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Staphylinoidea, Staphylinidae (Coleoptera)*

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Staphylinoidea, Staphylinidae (Coleoptera)* SYSTEMATIC CATALOGUE OF THE ENTOMOFAUNA OF THE MADEIRA ARCHIPELAGO AND SELVAGENS ISLANDS STAPHYLINOIDEA, STAPHYLINIDAE (COLEOPTERA)* Vol. II, Part 2 1 2 By VOLKER ASSING & MICHAEL SCHÜLKE With 59 figures ABSTRACT. Based on a revision of the relevant taxonomic and faunistic literature, as well as of previously published and unpublished material, a critical updated catalogue of the Staphylinidae of Madeira and the Selvagens is compiled. 25 species are deleted from the list of Madeiran Staphylinidae, 10 species are recorded from the archipelago for the first time. Seven new synonymies are proposed: Stenus brunneus Puthz, 1978 = S. maderensis Puthz, 1980, syn. n.; Pseudomedon obscurellus (Erichson, 1840) = Lithocharis brevipes Wollaston, 1860, syn. n.; Ocypus pedemontanus (Müller, 1924) = Staphylinus caroli Jarrige, 1943, syn. n.; Tasgius winkleri (Bernhauer, 1906) = Staphylinus maderae Jarrige, 1943, syn. n.; Quedius nigriceps Kraatz, 1857 = Q. nigriceps maderensis Smetana, 1963, syn. n.; Atheta immucronata Pace, 1999 = A. pseudolaticollis Erber & Hinterseher, 1992, syn. n., = A. atlantidum Smetana, 2004, syn. n. For all the species recorded from the archipelagos, the references are listed, additional (i. e. previously unpublished) records, if available, are reported, the distribution in the Atlantic Islands and elsewhere is summarised, and bionomic data are compiled. Disregarding 5 names of doubtful taxonomic status, a total of 210 species and subspecies are reported from the Madeiras, 198 of them from Madeira proper, 52 from Porto Santo, and 11 from the Desertas. Four species, two of them endemic, are recorded from the Selvagens. A zoogeographic analysis revealed that the Madeiran staphylinid fauna is characterised by a high proportion (30%) of endemic species; the remainder is composed mainly of species with West Palaearctic (25%), Cosmopolitan (18%), Holarctic (11%), Palaearctic (4%), and Western Mediterranean (4%) distributions. 9% have not been recorded again since 1900, suggesting that their (Madeiran) populations may have gone extinct. The distributions of various Madeiran endemics are mapped. The occurrence, distributions, and wing development of the species present in the archipelagos are summarised in a checklist. KEY WORDS: Coleoptera, Staphylinidae, Madeira, Selvagens, islands, taxonomy, catalogue, new synonyms, new records, endemism. * We dedicate this contribution to our dear friend and colleague, the late Dieter Erber, in gratitude for his inspiration and for his devotion to the study of the Coleoptera of the Atlantic Islands. 1 Gabelsbergerstr. 2, D-30163 Hannover, Germany. E-mail: [email protected] 2 Rue Ambroise Paré 11, D-13405 Berlin, Germany. E-mail: [email protected] Bol. Mus. Mun. Funchal, Sup. N.º 11: 5-167, 2006 ISSN 0870-3876 6 Boletim do Museu Municipal do Funchal (História Natural) Supl. No. XI RESUMO. O presente trabalho consiste num catálogo critico e actualizado dos Staphylinidae da Madeira e das Selvagens, baseado na revisão da literatura relevante de âmbito taxonómico e faunistico acerca desta família. São retiradas da lista de Staphylinidae da Madeira 25 espécies e 10 são referidas para o arquipélago pela primeira vez. Sete novas sinonímias são propostas: Stenus brunneus Puthz, 1978 = S. maderensis Puthz, 1980, syn. n.; Pseudomedon obscurellus (Erichson, 1840) = Lithocharis brevipes Wollaston, 1860, syn. n.; Ocypus pedemontanus (Müller, 1924) = Staphylinus caroli Jarrige, 1943, syn. n.; Tasgius winkleri (Bernhauer, 1906) = Staphylinus maderae Jarrige, 1943, syn. n.; Quedius nigriceps Kraatz, 1857 = Q. nigriceps maderensis Smetana, 1963, syn. n.; Atheta immucronata Pace, 1999 = A. pseudolaticollis Erber & Hinterseher, 1992, syn. n., = A. atlantidum Smetana, 2004, syn. n. Para todas as espécies abrangidas pelo catálogo, são indicadas as referências bibliográficas, novas adições para a fauna da Madeira (i. e. por publicar), se disponíveis, são referidas, a sua distribuição nas Ilhas Atlânticas e noutros locais é sumariada e compilados dados sobre a sua biologia. Excluindo 5 nomes de estatuto taxonómico duvidoso, são referidas para o arquipélago um total de 210 espécies e subespécies, 1987 das quais da Madeira, 52 do Porto Santo e 11 das Desertas. Quatro espécies, duas das quais endémicas, são referidas para as Selvagens. Uma análise zoogeográfica revelou que a fauna de Staphylinidae da Madeira é caracterizada por uma elevada proporção (30%) de espécies endémicas, sendo o restante composto por espécies de distribuição Oeste – Paleárctica (25%), Cosmopolita (18%), Holárctica (11%), Paleárctica (4%), e Oeste – Mediterrânea 4%. 