Deyrup: List of Florida 43

AN UPDATED LIST OF FLORIDA ANTS (: FORMICIDAE)

MARK DEYRUP Archbold Biological Station, P.O. Box 2057, Lake Placid, FL 33862

ABSTRACT

A list of ants of Florida published in 1989 is replaced to accommodate 49 additional now known from Florida, and 34 name changes in species already on the 1989 list. Cur- rently, 218 species of ants are reliably reported from Florida.

Key Words: exotic species, faunistics

RESUMEN

Se presenta una nueva lista de las hormigas de Florida para reemplazar una lista publicada en 1989. La nueva lista incluya 49 mas especies y 34 cambios de nomenclatura. Al presente, 218 especies de hormigas son sabidas a hallarse en Florida.

In 1989 Deyrup et al. published a list of the nine exotic species that have been added to the ants of Florida. In the ensuing thirteen years list. Most of the added Florida records are either there has been considerable myrmecological ac- native species whose Florida populations have tivity, both survey work in Florida, and taxonomic been recently discovered, or native species that descriptions and revisions. Consequently, the have recently been described or are awaiting de- 1989 list is drastically out of date: there are 49 ad- scription. ditional species to be added to the list, 34 name The number of species listed below, 218, is the changes that apply to species already on the 1989 largest fauna known from any state in eastern list, and 4 species that have been removed from North America, and is likely to remain so, even af- the list because the records are probably based on ter other large states have received as much at- misidentifications. tention as Florida. The reasons for this lie in the These advances do not mean that there is no convergence of various faunal elements. There is need for further work in the inventory and taxon- a set of Antillean species, such as Leptothorax al- omy of Florida ants. Included in the list are a lardycei and L. torreyi in the southern Peninsula. number of species that are awaiting description There are many tropical exotics, including both by various specialists. The species epithets in the Old World species, such as Technomyrmex albipes entire genus Brachymyrmex are suspect, al- and Strumigenys emmae, and New World species, though it is clear that at least five species of Bra- such as Wasmannia auropunctata and chymyrmex occur in Florida. It is probable that gracilis. There are southeastern there are additional species of ants still to be coastal plain species such as Camponotus socius found in Florida. This can be deduced from the and Paratrechina arenivaga. There are, mostly in fact that there are a number of species on the list north Florida, species probably derived from the that are known from only one or two collections, or southern Appalachians, such as Pyramica ros- from one or two sites; there are probably other trata and P. pulchella. There are species repre- equally rare species that nobody has been lucky senting western lineages, such as Pogonomyrmex enough to find. There is reason to suppose that ex- badius and Neivamyrmex texanus. All this diver- otic ants will continue to become established in sity, however, still cannot compare with that of Florida (Deyrup et al. 2000); even now there are southwestern states such as Arizona, where an al- likely to be some species of localized exotics that most intact, pre-ice age, dryland fauna (Madroter- have not yet been reported. On the other hand, tiary) is augmented at higher elevations by north there are four species of ants on the list that have temperate species. not been found for many years and may have been The following alphabetical list generally fol- extirpated. These are the native species Formica lows the nomenclature in Barry Bolton’s catalog subsericea and Solenopsis xyloni, and the exotic (1995), combined with the nomenclature in his re- species Myrmelachista ramulorum and Tetramor- vision of the Dacetini (2000). These two works are ium lanuginosum. currently the foundations of North American ant The 49 species that have been added to the list nomenclature. Numbers in parentheses following were overlooked before for various reasons. They a name in the list usually refer to variances from are not, fortunately, primarily exotics that have the nomenclature or from the lists of species in invaded Florida since 1989, although there are Bolton 1995 and 2000. These variances are ex-

