Digestive Structures in Ordovician Trilobites Colpocoryphe and Flexicalymene from the Barrandian Area of Czech Republic
Total Page:16
File Type:pdf, Size:1020Kb
Load more
Recommended publications
-
Biostratigraphic Precision of the Cruziana Rugosa Group: a Study from the Ordovician Succession of Southern and Central Bolivia
Geol. Mag. 144 (2), 2007, pp. 289–303. c 2007 Cambridge University Press 289 doi:10.1017/S0016756807003093 First published online 9 February 2007 Printed in the United Kingdom Biostratigraphic precision of the Cruziana rugosa group: a study from the Ordovician succession of southern and central Bolivia SVEN O. EGENHOFF∗, BERND WEBER†, OLIVER LEHNERT‡ &JORG¨ MALETZ§ ∗Colorado State University, Department of Geosciences, 322 Natural Resources Building, Fort Collins, CO 80523-1482, USA †Freie Universitat¨ Berlin, Institut fur¨ Geologische Wissenschaften, Fachrichtung Geologie, Malteserstrasse 74-100, D-12249 Berlin, Germany ‡University of Erlangen, Institute of Geology and Mineralogie, Schlossgarten 5, D-91054 Erlangen, Germany §Department of Geology, State University of New York at Buffalo, 772 Natural Sciences and Mathematics Complex, Buffalo, New York 14260-3050, USA (Received 10 October 2005; revised version received 1 May 2006; accepted 22 May 2006) Abstract – Cruziana ichnospecies have been repeatedly reported to have biostratigraphic significance. This study presents a re-evaluation of the arthropod ichnotaxa of the Cruziana rugosa Group from bio- and/or lithostratigraphically well-defined Lower to Upper Ordovician siliciclastic sections of southern and central Bolivia. With the exception of Cruziana rouaulti, the ichnofaunas contain all the members of the Cruziana rugosa Group throughout the Ordovician (Arenig to Caradoc) successions in Bolivia. The Bolivian material therefore indicates that these arthropod ichnofossil assemblages are suitable for recognizing Ordovician strata in Bolivia. These findings cast doubt on their use as reliable indicators for a global intra-Ordovician (Arenig to Caradoc) biozonation of Peri-Gondwanan sedimentary successions. Keywords: Cruziana, biostratigraphy, Bolivia, Ordovician. 1. Introduction to the present study. -
Sequence of Post-Moult Exoskeleton Hardening Preserved in a Trilobite Mass Moult Assemblage from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco
Editors' choice Sequence of post-moult exoskeleton hardening preserved in a trilobite mass moult assemblage from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco HARRIET B. DRAGE, THIJS R.A. VANDENBROUCKE, PETER VAN ROY, and ALLISON C. DALEY Drage, H.B., Vandenbroucke, T.R.A., Van Roy, P., and Daley, A.C. 2019. Sequence of post-moult exoskeleton hardening preserved in a trilobite mass moult assemblage from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco. Acta Palaeontologica Polonica 64 (2): 261–273. Euarthropods have a tough exoskeleton that provides crucial protection from predation and parasitism. However, this is restrictive to growth and must be periodically moulted. The moulting sequence is well-known from extant arthropods, consisting of: (i) the long inter-moult stage, in which no changes occur to the hardened exoskeleton; (ii) the pre-moult stage where the old exoskeleton is detached and the new one secreted; (iii) exuviation, when the old exoskeleton is moulted; and (iv) the post-moult stage during which the new exoskeleton starts as soft, thin, and partially compressed and gradually hardens to the robust exoskeleton of the inter-moult stage. Trilobite fossils typically consist of inter-moult carcasses or moulted exuviae, but specimens preserving the post-moult stage are rare. Here we describe nine specimens assigned to Symphysurus ebbestadi representing the first group of contemporaneous fossils collected that preserve all key stages of the moulting process in one taxon, including the post-moult stage. They were collected from a single lens in the Tremadocian part of the Fezouata Shale Formation, Morocco. Based on cephalic displacement and comparison to other trilobite moults, one specimen appears to represent a moulted exoskeleton. -
The Case of the Diminutive Trilobite Flexicalymene Retrorsa Minuens from the Cincinnatian Series (Upper Ordovician), Cincinnati Region
