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Íîâûé Ýîöåíîâûé Ipelates (Coleoptera, Agyrtidae) Èç Ñàêñîíñêîãî ßíòàðß

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Íîâûé Ýîöåíîâûé Ipelates (Coleoptera, Agyrtidae) Èç Ñàêñîíñêîãî ßíòàðß Vestnik zoologii, 41(4): 365–367, 2007 © Å. Ý. Ïåðêîâñêèé, 2007 ÓÄÊ 565.763(430) ÍÎÂÛÉ ÝÎÖÅÍÎÂÛÉ IPELATES (COLEOPTERA, AGYRTIDAE) ÈÇ ÑÀÊÑÎÍÑÊÎÃÎ ßÍÒÀÐß Å. Ý. Ïåðêîâñêèé Èíñòèòóò çîîëîãèè èì. È. È. Øìàëüãàóçåíà ÍÀÍ Óêðàèíû, óë. Á. Õìåëüíèöêîãî, 15, Êèåâ, 01601 Óêðàèíà Ïðèíÿòî 12 ôåâðàëÿ 2007 Íîâûé ýîöåíîâûé Ipelates (Coleoptera, Agyrtidae) èç ñàêñîíñêîãî ÿíòàðÿ. Ïåðêîâñêèé Å. Ý. — Âòîðîé âèä ïîçäíåýîöåíîâûõ àãèðòèä — Ipelates weitshati Perkovsky, sp. n. îïèñàí èç ñàêñîíñêîãî ÿíòàðÿ. Îí ëåãêî îòëè÷èì îò I. kerneggeri Perkovsky èç áàëòèéñêîãî ÿíòàðÿ ïî ïî÷òè ïðÿìûì çàäíèì óãëàì ïðîíîòóìà è áîëåå äëèííûì óñèêàì ñ óçêîé áóëàâîé (äëèíà 7-ãî ÷ëåíèêà óñèêîâ â 1,9 ðàçà áîëüøå øèðèíû, 11-ãî — â 2,5 ðàçà áîëüøå øèðèíû). Îò ñîâðåìåííûõ I. striatipennis (Lewis) è I. castaneicolor (Champion) ñ ïðÿìûìè çàäíèìè óãëàìè ïðîíîòóìà íîâûé âèä îòëè÷àåòñÿ ìåëêèìè ðàçìåðàìè, õàðàêòåðîì ïóíêòèðîâêè è îòñóòñòâèåì ìèêðîñêóëüïòóðû íà ïðîíîòóìå. Êëþ÷åâûå ñëîâà: Coleoptera, Agyrtidae, Ipelates, òàêñîíîìèÿ, ýîöåí, ÿíòàðü. The New Eocene Ipelates (Coleoptera, Agyrtidae) from the Bitterfeldian Amber. Perkovsky E. E. — Second Late Eocene agyrtid species — Ipelates weitshati Perkovsky, sp. n. is described from the Bitterfeldian amber. It is strongly differs from the I. kerneggeri Perkovsky from Baltic amber by right hind angles of pronotum and longer antennae with narrower segments of the club (7th antennal segment 1.9 times as long as wide, 11th antennal segment 2.5 times as long as wide). It differs from the recent species of Ipelates with right hind angles of pronotum, I. striatipennis (Lewis) and I. castaneicolor (Champion), by the smaller size and sculpture of pronotum. Key words: Coleoptera, Agyrtidae, Ipelates, taxonomy, Eocene, amber. Ñàêñîíñêèì òðàäèöèîííî íàçûâàþò ÿíòàðü, êîòîðûé äîáûâàëñÿ íà îòêðûòîé óãîëüíîé øàõòå â Ãîéòøå (Goitsche) áëèç ã. Áèòòåðôåëüä (çåìëÿ Ñàêñîíèÿ-Àíõàëüò). Ñåé÷àñ ýòà øàõòà çàòîïëåíà. Âîçðàñò ñëþäû â ïåðåêðûâàþùèõ îòëîæåíèÿõ (Ritzkowski, 1999) äîêàçûâàåò, ÷òî ýòîò ÿíòàðü áûë ïåðåìåùåí â Ñàêñîíèþ-Àíõàëüò èç òîãî æå ðåãèîíà, ÷òî è áàëòèéñêèé ÿíòàðü; àíàëèç ñîñòàâà âêëþ÷åíèé (Weitshat, Wichard, 2002) ïîäòâåðæäàåò åãî îäíîâîçðàñòíîñòü ñ áàëòèéñêèì ÿíòàðåì è îáùíîñòü èõ ïðîèñõîæäåíèÿ (Weitshat, 1997). Ïåðâûé ýîöåíîâûé âèä àãèðòèä — Ipelates kerneggeri Perkovsky áûë îïèñàí íàìè èç áàëòèéñêîãî ÿíòàðÿ, äîáûòîãî íà Ñàíáèéñêîì ï-îâå, ï. ßíòàðíûé (Ïåðêîâñêèé, 2005).  