44 Florida Entomologist 86(1) March 2003

plained in the section following the list. In cases lands Cos. Records in 1989 list under sayi Emery in which a name appearing in the 1989 list has apparently refer to this species. been changed and the changes are referenced in Camponotus floridanus (Buckley). Wide- one of the two works by Bolton mentioned above, spread. the 1989 usage is noted but the original source of Camponotus impressus (Roger). Widespread. the change in nomenclature may be omitted. Listed under Colobopsis in 1989 list. An asterisk (*) denotes a species that was ab- Camponotus nearcticus Emery. Widespread. sent from the 1989 list. Camponotus pennsylvanicus (DeGeer). North- Florida specimens of all but seven species are ernmost FL, including Panhandle. deposited in the collection of Archbold Biological Camponotus planatus Roger. South FL, north Station. into Hillsborough and Orange Cos. Introduced. Camponotus pylartes Wheeler.* North-central LIST OF FLORIDA ANTS Peninsula. Camponotus sexguttatus (Fabricius).* Dade Acanthomyops claviger (Roger).* Okaloosa Co. Co. Rare. Introduced. Rare. Camponotus snellingi Bolton. Widespread. Acanthomyops interjectus (Mayr).* Liberty Co. Records under C. pavidus Wheeler in 1989 list re- Rare. fer to this species. Amblyopone pallipes (Haldeman). Widespread. Camponotus socius Roger. Widespread, south Anochetus mayri Emery. Dade and Palm Beach into Broward Co. Cos. Introduced. Camponotus tortuganus Emery. South FL, ashmeadi (Emery). Widespread north into Volusia Co. in north FL, south into Highlands Co. Cardiocondyla emeryi Forel. Widespread. In- Aphaenogaster carolinensis Wheeler. North troduced. FL; distribution unclear: confounded with miami- Cardiocondyla nuda (Mayr). Widespread. In- ana. (1) troduced. Aphaenogaster flemingi M. R. Smith. Wide- Cardiocondyla venustula Wheeler. Wide- spread. spread. Introduced. Aphaenogaster floridana M. R. Smith. Wide- Cardiocondyla wroughtonii (Forel). Wide- spread in north FL, south into Highlands Co. spread in peninsular FL. Introduced. Aphaenogaster fulva Roger. Widespread in Cardiocondyla sp.* Widespread in peninsular north FL, south into Orange and Volusia Cos. FL. Introduced. (3) Aphaenogaster lamellidens Mayr. Widespread Cephalotes varians (F.Smith). Dade and Mon- in north FL, south into Highlands and St. Lucie roe Cos. (4) Cos. Crematogaster agnita Wheeler.* Monroe Co. Aphaenogaster mariae Forel. Liberty Co. Rare. Rare. Introduced. Aphaenogaster miamiana Wheeler.* South FL; Crematogaster ashmeadi Mayr. Widespread. distribution unclear: confounded with carolinen- Crematogaster atkinsoni Wheeler. Widespread. sis. Crematogaster cerasi (Fitch). Widespread in Aphaenogaster tennesseensis (Mayr). North- north FL, south into Highlands Co. ernmost counties of peninsular Florida. Crematogaster lineolata (Say). Widespread in Aphaenogaster treatae Forel. Widespread. north FL, south into Hernando and Sumter Cos. Aphaenogaster umphreyi Deyrup & Davis.* Crematogaster minutissima Mayr. Wide- Scattered sites in north FL, south into Highlands spread. Co. (2). Crematogaster missuriensis Emery.* Panhan- Brachymyrmex brevicornis Emery?* (Genus in dle. (5) disarray). Columbia Co. Rare. Introduced. Crematogaster pilosa Emery. Widespread. Brachymyrmex depilis Emery? (Genus in dis- Crematogaster vermiculata Emery. Scattered array). Widespread. sites in north FL, south into Hillsborough Co. Brachymyrmex minutus Forel?* (Genus in dis- Crematogaster sp. A (pine species).* Wide- array). Dade and Monroe Cos. Introduced. spread in north and central FL, south into Lee Brachymyrmex musculus Forel?* (Genus in and Palm Beach Cos. disarray). Widespread. Introduced. Crematogaster sp. B (large species in man- Brachymyrmex obscurior Forel? (Genus in dis- groves).* Monroe Co. Rare. array). Widespread. Cryptopone gilva (Roger). Widespread in north Camponotus caryae (Fitch). Liberty Co. Rare. FL, south into Highlands Co. Camponotus castaneus (Latreille). Wide- Cyphomyrmex minutus Mayr. Widespread in spread. south FL, north into Hillsborough and Volusia Camponotus decipiens Emery. Widespread. Cos. Camponotus discolor (Buckley).* Scattered Cyphomyrmex rimosus (Spinola). Widespread. sites in peninsular Florida, Alachua into High- Introduced.

