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Zootaxa,Montealtosuchus Arrudacamposi, a New Peirosaurid
Zootaxa 1607: 35–46 (2007) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ ZOOTAXA Copyright © 2007 · Magnolia Press ISSN 1175-5334 (online edition) Montealtosuchus arrudacamposi, a new peirosaurid crocodile (Mesoeucrocodylia) from the Late Cretaceous Adamantina Formation of Brazil ISMAR DE SOUZA CARVALHO1, FELIPE MESQUITA DE VASCONCELLOS1 & SANDRA APARE- CIDA SIMIONATO TAVARES2 1Universidade Federal do Rio de Janeiro. Departamento de Geologia, CCMN/IGEO. 21.949-900 Cidade Universitária - Ilha do Fundão. Rio de Janeiro - RJ. Brasil. E-mail: [email protected]; [email protected] 2Museu de Paleontologia de Monte Alto. Praça do Centenário, Centro de Artes s/no. 15.910-000, MonteAlto-Brazil. E-mail: [email protected] Abstract We describe a new species of Peirosauridae (Crocodyliformes, Mesoeucrocodylia), Montealtosuchus arrudacamposi gen. nov. et sp. nov., from the Late Cretaceous (Turonian-Santonian) strata of the Bauru Basin, Brazil. Montealtosuchus was found at the outskirts of Monte Alto County in reddish sandstones of the Adamantina Formation. This specimen is exquisitely preserved with skull, mandible, postcranial and exoskeletal elements in articulation that provides critical information of the anatomy of this group. The occurrence of Peirosauridae in the Adamantina Formation (Turonian-San- tonian) widens the chronostratigraphic range of this Mesoeucrocodylia taxon in Brazil. Recent analysis suggests that the Peirosauridae is restricted to the Late Cretaceous deposits of South America. Key words: Montealtosuchus arrudacamposi gen. nov. et sp. nov.; Peirosauridae; Upper Cretaceous; Adamantina For- mation; Bauru Basin Introduction The Bauru Basin comprises an area between latitudes 18o S and 24o S, and longitudes 47o W and 56o W, and covers an area over 370.000 km2 in the southeast interior of Brazil, with outcrops in São Paulo, Minas Gerais, Mato Grosso do Sul and Goiás states. -
Taxonomic Reappraisal of the Sphagesaurid Crocodyliform Sphagesaurus Montealtensis from the Late Cretaceous Adamantina Formation of São Paulo State, Brazil
TERMS OF USE This pdf is provided by Magnolia Press for private/research use. Commercial sale or deposition in a public library or website is prohibited. Zootaxa 3686 (2): 183–200 ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Article ZOOTAXA Copyright © 2013 Magnolia Press ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.3686.2.4 http://zoobank.org/urn:lsid:zoobank.org:pub:9F87DAC0-E2BE-4282-A4F7-86258B0C8668 Taxonomic reappraisal of the sphagesaurid crocodyliform Sphagesaurus montealtensis from the Late Cretaceous Adamantina Formation of São Paulo State, Brazil FABIANO VIDOI IORI¹,², THIAGO DA SILVA MARINHO3, ISMAR DE SOUZA CARVALHO¹ & ANTONIO CELSO DE ARRUDA CAMPOS² 1UFRJ, Departamento de Geologia, CCMN/IGEO, Cidade Universitária – Ilha do Fundão, 21949-900. Rio de Janeiro, Brazil. E-mail: [email protected]; [email protected] 2Museu de Paleontologia “Prof. Antonio Celso de Arruda Campos”, Praça do Centenário s/n, Centro, 15910-000 – Monte Alto, Brazil 3Instituto de Ciências Exatas, Naturais e Educação (ICENE), Universidade Federal do Triângulo Mineiro (UFTM), Av. Dr. Randolfo Borges Jr. 1700 , Univerdecidade, 38064-200, Uberaba, Minas Gerais, Brasil. [email protected] Abstract Sphagesaurus montealtensis is a sphagesaurid whose original description was based on a comparison with Sphagesaurus huenei, the only species of the clade described to that date. Better preparation of the holotype and the discovery of a new specimen have allowed the review of some characteristics and the identification -
71St Annual Meeting Society of Vertebrate Paleontology Paris Las Vegas Las Vegas, Nevada, USA November 2 – 5, 2011 SESSION CONCURRENT SESSION CONCURRENT