EVOLUTION & DEVELOPMENT 9:5, 483–498 (2007) Evaluating paedomorphic heterochrony in trilobites: the case of the diminutive trilobite Flexicalymene retrorsa minuens from the Cincinnatian Series (Upper Ordovician), Cincinnati region Brenda R. Hundaa,Ã and Nigel C. Hughesb aCincinnati Museum Center, 1301 Western Avenue, Cincinnati, OH 45203, USA bDepartment of Earth Sciences, University of California, Riverside, CA 92521, USA ÃAuthor for correspondence (email: [email protected]) SUMMARY Flexicalymene retrorsa minuens from the upper- rate of progress along a common ontogenetic trajectory with most 3 m of the Waynesville Formation of the Cincinnatian respect to size, coupled with growth cessation at a small size, Series (Upper Ordovician) of North America lived ‘‘sequential’’ progenesis, or non-uniform changes in the rate of approximately 445 Ma and exhibited marked reduction in progress along a shared ontogenetic trajectory with respect to maximum size relative to its stratigraphically subjacent sister size, can also be rejected. Rather, differences between these subspecies, Flexicalymene retrorsa retrorsa. Phylogenetic subspecies are more consistent with localized changes in analysis is consistent with the notion that F. retrorsa retrorsa rates of character development than with a global hetero- was the ancestor of F. retrorsa minuens. F. retrorsa minuens chronic modification of the ancestral ontogeny. The evolution has been claimed to differ from F. retrorsa retrorsa ‘‘in size of F. retrorsa minuens from F. retrorsa retrorsa was largely alone,’’ and thus presents a plausible example of global dominated by modifications of the development of characters paedomorphic evolution in trilobites. Despite strong similarity already evident in the ancestral ontogeny, not by the origin of in the overall form of the two subspecies, F. -
Available Generic Names for Trilobites
AVAILABLE GENERIC NAMES FOR TRILOBITES P.A. JELL AND J.M. ADRAIN Jell, P.A. & Adrain, J.M. 30 8 2002: Available generic names for trilobites. Memoirs of the Queensland Museum 48(2): 331-553. Brisbane. ISSN0079-8835. Aconsolidated list of available generic names introduced since the beginning of the binomial nomenclature system for trilobites is presented for the first time. Each entry is accompanied by the author and date of availability, by the name of the type species, by a lithostratigraphic or biostratigraphic and geographic reference for the type species, by a family assignment and by an age indication of the type species at the Period level (e.g. MCAM, LDEV). A second listing of these names is taxonomically arranged in families with the families listed alphabetically, higher level classification being outside the scope of this work. We also provide a list of names that have apparently been applied to trilobites but which remain nomina nuda within the ICZN definition. Peter A. Jell, Queensland Museum, PO Box 3300, South Brisbane, Queensland 4101, Australia; Jonathan M. Adrain, Department of Geoscience, 121 Trowbridge Hall, Univ- ersity of Iowa, Iowa City, Iowa 52242, USA; 1 August 2002. p Trilobites, generic names, checklist. Trilobite fossils attracted the attention of could find. This list was copied on an early spirit humans in different parts of the world from the stencil machine to some 20 or more trilobite very beginning, probably even prehistoric times. workers around the world, principally those who In the 1700s various European natural historians would author the 1959 Treatise edition. Weller began systematic study of living and fossil also drew on this compilation for his Presidential organisms including trilobites. -
Copertina Guida Ai TRILOBITI V3 Esterno
Enrico Bonino nato in provincia di Bergamo nel 1966, Enrico si è laureato in Geologia presso il Dipartimento di Scienze della Terra dell'Università di Genova. Attualmente risiede in Belgio dove svolge attività come specialista nel settore dei Sistemi di Informazione Geografica e analisi di immagini digitali. Curatore scientifico del Museo Back to the Past, ha pubblicato numerosi volumi di paleontologia in lingua italiana e inglese, collaborando inoltre all’elaborazione di testi e pubblicazioni scientifiche a livello nazonale e internazionale. Oltre alla passione per questa classe di artropodi, i suoi interessi sono orientati alle forme di vita vissute nel Precambriano, stromatoliti, e fossilizzazioni tipo konservat-lagerstätte. Carlo Kier nato a Milano nel 1961, Carlo si è laureato in Legge, ed è attualmente presidente della catena di alberghi Azul