êîëëåêöèè ñàêñîíñêîãî ÿíòàðÿ Åñòåñòâåííî-èñòîðè÷åñêîãî ìóçåÿ óíèâåðñèòåòà Ãóìáîëüäòà â Áåðëèíå (Humboldt-Universitat zu Berlin, Museum fur Naturkunde, êóðàòîð — Dr. Christian Neumann) áûë îáíàðóæåí âòîðîé âèä ýîöåíîâûõ àãèðòèä, îïèñûâàåìûé â íàñòîÿùåé ñòàòüå. Ipelates weitshati Perkovsky, sp. n. (ðèñ. 1 íà 3-é ñòð. îáëîæêè) Ìàòåðèàë. Ãîëîòèï { â ñàêñîíñêîì ÿíòàðå; øàõòà Ãîéòøå áëèç ã. Áèòòåðôåëüä; èíâ. íîìåð Col. 8/155 èç êîëëåêöèè ñàêñîíñêîãî ÿíòàðÿ Åñòåñòâåííî-èñòîðè÷åñêîãî ìóçåÿ óíèâåðñèòåòà Ãóìáîëüäòà â Áåðëèíå (coll. Humboldt-Universitat zu Berlin, Museum fur Naturkunde, Col. 8/155), ïîçäíèé ýîöåí. Õðàíèëñÿ â áóìàæíîì êîíâåðòèêå, ïîäïèñàííîì «Sachs. Bernstein [Col. 8/155] Serie 8 (1984) Col.: Nitidulidae (?)». Öåëûé æóê õîðîøåé ñîõðàííîñòè. Äëèíà òåëà ñáîêó (ñ ïîäîãíóòîé ãîëîâîé) 3,3 ìì. Âåñü æóê òåìíî-êîðè÷íåâûé. Ãëàçà áîëåå èëè ìåíåå êðóïíûå, ãîëîâà çà ãëàçàìè áåç âèñêîâ; ðàññòîÿíèå ìåæäó îñíîâàíèÿìè óñèêîâ â 2,5 ðàçà ïðåâûøàåò äëèíó 3-ãî ÷ëåíèêà óñèêîâ; ðàññòîÿíèå ìåæäó ïåðåäíèì êðàåì ãëàçà è îñíîâàíèåì ÿâñòâåííî ìåíüøå øèðè- íû ñêàïóñà. Ãîëîâà â äâîéíîé ïóíêòèðîâêå; êðóïíûå òî÷êè â îñíîâíîì íà ëáó ìåæäó ãëàçàìè, íå ìåëü÷å êðóïíûõ òî÷åê ïðîíîòóìà ðàññòîÿíèå ìåæäó òî÷êàìè 366 Å. Ý. Ïåðêîâñêèé ðàâíî 1–2 äèàìåòðàì òî÷åê. Ïîñëåäíèé ÷ëåíèê ÷åëþñòíûõ ùóïèêîâ â 3 ðàçà äëèííåå ïðåäïîñëåäíåãî, óçêèé, íà òðåòü êîðî÷å 3-ãî ÷ëåíèêà óñèêîâ, îò ñåðåäè- íû ñóæåí ê âåðøèíå. Óñèêè ñî ñëàáîé 5-÷ëåíèêîâîé áóëàâîé, çàõîäÿò çà çàäíèé êðàé ïðîíîòóìà, äëèíà óñèêà â 2,3 ðàçà áîëüøå øèðèíû ãîëîâû; 3-é ÷ëåíèê â 1,3 ðàçà äëèííåå 2-ãî, åãî äëèíà â 4 ðàçà áîëüøå øèðèíû, â 1,4 ðàçà ìåíüøå äëèíû äëèíû 4-ãî è 5-ãî, âìåñòå âçÿòûõ; äëèíà 4-ãî è 5-ãî ÷ëåíèêîâ â 3 ðàçà áîëüøå øèðèíû; äëèíà 6-ãî ÷ëåíèêà â 2,4 ðàçà áîëüøå øèðèíû, 7-é ÷ëåíèê ïðîäîëüíûé, â äèñòàëüíîé òðåòè ñëàáî ðàñøèðåí, åãî äëèíà â 1,9 ðàçà áîëüøå øèðèíû, 8-é ÷ëåíèê â 1,2 ðàçà øèðå 7-ãî, ðàâíîìåðíî ðàñøèðåí îò îñíîâàíèÿ; åãî äëèíà â 1,4 ðàçà áîëüøå øèðèíû, 9-é è 10-é ÷ëåíèêè áîëåå èëè ìåíåå òðàïå- öèåâèäíûå, èõ äëèíà â 1,3 ðàçà áîëüøå øèðèíû, äëèíà 11-ãî ÷ëåíèêà â 2,5 ðàçà áîëüøå øèðèíû. Øèðèíà ãîëîâû 0,7 ìì, äëèíà óñèêà 1,65 ìì. Ïðîíîòóì íàèáîëüøåé øèðèíû äîñòèãàåò ïåðåä îñíîâàíèåì, åãî äëèíà âäâîå ìåíüøå øèðèíû, îñíîâàíèå ïðîíîòóìà áåç âäàâëåíèé, çàäíèå óãëû ïî÷òè ïðÿìûå, êàê ó I. striatipennis (Lewis). Ïóíêòèðîâêà êðàåâ ïðîíîòóìà êðóïíàÿ, ðàñ- ñòîÿíèå ìåæäó òî÷êàìè ðàâíî 0,5–1 äèàìåòðàì òî÷åê; ïóíêòèðîâêà äèñêà ïðîíî- òóìà äâîéíàÿ, â öåíòðå äèñêà áîëåå ðåäêàÿ è íåðàâíîìåðíàÿ, ÷åì ïî êðàÿì, ðàññòîÿíèå ìåæäó òî÷êàìè ðàâíî 1–2 äèàìåòðàì òî÷åê: êðóïíûå òî÷êè êðóïíåå, ÷åì íà ãîëîâå. Ïðîñòåðíóì óçêèé, ðàññòîÿíèå îò ïåðåäíèõ òàçèêîâûõ âïàäèí äî ïåðåäíåãî êðàÿ ïðîñòåðíóìà â 5 ðàç ìåíüøå äëèíû ïåðåäíèõ òàçèêîâ Òðåóãîëü- íûé âûðîñò ïðîïëåâð çàõîäèò äàëåêî çà ñðåäèíó ïåðåäíèõ òàçèêîâ. Øèðèíà ïðîíîòóìà 1,6 ìì, äëèíà ïðîíîòóìà 0,8 ìì. Íàäêðûëüÿ ïàðàëëåëüíîñòîðîííèå, òî÷å÷íûå áîðîçäêè ñèëüíî âäàâëåííûå: ïåðâàÿ–÷åòâåðòàÿ áîðîçäêè äîõîäÿò äî âåðøèíû íàäêðûëüÿ, òðåòüÿ è ÷åòâåðòàÿ ó âåðøèíû îáðàçóþò äâîéíîé ðÿä òî÷åê. Òî÷êè âî âñåõ òî÷å÷íûõ áîðîçäêàõ ðàâ- íîé âåëè÷èíû. Ìåæäóðÿäüÿ íåðàâíîìåðíî ðåäêî è íåæíî ïóêíêòèðîâàíû. Ïðî- ìåæóòîê ìåæäó 9-é áîðîçäêîé è ýïèïëåâðàëüíûè êèëåì ñ äåñÿòêîì êðóïíûõ òî÷åê; øèðèíà ýòîãî ïðîìåæóòêà íå ïðåâûøàåò äâîéíîãî äèàìåòðà òî÷êè. Ýïè- ïëåâðàëüíûé êèëü â ïåðåäíåé ÷åòâåðòè íàäêðûëèé óçêèé, áåç çàìåòíûõ (ïðåâû- øàþùèõ äèàìåòð òî÷êè) çóá÷èêîâ. Ýïèïëåâðû íàäêðûëèé ñëàáîâîãíóòûå. Âåð- øèíû íàäêðûëèé íå îáðóáëåíû, âåðøèííûé óãîë ïî÷òè ïðÿìîé. Äëèíà íàäêðûëèé 2,3 ìì. Êðûëüÿ ðàçâèòû. 4-é ñòåðíèò áðþøêà áåç âäàâëåíèé. Ñòåðíèòû áðþøêà øàãðåíèðîâàíû, â äâîéíîé ïóíêòèðîâêå. Ïåðåäíèå ãîëåíè óçêèå, èõ äëèíà â 10 ðàç áîëüøå øèðèíû, ïî âåðõíåìó êðàþ ñ âíóòðåííåé ñòîðîíû ñ óçêèì âàëèêîîáðàçíûì ïîäíÿòèåì, äîõîäÿùèì äî îñíîâàíèÿ ãîëåíè. Çàäíèå ãîëåíè òîíêèå — èõ äëèíà â 12 ðàç áîëüøå øèðèíû. Ïåðåäíèå ëàïêè ñëàáî ðàñøèðåíû; ÷ëåíèêè ñðåäíèõ ëàïîê íå ðàñøèðåíû. Êîãîòêè ó îñíîâàíèÿ ñ òóïûì çóáöîì. Èç-ïîä ñäâèíóòîãî êâåðõó ïîñëåäíåãî ñòåðíèòà âèäíà óçêîîêðóãëåííàÿ âåð- øèíà ýäåàãóñà; ïðîêñèìàëüíåå ýäåàãóñ êàæåòñÿ óçêèì è ïàðàëëåëüíîñòîðîííèì. Cðàâíåíèå. Âåðøèíû ìàíäèáóë áåç çóáöà, êàê ó âñåõ àãèòèä, êðîìå Pte- rolominae. Íîâûé âèä ñîîòâåòñòâóåò äèàãíîçó ðîäà Ipelates; ïî ôîðìå òåëà è ôîðìå çàäíèõ óãëîâ ïðîíîòóìà íîâûé âèä ñõîäåí ëèøü ñ ÿïîíñêèì I. striatipen- nis è ãèìàëàéñêèì I. castaneicolor (Champion) (Shawaller, 1983); îáà âèäà çàìåòíî êðóïíåå: äëèíà òåëà I. striatipennis — 5–5,5 ìì, I. castaneicolor — 6–7 ìì; ýïèïëåâðàëüíûé êèëü I. striatipennis è I. castaneicolor ñíàáæåí 8–11 ÿâñòâåííûìè çóá÷èêàìè, à ïðîíîòóì øàãðåíèðîâàí. Ýòè âèäû îáíàðóæèâàþò è äðóãèå îòëè- ÷èÿ: ó I. striatipennis ÿâñòâåííî øàãðåíèðîâàíû íàäêðûëüÿ, 3-ÿ òî÷å÷íàÿ áîðîçäêà óäàëåíà îò 4-é; ó I. castaneicolor äèñê ïðîíîòóìà — ãëàäêèé, à êðóïíàÿ ïóíêòè- ðîâêà ãîëîâû âäâîå ìåëü÷å ïóíêòèðîâêè áîêîâûõ êðàåâ ïðîíîòóìà. Íîâûé ýîöåíîâûé Ipelates (Coleoptera, Agyrtidae)… 367 Îò I. kerneggeri Perkovsky èç áàëòèéñêîãî ÿíòàðÿ íîâûé âèä ëåãêî îòëè÷àåòñÿ ïî ôîðìå çàäíèõ óãëîâ ïðîíîòóìà, íàèáîëüøåé øèðèíû äîñòèãàþùåãî ïåðåä îñíîâàíèåì (ó I. kerneggeri çàäíèå óãëû ïðîíîòóìà øèðîêîîêðóãëåíû, ïðîíîòóì íàèáîëüøåé øèðèíû äîñòèãàåò ó îñíîâàíèÿ). Îò I. kerneggeri (Ïåðêîâñêèé, 2005) íîâûé âèä îòëè÷àåòñÿ òàêæå áîëåå äëèííûìè óñèêàìè (3-é ÷ëåíèê óñèêà I. ker- neggeri ëèøü â 1,25 ðàçà êîðî÷å 4-ãî è 5-ãî, âìåñòå âçÿòûõ) ñ áîëåå óçêèìè ÷ëåíèêàìè áóëàâû. Òàê, 7-é ÷ëåíèê