolomvang in cm. % De maat is de omtrek van de bol direct na het rooien. Door vocht- verlies kunnen de bollen krimpen naarmate de tijd in de herfst vordert. De grootte wordt gemeten met een bollenmaat: Wanneer er grootte I. wordt aangegeven dan ontvangt u de voor dit seizoen grootst leverbare bollen, die onder normale omstandigheden tot bloei komen. Voorkeurplaatsing: Z Voor een zonnige standplaats. CATALOGUS 2018 Voor een licht beschaduwde L standplaats. NIJSSEN BULBS HEEMSTEDE H Voor in de halfschaduw. WWW.NIJSSENBULBS.NL S Verdraagt volledige schaduw. 60180557_Omslag_2018.indd 1 01-08-18 10:48 Afbeelding omslag: Allium amethystinum ‘Red Mohican’, zie pagina 6. Fotografie: Gert-Pieter Nijssen en vele enthousiaste kwekers, waarvoor mijn dank. LEGENDA Deze uitgave kwam tot stand door samenwerking tussen: Tekst: Gert-Pieter Nijssen en archief Peter C. Nijssen Verklaring van de gebruikte Lithografie/Produktie: LNO drukkerij, Zierikzee. symbolen: Niets uit deze uitgave mag worden vermenigvuldigd en/of openbaar gemaakt door middel van druk, (Her)introductie in 2018. fotokopie, microfilm of op welke wijze ook, zonder voorafgaande toestemming van Nijssen Bulbs Heemstede.
    [Show full text]
  • Cyclamen Hederifolium Cyclamen Hederifolium (Ivy-Leaved Cyclamen Or Sowbread) Is a Species of Flowering Plant in the Genus Cyclamen, of the Family Primulaceae
    Cyclamen hederifolium Cyclamen hederifolium (ivy-leaved cyclamen or sowbread) is a species of flowering plant in the genus Cyclamen, of the family Primulaceae. It is the most widespread cyclamen species, the most widely cultivated after the florist's cyclamen (Cyclamen persicum), and the most hardy and vigorous in oceanic climates. It is native to woodland, shrubland, and rocky areas in the Mediterranean region from southern France to western Turkey and on Mediterranean islands, and naturalized farther north in Europe and in the Pacific Northwest. Names The word "cyclamen" comes through Latin from the Greek cyclamīnos or κυκλάμινος (also κυκλαμίς) meaning "circle", with reference to the shape of the tuber. The species name hederifolium comes from the Latin hedera (ivy) and folium (leaf), because of the shape and patterning of the leaves. The older species name, neapolitanum, refers to Naples, where the species grows. Description Cyclamen hederifolium is a tuberous perennial that blooms and sprouts leaves in autumn, grows through the winter, and goes dormant before summer, when the seed pods ripen and open. Tuber The tuber is round-flattened and produces roots from the top and sides, leaving the base bare. In the florist's cyclamen (Cyclamen persicum), roots come from the bottom, leaving the top and sides bare. The tuber becomes larger with age; older specimens commonly become more than 25 cm (10 in) across. In other species, tubers do not grow as large; Cyclamen coum usually does not reach more than 6.5 cm (2.6 in) across. Leaves and flowers grow from buds on top. Leaves The leaves are variably shaped and colored.
