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Goldfish Morphology As a Model for Evolutionary Developmental Biology
Overview Goldfish morphology as a model for evolutionary developmental biology Kinya G. Ota* and Gembu Abe Morphological variation of the goldfish is known to have been established by artificial selection for ornamental purposes during the domestication process. Chinese texts that date to the Song dynasty contain descriptions of goldfish breeding for ornamental purposes, indicating that the practice originated over one thousand years ago. Such a well-documented goldfish breeding process, combined with the phylogenetic and embryological proximities of this species with zebrafish, would appear to make the morphologically diverse goldfish strains suitable models for evolutionary developmental (evodevo) studies. How- ever, few modern evodevo studies of goldfish have been conducted. In this review, we provide an overview of the historical background of goldfish breed- ing, and the differences between this teleost and zebrafish from an evolutionary perspective. We also summarize recent progress in the field of molecular devel- opmental genetics, with a particular focus on the twin-tail goldfish morphology. Furthermore, we discuss unanswered questions relating to the evolution of the genome, developmental robustness, and morphologies in the goldfish lineage, with the goal of blazing a path toward an evodevo study paradigm using this tel- eost species as a new model species. © 2016 The Authors. WIREs Developmental Biology pub- lished by Wiley Periodicals, Inc. How to cite this article: WIREs Dev Biol 2016, 5:272–295. doi: 10.1002/wdev.224 INTRODUCTION processes of goldfish strains have been documented by authors in many different countries using different fi – he gold sh (Carassius auratus) is a well-known, languages.1 9 Of these reports, the descriptions by Tornamental, domesticated teleost species, which Smartt2 are the most up-to-date and cover the widest consists of a number of morphologically divergent range of the literature. -
TOP-VIEW GOLDFISH: the OTHER PERSPECTIVE Steve Hopkins
TOP-VIEW GOLDFISH: THE OTHER PERSPECTIVE Steve Hopkins By some accounts, there are over three hundred varieties of goldfish. These can be grouped in various ways such as by tail type, presence or absence of head growth, presence or absence of dorsal fin, eye shape, etc. They can also be grouped based on a whether they were bred and selected to be viewed from the top or viewed from the side. Originally, all goldfish were kept in shallow ponds, ceramic bowls or other containers and viewed from the top. Considering the thousand-year history of goldfish keeping, the glass aquarium is a relatively new innovation which did not come into use until about 150 years ago. However, being able to easily view goldfish from the side through glass has undoubtedly influenced what characteristics are selected for and impacted the development of new varieties. Today, the goldfish hobbyists are a diverse group. While most goldfish are destined for the home aquarium and represent an indoor diversion, goldfish ponds, tubs and goldfish in the water garden continue to increase in popularity. When choosing a goldfish, it is important to consider how it will be viewed and select a variety which is appropriate for the setting in which it will be displayed. In selecting a top-view goldfish, remember that they are typically seen against a dark background. It does not matter what color your tub or pond was when it was new, over time the surfaces will become covered with algae and other growth and appear dark green to black. Without doubt, red and white metallic-scale goldfish provide the contrast to display best against a dark background. -
Pond Department Fish Tips