9% das espécies não voltaram a ser encontradas desde 1900, o que sugere que as suas populações (madeirenses) podem ter-se extinguido. Apresentam-se mapas de distribuição de vários endemismos da Madeira. A ocorrência, distribuição e desenvolvimento alar das espécies presentes nos arquipélagos encontra-se sumariada numa checklist. PALAVRAS CHAVE: Coleoptera, Staphylinidae, Madeira, Selvagens, ilhas, taxonomia, catálogo, novos sinónimos, novas referências, endemismo. 2006 Assing & Schülke, Systematic catalogue of the Entomofauna 7 CONTENTS CONTENTS .......................................................................................................................................7 INTRODUCTION ............................................................................................................................8 Depositories of types and non-type material .......................................................................................10 The Staphylinidae of the Madeiras and the Selvagens ........................................................................11 Checklist of the Staphylinidae of the Madeira and Selvagens archipelagos .......................................15 Catalogue .............................................................................................................................................20 Subfamily Proteininae ...................................................................................................................21 Subfamily Pselaphinae...................................................................................................................23 Subfamily Omaliinae .....................................................................................................................26 Subfamily Osoriinae ......................................................................................................................31 Subfamily Oxytelinae.....................................................................................................................32 Subfamily Steninae ........................................................................................................................41 Subfamily Paederinae....................................................................................................................49 Subfamily Staphylininae ...............................................................................................................56 Subfamily Tachyporinae ...............................................................................................................79 Subfamily Trichophyinae..............................................................................................................93 Subfamily Habrocerinae ...............................................................................................................94 Subfamily Aleocharinae ................................................................................................................95 Acknowledgments..............................................................................................................................137 References..........................................................................................................................................138 Index to scientific names ...................................................................................................................153 8 Boletim do Museu Municipal do Funchal (História Natural) Supl. No. XI INTRODUCTION Insects were first reported from Madeira in the middle of the 19th century. The real starting point for the study of Madeiran beetles is the year 1847 when T. V. Wollaston organised his first trip to the archipelago. During the period from 1847 to 1871 he visited the islands seven times, each time for some months. In several longer papers (WOLLASTON, 1854, 1857, 1865, 1871A) and numerous shorter contributions he provided a synopsis of the Madeiran Coleoptera as a whole, including the Staphylinidae, based on material collected by himself as well as by Bewicke, Lowe, Moniz, Anderson, Paiva, Park and others. The Wollaston collection is today divided into three major fractions housed at the Natural History Museum, London, the Hope Museum, Oxford, and the collection of the California Academy of Sciences, San Francisco (KAVANAUGH, 1979). Additional material, including types, is deposited in various other collections all over the world, e. g. in the collections of the Deutsches Entomologisches Institut, Müncheberg. At the turn of the century, FAUVEL (1897A, B) published catalogues of the Staphylinidae of Madeira, Porto Santo, the Desertas, and the Selvagens, which he subsequently included in the fourth and fifth editions of his catalogue of the Staphylinidae of North Africa (FAUVEL, 1897C, 1902). In the 20th century, the study of Madeiran Staphylinidae was continued mainly through
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