óñèêà íîâîãî âèäà â 1,9 ðàçà (I. kerneggeri — â 1,5 ðàçà) äëèííåå ñâîåé øèðèíû, 11-é ÷ëåíèê â 2,5 ðàçà äëèííåå ñâîåé øèðèíû (I. kerneggeri — â 2 ðàçà). Íîâûé âèä îòëè÷àåòñÿ îò I. kerneggeri òàêæå êðóïíîïóíêòèðîâàííûì äèñêîì ïðîíîòóìà è áîëåå êðóïíûìè ãëàçàìè, ïåðåäíèé êðàé êîòîðûõ çàìåòíî áëèæå ê îñíîâàíèÿì óñèêîâ, ÷åì ó I. kerneggeri. Ãîëîòèï íîâîãî âèäà ÷óòü ìåëü÷å ãîëîòèïà I. kerneggeri (â ïåðâîîïèñàíèè ýòîãî âèäà äëèíà ïîäîãíóòîé ãîëîâû íå ó÷òåíà â îáùåé äëèíå òåëà). Êîãîòêè I. kerneggeri ïðîñòûå, êàê è ó ñåâåðîàìåðèêàíñêèõ I. latus è Agyrtes longulus (LeConte) — â òî âðåìÿ êàê ó åâðàçèéñêèõ Agyrtes castaneus (F.) è A. ferrugineus Solsky êîãîòêè, êàê è ó íîâîãî âèäà, ñ çóáöîì; ðàñïðîñòðàíåíèå ýòîãî ïðèçíàêà ó àãèðòèä íèêîãäà ñïåöèàëüíî íå èçó÷àëîñü, íî åãî âèäîñïåöèôè÷íîñòü ïðåäñòàâ- ëÿåòñÿ äîñòàòî÷íî âåðîÿòíîé. Âåðõ ãîëîòèïà íîâîãî âèäà îäíîãî öâåòà ñ íîãàìè (ó I. latus è I. kerneggeri îí çàìåòíî òåìíåå — ãîëîòèï áàëòèéñêîãî âèäà òåìíî-÷åðíî- êîðè÷íåâûé), íî, âîçìîæíî, ÷òî ýòî — íåäàâíî âûøåäøèé èç êóêîëêè ýêçåìïëÿð. Ó íîâîãî âèäà, êàê è ó I. kerneggeri, íåò õàðàêòåðíûõ ùåòèíîê íà ëàïêàõ, îáíàðóæåííûõ íà 3–4-ì ÷ëåíèêàõ âñåõ ëàïîê îáîèõ ïîëîâ I. indicus è I. sikkimen- sis (Ãèìàëàè, Þãî-Âîñòî÷íàÿ Àçèÿ) À. Íüþòîíîì (Newton, 1997: 118). Âèä íàçâàí â ÷åñòü âûäàþùåãî îðãàíèçàòîðà ÿíòàðíûõ èññëåäîâàíèé Â. Âàéòøàòà (Dr. Wolfgang Weitschat) èç Ãàìáóðãñêîãî óíèâåðñèòåòà. Àâòîð èñêðåííå ïðèçíàòåëåí Êðèñòèàíó Íîéìàíó (Dr. Christian Neumann, Humboldt-Universitat zu Berlin, Museum fur Naturkunde) çà ïðåäîñòàâëåííûé ìàòåðèàë; À. Ï. Ðàñíèöûíó (Ïàëåîíòîëî- ãè÷åñêèé èíñòèòóò ÐÀÍ) è C. À. Cèìóòíèêó (Èíñòèòóò çîîëîãèè èì. è. È. Øìàëüãàóçåíà ÍÀÍ Óêðàèíû) çà ôîòîãðàôèè ãîëîòèïîâ ÿíòàðíûõ àãèðòèä. Ðàáîòà áûëà ïîääåðæàíà ãðàíòàìè DAAD A/02/16230 è Sepkoski Grant-2002 Ïàëåîíòîëî- ãè÷åñêîãî îáùåñòâà (The Paleontological Society). Ïåðêîâñêèé Å. Ý. Ïåðâûé
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    The evolution and genomic basis of beetle diversity Duane D. McKennaa,b,1,2, Seunggwan Shina,b,2, Dirk Ahrensc, Michael Balked, Cristian Beza-Bezaa,b, Dave J. Clarkea,b, Alexander Donathe, Hermes E. Escalonae,f,g, Frank Friedrichh, Harald Letschi, Shanlin Liuj, David Maddisonk, Christoph Mayere, Bernhard Misofe, Peyton J. Murina, Oliver Niehuisg, Ralph S. Petersc, Lars Podsiadlowskie, l m l,n o f l Hans Pohl , Erin D. Scully , Evgeny V. Yan , Xin Zhou , Adam Slipinski , and Rolf G. Beutel aDepartment of Biological Sciences, University of Memphis, Memphis, TN 38152; bCenter for Biodiversity Research, University of Memphis, Memphis, TN 38152; cCenter for Taxonomy and Evolutionary Research, Arthropoda Department, Zoologisches Forschungsmuseum