ISSN 1937-2809 online Journal of Supplement to the November 2011 Vertebrate Paleontology Vertebrate Society of Vertebrate Paleontology Society of Vertebrate 71st Annual Meeting Paleontology Society of Vertebrate Las Vegas Paris Nevada, USA Las Vegas, November 2 – 5, 2011 Program and Abstracts Society of Vertebrate Paleontology 71st Annual Meeting Program and Abstracts COMMITTEE MEETING ROOM POSTER SESSION/ CONCURRENT CONCURRENT SESSION EXHIBITS SESSION COMMITTEE MEETING ROOMS AUCTION EVENT REGISTRATION, CONCURRENT MERCHANDISE SESSION LOUNGE, EDUCATION & OUTREACH SPEAKER READY COMMITTEE MEETING POSTER SESSION ROOM ROOM SOCIETY OF VERTEBRATE PALEONTOLOGY ABSTRACTS OF PAPERS SEVENTY-FIRST ANNUAL MEETING PARIS LAS VEGAS HOTEL LAS VEGAS, NV, USA NOVEMBER 2–5, 2011 HOST COMMITTEE Stephen Rowland, Co-Chair; Aubrey Bonde, Co-Chair; Joshua Bonde; David Elliott; Lee Hall; Jerry Harris; Andrew Milner; Eric Roberts EXECUTIVE COMMITTEE Philip Currie, President; Blaire Van Valkenburgh, Past President; Catherine Forster, Vice President; Christopher Bell, Secretary; Ted Vlamis, Treasurer; Julia Clarke, Member at Large; Kristina Curry Rogers, Member at Large; Lars Werdelin, Member at Large SYMPOSIUM CONVENORS Roger B.J. Benson, Richard J. Butler, Nadia B. Fröbisch, Hans C.E. Larsson, Mark A. Loewen, Philip D. Mannion, Jim I. Mead, Eric M. Roberts, Scott D. Sampson, Eric D. Scott, Kathleen Springer PROGRAM COMMITTEE Jonathan Bloch, Co-Chair; Anjali Goswami, Co-Chair; Jason Anderson; Paul Barrett; Brian Beatty; Kerin Claeson; Kristina Curry Rogers; Ted Daeschler; David Evans; David Fox; Nadia B. Fröbisch; Christian Kammerer; Johannes Müller; Emily Rayfield; William Sanders; Bruce Shockey; Mary Silcox; Michelle Stocker; Rebecca Terry November 2011—PROGRAM AND ABSTRACTS 1 Members and Friends of the Society of Vertebrate Paleontology, The Host Committee cordially welcomes you to the 71st Annual Meeting of the Society of Vertebrate Paleontology in Las Vegas. -
A Troodontid Dinosaur from the Latest Cretaceous of India
ARTICLE Received 14 Dec 2012 | Accepted 7 Mar 2013 | Published 16 Apr 2013 DOI: 10.1038/ncomms2716 A troodontid dinosaur from the latest Cretaceous of India A. Goswami1,2, G.V.R. Prasad3, O. Verma4, J.J. Flynn5 & R.B.J. Benson6 Troodontid dinosaurs share a close ancestry with birds and were distributed widely across Laurasia during the Cretaceous. Hundreds of occurrences of troodontid bones, and their highly distinctive teeth, are known from North America, Europe and Asia. Thus far, however, they remain unknown from Gondwanan landmasses. Here we report the discovery of a troodontid tooth from the uppermost Cretaceous Kallamedu Formation in the Cauvery Basin of South India. This is the first Gondwanan record for troodontids, extending their geographic range by nearly 10,000 km, and representing the first confirmed non-avian tetanuran dinosaur from the Indian subcontinent. This small-bodied maniraptoran dinosaur is an unexpected and distinctly ‘Laurasian’ component of an otherwise typical ‘Gondwanan’ tetrapod assemblage, including notosuchian crocodiles, abelisauroid dinosaurs and gondwanathere mammals. This discovery raises the question of whether troodontids dispersed to India from Laurasia in the Late Cretaceous, or whether a broader Gondwanan distribution of troodontids remains to be discovered. 1 Department of Genetics, Evolution, and Environment, University College London, London WC1E 6BT, UK. 2 Department of Earth Sciences, University College London, London WC1E 6BT, UK. 3 Department of Geology, Centre for Advanced Studies, University of Delhi, New Delhi 110 007, India. 4 Geology Discipline Group, School of Sciences, Indira Gandhi National Open University, New Delhi 110 068, India. 5 Division of Paleontology and Richard Gilder Graduate School, American Museum of Natural History, New York, New York 10024, USA. -