Deyrup: List of Florida Ants 45

Discothyrea testacea Roger. Widespread. Leptothorax schaumii Roger. North FL, south Dolichoderus mariae Forel.* Leon Co. Rare. into Highlands Co. Dolichoderus pustulatus Mayr.* Scattered Leptothorax smithi Baroni Urbani. Scattered sites throughout FL. Records in 1989 list under D. sites in north FL, south into Highlands Co. plagiatus Mayr refer to this species. Records under L. wheeleri in 1989 list refer to this Dorymyrmex bossutus (Trager).* Widespread species. in peninsular FL, west into Leon Co. (6) Leptothorax texanus Wheeler. Widespread. (7) Dorymyrmex bureni (Trager).* Widespread. Leptothorax torrei (Aguayo). Dade, Monroe, Records in 1989 list under Conomyrma insana Martin Cos. Listed under Macromischa in 1989 (Buckley) refer to this species. (6) list. Dorymyrmex elegans (Trager).* Central penin- Leptothorax sp.* Undescribed. Liberty and sular ridges. (6) Leon Cos. (7) Dorymyrmex flavopectus M. R. Smith. Central Linepithema humile (Mayr). Scattered sites peninsular FL, south Highlands Co. into Marion throughout FL. Introduced. Listed under Iri- Co. Listed under Conomyrma in1989 list. (6) domyrmex in 1989 list. Dorymyrmex grandulus (Forel).* North FL, Monomorium destructor (Jerdon). Scattered south into Citrus Co. (6) sites in south FL, chiefly in Key West. Introduced. Dorymyrmex medeis (Trager).* Scattered sites Monomorium ebeninum Forel. Monroe Co. In- in north FL, south into Highlands Co. (6) troduced. Dorymyrmex reginicula (Trager).* Central Monomorium floricola (Jerdon). South FL, peninsular FL, southern Highlands Co. north into north into Pinellas Co. Introduced. Volusia Co. (6) Monomorium pharaonis (Linnaeus). Wide- Eurhopalothrix floridana Brown & Kempf. spread. Introduced. Widespread in peninsular FL. Monomorium trageri DuBois.* Scattered sites Forelius pruinosus (Roger). Widespread. throughout FL. Records listed under M. mini- Forelius sp.* Scattered sites in north FL, south mum (Buckley) in 1989 list refer to this species. into Citrus Co. Monomorium viride Brown. Widespread. Formica archboldi M. R. Smith. Widespread in Myrmecina americana Emery. Widespread. peninsular FL, west into Liberty Co. Myrmecina sp.* Undescribed. Bradford Co. Formica pallidefulva Latreille. Widespread in Rare. north FL, south into Highlands Co. Myrmelachista ramulorum Wheeler. Polk Co. Formica schaufussi dolosa Wheeler. North FL, Rare or extirpated. Introduced. south into Lake Co. Myrmica punctiventris Roger.* Walton, Sta. Formica subsericea Say. Liberty Co. Rare, not Rosa, Escambia Cos. Rare. seen in recent decades. Neivamyrmex carolinensis (Emery). Scattered Gnamptogenys triangularis (Mayr). Isolated sites in north FL, south into Highlands Co. records from Dade, Broward and Escambia Cos. Neivamyrmex opacithorax (Emery). Scattered Rare. Introduced. Record in 1989 list under G. ac- sites throughout FL. uleaticoxae (Santschi) refers to this species. Neivamyrmex texanus Watkins. Scattered sites Hypoponera inexorata (Wheeler). Widespread. in north FL, south into Highlands Co. Hypoponera opaciceps (Mayr). Widespread. Ochetellus glaber (Mayr). Orange and Volusia Hypoponera opacior (Forel). Widespread. Cos. Introduced. Listed under Iridomyrmex in Hypoponera punctatissima (Roger). South FL, 1989 list. north into Alachua and Bradford Cos. Introduced. Odontomachus brunneus (Patton). Widespread Lasius alienus (Foerster). North FL, south into in Peninsula, in Panhandle west into Leon Co. Marion Co. Odontomachus ruginodis M. R. Smith. South Lasius neoniger Emery. Scattered sites in FL, north into Orange and Co. Introduced. north FL. Odontomachus sp. Undescribed. Highlands Lasius umbratus (Nylander). A few sites in into Orange, Citrus Cos. Records listed under O. Panhandle. clarus in 1989 list refer to this species. Leptogenys manni Wheeler. Highlands Co., Pachycondyla stigma (Fabricius). South FL, north and west into Leon Co. Records in 1989 list north into Orange and Volusia Cos. Introduced. under L. elongata (Buckley) refer to this species. Paratrechina arenivaga (Wheeler). Wide- Leptothorax allardycei (Mann). Monroe and spread. Lee Cos. Listed under Macromischa in 1989 list. Paratrechina bourbonica (Forel). South FL, Leptothorax bradleyi Wheeler. North FL, south north into Orange and Hillsborough Cos. Intro- to Highlands Co. duced. Leptothorax curvispinosus Mayr. Northern- Paratrechina concinna Trager. Widespread. most FL. Paratrechina faisonensis (Forel). Widespread Leptothorax pergandei Emery. Widespread. in north FL, south into Highlands Co. 46 Florida Entomologist 86(1) March 2003

