Genus Pseudosageceras Diener, 1895 Type Species

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Genus Pseudosageceras Diener, 1895 Type Species 140 Yasunari Shigeta et al. (eds.) Genus Pseudosageceras Diener, 1895 28) described as Pseudosageceras sp. indet from the Early Triassic of the Shamara Bay Type species: Pseudosageceras sp. indet. area, South Primorye is only a partial Diener, 1895. phargmocone, but it displays the same shell shape and suture as Pseudosageceras multilo- Pseudosageceras multilobatum batum Noetling, and it is probably conspecific. Noetling, 1905 P. multilobatum is one of the most cosmopoli- Figs. 129, 130 tan species of the Early Triassic. Pseudosageceras sp. indet. Diener, 1895, p. 28, pl. 1, fig. 8. Pseudosageceras multilobatum Noetling, 1905a, p. 181, Scaphopods (by Y. Shigeta) pls. 19–27; Noetling, 1905b, pl. 23, fig. 4, pl. 25, fig. Systematic descriptions basically follow the 1, pl. 26, fig. 3; Krafft and Diener, 1909, p. 145, pl. classification by Palmer (1974), Skelton and 21, fig. 5; Smith, 1932, p. 87, pl. 4, figs. 1–3, pl. 5, Benton (1993), and Steiner and Kabat (2001). figs. 1–6, pl. 25, figs. 7–16, pl. 60, fig. 32, pl. 63, figs. Abbreviations for shell dimensions: Lϭshell 1–6; Collignon, 1933, p. 56, pl. 11, fig. 2; Spath, ϭ 1934, p. 54, text-fig. 6a; Kiparosova, 1947, p. 127, pl. length; D diameter of anterior aperture. 25, figs. 3, 4; Chao, 1959, p. 183, pl. 1, figs. 9, 12; Institution abbreviations: NSMϭNational Kummel and Steele, 1962, p. 701, pl. 102, figs. 1, 2; Museum of Nature and Science, Tokyo. Hada, 1966, pl. 4, fig. 6; Kummel and Erben, 1968, p. 112, pl. 19, fig. 9; Shevyrev, 1968, p. 79, pl. 1, figs. Class Scaphopoda Bronn, 1862 1, 2; Weitschat and Lehmann, 1978, p. 95, pl. 10, fig. Order Dentalioida Palmer, 1974 2; Tozer, 1994, p. 83, pl. 18, fig. 1, text-fig. 17; Bra- yard and Bucher, 2008, p. 70, pl. 37, figs. 1–5. Family Laevidentaliidae Palmer, 1974 Genus Laevidentalium Cossmann, 1888 Lectotype: Designated by Spath (1934, p. 54), is original of Noetling (1905a, p. 181, pl. Type species: Dentalium incertum De- 19, fig. 1, pl. 24, fig. 12) from the Ceratite shayes, 1825. Marls of Salt Range, Pakistan. Material examined: One specimen, NSM PM23289, from AB1024, two specimens, Laevidentalium? sp. indet. NSM PM23290, 23291, from AB1022. Fig. 131 Description: Very involute, very compressed oxycone with very narrow, bicarinate venter Material examined: NSM PM23344 from and weakly convex flanks, convergent from AB1014, and NSM PM23345 from AB1016. occluded umbilicus to venter. Surface smooth Description: Shell slender, tapering, moder- without ornamentation. Suture line ceratitic ately curved, with circular cross section. Shell with many adventitious elements. Lateral wall thin to moderately thick. Ornamentation lobes trified, other lobes bified. consists of oblique, circular annulations of Measurements (mm): various strength as well as fine growth lines. Specimen no. D U H W W/H Internal mould smooth. NSM PM23289 30.2 0.0 18.7 6.2 0.33 Measurements (mm): NSM PM23291 43.8 0.0 28.0 10.0 0.36 Specimen no. L D NSM PM23344 12.4 2.1 Occurrence: Described specimens from NSM PM23345 9.6 1.4 AB1022, AB1024 within the Clypeoceras tim- orense Zone (early Early Olenekianϭearly Occurrence: Described specimens from Smithian) in the main part of the Zhitkov For- AB1014 within the Clypeoceras spitiense ϭ mation, Abrek Bay area, South