Hotel. Risiede a Cancun, Messico, dove si dedica ad attività legate all'ambiente marino. All'età di 16 anni, ha iniziato una lunga collaborazione con il Museo di Storia Naturale di Milano, ed è a partire dal 1970 che prese inizio la vera passione per i trilobiti, dando avvio a quella che oggi è diventata una delle collezioni paleontologiche più importanti al mondo. La sua instancabile attività di ricerca sul terreno in varie parti del globo e la collaborazione con professionisti del settore, ha permesso la descrizione di nuove specie di trilobiti ed artropodi. Una forte determinazione e la costruzione di un nuovo complesso alberghiero (AZUL Sensatori) hanno infine concretizzzato la realizzazione -
Enrollment and Coaptations in Some Species of the Ordovician Trilobite Genus Placoparia
Enrollment and coaptations in some sp eeies of the Ordovician trilobite genus Placoparia JEAN-LOUIS HENRY AND EUAN N.K. CLARKSON Henry, J.-L. & Clarkson, E.N.K. 1974 12 15: Enrollment and coaptations in some species of the Ordovician trilobite genus Placoparia. Fossils and Strata, No. 4, pp. 87-95. PIs. 1-3. Oslo ISSN 0300-949l. ISBN 82-00-04963-9. The species Pl. (Placoparia) cambriensis, Pl. (Coplacoparia ) tournemini and Pl. (Coplacoparia ) borni, described by Hammann (1971) from the Spanish Ordovician, are found in Brittany in formations attributed to the Llanvirn and the LIandeilo. The lateral borders of the librigenae of Pl. (Coplacoparia) tournemini and Pl. (Coplacoparia ) borni bear depressions into which the distal ends of the thoracic pleurae and the tips of the first pair of pygidial ribs come and fit during enrollment. These depressions, or coaptative structures sensu Cuenot (1919), are also to be observed in Pl. (Placoparia) zippei and Pl. (Hawleia) grandis, from the Ordovician of Bohemia. However, in the species tournemini and borni, additional depressions appear on the anterior cephalic border, and the coaptations evolve towards an increasing complexity. From Pl. (Placoparia) cambriensis, an ancestrai form with a wide geographic distribution, two distinct populations se em to have individu· alised by a process of allopatric speciation; one of these, probably neotenous, is represented by Pl. (Coplacoparia ) tournemini (Massif Armoricain and Iberian Peninsula), the other by Pl. (Placoparia ) zippei (Bohemia). Geo graphic isolation may be indirectly responsible for the appearance of new coaptative devices. jean-Louis Henry, Laboratoire de Paleontologie et de Stratigraphie, Institut de Geologie de l'Universite, B.P. -
001-012 Primeras Páginas
PUBLICACIONES DEL INSTITUTO GEOLÓGICO Y MINERO DE ESPAÑA Serie: CUADERNOS DEL MUSEO GEOMINERO. Nº 9 ADVANCES IN TRILOBITE RESEARCH ADVANCES IN TRILOBITE RESEARCH IN ADVANCES ADVANCES IN TRILOBITE RESEARCH IN ADVANCES planeta tierra Editors: I. Rábano, R. Gozalo and Ciencias de la Tierra para la Sociedad D. García-Bellido 9 788478 407590 MINISTERIO MINISTERIO DE CIENCIA DE CIENCIA E INNOVACIÓN E INNOVACIÓN ADVANCES IN TRILOBITE RESEARCH Editors: I. Rábano, R. Gozalo and D. García-Bellido Instituto Geológico y Minero de España Madrid, 2008 Serie: CUADERNOS DEL MUSEO GEOMINERO, Nº 9 INTERNATIONAL TRILOBITE CONFERENCE (4. 2008. Toledo) Advances in trilobite research: Fourth International Trilobite Conference, Toledo, June,16-24, 2008 / I. Rábano, R. Gozalo and D. García-Bellido, eds.- Madrid: Instituto Geológico y Minero de España, 2008. 448 pgs; ils; 24 cm .- (Cuadernos del Museo Geominero; 9) ISBN 978-84-7840-759-0 1. Fauna trilobites. 2. Congreso. I. Instituto Geológico y Minero de España, ed. II. Rábano,I., ed. III Gozalo, R., ed. IV. García-Bellido, D., ed. 562 All rights reserved. No part of this publication may be reproduced or transmitted in any form or by any means, electronic or mechanical, including photocopy, recording, or any information storage and retrieval system now known or to be invented, without permission in writing from the publisher. References to this volume: It is suggested that either of the following alternatives should be used for future bibliographic references to the whole or part of this volume: Rábano, I., Gozalo, R. and García-Bellido, D. (eds.) 2008. Advances in trilobite research. Cuadernos del Museo Geominero, 9. -
United States