Paratrechina guatemalensis (Forel). South FL, Proceratium crassicorne Emery.* Sta. Rosa north to Sarasota and Indian River Cos. Intro- and Liberty Cos. Rare. (9) duced. Proceratium croceum (Roger). Scattered sites Paratrechina longicornis (Latreille). South FL, in north FL, south into Levy Co. north to St. Johns and Columbia Cos. Introduced. Proceratium pergandei (Emery). North FL, Paratrechina parvula (Mayr). North FL, south south into Pinellas and Highlands Cos. to Volusia Co. Proceratium silaceum Roger. North FL, south Paratrechina phantasma Trager. Peninsular into Highlands Co. FL, Alachua Co. south into Palm Beach Co. Proceratium sp. A.* Undescribed. Leon and Paratrechina pubens (Forel). Dade, Sarasota Lafayette Cos. Rare. and Palm Beach Cos. Introduced. Proceratium sp. B.* Undescribed. Liberty Co. Paratrechina vividula (Nylander). North FL, Rare. south into Highlands Co. Pseudomyrmex cubaensis (Forel). South FL, Paratrechina wojciki Trager. Widespread. north into Polk Co. Paratrechina sp. A.* Undescribed. Workerless Pseudomyrmex ejectus (F. Smith). Widespread. parasite of wojciki. Highlands Co. Pseudomyrmex elongatus (Mayr). South FL, Paratrechina sp. B.* Undescribed. Workerless north to Highlands Co. parasite of faisonensis. Hamilton Co. Pseudomyrmex gracilis (Fabricius). South FL, Pheidole adrianoi Naves. Widespread. north into Alachua Co. Introduced. Records listed Pheidole bicarinata vinelandica Forel.* North under P. mexicanus in 1989 list refer to this spe- FL, south into Citrus Co. cies. Pheidole carrolli Naves. Scattered sites in Pseudomyrmex leptosus Ward. A few sites in north FL, south into Citrus Co. Rare. Peninsula, north into Alachua Co. Pheidole crassicornis Emery. Scattered sites in Pseudomyrmex pallidus (F. Smith). Wide- north FL, south into Alachua Co. spread. Pheidole dentata Mayr. Widespread. Pseudomyrmex seminole Ward. Widespread in Pheidole dentigula M. R. Smith. Widespread. south FL, Orange and Bay Cos. in north FL. Pheidole diversipilosa Wheeler.* A few sites in Pseudomyrmex simplex (F. Smith). South FL, Panhandle, east into Columbia Co. Rare. north into Orange Co. Pheidole flavens Roger.* South FL, north into Pyramica abdita (Wesson & Wesson).* Alachua Martin Co. Introduced. and Leon Cos. Rare. Pheidole floridana Emery. Widespread. Pyramica angulata (M. R. Smith).* A few sites Pheidole lamia Wheeler. Leon and Jackson in north FL. Rare. Cos. Rare. Pyramica apalachicolensis Deyrup & Luber- Pheidole littoralis Cole. Peninsular FL, west tazzi.* Leon Co. Rare. (10) into Franklin Co., south into Highlands Co. Pyramica archboldi (Deyrup & Cover).* North Pheidole megacephala (Fabricius). Scattered FL, south into Volusia Co., west into Jefferson Co. sites in south FL, north into Hillsborough Co. In- Pyramica bunki (Brown). Scattered sites in troduced. north FL, south into Highlands Co. Under Smith- Pheidole metallescens Emery. Widespread in istruma in 1989 list. north FL, south into Highlands Co. Pyramica carolinensis (Brown). A few sites in Pheidole moerens Wheeler. Widespread. Intro- north FL. Rare. Under Smithistruma in 1989 list. duced. Pyramica cloydi (Pfitzer).* Lake Co. Rare. Pheidole morrisi Forel. Widespread. Pyramica clypeata (Roger). North FL, south Pheidole obscurithorax Naves*. Panhandle, into Highlands Co. Under Smithistruma in 1989 east into Leon Co. Introduced. (8) list. Pheidole tysoni Forel. Alachua and Madison Pyramica creightoni (M. R. Smith). North pen- Cos. insular FL, south into Highlands Co. Under Platythyrea punctata (F. Smith). South FL, Smithistruma in 1989 list. north into Highlands Co. Pyramica deyrupi Bolton.* Marion Co. Pogonomyrmex badius (Latreille). Widespread. Pyramica dietrichi (M. R. Smith). Widespread. Polyergus lucidus Mayr. Alachua, Columbia, Under Smithistruma in 1989 list. Leon, Sta. Rosa Cos. Rare. Pyramica eggersi (Emery). South FL, north Ponera exotica M. R. Smith. Scattered sites in into Alachua Co. Introduced. Under Strumigenys north FL, south into Highlands Co. in 1989 list. Ponera pennsylvanica Buckley. North FL, Pyramica gundlachi Roger. Dade, Monroe, Col- south into Marion Co. lier Cos. Introduced. Under Strumigenys in 1989 Prenolepis imparis (Say). North FL, south into list. Orange Co. Pyramica hexamera (Brown). Marion and Her- Prionopelta antillana Forel. Marion Co. Intro- nando Cos. Rare. Introduced. Under Epitritus in duced. 1989 list. Deyrup: List of Florida Ants 47