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DOI 10.5935/0100-929X.20120007 Revista do Instituto Geológico, São Paulo, 33 (2), 13-29, 2012 DESCRIÇÃO DE UM ESPÉCIME JUVENIL DE BAURUSUCHIDAE (CROCODYLIFORMES: MESOEUCROCODYLIA) DO GRUPO BAURU (NEOCRETÁCEO): CONSIDERAÇÕES PRELIMINARES SOBRE ONTOGENIA Caio Fabricio Cezar GEROTO Reinaldo José BERTINI RESUMO Entre os táxons de Crocodyliformes do Grupo Bauru (Neocretáceo), grande quan- tidade de morfótipos, incluindo materiais cranianos e pós-cranianos, vem sendo des- crita em associação a Baurusuchus pachecoi. Porém, a falta de estudos ontogenéticos, como os realizados para Mariliasuchus amarali, levou à atribuição de novos gêneros e espécies aos poucos crânios completos encontrados. A presente contribuição traz a descrição de um espécime juvenil de Baurusuchidae depositado no acervo do Museu de Ciências da Terra no Rio de Janeiro sob o número MCT 1724 - R. Trata-se de um rostro e mandíbula associados e em oclusão, com o lado esquerdo melhor preservado que o direito, e dentição zifodonte extremamente reduzida. O fóssil possui 125,3 mm de comprimento preservado da porção anterior do pré-maxilar até a extremidade pos- terior do dentário; 117,5 mm de comprimento preservado do pré-maxilar aos palatinos e altura lateral de 51,4 mm. Entre as informações de caráter ontogenético identificadas destacam-se: ornamentação suave composta de estrias vermiformes muito espaçadas e largas, linha ventral do maxilar mais reta, dentário levemente inclinado dorsalmente na porção mediana e sínfise mandibular menos vertical que em outros baurussúqui- dos de tamanho maior. A maioria das características rostrais e dentárias, diagnósticas para Baurusuchus pachecoi, foi identificada no exemplar MCT 1724 - R: rostro alto e comprimido lateralmente, além de dentição zifodonte com forte redução dentária, que culmina em quatro dentes pré-maxilares e cinco maxilares. -
Craniofacial Morphology of Simosuchus Clarki (Crocodyliformes: Notosuchia) from the Late Cretaceous of Madagascar
Society of Vertebrate Paleontology Memoir 10 Journal of Vertebrate Paleontology Volume 30, Supplement to Number 6: 13–98, November 2010 © 2010 by the Society of Vertebrate Paleontology CRANIOFACIAL MORPHOLOGY OF SIMOSUCHUS CLARKI (CROCODYLIFORMES: NOTOSUCHIA) FROM THE LATE CRETACEOUS OF MADAGASCAR NATHAN J. KLEY,*,1 JOSEPH J. W. SERTICH,1 ALAN H. TURNER,1 DAVID W. KRAUSE,1 PATRICK M. O’CONNOR,2 and JUSTIN A. GEORGI3 1Department of Anatomical Sciences, Stony Brook University, Stony Brook, New York, 11794-8081, U.S.A., [email protected]; [email protected]; [email protected]; [email protected]; 2Department of Biomedical Sciences, Ohio University College of Osteopathic Medicine, Athens, Ohio 45701, U.S.A., [email protected]; 3Department of Anatomy, Arizona College of Osteopathic Medicine, Midwestern University, Glendale, Arizona 85308, U.S.A., [email protected] ABSTRACT—Simosuchus clarki is a small, pug-nosed notosuchian crocodyliform from the Late Cretaceous of Madagascar. Originally described on the basis of a single specimen including a remarkably complete and well-preserved skull and lower jaw, S. clarki is now known from five additional specimens that preserve portions of the craniofacial skeleton. Collectively, these six specimens represent all elements of the head skeleton except the stapedes, thus making the craniofacial skeleton of S. clarki one of the best and most completely preserved among all known basal mesoeucrocodylians. In this report, we provide a detailed description of the entire head skeleton of S. clarki, including a portion of the hyobranchial apparatus. The two most complete and well-preserved specimens differ substantially in several size and shape variables (e.g., projections, angulations, and areas of ornamentation), suggestive of sexual dimorphism. -
Cranial Features of Baurusuchus Salgadoensis