Primorye. “bed” (early Late Induan early Dienerian) Discussion: The specimen Diener (1895, p. and from AB1016 within the Paranorites vari- Lower Triassic System in the Abrek Bay area, South Primorye 141 the glossary of malacological terms by Arnold (1965). Abbreviations for shell dimensions: For bellerophontids (according to Yochelson, 1960): Lϭlength, Wϭwidth, Tϭthickness; all others: Hϭshell height; Dϭshell diameter. Institution abbreviation: NSMϭNational Museum of Nature and Science, Tokyo. Class Gastropoda Cuvier, 1798 Order Amphigastropoda Simroth, 1906 Superfamily Bellerophontoidea McCoy, 1852 Family Euphemitidae Knight, 1956 Genus Warthia Waagen, 1880 Type species: Warthia brevisinuata Waagen, 1880. Warthia zakharovi Kaim sp. nov. Figs. 132–134 Type specimens: Holotype, NSM PM23322; four paratypes, NSM PM23323–23326, all from AB1016. Diagnosis: Warthia with wide shell in com- Fig. 131. Laevidentalium? sp. indet. 1, NSM parison to its thickness and weakly depressed PM23344, from AB1014. 2, NSM PM23345, from AB1016. Scale barϭ2mm. selenizone. Etymology: In honor of Yuri D. Zakharov. Description: Shell globular, almost as long as wide and lacking ornamentation. Slit short and broad at base of U-shaped sinus. Seleni- ans Zone (Late InduanϭDienerian) in the zone weakly depressed. Umbilicus absent. Zhitkov Formation, Abrek Bay area, South Measurements (mm): Primorye. Specimen no. L W T Discussion: The assignment of the speci- NSM PM23322 10.3 10.1 8.9 mens to Laevidentalium is uncertain and is NSM PM23323 10.8 10.0 7.7 based only on the similarity of their morpholo- NSM PM23324 14.3 12.7 11.2 gy with Laevidentalium. The described speci- NSM PM23325 14.5 10.8 9.8 mens are somewhat similar to L. bangtoupoen- NSM PM23326 9.2 9.4 7.5 NSM PM23329 13.0 12.1 11.1 sis Stiller (2001, p. 620) from the Upper NSM PM23330 17.6 17.5 15.0 Anisian of Qingyan, South-western China. Occurrence: Type specimens from AB1016 Gastropods (by A. Kaim) within the Paranorites varians Zone (Late Ind- Systematic descriptions basically follow the uanϭDienerian) in the lower main part of the classification by Bouchet et al. (2005). Mor- Zhitokv Formation, Abrek Bay area, South phological terms are those used in the Treatise Primorye, where this species locally forms a on Invertebrate Paleontology (Cox, 1960) and coquina composed mostly of Warthia shells. 142 Yasunari Shigeta et al. (eds.) Fig. 132. Warthia zakharovi Kaim sp. nov. from AB1016. 1–5, NSM PM23323, paratype. 6–12, NSM PM23322, holotype. All SEM images. Lower Triassic System in the Abrek Bay area, South Primorye 143 Fig. 133. Warthia zakharovi Kaim sp. nov. from AB1016. 1–4, NSM PM23324, paratype. 5–9, NSM PM23325, paratype. 10–14, NSM PM23322, holotype. 15–19, NSM PM23323, paratype. 20–23, NSM PM23326, paratype. All light images by Y. Shigeta. 144 Yasunari Shigeta et al. (eds.) Fig. 134. Warthia zakharovi Kaim sp. nov. 1–4, NSM PM23330, from AB1021. 5–9, NSM PM23329, from AB1008. 10–13, NSM PM23327, from AB1004. 14–17, NSM PM23328, from AB1004. All light images by Y. Shigeta. Lower Triassic System in the Abrek Bay area, South Primorye 145 Fig. 135. Bellerophon abrekensis Kaim sp. nov. from AB1010. 1–6, 10, NSM PM23332, paratype. 7–9, 11–12, NSM PM23333, paratype. All SEM images. 146 Yasunari Shigeta et al. (eds.) Fig. 136. Bellerophon abrekensis Kaim sp. nov. from AB1010. 1–4, NSM PM23331, holotype. 5–7, NSM PM23343, inner mould. 8, NSM PM23334, paratype. 