The Digital Atlas of Ancient Life: Delivering Information on Paleontology and Biogeography
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Sequence of Post-Moult Exoskeleton Hardening Preserved in a Trilobite Mass Moult Assemblage from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco
Editors' choice Sequence of post-moult exoskeleton hardening preserved in a trilobite mass moult assemblage from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco HARRIET B. DRAGE, THIJS R.A. VANDENBROUCKE, PETER VAN ROY, and ALLISON C. DALEY Drage, H.B., Vandenbroucke, T.R.A., Van Roy, P., and Daley, A.C. 2019. Sequence of post-moult exoskeleton hardening preserved in a trilobite mass moult assemblage from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco. Acta Palaeontologica Polonica 64 (2): 261–273. Euarthropods have a tough exoskeleton that provides crucial protection from predation and parasitism. However, this is restrictive to growth and must be periodically moulted. The moulting sequence is well-known from extant arthropods, consisting of: (i) the long inter-moult stage, in which no changes occur to the hardened exoskeleton; (ii) the pre-moult stage where the old exoskeleton is detached and the new one secreted; (iii) exuviation, when the old exoskeleton is moulted; and (iv) the post-moult stage during which the new exoskeleton starts as soft, thin, and partially compressed and gradually hardens to the robust exoskeleton of the inter-moult stage. Trilobite fossils typically consist of inter-moult carcasses or moulted exuviae, but specimens preserving the post-moult stage are rare. Here we describe nine specimens assigned to Symphysurus ebbestadi representing the first group of contemporaneous fossils collected that preserve all key stages of the moulting process in one taxon, including the post-moult stage. They were collected from a single lens in the Tremadocian part of the Fezouata Shale Formation, Morocco. Based on cephalic displacement and comparison to other trilobite moults, one specimen appears to represent a moulted exoskeleton. -
22 April 2013 the Note from CONE the Editor COLLECTOR Dear Friends
THE CONE COLLECTOR #22 April 2013 THE Note from CONE the Editor COLLECTOR Dear friends, Editor The project “The Cone Collector” is still under seven years old António Monteiro and yet when I look at all we have achieved so far I cannot help thinking that we have probably exceeded expectations. Layout André Poremski We started modestly – as becomes any serious project – back in Contributors October 2006, with our newsletter aimed at all those who are Carlos Afonso interested in studying or collecting Cones, from professional Jim Cootes biologists to amateur collectors. Today we can proudly display Remy Devorsine a total of twenty-four numbers of TCC, two hugely successful Sébastien Dutertre international meetings and a website that brings together an Günther Herndl unparalleled wealth of information on Cones. Joaquin M. Inchaustegui Bruce Livett As a matter of fact, after the uploading in our website (at www. Philippe Quiquandon Christopher Roux theconecollector.com ) of the important and vastly updated Manuel Jiménez Tenorio and augmented work by Mike Filmer’s involving taxonomy and Will van Damme nomenclature, we now have at the same address Paul Kersten’s Alessandro Zanzi extremely useful and well-known Checklist, enriched with new images and much more detailed information than before. This is the work of a team – the names of Manuel Jimenez Tenorio, Bill Fenzan, John Tucker, Gavin Malcolm, Mike Filmer, Paul Kersten and André Poremski readily come to my mind as front row collaborators of TCC, but all others who have contributed with articles, photos, opinions, suggestions and unfailing support deserve equal credit! The project belongs to all and can only survive with the continued support of all. -
The Case of the Diminutive Trilobite Flexicalymene Retrorsa Minuens from the Cincinnatian Series (Upper Ordovician), Cincinnati Region