ISBN 978-85-7193-184-8 – Editora Interciência 2007 Paleontologia: Cenários de Vida CRANIAL FEATURES OF BAURUSUCHUS SALGADOENSIS CARVALHO, CAMPOS & NOBRE 2005, A BAURUSUCHIDAE ΈMESOEUCROCODYLIAΉ FROM THE ADAMANTINA FORMATION, BAURU BASIN, BRAZIL: PALEOICHNOLOGICAL, TAXONOMIC AND SYSTEMATIC IMPLICATIONS Felipe Mesquita de Vasconcellos & Ismar de Souza Carvalho Universidade Federal do Rio de Janeiro. Departamento de Geologia, CCMN/IGEO. 21.949-900 Cidade Universitária - Ilha do Fundão. Rio de Janeiro - RJ. Brasil E-mail: [email protected], [email protected] ABSTRACT Some features of the skull of Baurusuchus salgadoensis Carvalho, Campos & Nobre 2005, a baurusuchid Mesoeucrocodylia from the Adamantina Formation of Bauru Basin, are described, discussed and reinterpreted. The punctures and perforations of the skull of B. salgadoensis, one of them previously described as the antobital fenestrae, were interpreted as tooth-marks. The probable producer is a medium or large ziphodont terrestrial archosaur, possibly a baurusuchid or Abelisauridae. The choanae of B. salgadoensis bears some similarities with Stratiotosuchus. The choanae and the palatal surfaces seem to be similar among baurusuchids, notosuchids and sphagesaurids with minor differences. This similarity is congruent with recent phylogenetic hypotheses, supporting a closer relationship among these Creataceous Mesoeucrodylia taxa. Key-words: Baurusuchus salgadoensis, Upper Cretaceous, Bauru Basin RESUMO Algumas características do crânio de Baurusuchus salgadoensis Carvalho, Campos & Nobre 2005, um baurussuquídeo Mesoeucrocodylia proveniente da Formação Adamantina da Bacia Bauru, são descritas, discutidas e reinterpretadas. As perfurações e depressões presentes no crânio de B. salgadoensis, uma delas descrita anteriormente como a fenestra antorbital, foram interpretadas como marcas de dentes. O provável produtor destas marcas é um arcossauro terrestre de médio à grande porte com dentes zifodontes, possivelmente um baurussuquídeo ou Abelisauridae. -
A New Pissarrachampsinae Specimen from the Bauru Basin, Brazil, Adds
YCRES104969_proof ■ 29 July 2021 ■ 1/13 Cretaceous Research xxx (xxxx) xxx 55 Contents lists available at ScienceDirect 56 57 58 Cretaceous Research 59 60 journal homepage: www.elsevier.com/locate/CretRes 61 62 63 64 65 1 A new Pissarrachampsinae specimen from the Bauru Basin, Brazil, 66 2 67 3 adds data to the understanding of the Baurusuchidae 68 4 (Mesoeucrocodylia, Notosuchia) distribution in the Late Cretaceous of 69 5 70 6 Q7 South America 71 7 72 8 a, * b, c d Q6 Gustavo Darlim , Ismar de Souza Carvalho , Sandra Aparecida Simionato Tavares , 73 9 Max Cardoso Langer a 74 10 75 11 a Universidade de Sao~ Paulo, Faculdade de Filosofia, Ci^encias e Letras de Ribeirao~ Preto, Laboratorio de Paleontologia, Av. Bandeirantes, 3900, Ribeirao~ Preto, 76 12 SP, Brazil 77 b ^ 13 Q2 Universidade Federal do Rio de Janeiro, Instituto de Geociencias, CCMN, 21.910-200 Av. Athos da Silveira Ramos 273, Rio de Janeiro, RJ, Brazil c Centro de Geoci^encias da Universidade de Coimbra, Portugal 78 14 d Museu de Paleontologia “Prof. Antonio^ Celso de Arruda Campos”, Praça do Centenario, Centro, Monte Alto, SP, Brazil 79 15 80 16 81 17 article info abstract 82 18 83 19 Article history: Baurusuchidae is one of the most diverse notosuchian groups, represented by ten formally described 84 20 Received 7 January 2021 species from the Upper Cretaceous deposits of the Bauru and Neuquen basins, respectively in Brazil and 85 21 Received in revised form Argentina. Among these, recent phylogenetic analyses placed Wargosuchus australis, Campinasuchus 86 18 July 2021 22 dinizi, and Pissarrachampsa sera within Pissarrachampsinae, whereas Baurusuchinae is composed by 87 Accepted in revised form 18 July 2021 23 Aphaurosuchus escharafacies, Aplestosuchus sordidus, Baurusuchus albertoi, Baurusuchus pachecoi, Baur- Available online xxx 88 24 usuchus salgadoensis, and Stratiotosuchus maxhechti. -
The Baurusuchidae Vs Theropoda Record in the Bauru Group (Upper Cretaceous, Brazil): a Taphonomic Perspective
Journal of Iberian Geology https://doi.org/10.1007/s41513-018-0048-4 RESEARCH PAPER The Baurusuchidae vs Theropoda record in the Bauru Group (Upper Cretaceous, Brazil): a taphonomic perspective Kamila L. N. Bandeira1 · Arthur S. Brum1 · Rodrigo V. Pêgas1 · Giovanne M. Cidade2 · Borja Holgado1 · André Cidade1 · Rafael Gomes de Souza1 Received: 31 July 2017 / Accepted: 23 January 2018 © Springer International Publishing AG, part of Springer Nature 2018 Abstract Purpose The Bauru Group is worldwide known due to its high diversity of archosaurs, especially that of Crocodyliformes. Recently, it has been suggested that the Crocodyliformes, especially the Baurusuchidae, were the top predators of the Bauru Group, based on their anatomical convergence with theropods and the dearth of those last ones in the fossil record of this geological group. Methods Here, we erect the hypothesis that assumption is taphonomically biased. For this purpose, we made a literature survey on all the published specimens of Theropoda, Baurusuchidae and Titanosauria from all geological units from the Bauru Group. Also, we gathered data from the available literature, and we classifed each fossil fnd under a taphonomic class proposed on this work. Results We show that those groups have diferent degrees of bone representativeness and diferent qualities of preservation pattern. Also, we suggest that baurusuchids lived close to or in the abundant food plains, which explains the good preserva- tion of their remains. Theropods and titanosaurs did not live in association with such environments and the quality of their preservation has thus been negatively afected. Conclusions We support the idea that the Baurusuchidae played an important role in the food chain of the ecological niches of the Late Cretaceous Bauru Group, but the possible biases in their fossil record relative to Theropoda do not support the conclusion that baurusuchids outcompeted theropods. -
I – Introdução
UNIVERSIDADE ESTADUAL PAULISTA Instituto de Geociências e Ciências Exatas Campus de Rio Claro REVISÃO SISTEMÁTICA E TAXONÔMICA DOS NOTOSUCHIA (METASUCHIA, CROCODYLOMORPHA) Marco Brandalise de Andrade Orientador: Prof. Dr. Reinaldo José Bertini Dissertação de Mestrado elaborada junto ao Programa de Pós-Graduação em Geologia - Área de concentração em Geologia Regional, para a obtenção do título de Mestre em Geociências Rio Claro (SP) 2005 Comissão Examinadora _____________________________________ Prof. Dr. Reinaldo José Bertini _____________________________________ Prof. Dr. Alexander Wilhelm Armin Kellner _____________________________________ Prof. Dr. Antonio Roberto Saad _____________________________________ Marco Brandalise de Andrade - aluno - Rio Claro, ____ de ________________ de 2005 Resultado: _____________________________________________________________ AGRADECIMENTOS Ao longo do desenvolvimento deste Mestrado, em seu caminho paradoxalmente tão longo e tão breve, contei com o apoio de um grande número de pessoas. Algumas destas contribuíram de forma direta, enquanto outras sequer estavam cientes de sua importância ao longo deste processo, em uma grande diversidade de aspectos. Agradeço em primeiro lugar ao Prof. Dr. Reinaldo José Bertini, por sua orientação e pela grande atenção e paciência a este estudante, me recebendo sem reservas na cidade de Rio Claro e na UNESP, bem como na comunidade científica. Agradeço à Universidade Estadual Paulista, ao Instituto de Geociências e Ciências Exatas e ao Departamento de Geologia Aplicada, em cujas dependências o estudo encontrou condições para o seu desenvolvimento. De fundamental importância foi o suporte fornecido pela Coordenação de Aperfeiçoamento de Pessoal de Nível Superior (CAPES), que forneceu uma Bolsa de Mestrado que em muito ajudou a ampliar o potencial e o alcance deste trabalho. Agradeço ao Prof. Dr. Antonio Roberto Saad e a Profa. -