Pyramica inopina (Deyrup & Cover).* Ala- Strumigenys rogeri Emery. South FL, north chua, Marion, Putnam Cos. Rare. into Orange Co. Introduced. Pyramica laevinasis (M. R. Smith). Walton Co. Strumigenys silvestrii Emery. A few sites, from Rare. Under Smithistruma in 1989 list. Monroe into Gadsden Cos. Rare. Introduced. Pyramica margaritae (Forel). Scattered sites litorale Wheeler. South FL, north in north FL, south into Marion Co. Introduced. into Pinellas Co. Under Smithistruma in 1989 list. Tapinoma melanocephalum (Fabricius). South Pyramica membranifera (Emery). Widespread. FL, north into Brevard Co. Introduced. Introduced. Under Trichoscapa in 1989 list. Tapinoma sessile (Say). Widespread. Pyramica missouriensis (M. R. Smith).* Ala- Technomyrmex albipes (F. Smith).* Wide- chua and Highlands Cos. Rare. spread. Introduced. Pyramica ohioensis (Kennedy & Schramm). Tetramorium bicarinatum (Nylander). Wide- Scattered sites in north FL, south into Alachua spread. Introduced. Co. Under Smithistruma in 1989 list. Tetramorium caldarium (Roger). South FL, Pyramica ornata (Mayr). North FL, south into north into Lake Co. Introduced. Highlands and Sarasota Cos. Under Smithis- Tetramorium lanuginosum Mayr. Hernando, truma in 1989 list. Holmes, Jackson Cos. Rare, extirpated? Intro- Pyramica pilinasis (Forel).* Scattered sites in duced. Under Triglyphothrix in 1989 list. north FL, south into Highlands Co. Tetramorium simillimum (F. Smith). South Pyramica pulchella (Emery). North FL, west FL, north into St. Johns Co. Introduced. into Bay Co., south into Highlands Co. Under Trachymyrmex jamaicensis (André). Martin Smithistruma in 1989 list. and Monroe Cos. Rare. Pyramica reflexa (Wesson & Wesson). North Trachymyrmex septentrionalis (McCook). FL, south into Polk Co. Under Smithistruma in Widespread. 1989 list. Wasmannia auropunctata (Roger). South FL, Pyramica rostrata (Emery).* Northern tier of north into Orange Co. Introduced. FL counties. Under Smithistruma in 1989 list. Xenomyrmex floridanus Emery. South FL, Pyramica talpa (Weber). North FL, south into north into Orange Co. Highlands Co. Under Smithistruma in 1989 list. Pyramica wrayi (Brown).* Leon and St. Johns (1). Aphaenogaster carolinensis is listed as a Cos. Rare. subspecies of A. texana Wheeler in Bolton 1995. It Solenopsis abdita Thompson.* Widespread. was raised to species level by Gary Umphrey Solenopsis carolinesis Forel. Widespread. (1996). Aphaenogaster carolinensis is difficult to Solenopsis corticalis Forel. Monroe Co. separate from A. miamiana. Solenopsis geminata (Fabricius). Widespread. (2). Aphaenogaster umphreyi is a recently de- Solenopsis globularia littoralis Creighton. scribed species (Deyrup and Davis 1998). South FL, north into Alachua and Franklin Cos. (3). There are at least five species of Cardio- Solenopsis invicta Buren. Widespread. (11) condyla in Florida, differentiated by structural Solenopsis nickersoni Thompson. Collier Co. character states, coloration and habitat prefer- north into Leon Co. ences. There is also some variation in color and Solenopsis pergandei Forel. Widespread. sculpture within apparent species. This could de- Solenopsis picta Emery. Widespread. note additional species, but I have kept in mind Solenopsis tennesseensis M. R. Smith. Wide- that members of this genus are fully capable of spread. founding inbred populations from the introduc- Solenopsis tonsa Thompson.