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ft elong to unusulomintion of feturesD suggesting the presene of yet unE reported notosuhin txon in the edritiEhinri gronte ltE theropod dinosur @F gurrieD persF ommFAF es dinosur form loted in the ethys etween the efrorin ontinent nd evidene is rre in stlyD it would e worth of noteF sn the xorth iuropen lndmss during vte greteous timesF xotosuE the mentimeD the tooth hs een reently mentioned in hins were typilly terrestril roodyliformsD supporting the preE literture s evidene of the genus hortodon in the sene of emergent res on the ronte pltformF pper greteous of stly @helfino PHHIY i PHHVAD issuntoF n dente munito di rene dentiolte rinvenuto nel with potentil pleogeogrphi implitionsD despite the sito fossilifero di olzzo @grsoD stli nordorientleA e dtile l ft tht its identifition in previous ppers ws lerly goniinoEntonino @gretio uperioreA eÁl prim testimoninz tenttive nd preliminryF di un oodrilliforme notosuo in stliF eene ondivid sinpoE morfie on denti ttriuiti lgenere europeo hortodon e on ilgondE wnino erripesuhusD esso si distingue per l9inusule ominzione di rtteriD he suggerise l presenz di un speie di notosuo non erminology nd methods nor identifit nell ittform gronti edritioEhinriD lolizzt tr il ontinente efrorio e quello xord iuropeoF s he odontologil terminology nd vriles used in this pper notosuhi erno oodrilliformi prettmente terrestri e questo eÁindiE re those of mith 8 hodson @PHHQA nd mith et lF @PHHSAF he terms tivo dell presenz di ree emerse sull pittform rontiF elle9 nd ude9 re used in the sense of eler @IWWPAF rmeters of hortodon teeth from eustriD omni nd rungry re sed on mesurements tken from photogrphs pulished in fuffetut @IWUWAD wrtin et lF @PHHTAD nd ourtesy of wrton iD respetivelyF sntrodution he verostrn theropods re onsidered ording to izurr huring the IWWU field work in pper greteous @PHHTA s those elonging to the lest inlusive lde inluding neoerE tosurs nd tetnurns @pigF PeAF he roodyliform phylogeneti olzzo fossil site @poglinoEedipugli wuniiplityD frmework nd txonomy followed herein is tht of ereno 8 vrsson qorizi rovineD xi stlyY pigF IAD n unusul serrted @PHHWA nd ereno et lF @PHHIAF eordinglyD the groodyliformes re tooth ws found @igo IWWUAF he speimen ws preliE the lest inlusive lde inluding protosuhids nd rownEgroup roE minrily nd tenttively referred to the ziphodont roE odiles @groodyliAD the wesoeuroodyli is the most inlusive roE odyliform hortodon in some overviews of the olzE odyliform lde exluding protosuhidsD nd the xotosuhi is the zo fossil ssemlge @eFgFD hll ehi et lF PHHIY most inlusive lde inluding xotosuhusD ut exluding rownEgroup hll ehi 8 entor PHHRAD ut remined undeE roodiles @pigF PfAF I snstitut gtlÁde leontologi @sgAD ere de reer del wesozoiD isol sndustril PQD iEHVPHI dellD pinF P wuseo qeologio qiovnni gpelliniD vi moni TQD sERHIPT folognD stlyF QIH hll ehi pFwF 8 gu eF seÂum xtionldu xigerD ximeyD eÂpulique de xigerY wgwD wuE seo dell o di wonflone ± ezione qeopleontologi @former wuseo leontologio gittdino di wonfloneAD qoriziD stlyY D leontologil snstituteD niversity of iennD eustriF qeologil nd pleontologil notes he olzzo fossil site ours in the western porE tion of the stlin setor of the urst lteu nmed grso ssontino9 in stlinF st tully inludes two sitesD nmed site e nd site f nd proly representing two distintD ut loseD strtigrphi setions @see hll eE hi et lF PHHI nd hll ehi 8 entor PHHR for more informtionAF pieldwork t site f where wgw IIUPH ws found hs een undertken systemtilly sine IWWT y the wuseo leontologio gittdino of wonflone @now leontologil etion of the wuseo dell o di wonfloneAF foth site e nd f hve yielded peulir fossil ssemlge omposed minly of perfetly rtiulted pigF I E votion of the olzzo site @poglinoEedipugliD stlyA nd wellEpreserved osteihthyn fishesF he ihthyofuE where wgw IIUPH ws foundF n inludes the pynodontiform olzzodus orontus oytoEerizD PHIHD the lopiform hynhoderetisD smll lupeomorphsD rre ghnideD inhodus spFDn nguilliformD nd some smllEsized eryiform nthopterygins @quidotti IWVQY hll ehi 8 entor PHHRY oytoEeriz PHIHAF errestrilplntreE mins @prenelopsis spFD frhiphyllum spFD giophyllum spFD fF phenolepisD gunninghmites fF elegnsD some prole ginkoles nd indeterminte ngiospermsA re reltively ommon t site eD nd re muh rrer t site f @hll ehi 