Razanandrongobe Sakalavae, a Gigantic Mesoeucrocodylian from the Middle Jurassic of Madagascar, Is the Oldest Known Notosuchian
Razanandrongobe sakalavae, a gigantic mesoeucrocodylian from the Middle Jurassic of Madagascar, is the oldest known notosuchian peerj.com /articles/3481/ Introduction A decade ago, Maganuco, Dal Sasso & Pasini (2006) described the fragmentary remains of a very large predatory archosaur from the Middle Jurassic (Bathonian) of the Mahajanga Basin, Madagascar. The material included a fragmentary right maxilla bearing three teeth, and seven peculiar isolated teeth clearly belonging to the same taxon. In spite of the scanty remains, the presence of a unique combination of features, which included a well-developed bony palate on the maxilla, mesial and lateral teeth respectively U-shaped and sub-oval in cross-section, and very large tooth denticles (1 per mm) on the carinae, allowed the authors to erect the new taxon Razanandrongobe sakalavae Maganuco, Dal Sasso & Pasini, 2006. However, the systematic position of the new species remained uncertain: indeed, besides the autapomorphic denticle size, R. sakalavae shared a mix of potential autapomorphic, synapomorphic, and homoplasic features with crocodylomorphs and theropods. Therefore, the species was referred to Archosauria incertae sedis. Here we describe new cranial material referable to R. sakalavae and consisting of an almost complete right premaxilla, the rostral half of a left dentary, a maxillary fragment with diagnostic teeth, and a very large isolated tooth crown. In addition, we tentatively refer to the same taxon five cranial fragments that were likely collected at the same locality. -
Chubut Province, Argentina) and Its Phylogenetic Position Within Basal Mesoeucrocodylia
Cretaceous Research 30 (2009) 1376–1386 Contents lists available at ScienceDirect Cretaceous Research journal homepage: www.elsevier.com/locate/CretRes The first crocodyliform from the Chubut Group (Chubut Province, Argentina) and its phylogenetic position within basal Mesoeucrocodylia Juan Martı´n Leardi a,c,*, Diego Pol b,c a Laboratorio de Paleontologı´a de Vertebrados, Departamento de Ciencias Geolo´gicas, Facultad de Ciencias Exactas y Naturales, Pabello´n II, Universidad de Buenos Aires, Intendente Gu¨iraldes 2160, Ciudad Universitaria (C1428EGA), Buenos Aires, Argentina b Museo Paleontolo´gico Egidio Feruglio, Avenue Fontana 140 (9100), Trelew, Chubut, Argentina c CONICET- Consejo Nacional de Investigaciones Cientı´ficas y Te´cnicas, Avenue Rivadavia 1917, Buenos Aires, Argentina article info abstract Article history: A new crocodyliform specimen is presented here found in the Cerro Castan˜o Member of the Cerro Received 27 February 2009 Barcino Formation (Chubut Group). The material consists of cranial and postcranial remains that Accepted in revised form 4 August 2009 represent a new taxon that has strong affinities with Peirosauridae, but also shares derived features Available online 11 August 2009 present in Araripesuchus. The phylogenetic relationships of this new taxon were tested through a cladistic analysis depicting it as a member of the Peirosauridae. The inclusion of Barcinosuchus within this clade of Keywords: basal mesoeucrocodylians is supported by the presence of hypapophyses up to the third or fourth dorsal Crocodyliformes vertebrae, anterolateral facing edge on postorbital, quadrate dorsal surface divided in two planes by Peirosauridae Lower Cretaceous a ridge; mandibular symphysis tapering anterirorly in ventral view, lateral surface of dentary convex Central Patagonia anterior to mandibular fenestra, distal body of quadrate well developed, anteroposteriorly thin and Argentina lateromedially broad.