Pond Department Fish Tips 1. Don’t overstock your pond! Without adding excess filtration and maintenance headaches, the arguable rule of thumb is 5 gallons per inch of fish for a maximum load. This equates to approximately 150 gallons for Koi and 40-60 gallons for smaller fish like Comets and Shubunkin. Donate or sell your unwanted babies. Save spending money on dozens and dozens of smaller fish and use it on some nice large ones instead. 2. Koi are living art. Consider your choices in breeds and colors to compliment your surrounding landscape and home. Think of them as a moving painting or sculpture. 3. Fish are social creatures, always buy and introduce them into your ponds in quantities of two to three of the same types. 4. If you introduce too many fish into your pond at once, you may cause undue stress on the ecosystem AND the fish! Ammonia spikes are a common result. Introduce your fish one to two weeks apart 5. Can’t accommodate the mature growth of koi in your little pond? Consider more exotic breeds of goldfish for the more interesting fish in a pond. Oranda, Ranchu and Ryukin are a few good eye-catchers to go with your run-of-the-mill Comets and Shubunkin. 6. Gambusias, better known as mosquito fish, are a great and cheap way to control mosquitoes in your water garden or ecosystem pond. Also try mosquito dunks. 7. If you leave room in your budget for larger Koi (12-16”) you won’t believe what those fish will do to string algae. -
4-22-19 Blue Ridge Koi Availability
Premium Select Koi Sale 15% off 6-8” Premium Select Koi $20.49 each after discount 15% off 12-14” Premium Select Koi $86.28 each after discount * Pictures representative of quality available. Exact fish not pictured. Blue Ridge Koi's 2019 UPS Fish Rates # of Full Boxes of Koi, Butterfly Koi, Goldfish, Catfish, Snails, Tadpoles, Crayfish and Orfe 1 to 3 4 or more 1 to 3 4 or more Price Per Box For All Boxes In Shipment Price Per Box For All Boxes In Shipment AL $60.00 $35.00 NC $35.00 $25.00 AR $60.00 $35.00 ND $70.00 $40.00 AZ $70.00 $40.00 NE $70.00 $40.00 CA $70.00 $40.00 NH $60.00 $35.00 CO $60.00 $35.00 NJ $60.00 $35.00 CT $60.00 $35.00 NM $60.00 $35.00 DE $60.00 $35.00 NV $70.00 $40.00 FL $60.00 $35.00 NY $60.00 $35.00 GA $60.00 $35.00 OH $60.00 $35.00 IA $60.00 $35.00 OK $60.00 $35.00 ID $70.00 $40.00 OR $70.00 $40.00 IL $60.00 $35.00 PA $60.00 $35.00 IN $60.00 $35.00 RI $60.00 $35.00 KS $60.00 $35.00 SC $35.00 $25.00 KY $35.00 $25.00 SD $60.00 $35.00 LA $60.00 $35.00 TN $35.00 $25.00 MA $60.00 $35.00 TX $60.00 $35.00 MD $60.00 $35.00 UT $70.00 $40.00 ME $60.00 $35.00 VA $35.00 $25.00 MI $60.00 $35.00 VT $65.00 $35.00 MN $60.00 $35.00 WA $70.00 $40.00 MO $60.00 $35.00 WI $60.00 $35.00 MS $60.00 $35.00 WV $35.00 $25.00 MT $70.00 $40.00 WY $70.00 $40.00 Please note - 1. -
FL 57. Care of Goldfish
UNITED STATES DEPARTMENT OF THE INTERIOR FISH AND WILDLIFE SERVICE BUREAU OF SPORT FISHERIES AND WILDLIFE Branch of Fish Hatcheries Leaflet FL- 57 Washington 25, D. C. Revised February 1962 CARE OF GOLDFISH HISTlW- Though now found in almost every country in the world, i n captivity and out, the goldfish, Carassius auratus, originated in easter n China as a red mutation of a normally greenish-silvery fish closely rel a ted to and generally resembling the carp. With the carp and many other smaller freshwater fishes including the "barbs" so well-known to t ropical fiSh hobbyists, the goldfish belongs to the great family Cyprinida e which, oddly enough, is found much more abundantly in the Northern Hemisphere than in the Southern. Xanthic, or orange, color mutations occasionally occur among a variety of wild fishes including the carp its elf, but a s i n the case of albinos , or pure white fishes, these do not oft en live to reproduce their kind under natural conditions, since t heir conspicuous appearance makes them easy t argets for predat ors. In captivity, on the other hand, goldfishes have been bred for many centuries and, by t aking advantage of certain additional mutations of a hereditary character, the breeders have produced a number of odd and colorful varieties , i ncl uding some with odd scales, double tails and fins, no s cales at all, or with certain fins missing altogether, shortened backbones , and enlarged eyes. Though known in the Orient for centuries, the goldfish is a com paratively recent arriv ~ l to the western world. -