EVOLUTION & DEVELOPMENT 9:5, 483–498 (2007) Evaluating paedomorphic heterochrony in trilobites: the case of the diminutive trilobite Flexicalymene retrorsa minuens from the Cincinnatian Series (Upper Ordovician), Cincinnati region Brenda R. Hundaa,Ã and Nigel C. Hughesb aCincinnati Museum Center, 1301 Western Avenue, Cincinnati, OH 45203, USA bDepartment of Earth Sciences, University of California, Riverside, CA 92521, USA ÃAuthor for correspondence (email: [email protected]) SUMMARY Flexicalymene retrorsa minuens from the upper- rate of progress along a common ontogenetic trajectory with most 3 m of the Waynesville Formation of the Cincinnatian respect to size, coupled with growth cessation at a small size, Series (Upper Ordovician) of North America lived ‘‘sequential’’ progenesis, or non-uniform changes in the rate of approximately 445 Ma and exhibited marked reduction in progress along a shared ontogenetic trajectory with respect to maximum size relative to its stratigraphically subjacent sister size, can also be rejected. Rather, differences between these subspecies, Flexicalymene retrorsa retrorsa. Phylogenetic subspecies are more consistent with localized changes in analysis is consistent with the notion that F. retrorsa retrorsa rates of character development than with a global hetero- was the ancestor of F. retrorsa minuens. F. retrorsa minuens chronic modification of the ancestral ontogeny. The evolution has been claimed to differ from F. retrorsa retrorsa ‘‘in size of F. retrorsa minuens from F. retrorsa retrorsa was largely alone,’’ and thus presents a plausible example of global dominated by modifications of the development of characters paedomorphic evolution in trilobites. Despite strong similarity already evident in the ancestral ontogeny, not by the origin of in the overall form of the two subspecies, F. -
001-012 Primeras Páginas
PUBLICACIONES DEL INSTITUTO GEOLÓGICO Y MINERO DE ESPAÑA Serie: CUADERNOS DEL MUSEO GEOMINERO. Nº 9 ADVANCES IN TRILOBITE RESEARCH ADVANCES IN TRILOBITE RESEARCH IN ADVANCES ADVANCES IN TRILOBITE RESEARCH IN ADVANCES planeta tierra Editors: I. Rábano, R. Gozalo and Ciencias de la Tierra para la Sociedad D. García-Bellido 9 788478 407590 MINISTERIO MINISTERIO DE CIENCIA DE CIENCIA E INNOVACIÓN E INNOVACIÓN ADVANCES IN TRILOBITE RESEARCH Editors: I. Rábano, R. Gozalo and D. García-Bellido Instituto Geológico y Minero de España Madrid, 2008 Serie: CUADERNOS DEL MUSEO GEOMINERO, Nº 9 INTERNATIONAL TRILOBITE CONFERENCE (4. 2008. Toledo) Advances in trilobite research: Fourth International Trilobite Conference, Toledo, June,16-24, 2008 / I. Rábano, R. Gozalo and D. García-Bellido, eds.- Madrid: Instituto Geológico y Minero de España, 2008. 448 pgs; ils; 24 cm .- (Cuadernos del Museo Geominero; 9) ISBN 978-84-7840-759-0 1. Fauna trilobites. 2. Congreso. I. Instituto Geológico y Minero de España, ed. II. Rábano,I., ed. III Gozalo, R., ed. IV. García-Bellido, D., ed. 562 All rights reserved. No part of this publication may be reproduced or transmitted in any form or by any means, electronic or mechanical, including photocopy, recording, or any information storage and retrieval system now known or to be invented, without permission in writing from the publisher. References to this volume: It is suggested that either of the following alternatives should be used for future bibliographic references to the whole or part of this volume: Rábano, I., Gozalo, R. and García-Bellido, D. (eds.) 2008. Advances in trilobite research. Cuadernos del Museo Geominero, 9. -
Introduction to the Trilobites: Morphology, Ecology, Macroevolution and More by Michelle M
Introduction to the Trilobites: Morphology, Ecology, Macroevolution and More By Michelle M. Casey1, Perry Kennard2, and Bruce S. Lieberman1, 3 1Biodiversity Institute, University of Kansas, Lawrence, KS, 66045, 2Earth Science Teacher, Southwest Middle School, USD497, and 3Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, KS 66045 Middle level laboratory exercise for Earth or General Science; supported provided by National Science Foundation (NSF) grants DEB-1256993 and EF-1206757. Learning Goals and Pedagogy This lab is designed for middle level General Science or Earth Science classes. The learning goals for this lab are the following: 1) to familiarize students with the anatomy and terminology relating to trilobites; 2) to give students experience identifying morphologic structures on real fossil specimens 3) to highlight major events or trends in the evolutionary history and ecology of the Trilobita; and 4) to expose students to the study of macroevolution in the fossil record using trilobites as a case study. Introduction to the Trilobites The Trilobites are an extinct subphylum of the Arthropoda (the most diverse phylum on earth with nearly a million species described). Arthropoda also contains all fossil and living crustaceans, spiders, and insects as well as several other extinct groups. The trilobites were an extremely important and diverse type of marine invertebrates that lived during the Paleozoic Era. They only lived in the oceans but occurred in all types of marine environments, and ranged in size from less than a centimeter to almost a meter across. They were once one of the most successful of all animal groups and in certain fossil deposits, especially in the Cambrian, Ordovician, and Devonian periods, they are extremely abundant. -