Alexander Koenig, 53113 Bonn, Germany; dBavarian State Collection of Zoology, Bavarian Natural History Collections, 81247 Munich, Germany; eCenter for Molecular Biodiversity Research, Zoological Research Museum Alexander Koenig, 53113 Bonn, Germany; fAustralian National Insect Collection, Commonwealth Scientific and Industrial Research Organisation, Canberra, ACT 2601, Australia; gDepartment of Evolutionary Biology and Ecology, Institute for Biology I (Zoology), University of Freiburg, 79104 Freiburg, Germany; hInstitute of Zoology, University of Hamburg, D-20146 Hamburg, Germany; iDepartment of Botany and Biodiversity Research, University of Wien, Wien 1030, Austria; jChina National GeneBank, BGI-Shenzhen, 518083 Guangdong, People’s Republic of China; kDepartment of Integrative Biology, Oregon State
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  • Ofcanada Part13
    THE INSECTS ANDARAOHNIDS OFCANADA PART13 The ofca,.m'ffitrslP; Coleo r* SgHHy'" THE INSECTS ANDARACHNIDS OFCANADA t%RT13 The Carrion Beetles of Canada and Alaska Coleoptera Silphidae and Agyrtidae Robert S. Andersonl and Stewart B. Peck2 Biosystematics Research Institute Ottawa, Ontario Research Branch Agriculture Canada Publication 1778 1985 rUniyersity of Alberta, Edmonton, Alberta 2Carleton University, Ottawa, Ontario oMinister of Supply and Services Canada 1985 Available in Canada through Authorized Bookstore Agents and other bookstores or by mail from Canadian Government Publishing Centre Supply and Services Canada Ottawa, Canada KIA 0S9 Catalogue No. A42-42,21985-l3E Canada: $7.00 ISBN 0-662-11752-5 Other Countries: $8.40 Price subject to change without notice Canadian Cataloguing in Publication Data Anderson, Robert Samuel The carrion beetles of Canada and Alaska (Coleoptera: Silphidae and Agyrtidae) (The Insects and arachnids of Canada, ISSN 0706-7313 ; pt. 13) (Publication ;1778) Includes bibliographical references and index. l. Silphidae. 2. Beetles - Canada. 3. Beetles -- Alaska. I. Peck, Stewart B. II. Canada. Agricul- ture Canada. Research Branch. III. Title. IV. Series. V. Series: Publication (Canada. Agri- culture Canada). English ; 1778. QL596.S5A5 1985 595.76 C85-097200-0 The Insects and Arachnids of Canada Part l. Collecting, Preparing, and Preserving Insects, Mites, and Spiders, compiled by J. E. H. Martin, Biosystematics Research Institute, Ottawa, 1977. 182 p. Price: Canada $3.50, other countries $4.20 (Canadian funds). Cat. No. A42-42/1977 -1. Partie 1. R6colte, prdparation et conservation des Insectes, des Acariens et des Araign6es, compil6 par J.E.H. Martin, Institut de recherches biosyst6- matiques, Ottawa, 1983.