Goldfish Varieties Poster
m Indu riu str ua ie q s GOLDFISH VARIETIES - (Carassius auratus) A STRAIGHT TAILS Common Goldfish FANTAILS Redcap Fantail PEARLSCALES Most fantail varieties have short globular bodies. Tail and Top of the head deep red, body Have the general characteristics of a fantail with a softer (ALSO KNOWN AS SINGLE TAILS) Body not as long or slender more globular body and characteristic, raised, convex as that of a comet, tail fin is other fins paired except for dorsal fin, which is single. and fins pure white. ECCTTOORRSS EEDDIITTI Common goldfish, comets and shubunkins have relatively (domed) scales. CCOOLLLLE IOONN long slender bodies. Tail fin is single. relatively short. Veiltail Pearlscale Ryukin Body short and globular. Tail fin As described above. Comet Fantail Body short and deep (a depth ¾ double, very broad, with straight-cut Redcap Comet (Tancho trailing edges. Length 1 to 1.5 times Body long and slender, tail fin is As described above. or more than body length) with Comet in Japan) body length. To date this variety has long and well spread. characteristic hump contour on the Top of the head, deep red, body back. The magnitude of the hump not been produced commercially. and fins pure white. increases as the fish matures. Tail is approximately half the length of the body length. Ping Pong Pearlscale Calico The name Ping Pong is used Mirrorscale Comet where the pearlscale’s body shape Scales mainly transparent Tail fin is long and well spread. Shubunkin is extremely round. with many colours same as A row of prominent large scales Scales mainly transparent. -
One of the Characteristics in the Body of the Gold-Fish Is the Caudal and Anal Fins
No. 10.] 655 187 . Preliminary Note on the Inheritance of Caudal and Anal Fins in Gold-fish of Japan . By Yoshiichi MATSUI. Picsicultural Branch of Imperial Fisheries Experimental Station , Toyahashi. (Comm. by C. ISHIKAWA, M.LA., Dec. 12, 1933.) One of the characteristics in the body of the gold-fish is the caudal and anal fins. In a fish the caudal and the anal fins are , like the dorsal fin, simple and vertical, but in some gold-fishes thecaudal fin is more or less horizontal and the anal fins are paired . The caudal fins of the gold-fish are classified into several sorts: the Funa-wo (simple tail), the Tsumami-wo (imperfect shaped tail of Mitsu-wo), the Mitsuo-wo (tri-lobed tail), the Sakura-wo (intermediate shape between Mitsu-wo and Yotsu-wo), the Yotsu-wo (tetra-lobed tail) , the Kujaku-wo (X-shaped tail), etc.Except the Funa-wo, which is vertical, all sorts of the caudal fin of the gold-fishes are horizontal and paired. The anal fin is also of either single or double, with the intermediate formsof y-shaped and v-shaped, and in rare cases anal fin is not present. The materials for the experiments were a number of Wakin, Ryukin, Ranchu, Oranda-shishigashira, allsorts of Demekin, Jikin, Shukin, Hibuna and Funa. The results of many experiments show that the paired caudal fins are apparently homozygous for the most part, with some few excep tions. And it is worthy of attention that they sometimes segregate the Funa-wo. The Funa-wo of gold-fish shown in following tables seems to be heterozygous in nature. -
Bulletin of the United States Fish Commission Seattlenwf V.24
THE CULTIVATION OF MARINE AND FRESH-WATER ANIMALS IN JAPAN. By I'C. MITSUI'CURI, Ph. D., Professor of Zoology. Imperial U1tiversiIJI. Tokyo, Japa», 257 B. B. F.190'1-17 CONTENTS. Page, Introduction . 261 The snapping turtle ' . 262 The gold·fish .................................................•••........... ' . 268 The carp - : .. 275 The eel ' . 276 The gray mullet '"'''' , . 277 The.salmon and trout . 278 Pisciculture in Formosa , '''''''''''''' . 279 The oyster. .......... .. , . 280 The pearl oyster , , . 285 The ark-shell . 286/ The razor clam........•................................................................... 288 Barnacles. .... ........ .. .................... .. ...... .. 