* North FL, Leon tion of a single female, so variation could result Co. east to Alachua Co., south to Orange Co. from the introduction of a species from more than Solenopsis truncorum Forel.* Alachua Co. one source. If this occurred, there might be vari- Rare. ant forms, even sympatric variant forms, that re- Solenopsis xyloni McCook. Western Panhan- flected geographic variation within the natural dle. Not seen in recent decades, possibly extir- range of the species, or genetic drift. The persis- pated. tence of these forms would reflect the degree of in- Solenopsis sp.* Undescribed parasite of breeding in the population. I have resorted to Pheidole dentata. Gilchrist Co. Rare. using Creighton (1950), which leaves one species Stenamma foveolocephalum M. R. Smith.* without a name. There are several names that Walton Co. Rare. might be applied to this species, or to other Flor- Strumigenys emmae (Emery). South FL, north ida species that I have identified using the into Hillsborough and Volusia Cos. Under Creighton key. These names include obscurior Quadristruma in 1989 list. Wheeler, ectopia Snelling, mauritanica Forel, and Strumigenys lanuginosa Wheeler. Dade, Lee minutior Forel. Inconvenient though it might and Monroe Cos. Rare. Introduced. seem, the most sensible approach to understand- Strumigenys louisianae Roger. Widespread. 48 Florida Entomologist 86(1) March 2003 ing the of exotic Cardiocondyla would series of identified Florida specimens of both spe- be to begin by looking at specimens from the na- cies that seem to justify this treatment. tive ranges of the species. (10). Pyramica apalachicolensis is a recently (4). Cephalotes varians was, until recently, described species (Deyrup and Lubertazzi 2001). placed in the genus Zacryptocerus, now synony- (11). Solenopsis invicta is listed as S. wagneri mized with Cephalotes (Andrade and Baroni Ur- Santschi in the 1995 catalog; the former name has bani 1999). been conserved. (5). Crematogaster missuriensis (the original description lacks the “o” in Missouri), was consid- ACKNOWLEDGMENTS ered a subspecies of C. minutissima by Creighton (1950). Although the two species are similar, they Many of the species added to the 1989 list were found for the first time in Florida by various entomolo- differ in nest site, in the sculpture of the me- gists, who recognized them as new records and kindly sopleuron (Creighton 1950), and are sympatric in sent or brought specimens to me. These collectors I a large part of their ranges. A paper on the subge- gratefully list below; the parenthesis after each name nus Orthocrema in eastern North America is in denotes the number of species that each person discov- preparation. ered: Lloyd Davis (7), Stefan Cover (3), Clifford Johnson (6). Species of Dorymyrmex were