DEPARTMENT OF THE INTERIOR BULLETIN OF THE UNITED STATES ISTo. 146 WASHINGTON GOVERNMENT Pit IN TING OFFICE 189C UNITED STATES GEOLOGICAL SURVEY CHAKLES D. WALCOTT, DI11ECTOK BIBLIOGRAPHY AND INDEX NORTH AMEEICAN GEOLOGY, PALEONTOLOGY, PETEOLOGT, AND MINERALOGY THE YEA.R 1895 FEED BOUGHTON WEEKS WASHINGTON Cr O V E U N M K N T P K 1 N T I N G OFFICE 1890 CONTENTS. Page. Letter of trail smittal...... ....................... .......................... 7 Introduction.............'................................................... 9 List of publications examined............................................... 11 Classified key to tlio index .......................................... ........ 15 Bibliography ............................................................... 21 Index....................................................................... 89 LETTER OF TRANSMITTAL DEPARTMENT OF THE INTEEIOE, UNITED STATES GEOLOGICAL SURVEY, DIVISION OF GEOLOGY, Washington, D. 0., June 23, 1896. SIR: I have the honor to transmit herewith the manuscript of a Bibliography and Index of North American Geology, Paleontology, Petrology, and Mineralogy for the year 1895, and to request that it be published as a bulletin of the Survey. Very respectfully, F. B. WEEKS. Hon. CHARLES D. WALCOTT, Director United States Geological Survey. 1 BIBLIOGRAPHY AND INDEX OF NORTH AMERICAN GEOLOGY, PALEONTOLOGY, PETROLOGY, AND MINER ALOGY FOR THE YEAR 1895. By FRED BOUGHTON WEEKS. INTRODUCTION. The present work comprises a record of publications on North Ameri can geology, paleontology, petrology, and mineralogy for the year 1895. It is planned on the same lines as the previous bulletins (Nos. 130 and 135), excepting that abstracts appearing in regular periodicals have been omitted in this volume. Bibliography. The bibliography consists of full titles of separate papers, classified by authors, an abbreviated reference to the publica tion in which the paper is printed, and a brief summary of the con tents, each paper being numbered for index reference. -
Introduction to the Trilobites: Morphology, Ecology, Macroevolution and More by Michelle M
Introduction to the Trilobites: Morphology, Ecology, Macroevolution and More By Michelle M. Casey1, Perry Kennard2, and Bruce S. Lieberman1, 3 1Biodiversity Institute, University of Kansas, Lawrence, KS, 66045, 2Earth Science Teacher, Southwest Middle School, USD497, and 3Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, KS 66045 Middle level laboratory exercise for Earth or General Science; supported provided by National Science Foundation (NSF) grants DEB-1256993 and EF-1206757. Learning Goals and Pedagogy This lab is designed for middle level General Science or Earth Science classes. The learning goals for this lab are the following: 1) to familiarize students with the anatomy and terminology relating to trilobites; 2) to give students experience identifying morphologic structures on real fossil specimens 3) to highlight major events or trends in the evolutionary history and ecology of the Trilobita; and 4) to expose students to the study of macroevolution in the fossil record using trilobites as a case study. Introduction to the Trilobites The Trilobites are an extinct subphylum of the Arthropoda (the most diverse phylum on earth with nearly a million species described). Arthropoda also contains all fossil and living crustaceans, spiders, and insects as well as several other extinct groups. The trilobites were an extremely important and diverse type of marine invertebrates that lived during the Paleozoic Era. They only lived in the oceans but occurred in all types of marine environments, and ranged in size from less than a centimeter to almost a meter across. They were once one of the most successful of all animal groups and in certain fossil deposits, especially in the Cambrian, Ordovician, and Devonian periods, they are extremely abundant. -
Contributions in BIOLOGY and GEOLOGY