    [Show full text]
  • THE CYTOLOGY of the GENUS CYCLAMEN by I
    582.918.3 :581.1 7 MEDEDELINGEN VAN DE LANDBOUWHOGESCHOOL TE WAGENINGEN/NEDERLAND-DEEL51,VERH. 7 (pp. 151-166), 1951 THE CYTOLOGY OF THE GENUS CYCLAMEN by I. DE HAAN andJ . DOORENBOS Publicatie no 100 van het Laboratorium voor Tuinbouwplantenteelt, Landbouwhogeschool, Wageningen (Received 25.8:51) CONTENTS 1. Introduction 151 2. Material 152 2.1. Nomenclature and taxonomy 152 2.2. Origin of the material 153 2.3. Forms not studied 153 3. Methods 154 4. Results 154 5. Discussion 158 6. Summary 164 7. Literature 165 Plates 1-4 with figures 1-25. I. INTRODUCTION This investigation of the cytology of the genus Cyclamen was started to find out the possibility of crossing different species,i n order to use thesefo r breeding purposes. This made it necessary to determine at first the chromosome numbers of the different species and varieties, see KAPPERT (9). It was self-evident that simultaneous with this investigation the systematic relationships of the different species were studied (DOORENBOS). Taxonomists chiefly use easily distinguishable macroscopical characters for the classification of species and genera. After the discovery, however, that chromosomes are bearers of genetical characteristics, these also have been used in the study of relationships. Especially after it became clear that in many cases differences in species can be traced back to differences in chromosome arrange­ ment, and none the less after new species were bred by intential species cros­ sings, more and more attention was paid to chromosome number and chromo­ some morphology as an aid to classification. In the different Cyclamen species many different chromosome numbers are found.
    [Show full text]
  • Tez Yazım Klavuz Ve Şablonu
    T.C. PAMUKKALE ÜNİVERSİTESİ FEN BİLİMLERİ ENSTİTÜSÜ BİYOLOJİ ANABİLİM DALI FARKLI GEOFİT CİNSLERİNDEN SEÇİLMİŞ TAKSON EKSTRAKTLARININ BİYOLOJİK AKTİVİTELERİ ÜZERİNE ARAŞTIRMALAR DOKTORA TEZİ MURAT TURAN DENİZLİ, TEMMUZ 2020 T.C. PAMUKKALE ÜNİVERSİTESİ FEN BİLİMLERİ ENSTİTÜSÜ BİYOLOJİ ANABİLİM DALI FARKLI GEOFİT CİNSLERİNDEN SEÇİLMİŞ TAKSON EKSTRAKTLARININ BİYOLOJİK AKTİVİTELERİ ÜZERİNE ARAŞTIRMALAR DOKTORA TEZİ MURAT TURAN DENİZLİ, TEMMUZ 2020 Bu tez çalışması Pamukkale Üniversitesi Bilimsel Araştırma Projeleri Koordinasyon Birimi tarafından 2019FEBE001 nolu proje ile desteklenmiştir. Bu tezin tasarımı, hazırlanması, yürütülmesi, araştırmalarının yapılması ve bulgularının analizlerinde bilimsel etiğe ve akademik kurallara özenle riayet edildiğini; bu çalışmanın doğrudan birincil ürünü olmayan bulguların, verilerin ve materyallerin bilimsel etiğe uygun olarak kaynak gösterildiğini ve alıntı yapılan çalışmalara atfedildiğine beyan ederim. Murat TURAN ÖZET FARKLI GEOFİT CİNSLERİNDEN SEÇİLMİŞ TAKSON EKSTRAKTLARININ BİYOLOJİK AKTİVİTELERİ ÜZERİNE ARAŞTIRMALAR DOKTORA TEZİ MURAT TURAN PAMUKKALE ÜNİVERSİTESİ FEN BİLİMLERİ ENSTİTÜSÜ BİYOLOJİ ANABİLİM DALI (TEZ DANIŞMANI: PROF. DR. RAMAZAN MAMMADOV) DENİZLİ, TEMMUZ 2020 Bu çalışmada, Arum rupicola var. virescens (Stapf) P.C. Boyce, Cyclamen cilicium Boiss. & Heldr, Gagea bohemica (Zauschn.) Schult. & Schult.f., Narcissus tazetta L. subsp. tazetta, Paeonia kesrouanensis J. Thiébaut türlerinin yer üstü ve yer altı kısımlarının antioksidan aktiviteleri, total fenolik, flavonoid ve tanen madde miktar
    [Show full text]