listed under (3), David Lubertazzi (3), Zachary Prusak (1), Vincent the genus Conomyrma in the 1989 list. In the Golia (1), Paul Skelley (1). I thank Lloyd Davis and Hi- 1989 list, several species of Dorymyrmex reported lary Swain for detailed and thoughtful comments on the from Florida by James Trager (1988) were ex- first draft of this paper. cluded from the list. This was a mistake, and an REFERENCE CITED injustice to Trager’s excellent work. Of the seven species known to occur in Florida, four made their ANDRADE, M. L. DE, AND C. BARONI URBANI. 1999. Di- way into Bolton’s catalog, three did not. Extensive versity and adaptation in the ant genus Cephaplotes, field work in Florida since 1989 has confirmed past and present. Stuttgarter Beit. Naturkunde, Ser. that there are at least seven species in Florida. B 271: 1-889. Whether any of these are identical with species BOLTON, B.1995. A new general catalogue of the ants. known from the southwestern U.S. is an open Harvard University Press, Cambridge. 504 pp. BOLTON, B. 2000. The ant tribe Dacetini. Mem. Ameri- question until the southwestern species have can Entomol. Inst. 65: 1-1028. been studied as intensively as those in the South- CREIGHTON, W. S. 1950. The ants of North America. east. In 2001, I did a little collecting of Dory- Bull. Mus. Comparative Zool. Harvard 104: 1-585, + myrmex during a short stay in Arizona. The 57 plates. results of even so brief an exposure have con- DEYRUP, M., AND L. DAVIS. 1998. A new species of Aph- vinced me that the southwestern Dorymyrmex are aenogaster from upland habitats in Florida. Ento- as complex and challenging as they are fascinat- mol. News 109: 88-94. ing. DEYRUP, M., L. DAVIS, AND S. COVER. 2000. Exotic ants (7). The subspecies Leptothorax texanus davisi in Florida. Trans. American Entomol. Soc. 126: 293- 326. Wheeler, listed in the 1989 list, was raised to spe- DEYRUP, M., C. JOHNSON, G. C. WHEELER, AND J. cies rank by Mackay (2000). In Florida and else- WHEELER. 1989. A preliminary list of the ants of where davisi does not seem to be recognizable Florida. Florida Entomol. 72: 91-101. either as a species or subspecies; a paper dealing DEYRUP, M., AND D. LUBERTAZZI. 2001. A new species of with this and the undescribed Florida Leptotho- ant (Hymenoptera: Formicidae) from North Florida. rax listed above is in preparation. Entomol. News 112: 15-21. (8). Pheidole obscurithorax is listed in Bolton TRAGER, J. C. 1988. A revision of Conomyrma (Hy- (1995) as a subspecies of P. fallax Mayr. Deyrup et menoptera: Formicidae) from the southeastern al. (2000) treated obscurithorax as a species. United States, especially Florida, with keys to the species. Florida Entomol. 71: 11-29. (9). Proceratium crassicorne was synonymized UMPHREY, G. J. 1996. Morphometric discrimination with P. silaceum by Creighton (1950), but Maria among sibling species in the fulva-rudis-texana com- de Andrade will be reviving this species in a forth- plex in the ant genus Aphaenogaster (Hymenoptera: coming revision of the genus; she has sent me a Formicidae). Canadian J. Zool. 74: 528- 559.