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Kenai National Wildlife Refuge Species List, Version 2017-01-23

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Kenai National Wildlife Refuge Species List, Version 2017-01-23 Kenai National Wildlife Refuge Species List, version 2017-01-23 Kenai National Wildlife Refuge biology staff January 23, 2017 2 Cover images represent changes to the checklist. Top left: Rhagoletis ba- siola by Nordic Lake, August 9, 2017. (http://arctos.database.museum/ media/10562924). Image CC0 Matt Bowser. Top right: Rhytisma ar- buti on leaf of Menziesia ferruginea, September 5, 2014 (http://www. inaturalist.org/observations/863266). Image CC BY Matt Bowser. Bot- tom left: Leaf gall caused by Pontania sp. bold:acl3478 on Salix bar- clayi, Soldotna, Ski Hill Road, June 27, 2016 (http://arctos.database. museum/media/10548664). Image CC0 Matt Bowser. Bottom right: In- ostemma sp. collected south of Tustumena Lake, June 24, 2006 (http: //arctos.database.museum/media/10508672). Image CC0 Matt Bowser. Contents Contents 3 Introduction 5 Purpose............................................................ 5 About the list......................................................... 5 Acknowledgments....................................................... 5 Native species 7 Vertebrates .......................................................... 7 Phylum Chordata.................................................... 7 Invertebrates ......................................................... 24 Phylum Annelida.................................................... 24 Phylum Arthropoda .................................................. 24 Phylum Cnidaria.................................................... 45 Phylum Mollusca.................................................... 45 Phylum Platyhelminthes................................................ 46 Phylum Tardigrada................................................... 46 Vascular Plants........................................................ 46 Division Tracheophyta................................................. 46 Bryophytes .......................................................... 58 Division Bryophyta................................................... 58 Fungi ............................................................. 62 Division Ascomycota.................................................. 62 Division Basidiomycota ................................................ 69 Division Chytridiomycota ............................................... 70 Division Myxomycota ................................................. 70 Unicellular Organisms.................................................... 70 Phylum Cercozoa.................................................... 70 Phylum Heterokonta.................................................. 70 Phylum Protozoa.................................................... 70 Non-native species 73 Vertebrates .......................................................... 73 Phylum Chordata.................................................... 73 Invertebrates ......................................................... 73 Phylum Annelida.................................................... 73 Phylum Arthropoda .................................................. 73 Phylum Mollusca.................................................... 74 Vascular Plants........................................................ 74 Division Tracheophyta................................................. 74 Extirpated species 77 Vertebrates .......................................................... 77 Phylum Chordata.................................................... 77 Vascular Plants........................................................ 77 Division Tracheophyta................................................. 77 Images 79 Change log 83 References 85 Index 87 3 Introduction Purpose names because our current system does not enable us to handle accepted and provisional names separately. We want A primary purpose for which the Kenai National Wildlife to avoid implying that any of the provisional names we use Refuge was established in the Alaska National Interest are formally accepted by formatting them as proper scien- Lands Conservation Act of 1980 was, \to conserve fish and tific names. Also, note that our database enforces lower case wildlife populations and habitats in their natural diver- letters in our specific epithets so that, for example, \Rab- sity. ," where the term “fish and wildlife" was defined dophaga sp. BOLD:ADA6450" becomes \Rabdophaga sp. as \any member of the animal kingdom, including without bold:ada6450." limitation any mammal, fish, bird. , amphibian, reptile, Synonymies we included do not necessarily reflect our mollusk, crustacean, arthropod