Th TRILO the Back to the Past Museum Guide to TRILO BITES
With regard to human interest in fossils, trilobites may rank second only to dinosaurs. Having studied trilobites most of my life, the English version of The Back to the Past Museum Guide to TRILOBITES by Enrico Bonino and Carlo Kier is a pleasant treat. I am captivated by the abundant color images of more than 600 diverse species of trilobites, mostly from the authors’ own collections. Carlo Kier The Back to the Past Museum Guide to Specimens amply represent famous trilobite localities around the world and typify forms from most of the Enrico Bonino Enrico 250-million-year history of trilobites. Numerous specimens are masterpieces of modern professional preparation. Richard A. Robison Professor Emeritus University of Kansas TRILOBITES Enrico Bonino was born in the Province of Bergamo in 1966 and received his degree in Geology from the Depart- ment of Earth Sciences at the University of Genoa. He currently lives in Belgium where he works as a cartographer specialized in the use of satellite imaging and geographic information systems (GIS). His proficiency in the use of digital-image processing, a healthy dose of artistic talent, and a good knowledge of desktop publishing software have provided him with the skills he needed to create graphics, including dozens of posters and illustrations, for all of the displays at the Back to the Past Museum in Cancún. In addition to his passion for trilobites, Enrico is particularly inter- TRILOBITES ested in the life forms that developed during the Precambrian. Carlo Kier was born in Milan in 1961. He holds a degree in law and is currently the director of the Azul Hotel chain. -
Introduction to the Trilobites: Morphology, Macroevolution and More by Michelle M
Introduction to the Trilobites: Morphology, Macroevolution and More By Michelle M. Casey1 and Bruce S. Lieberman1,2, 1Biodiversity Institute and 2Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, KS 66045 Undergraduate laboratory exercise for sophomore/junior level paleontology course Learning Goals and Pedagogy This lab is intended for an upper level paleontology course containing sophomores and juniors who have already taken historical geology or its equivalent; however it may be suitable for introductory biology or geology students familiar with geological time, phylogenies, and trace fossils. This lab will be particularly helpful to those institutions that lack a large teaching collection by providing color photographs of museum specimens. Students may find previous exposure to phylogenetic methods and terminology helpful in completing this laboratory exercise. The learning goals for this lab are the following: 1) to familiarize students with the anatomy and terminology relating to trilobites; 2) to give students experience identifying morphologic structures on real fossil specimens, not just diagrammatic representations; 3) to highlight major events or trends in the evolutionary history and ecology of Trilobita; and 4) to expose students to the study of macroevolution in the fossil record using trilobites as a case study. Introduction to the Trilobites The Trilobites are an extinct subphylum of the Arthropoda (the most diverse phylum on earth with nearly a million species described). Arthropoda also contains all fossil and living crustaceans, spiders, and insects as well as several other solely extinct groups. The trilobites were an extremely important and diverse type of marine invertebrates that lived during the Paleozoic Era. They were exclusively marine but occurred in all types of marine environments, and ranged in size from less than a centimeter to almost a meter. -
Ordovician Stratigraphy and Benthic Community Replacements in the Eastern Anti-Atlas, Morocco J