289 Miscellaneous: Pinna, egg-cases of gastropods, sea dam, round clam, basket clam, trepang, laver, "fllnori" . 289 258 BuII.U. 5 B.F.1904. P LAT E I. 1. VIE W OF A TURTLE FARM , FUKAGAWA, T OKYO,JA PAN . 2. VIEW IN A GOL D- FISH BREE DER'S ESTA BLI SHMENT . THE CULTIVATION OF MARINE AND FRESH-WATER ANIMALS IN JAPAN.a By K. MITSUKURI, Ph. D.• Professor of 7,ooZO{/!/, Imperial Univer.9ity, Tokyo, Japan. While the pasturage of cattle and the cultivation of plants marked very early steps in man's advancement toward civilization, the raising of aquatic animals and plants, on any extensive scale at all events, seems to belong to much later stages of human development. In fact, the cultivation of some marine animals has been ren dered possible only by utilizing the most recent discoveries and methods of science. I believe, however, the time is now fast approaching when the increase of population on the earth, and the question of food supply which must arise as a necessary conse quence, will compel us to pay most serious attention to the utilization for this purpose of what has been termed the "watery waste." For man to overfish and then to wait for the bounty of nature to replenish, or, failing that, to seek new fishing grounds, is, it seems to me, an act to be put in the same category with the doings of nomadic peoples wandoring from place to place in search of pastures. -
National Goldfish Standards & Technical Information
NATIONAL GOLDFISHSTANDARDS & TECHNICAL INFORMATION BOOKLET No: 4 EleventhEdition 2002 FederatioDof British Aquatic Societies '[i{milbnri Eir'ni^tp{(hrFid FOREWORD This is the eleventh repdnt and tlle third revision of the Federation's Goldish Standards.Fist pdflted in 1947they were unique in that many ol the feau.res tust appea.ringin the standads hav€ been adopted by other orglnjsatioDs both at home and thorEhout the world. The five hverty pointing system being but one ofthem. Th€ 1947st ndaralswerc subject to a major revision in 1954to recogllisethe advancementsthat had beenmade in gold6sh breeding. In 1973 when the last revision took place some adjustnents werc made to exrstingstandards, but pinaily the rcvision was to introduce rcw standardsto cater for severalnew vadetiesthat werebeins imporledftom the Far fast in quandry Somer\ /mry yearslatrer we ari aware of yet firther variation in some of the standards,most notable the fi$age of be Bistol Sh"bunkin, the Tancho CMet ^ d. Tatcho Orunda afld tbe eye sacsof the Subble-eft. To enableB to recogniseand cater for these alterations the Federation's Judges & Si:ndards Committee have lmdertakena major ovemll ofthe Goldfsh Shndards. The conunittee has sought opinion iom goldfsh keepels both within the Federahonaid odemally to il and whilst not claiming to have accepteda1i of the views put forward l'e have r]sed those, which we consideredwere best suited to our requirements,thjs has resultedin some modifcation of both some dfawings ard texl with a view ofrendedng them morc faciie in use ard to seek their acce?tability to the widest possible spectmm of goidfsh opinion. -
Aquarist's Guide
AQUARIST’S GUIDE TO FISH FEEDING GB PROFESSIONAL AQUARIUM TROPICAL – AQUARIST’S GUIDE TO FISH FEEDING Dear Aquarist, beautiful and healthy fish in the aquarium the highest quality natural raw materials. The formulation are the result of the combined influence of many factors. of each food is the distillation of many years’ experience, One of the most important of these is a well-balanced knowledge, and passion for aquarium fish. High quality diet that suits the fishes’ needs. This guide will help you and proven effectiveness are the quintessential virtues of to choose Tropical foods most suitable for your fish and Tropical foods. And the special feature that distinguishes teach you how to put them together to compose a com- them from other foods is the passion we put into their cre- plete healthy diet. Tropical foods are manufactured from ation. This makes Tropical products unique. Raw materials presented in the picture are the example of ingredients used for production of Tropical’s foods. 