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  • A Redescripton of Lyrosoma Pallidum (Eschscholtz) and Distributional Range Extension of Lyrosoma Mannerheim (Coleoptera, Agyrtidae)
    A peer-reviewed open-access journal ZooKeys 329: 23–32 (2013) Korean Lyrosoma 23 doi: 10.3897/zookeys.329.4957 RESEARCH ARTICLE www.zookeys.org Launched to accelerate biodiversity research A redescripton of Lyrosoma pallidum (Eschscholtz) and distributional range extension of Lyrosoma Mannerheim (Coleoptera, Agyrtidae) In-Seong Yoo1, Derek Sikes2, Kee-Jeong Ahn1 1 Department of Biology, Chungnam National University, Daejeon 305-764, Republic of Korea 2 University of Alaska Museum, 907 Yukon Dr., Fairbanks, Alaska, 99775, USA Corresponding author: Kee-Jeong Ahn ([email protected]) Academic editor: Michael Ivie | Received 4 March 2013 | Accepted 26 August 2013 | Published 5 September 2013 Citation: Yoo I-S, Sikes D, Ahn K-J (2013) A redescripton of Lyrosoma pallidum (Eschscholtz) and distributional range extension of Lyrosoma Mannerheim (Coleoptera, Agyrtidae). ZooKeys 329: 23–32. doi: 10.3897/zookeys.329.4957 Abstract A redescription with illustrations of the species Lyrosoma pallidum and a key to the Korean species of the family Agyrtidae are provided. New distributional data, including a range extension, of the two Lyrosoma Mannerheim species are presented. Lyrosoma pallidum (Eschscholtz) is recorded for the first time in Korea. Keywords Lyrosoma pallidum, L. opacum, distribution range, coastal, Agyrtidae Introduction The genusLyrosoma Mannerheim, containing two species worldwide, is confined to coastal habitats, such as under stones, seaweeds, and carcasses of various coastal ani- mals along the seashore. They have also been reported in nests of maritime birds, but little is known regarding their immature stages and bionomics. They can be recognized by the combination of the following characters: mandibles without subapical teeth; an- tennomeres 9–10 each with apical grooves including compact distribution of sensilla; elytron with 9 striae; hind wings absent; pro- and mesotarsi dilated in males; aedeagus without parameres (Newton 1997; Schawaller 1998).
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  • Download Full Article 514.1KB .Pdf File
    Memoirs of the Museum of Victoria 56(2):659-666 (1997) 28 February 1997 https://doi.org/10.24199/j.mmv.1997.56.67 BIODIVERSITY OF NEW ZEALAND BEETLES (INSECTA, COLEOPTERA) J. KLIMASZEWSK.I Manaaki Whenua — Landcare Research, Private Bag 92170, Auckland, New Zealand Present address: BC Research. 3650 Weshrook Mall, Vancouver V6S SLS, Canada Abstract Klimaszewski, J., 1 997. Biodiversity of New Zealand beetles (Insecta: Coleoptera). Memoirs of the Museum of Victoria 56(2): 659-666. Approximately 5235 species are described for New Zealand, including 354 introduced. They belong to 82 families in two suborders, Adephaga and Polyphaga. The New Zealand beetle fauna is distinguished by the absence of many major lineages, a high level of endem- ism. which in many groups is over 90% at the specific level and over 43% at the generic level (e.g.. Staphylinidae), and the radiation of many groups of genera and species. The origins of New Zealand's beetle fauna are still poorly understood. They are likely to be varied, includ- ing Gondwanan elements and elements which arrived here by short and long-distance dispersal recently and in the remote past. The size of the New Zealand beetle fauna is con- sistent with species number/land area relationships in other areas around the world. Introduction Zealand beetles is that of Kuschel (1990), in the suburb of Lynfield, Auckland, in which 982 The beetles are the largest order of organisms, beetle species were recorded in a diverse veg- with over 350 000 described species world- etation including remnant forest, pastureland, wide. and suburban garden.