9–10, NSM PM23333, paratype. 11, NSM PM23332, paratype. 12, NSM PM23335, paratype. All light images by Y. Shigeta. Additional specimens from AB1004, AB1008, selenizone. Poorly preserved shells of the pu- and AB1021. Two specimens, (NSM PM23327 tative euphemitid bellerophontid Euphemites and PM23328) from AB1004 (Early Induanϭ guizhouensis Pan, 1982 from the Lower Trias- Griesbachian), are poorly preserved and tenta- sic of China are much more compressed as is tively assigned to W. zakharovi. Fossiliferous also the case of the poorly constituted genus horizon at AB1008 (Early InduanϭGries- Stachella (for discussion see Yochelson & bachian) yielded only a single specimen, NSM Hongfu, 1985). PM23329. The largest shell, NSM PM23330, came from AB1021 within the Clypeoceras Family Bellerophontidae McCoy, 1852 timorense Zone (early Early Olenekianϭearly Genus Bellerophon Montfort, 1808 Smithian). Discussion: Warthia zakharovi sp. nov. dif- Type species: Bellerophon vasulites Mont- fers from other species of the genus in having fort, 1808. a strongly globular shell and weakly depressed Lower Triassic System in the Abrek Bay area, South Primorye 147 Bellerophon abrekensis Kaim sp. nov. which was later indentified as “Bellerophon Figs. 135, 136 asiaticus” by Kiparisova (1947a). I agree with Yochelson and Hongfu (1985) that specimens Bellerophon sp. indet. Bittner, 1899b, p. 28, pl. 4, figs. 26–28. discussed by Bittner (1899b) and Kiparisova non Bellerophon asiaticus Wirth, 1936, p. 441, fig. 14: 7a, (1947a) are not conspecific with Retispira asi- b. atica (Wirth, 1936), a species based on poorly Bellerophon asiaticus Wirth. Kiparisova 1947a, p. 121, pl. preserved material from Sichuan Province of 24, figs. 6, 7. China. Yochelson and Hongfu (1985) based Type specimens: Holotype, NSM PM23331; their concept of this species on a new material four paratypes, NSM PM23332–23335, all from Guizhou Province of China, which clear- from AB1010. ly displays spiral ornamentation that is diag- Diagnosis: Bellerophon with well developed nostic of the genus Retispira. I could not con- collabral ornamentation on adults composed firm such ornamentation on any of the well of sets of thicker ribs with numerous, smaller preserved specimens from Abrek Bay, and intercalated ribs. Selenizone slightly elevated. therefore, I retained the genus Bellerophon for Moderately thick inductura well developed on B. abrekensis. On the other hand I concur with adult specimens. No spiral ornamentation. the opinion of Yochelson and Hongfu (1985) Etymology: After type locality. that the specimens from Far East Russia could Description: Shell strongly globular, wider be synonimized with B. panxianensis Yu in than long. Shell ornamented with collabral Wang and Xi (1980). I examined a latex cast ribs bent posteriorly towards selenizone. In of the holotype of the latter species and it is adults, ribs differentiated into sets of stronger clearly an anomphalous form, while B. and weaker ribs. Selenizone long and slightly abrekensis sp. nov. has a well developed um- elevated. Umbilicus well developed, but par- bilicus. tially obscured in adults by well developed in- ductura extending over the parietal region of Order Vetigastropoda Salvini-Plawen, 1980 aperture. Superfamily Trochonematoidea Zittel, 1895 Measurements (mm): Family Lophospiridae Wenz, 1938 Specimen no.
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