Ordovician stratigraphy and benthic community replacements in the eastern Anti-Atlas, Morocco J. Javier Alvaro, Mohammed Benharref, Jacques Destombes, Juan Carlos Gutiérrez-Marco, Aaron Hunter, Bertrand Lefebvre, Peter van Roy, Samuel Zamora To cite this version: J. Javier Alvaro, Mohammed Benharref, Jacques Destombes, Juan Carlos Gutiérrez-Marco, Aaron Hunter, et al.. Ordovician stratigraphy and benthic community replacements in the eastern Anti- Atlas, Morocco. The Great Ordovician Biodiversification Event: Insights from the Tafilalt Biota, Morocco, 485, The Geological Society of London, pp.SP485.20, In press, Geological Society, London, Special Publication, 10.1144/SP485.20. hal-02405970 HAL Id: hal-02405970 https://hal.archives-ouvertes.fr/hal-02405970 Submitted on 13 Nov 2020 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. The Geological Society Special Publications Ordovician stratigraphy and benthic community replacements in the eastern Anti-Atlas, Morocco --Manuscript Draft-- Manuscript Number: GSLSpecPub2019-17R1 Article Type: Research article Full Title: Ordovician stratigraphy and benthic community replacements in the eastern Anti-Atlas, Morocco Short Title: Ordovician stratigraphy of the Anti-Atlas Corresponding Author: Javier Alvaro Instituto de Geociencias SPAIN Corresponding Author E-Mail: [email protected] Other Authors: MOHAMMED BENHARREF JACQUES DESTOMBES JUAN CARLOS GUTIÉRREZ-MARCO AARON W. -
Late Silurian Trilobite Palaeobiology And
LATE SILURIAN TRILOBITE PALAEOBIOLOGY AND BIODIVERSITY by ANDREW JAMES STOREY A thesis submitted to the University of Birmingham for the degree of DOCTOR OF PHILOSOPHY School of Geography, Earth and Environmental Sciences University of Birmingham February 2012 University of Birmingham Research Archive e-theses repository This unpublished thesis/dissertation is copyright of the author and/or third parties. The intellectual property rights of the author or third parties in respect of this work are as defined by The Copyright Designs and Patents Act 1988 or as modified by any successor legislation. Any use made of information contained in this thesis/dissertation must be in accordance with that legislation and must be properly acknowledged. Further distribution or reproduction in any format is prohibited without the permission of the copyright holder. ABSTRACT Trilobites from the Ludlow and Přídolí of England and Wales are described. A total of 15 families; 36 genera and 53 species are documented herein, including a new genus and seventeen new species; fourteen of which remain under open nomenclature. Most of the trilobites in the British late Silurian are restricted to the shelf, and predominantly occur in the Elton, Bringewood, Leintwardine, and Whitcliffe groups of Wales and the Welsh Borderland. The Elton to Whitcliffe groups represent a shallowing upwards sequence overall; each is characterised by a distinct lithofacies and fauna. The trilobites and brachiopods of the Coldwell Formation of the Lake District Basin are documented, and are comparable with faunas in the Swedish Colonus Shale and the Mottled Mudstones of North Wales. Ludlow trilobite associations, containing commonly co-occurring trilobite taxa, are defined for each palaeoenvironment. -
A Twist of Nature: a Left-Handed Bythinella Schmidti (Küster, 1852) (Caenogastropoda: Bythinellidae)
NATURA SLOVENIAE 20(2): 25-31 Prejeto / Received: 13. 8. 2018 SHORT COMMUNICATION Sprejeto / Accepted: 22. 10. 2018 A twist of nature: a left-handed Bythinella schmidti (Küster, 1852) (Caenogastropoda: Bythinellidae) Simona PREVORČNIK1, Andrzej FALNIOWSKI2 1Department of Biology, Biotechnical Faculty, University of Ljubljana, Jamnikarjeva 101, SI-1000 Ljubljana, Slovenia; E-mail: [email protected] 2Department of Malacology, Institute of Zoology, Jagiellonian University, Gronostajowa 9, PL-30-387, Kraków, Poland Abstract. As most extant snails, Bythinella schmidti is characterised by dextral (right-handed) coiling of the shell. Nevertheless, a small sinistral (left-handed) individual from the spring on a mountain pasture was sampled, together with its larger dextral conspecifics. In our report on this first case of sinistrality within the superfamily Truncatelloidea, we discuss its shell abnormalities and provide a review on chirality in snails. Key words: Bythinella, sinistrality, shell abnormalities, spring, Slovenia Izvleček. Zasuk narave: levosučna Bythinella schmidti (Küster, 1852) (Caenogastropoda: Bythinellidae) – Kot za večino obstoječih vrst je tudi za vrsto Bythinella schmidti