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  • New Distributional Records of the Flightless Primitive Carrion Beetle Necrophilus Pettitii Horn in Eastern North America (Coleoptera: Agyrtidae)
    University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln Center for Systematic Entomology, Gainesville, Insecta Mundi Florida March 2005 New distributional records of the flightless primitive carrion beetle Necrophilus pettitii Horn in eastern North America (Coleoptera: Agyrtidae) Stuart B. Peck Ottawa, Ontario, K1S 5B6, Canada Follow this and additional works at: https://digitalcommons.unl.edu/insectamundi Part of the Entomology Commons Peck, Stuart B., "New distributional records of the flightless primitive carrion beetle Necrophilus pettitii Horn in eastern North America (Coleoptera: Agyrtidae)" (2005). Insecta Mundi. 82. https://digitalcommons.unl.edu/insectamundi/82 This Article is brought to you for free and open access by the Center for Systematic Entomology, Gainesville, Florida at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Insecta Mundi by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. INSECTA MUNDI, Vol. 19, No. 1-2, March-June, 2005 123 New distributional records of the flightless primitive carrion beetle Necrophilus pettitii Horn in eastern North America (Coleoptera: Agyrtidae) Stewart B. Peck Department of Biology, Carleton University, 1125 Colonel By Drive Ottawa, Ontario, K1S 5B6, Canada ABSTRACT. New distribution records significantly expand the known range of flightless Necrophilus pettitii Horn in the eastern United States. Introduction 18.V.2002, J. Lewis and R. Burns, pitfall, 1 (SBPC); Molly Cave, Springs Valley, Hoosier National For- The only beetle in the family Agyrtidae known est, 18.V. 2002, J. Lewis and R. Burns, 2, (SBPC); from eastern North America is the flightless prim- Caves Cave, Paoli Country Club, 30.IV.1998, J. itive carrion beetle Necrophilus pettitii Horn, 1880.
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  • Mckenna2009chap34.Pdf
    Beetles (Coleoptera) Duane D. McKenna* and Brian D. Farrell and Polyphaga (~315,000 species; checkered beetles, Department of Organismic and Evolutionary Biology, 26 Oxford click beetles, A reP ies, ladybird beetles, leaf beetles, long- Street, Harvard University, Cambridge, MA 02138, USA horn beetles, metallic wood-boring beetles, rove beetles, *To whom correspondence should be addressed scarabs, soldier beetles, weevils, and others) (2, 3). 7 e ([email protected]) most recent higher-level classiA cation for living beetles recognizes 16 superfamilies and 168 families (4, 5). Abstract Members of the Suborder Adephaga are largely preda- tors, Archostemata feed on decaying wood (larvae) and Beetles are placed in the insect Order Coleoptera (~350,000 pollen (adults), and Myxophaga are aquatic or semi- described species). Recent molecular phylogenetic stud- aquatic and feed on green and/or blue-green algae ( 6). ies defi ne two major groups: (i) the Suborders Myxophaga Polyphaga exhibit a diversity of habits, but most spe- and Archostemata, and (ii) the Suborders Adephaga and cies feed on plants or dead and decaying plant parts Polyphaga. The time of divergence of these groups has (1–3). 7 e earliest known fossil Archostemata are from been estimated with molecular clocks as ~285–266 million the late Permian (7), and the earliest unequivocal fossil years ago (Ma), with the Adephaga–Polyphaga split at ~277– Adephaga and Polyphaga are from the early Triassic (1). 266 Ma. A majority of the more than 160 beetle families Myxophaga are not known from the fossil record, but are estimated to have originated in the Jurassic (200–146 extinct possible relatives are known from the Permian Ma).
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  • An Annotated List of Insects and Other Arthropods
    This file was created by scanning the printed publication. Text errors identified by the software have been corrected; however, some errors may remain. Invertebrates of the H.J. Andrews Experimental Forest, Western Cascade Range, Oregon. V: An Annotated List of Insects and Other Arthropods Gary L Parsons Gerasimos Cassis Andrew R. Moldenke John D. Lattin Norman H. Anderson Jeffrey C. Miller Paul Hammond Timothy D. Schowalter U.S. Department of Agriculture Forest Service Pacific Northwest Research Station Portland, Oregon November 1991 Parson, Gary L.; Cassis, Gerasimos; Moldenke, Andrew R.; Lattin, John D.; Anderson, Norman H.; Miller, Jeffrey C; Hammond, Paul; Schowalter, Timothy D. 1991. Invertebrates of the H.J. Andrews Experimental Forest, western Cascade Range, Oregon. V: An annotated list of insects and other arthropods. Gen. Tech. Rep. PNW-GTR-290. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Research Station. 168 p. An annotated list of species of insects and other arthropods that have been col- lected and studies on the H.J. Andrews Experimental forest, western Cascade Range, Oregon. The list includes 459 families, 2,096 genera, and 3,402 species. All species have been authoritatively identified by more than 100 specialists. In- formation is included on habitat type, functional group, plant or animal host, relative abundances, collection information, and literature references where available. There is a brief discussion of the Andrews Forest as habitat for arthropods with photo- graphs of representative habitats within the Forest. Illustrations of selected ar- thropods are included as is a bibliography. Keywords: Invertebrates, insects, H.J. Andrews Experimental forest, arthropods, annotated list, forest ecosystem, old-growth forests.