or other invertebrate." An opinions and are not intended as statements on correct tax- obvious first step toward fulfilling this purpose is to know onomy (we generally follow ITIS); rather, synonymns are what fish and wildlife, habitats, and natural diversity are to included in the index as an aid to the reader to help find be conserved. This checklist is intended to be a frequently- taxa for which names may have changed. updated document reflecting our current knowledge of which In the checklist non-native species are indicated with a living things call the Kenai National Wildlife Refuge home. dagger symbol (y) and species represented by both native and non-native populations are indicated with a double dag- ger symbol (z). About the list This list is dynamic and far from perfect with many changes and corrections yet to be made. Please The present list includes a total of 2,075 species. Of these, send comments and corrections to Matt Bowser at 1,982 are considered to be native, 89 species are consid- matt [email protected]. ered to be non-native, and four species are represented by both native and non-native populations. Eighteen non- native species are believed to have been extirpated from the Acknowledgments Refuge. Since the last version of this list (Kenai National Wildlife Thanks and credit are due to many people who have re- Refuge biology staff, 2016) where 2,053 species were listed, viewed this list, provided identifications, and helped in other there has been a net increase of 22 species added. In this ways. The list below is by no means exhaustive. draft records from eBird Cornell Lab of Ornithology(2016) James Bergdahl (Conservation Biology Center, Spokane, that fell on the Kenai National Wildlife Refuge were added. Washington) reviewed and made helpful comments on An image section was also added. See the change log (page the Refuge's list of Carabidae. 83) for details. The list was produced from an in-house database using Peter Hovingh provided a list of leeches (Hirudinea) col- ITIS (http://www.itis.gov) for its taxonomic backbone. Ac- lected on the Refuge. cordingly, names and hierarchies generally follow ITIS ex- cept where taxa are not represented in ITIS. John Hudson (USFWS, Juneau, Alaska) provided lists of Although accepted scientific names are used wherever Odonata observed on the Refuge. these are known, we have included provisional identifications Richard Payne (University of York, Heslington, UK) pro- (for example, \Typhula sp.") where these represent the best vided a list of protists from Jigsaw Lake from Payne information available. Species recognized only as molecu- et al.(2006). lar operational taxonomic units are also included, mainly entities recognized by Barcode Index Numbers (BINs, Rat- David Wartinbee (retired from Kenai Peninsula College, nasingham and Hebert, 2013). We have chosen not to fol- Soldotna, Alaska) shared a list of Chironomidae he had low the usual convention of italicizing genus and species collected on the Refuge. 5 Native species occurring on the Kenai National Wildlife Refuge Vertebrates Common names: Steelhead Literature records: Kyle(1992): iNaturalist observations: Phylum Chordata 811982, 1814201 Class Actinopterygii Oncorhynchus nerka Order Cypriniformes Common names: Sockeye salmon or kokanee Literature records: Kyle(1992) iNaturalist observations: Family Catostomidae 1021584 Catostomus catostomus Oncorhynchus tshawytscha Common names: Longnose sucker Common names: Chinook salmon or king salmon Image(s):1 Prosopium cylindraceum Taxonomic notes: Two forms have been documented from the Refuge: the typical form and a semi-dwarf form known from Wolf Common names: Round whitefish and Finger Lakes (Dean and Rickabough, 2005). Literature records: Kyle(1992) Distribution and trends: Widespread across Alaska, northern Salvelinus alpinus North America, and into Arctic Siberia (Page and Burr, 1991), This Common names: Arctic char species is present in the Kenai River and Swanson River watersheds on the Refuge (Dean and Rickabough, 2005). Salvelinus malma Biology: Catostomus catostomus is a bottom-feeder, consuming Common names: Dolly varden mainly invertebrates (Mansfield, 2004). Literature records: Kyle(1992) iNaturalist observations: : Dean and Rickabough(2005), Friedersdorff Literature records 811997 (1986), Friedersdorff and Jakubas(1984a), Friedersdorff and Jakubas (1984b) Salvelinus namaycush Common names: Lake trout Order Gasterosteiformes Literature records: Kyle(1992) Family Gasterosteidae Order Scorpaeniformes Gasterosteus aculeatus Common names: Threespine stickleback Family Cottidae Image(s):2 Cottus aleuticus Literature records: Friedersdorff and Jakubas(1984a), Kyle (1992), Sprehn et al.
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