8 entor PHHRAF snverterte evidene is represented only y few depod rustensD some rudist ivlve shells nd level with tre fossils @hll ehi 8 entor PHHRAF etrpod remins were found nerly exlusively t site e nd onsist of disrtiulted ones of nonEmrine helonins nd worn tooth of roodyliform @hll ehi 8 entor PHHRAF he fossiliferous setion of site f is only IPPFS m thik nd is omposed of thinly lmintedD light gryE pigF P E hylogeneti reltionships mong the min theropod @eA greenish limestone @hll ehi et lF PHHIY hll nd roodyliform @fA ldes mentioned in the textF ine ehi 8 entor PHHRAF wirofies nlysis suggests the position of the hlttosuhi mong the groodyliE tht deposition ws in restrited lgoonl environment formes is ontroversilD n unresolved polytomy is preE with intertidlepisodes @ndro enturiniD persF ommF sented hereF to pF wF hF FAF wgw IIUPH ws found t the oundE ry etween lyers f nd gD PHH nd IWH mm thik erevitions for vrilesF evD pillengthYgeD rown respetively @see hll ehi 8 entor PHHRD figF PUAF ngleY gfvD rown se lengthY gfD rown se widthY gfD rown foth lyers re omposed of sets of stromtolite lmiE se rtio @gfGgfvAY grD rown heightY grD rown height rtio @grGgfvAY heD distlpildentilesdensity @per mmAY hfD distl ne nd thinD orser storm levelsF he tooth ws deE sl dentiles density @per mmAY hgD distl midErown dentiles denE posited t the se of the lmin fT tht is the lowerE sity @per mmAY wfD mesilsldentilesdensity @per mmAY wgD mesil most storm level of lyer fY other fossils found in fT midErown dentiles density @per mmAF inlude the fish hynhoderetisF his setion is prt of thiker strtigrphi setion mde of pltyD wellE snstitutionl revitionsF pqq fD pultte de qeologie sÎi qeofiziD niversitte fuuresÎtiD fuhrestD omniY wxxD wuE edded limestone ropping out est of the villge of he first reord of notosuhin roodyliform from stly QII olzzoD elonging to the thik pper greteous rE onte pltform sequene of the urstF his sequene is dominted y limestone with rudist ivlves nd is known in literture under the nme of glri di wFte n wihele9 @wrtinis IWTPAF he lithostrtiE grphi unit ontining the olzzo setion is lled lolly glri di eurisin9 @known informlly lso s lri udiste9A nd its ge is onsidered uroninE erly enonin @entor et lF IWWRAF intori et lF @IWWQA referred site e to the lower enonin euse of the presene of the forminifer eordiell oni nd the lge grossoell prtenopei ove the fish levels nd the forminifer wonthrmonti pennini elowF wonthrmonti spF ws found lso t site fY he forE minifers wF penniniD seudoylmmin spheroiE deD eordiell spFD nd ndone smnitiD nd the lge grossoell spF were found in setion immediE tely elow the fossiliferous setion of site fF his sE semlge suggests n erly enonin ge @iFeFD goniE inEntoninA lso for site fD in greement with the strtigrphi frmework of the urst lteu @F entuE riniD persF ommF to pF wF hF FAF he urst lteu is formed minly y rontes generlly supposed to hve een deposited in shllow pigF Q E he western ethys during ntonin @vte greteousA mrine environment t tropilltitudesnd dominted timesD sed on ierli et lF @IWWQAD with lotion of the y rudist reefs9 during vte greteous timesF pollowE urst lteu @steriskA on the edritiEhinri groE ing reent pleogeogrphi reonstrution of western nte ltformF hrk gry olor represents deep se on ethys @seeD for exmpleD ierli et lF IWWQY gmoin et oeni rustD light gry epiontinentl se nd mrine lF IWWQY hilip et lF PHHHAD the urst lteu ws situE sins on ontinentlrustD white emergent lndDnd rikEpttern identifies the ronte pltformsF ted t the northern point of wideD isolted introeE ni ronte pltform of fhmin type @edritiEhiE nri ronte pltformA during enonin times @pigF hesriptionF wgw IIUPH is shed rownY the QAF imersion events ffeted lolly or extensively this root ws nerly ompletely reEdsored nd the sl pltform severl times during greteous timesY the side of the rown ppers onve in liolingul view most importnt ourred during the lte gmpninE @pigF ReEfAF foth mesilnd distlutting mrgins er wstrihtin when reltively lrge islnd formed @edriti sslndX ytoniÏr PHHUY hll ehi PHHVD serrted rineF sts ev is IPFPS mm nd gr is IIFQTY PHHWAF he emergent prts of the ronte pltform the gfv is SFVP mmF he gf is not signifitive eE elonged to n rhipelgo of islnds pled etween use the lil side is dmged nd the lingul side is the xorthern
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