MILWAUKEE PUBLIC MUSEUM Contributions In BIOLOGY and GEOLOGY Number 51 November 29, 1982 A Compendium of Fossil Marine Families J. John Sepkoski, Jr. MILWAUKEE PUBLIC MUSEUM Contributions in BIOLOGY and GEOLOGY Number 51 November 29, 1982 A COMPENDIUM OF FOSSIL MARINE FAMILIES J. JOHN SEPKOSKI, JR. Department of the Geophysical Sciences University of Chicago REVIEWERS FOR THIS PUBLICATION: Robert Gernant, University of Wisconsin-Milwaukee David M. Raup, Field Museum of Natural History Frederick R. Schram, San Diego Natural History Museum Peter M. Sheehan, Milwaukee Public Museum ISBN 0-893260-081-9 Milwaukee Public Museum Press Published by the Order of the Board of Trustees CONTENTS Abstract ---- ---------- -- - ----------------------- 2 Introduction -- --- -- ------ - - - ------- - ----------- - - - 2 Compendium ----------------------------- -- ------ 6 Protozoa ----- - ------- - - - -- -- - -------- - ------ - 6 Porifera------------- --- ---------------------- 9 Archaeocyatha -- - ------ - ------ - - -- ---------- - - - - 14 Coelenterata -- - -- --- -- - - -- - - - - -- - -- - -- - - -- -- - -- 17 Platyhelminthes - - -- - - - -- - - -- - -- - -- - -- -- --- - - - - - - 24 Rhynchocoela - ---- - - - - ---- --- ---- - - ----------- - 24 Priapulida ------ ---- - - - - -- - - -- - ------ - -- ------ 24 Nematoda - -- - --- --- -- - -- --- - -- --- ---- -- - - -- -- 24 Mollusca ------------- --- --------------- ------ 24 Sipunculida ---------- --- ------------ ---- -- --- - 46 Echiurida ------ - --- - - - - - --- --- - -- --- - -- - - --- -
In Search of a Trilobite (Shropshire)
IN SEARCH OF A TRILOBITE Iain Brown On a May weekend last year I travelled to Shropshire to try The rocks were wet crumbly shales laid down in deep and scratch an old itch of mine which was to find a water sometime in the Ordovician. Finds were scarce but I trilobite. I already have a couple of examples (Calymene did manage to find a rather distorted fossil. The distortion, and Eltrathia) which I received as a present bought from I assume, is due to folding activity sometime in the 460 the Lyme Regis fossil shop, but nothing compares to odd million years it’s been lying around waiting for actually finding the specimens yourself. Words like somebody to find it. However I was told subsequently that context and provenance spring to mind along with more it could have been caused by the fossilisation process personal ones like satisfaction and achievement. I had lived (compaction and lithification); one way to get further in Shropshire previous to moving to the Bath area and had useful information would be from a petrological even tried to find trilobites on my own a couple of times. examination in thin section. I’ve no resources for such an This involved locating disused quarries on the map from activity and as it was my first find, I had very little around the Wrekin area and then if possible having a look. motivation to destroy it even if I had. It may be distorted, Like most things, if I were looking for rusty old farm but I had a feeling that I hoped I looked that good after the machinery and sheep carcasses, I’d have found loads of best part of half a billion years. -
The Evolution of Trilobite Body Patterning
ANRV309-EA35-14 ARI 20 March 2007 15:54 The Evolution of Trilobite Body Patterning Nigel C. Hughes Department of Earth Sciences, University of California, Riverside, California 92521; email: [email protected] Annu. Rev. Earth Planet. Sci. 2007. 35:401–34 Key Words First published online as a Review in Advance on Trilobita, trilobitomorph, segmentation, Cambrian, Ordovician, January 29, 2007 diversification, body plan The Annual Review of Earth and Planetary Sciences is online at earth.annualreviews.org Abstract This article’s doi: The good fossil record of trilobite exoskeletal anatomy and on- 10.1146/annurev.earth.35.031306.140258 togeny, coupled with information on their nonbiomineralized tis- Copyright c 2007 by Annual Reviews. sues, permits analysis of how the trilobite body was organized and All rights reserved developed, and the various evolutionary modifications of such pat- 0084-6597/07/0530-0401$20.00 terning within the group. In several respects trilobite development and form appears comparable with that which may have charac- terized the ancestor of most or all euarthropods, giving studies of trilobite body organization special relevance in the light of recent advances in the understanding of arthropod evolution and devel- opment. The Cambrian diversification of trilobites displayed mod- Annu. Rev. Earth Planet. Sci. 2007.35:401-434. Downloaded from arjournals.annualreviews.org ifications in the patterning of the trunk region comparable with by UNIVERSITY OF CALIFORNIA - RIVERSIDE LIBRARY on 05/02/07. For personal use only. those seen among the closest relatives of Trilobita. In contrast, the Ordovician diversification of trilobites, although contributing greatly to the overall diversity within the clade, did so within a nar- rower range of trunk conditions.