1 2 3 4 5 6 7 8 9 10 1. egg • 2. algae • 3. spinach • 4. beta-carotene • 5. natural xanthophylls from marigold flowers • 6. shrimp meal 7. spirulina • 8. astaxanthin • 9. krill meal • 10. lysine 2 TROPICAL – AQUARIST’S GUIDE TO FISH FEEDING CONTENTS Foreword 4 Basic principles of fish feeding 6 Multi-ingredient foods 8 Vegetable foods 14 Colour-enhancing foods 18 Natural foods 21 Nano aquaria and foods for the fry 22 Foods for freshwater and marine crustaceans 24 Water treatment products 26 Accessories 28 Filtration media and sponge filters 29 Sample diets 30 TROPICAL – AQUARIST’S GUIDE TO FISH FEEDING FOREWORD We still continue in our efforts to improve the formulas of our foods. -
Single Tailed Goldfish Are Very Closely Related to the Common Goldfish, Or Wild Goldfish
How To Take Care Of Goldfish http://www.howtotakecareofgoldfish.com Page 1 How To Take Care Of Goldfish When You Reach The End Of This Book, You Will Know How To Take Care Of Goldfish Easily! Are You Ready? Let's Start! Legal Notice: This e-book is copyright protected. This is only for personal use. You cannot amend, distribute, sell, use, quote or paraphrase any part or the content within this e-book without the consent of the author or copyright owner. http://www.howtotakecareofgoldfish.com Page 2 How To Take Care Of Goldfish Table of Contents Chapter 1 Introduction All About Goldfish Types of Goldfish How To Select A Goldfish Chapter 2 What Is An Aquarist Selecting an Aquarium How To Setup Your Aquarium Circulation Filtration Lighting Plants – Artificial vs. Living Maintaining Your Aquarium Fishbowls and Tanks Chapter 3 Is A Pond Right For You Types of Ponds Maintaining Your Pond Pond Supplies Conditioning and Treating Water In A Pond Chapter 4 What Do Goldfish Eat Chapter 5 Sick Goldfish and How To Care For Their Illness Common Diseases, Symptoms and Treatments How Can You Tell If Your Goldfish Are Pregnant Caring for Pregnant Goldfish Dying Goldfish and Euthanasia Chapter 6 Goldfish Trivia http://www.howtotakecareofgoldfish.com Page 3 How To Take Care Of Goldfish CHAPTER ONE INTRODUCTION If you are reading this, then most likely you are one of the many people who love goldfish. You are in good company because goldfish make excellent pets. Actually, goldfish are the most popular domesticated aquatic life in the world. -
REMEMBERING Brian Bates
REMEMBERING Brian Bates LOCK DOWN FISH KEEPING- KEITH MULLIN BREEDING CALICO VEIL TAILS JUDGING VEILTAILS BY BRIAN YOUNG FILTER DESIGN -BY IAN MILDON MICK SMITH'S FROZEN FOOD RECIPE VIDEO SHOWS AND MORE! JUNE 2020 CONTENTS OF THIS ISSUE DISCLAIMER FOREWORD BY THE AMGK CHAIRMAN 3 Please note that the THIS MONTHS NEWS LETTER INTRO 4 views, opinions and LETTER FROM NATIONWIDE CHAIRMAN 5 individuals' comments expressed in this BREEDING CALICO VEIL TAILS - BY KEITH MULLIN 6 newsletter do not JUDGING THE VEIL TAILED GOLDFISH BY BRIAN YOUNG 11 necessarily express the views of the AMGK A GOLDFISH JUDGE'S VIEW BY BRIAN YOUNG 15 Society or its Committee REMEMBERING BRIAN BATES 18 and no liability will be accepted by AMGK about FISH HOUSE MODIFICATIONS BY BRIAN YOUNG 23 the information contained COVID 20 FILTER BY IAN MILDON 26 herein. MICK'S FROZEN FOOD RECIPE 28 BAS VIRTUAL SHOW SCHEDULE 29 JUNE 2020 As Featured on Club Chairman Phil Riley Club President Andrew Barton Club Treasurer Gary Malpas Club Secretary Linda Malpas CONTRIBUTE TO THE GOLDFISH STANDARD, W R I T E A N ARTICLE TODAY Send in your Letters and Photos and feature in the Next Issue! EDITORIAL SUPPORT [email protected] GUEST EDITORS Jackie Pedley Grahame Draper MEETING VENUE St. Bartholomew's Church, Binley, Brinklow Rd, Coventry CV3 2DT SHOW VENUE West Orchard Church Hall 83 Baginton Rd, Coventry CV3 6FP amgk.co.uk AMGK Chairman’s Report – Phil Riley Just one or two points for now in the absence of our usual meetings Firstly, I hope that all is well with you all during these disturbing times and that you are managing to breed and rear some excellent fish this season.