značilna desnosučnost, torej zavitost hišice v smeri urinega kazalca. A v izviru na planinskem pašniku smo med večjimi desnosučnimi sovrstniki našli tudi majhnega levosučneža te vrste. V poročilu o prvem primeru levosučnosti znotraj naddružine Truncatelloidea obravnavamo opažene nepravilnostih v zgradbi hišice in podajamo pregled objav o sučnosti polžev. Ključne besede: Bythinella, levosučnost, nepravilnosti hišice, izvir, Slovenija Introduction Handedness is the phenomenon relating to the ability to classify chiral objects into right- and left-handed. Most snail species in nature show either uniform right- or left-handedness. Sinistral species are rare, especially in the marine realm, where such taxa have independently originated only 19 times among Cenozoic gastropod clades (Vermeij 2002). -
Introduction to the Trilobites: Morphology, Macroevolution and More by Michelle M
Introduction to the Trilobites: Morphology, Macroevolution and More By Michelle M. Casey1 and Bruce S. Lieberman1,2, 1Biodiversity Institute and 2Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, KS 66045 Undergraduate laboratory exercise for sophomore/junior level paleontology course Learning Goals and Pedagogy This lab is intended for an upper level paleontology course containing sophomores and juniors who have already taken historical geology or its equivalent; however it may be suitable for introductory biology or geology students familiar with geological time, phylogenies, and trace fossils. This lab will be particularly helpful to those institutions that lack a large teaching collection by providing color photographs of museum specimens. Students may find previous exposure to phylogenetic methods and terminology helpful in completing this laboratory exercise. The learning goals for this lab are the following: 1) to familiarize students with the anatomy and terminology relating to trilobites; 2) to give students experience identifying morphologic structures on real fossil specimens, not just diagrammatic representations; 3) to highlight major events or trends in the evolutionary history and ecology of the Trilobita; and 4) to expose students to the study of macroevolution in the fossil record using trilobites as a case study. Introduction to the Trilobites The Trilobites are an extinct subphylum of the Arthropoda (the most diverse phylum on earth with nearly a million species described). Arthropoda also contains all fossil and living crustaceans, spiders, and insects as well as several other solely extinct groups. The trilobites were an extremely important and diverse type of marine invertebrates that lived during the Paleozoic Era. -
• Every Major Animal Phylum That Exists on Earth Today, As Well As A
• Every major animal phylum that exists on Earth today, as well as a few more that have since become ex:nct, appeared within less than 10 million years during the early Cambrian evolu:onary radiaon, also called the Cambrian explosion. Bristleworm • Phylum Annelida is represented by over 22,000 modern species of marine, freshwater, and terrestrial annelids, or segmented worms, including bristleworms, earthworms, and leeches. Earthworm These worms are classified separately from the roundworms in Phylum Nematoda and the flatworms in Phylum Platyhelminthes. Leech • The annelids may be further subdivided into Class Polychaeta, which includes the bristleworms (polychaetes), and Class Clitellata, which includes the earthworms (oligochaetes) and leeches (hirudineans). • The polychaetes, or bristleworms, are PlanKtonic Tomopteris the largest group of annelids and are mostly marine. They include species that live in the coldest ocean temperatures of the abyssal plain, to forms which tolerate the extreme high temperatures near hydrothermal vents. Polychaetes occur throughout the oceans at all depths, from planKtonic species such as Tomopteris that live near the surface to an unclassified benthonic form living at a depth of nearly 11,000 m at the boLom of the Challenger Deep in the Benthic sp. 1,200 m Mariana Trench. • Although many polychaetes are pelagic or benthic, others have adapted to burrowing, boring or tube-dwelling, and parasi:sm. • Each segment of a polychaete bears a pair of paddle-liKe and highly vascularized parapodia, which are used for locomo:on and, in many species, act as the worm’s primary respiratory surfaces. Bundles of chi:nous bristles, called chaetae, project from the parapodia.