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  • A CATALOG of the COLEÓPTERA of AMERICA NORTH of MEXICO FAMILY: SILPHIDAE R
    ÑaHPtk as A CATALOG OF THE COLEÓPTERA OF AMERICA NORTH OF MEXICO FAMILY: SILPHIDAE r: CO ;~~ in NAL Digitizing ProjectEll ah52928 ^^i. UNITED STATES AGRICULTURE PREPARED BY ñka DEPARTMENT OF HANDBOOK AGRICULTURAL ^S&?f AGRICULTURE NUMBER 529-28 RESEARCH ^^^^ SERVICE FAMILIES OF COLEóPTERA IN AMERICA NORTH OF MEXICO Fascicle' Family Year issued Fascicle' Family Year issued Fascicle' Family Year issued 1 Cupedidae 1979 46 Callirhipidae 102 Biphyllidae 2 Micromalthidae 1982 47 Heteroceridae 1978 103 Byturidae 1991 3 Carabidae 48 Limnichidae 1986 104 Mycetophagidae 4 Rhysodidae 1985 49 Dryopidae 1983 105 Ciidae ^. 1982 5 Amphizoidae 1984 50 Elmidae 1983 107 Prostomidae 6 Haliplidae 51 Buprestidae 109 Colydiidae 8 Noteridae 52 Cebrionidae 110 Monommatidae 9 Dytiscidae 53 Elateridae 111 Cephaloidae 10 Gyrinidae 54 Throscidae 112 Zopheridae 13 Sphaeriidae 55 Cerophytidae 115 Tenebrionidae 14 Hydroscaphidae 56 Perothopidae 116 Alleculidae 15 Hydraenidae 57 Eucnemidae 117 Lagriidae 16 Hydrophilidae 58 Telegeusidae 118 Salpingidae 17 Georyssidae 61 Phengodidae 119 Mycteridae 18 Sphaeritidae 62 Lampyridae 120 Pyrochroidae 1983 20 Histeridae 63 Cantharidae 121 Othniidae 21 Ptiliidae 64 Lycidae 122 Inopeplidae 22 Limulodidae 65 Derodontidae 1989 123 Oedemeridae 23 Dasyceridae 66 Nosodendridae 124 Melandryidae 24 Micropeplidae 1984 67 Dermestidae 125 Mordellidae 1986 25 Leptinidae 69 Ptinidae 126 Rhipiphoridae 26 Leiodidae 70 Anobiidae 1982 127 Meloidae 27 Scydmaenidae 71 Bostrichidae 128 Anthicidae 28 Silphidae 1993 72 Lyctidae 129 Pedilidae 29 Scaphidiidae
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  • Coleoptera: Agyrtidae) of Central Asia and the Himalayas with a New Synonymy
    Zootaxa 3974 (1): 093–105 ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Article ZOOTAXA Copyright © 2015 Magnolia Press ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.3974.1.6 http://zoobank.org/urn:lsid:zoobank.org:pub:D03EB6D8-D375-4E65-825F-8B1DD9118CDD New data on Apteroloma (Coleoptera: Agyrtidae) of central Asia and the Himalayas with a new synonymy JAN RŮŽIČKA1, LEONARDO LATELLA2 & WOLFGANG SCHAWALLER3 1Department of Ecology, Faculty of Environmental Sciences, Czech University of Life Sciences Prague, CZ-165 21 Praha 6, Czech Republic. E-mail: [email protected] 2Museo Civico di Storia Naturale of Verona, Lungadige Porta Vittoria 9, I-37129 Verona, Italy. E-mail: [email protected] 3Staatliches Museum für Naturkunde, Rosenstein 1, D-70191 Stuttgart, Germany. E-mail: [email protected] Abstract The distribution of Apteroloma anglorossicum (Semenov, 1890), A. harmandi (Portevin, 1903) and A. sillemi Jeannel, 1935 in central Asia and along the Himalayas is summarized, and the collecting circumstances and ecology of all three species from Gilgit District, Pakistan are described in detail. Revised diagnoses of all three species are provided, habitus and important morphological structures are illustrated, and available types have been examined. Apteroloma jankovskii Semenov and Znojko in Semenov, 1932 is confirmed as junior subjective synonym of A. anglorossicum. Apteroloma hein- zi Schawaller, 1991 is newly treated as a junior subjective synonym of A. harmandi. Key words: Coleoptera, Agyrtidae, Apteroloma, taxonomy, new synonymy, distribution, Palaearctic region Introduction Agyrtidae are a small family of staphylinoid beetles, with more than 60 valid extant species in eight genera, distributed in temperate areas of the northern hemisphere (with one genus known from New Zealand); most of the species display a relict or disjunct distribution pattern (Newton 2005).
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