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ivist stlin di leontologi e trtigrfi volume IIU noF P ppF QHWEQPI tuly PHII ri ps igyh yp e xyy grsex gygyhvspyw pyw sev I P pefsy wF hevve iggrse 8 exhie ge eeivedX heemer PWD PHIHY eptedX epril TD PHII uey wordsX iphodont toothD groodyliformesD xotosuhiD sriedF he tooth is stored in the olletions of the hortodonD erripesuhus wegeneriD pper greteousD olzzo fossil leontologil etion of the wuseo dell o di siteD urst lteuF wonflone with the speimen numer wgw IIUPH @the originlfieldnumer ws WUGIIVAF estrtF e serrted tooth from the goniinEntonin @ pper greteousA olzzo fossil site @urstD xi stlyA is the first he im of this work is to desrie the speimen reord of notosuhin roodyliform from stlyF elthough it shres in detilnd disuss its ffinityF istlishingthe txoE synpomorphies with teeth referred to the iuropen genus hortodon nomi sttus of wgw IIUPH is neessryD s there hve nd with the qondwnn genus erripesuhusD it is distint in the een suggestions tht the tooth ould in ft elong to unusulomintion of feturesD suggesting the presene of yet unE reported notosuhin txon in the edritiEhinri gronte ltE theropod dinosur @F gurrieD persF ommFAF es dinosur form loted in the ethys etween the efrorin ontinent nd evidene is rre in stlyD it would e worth of noteF sn the xorth iuropen lndmss during vte greteous timesF xotosuE the mentimeD the tooth hs een reently mentioned in hins were typilly terrestril roodyliformsD supporting the preE literture s evidene of the genus hortodon in the sene of emergent res on the ronte pltformF pper greteous of stly @helfino PHHIY i PHHVAD issuntoF n dente munito di rene dentiolte rinvenuto nel with potentil pleogeogrphi implitionsD despite the sito fossilifero di olzzo @grsoD stli nordorientleA e dtile l ft tht its identifition in previous ppers ws lerly goniinoEntonino @gretio uperioreA eÁl prim testimoninz tenttive nd preliminryF di un oodrilliforme notosuo in stliF eene ondivid sinpoE morfie on denti ttriuiti lgenere europeo hortodon e on ilgondE wnino erripesuhusD esso si distingue per l9inusule ominzione di rtteriD he suggerise l presenz di un speie di notosuo non erminology nd methods nor identifit nell ittform gronti edritioEhinriD lolizzt tr il ontinente efrorio e quello xord iuropeoF s he odontologil terminology nd vriles used in this pper notosuhi erno oodrilliformi prettmente terrestri e questo eÁindiE re those of mith 8 hodson @PHHQA nd mith et lF @PHHSAF he terms tivo dell presenz di ree emerse sull pittform rontiF elle9 nd ude9 re used in the sense of eler @IWWPAF rmeters of hortodon teeth from eustriD omni nd rungry re sed on mesurements tken from photogrphs pulished in fuffetut @IWUWAD wrtin et lF @PHHTAD nd ourtesy of wrton iD respetivelyF sntrodution he verostrn theropods re onsidered ording to izurr huring the IWWU field work in pper greteous @PHHTA s those elonging to the lest inlusive lde inluding neoerE tosurs nd tetnurns @pigF PeAF he roodyliform phylogeneti olzzo fossil site @poglinoEedipugli wuniiplityD frmework nd txonomy followed herein is tht of ereno 8 vrsson qorizi rovineD xi stlyY pigF IAD n unusul serrted @PHHWA nd ereno et lF @PHHIAF eordinglyD the groodyliformes re tooth ws found @igo IWWUAF he speimen ws preliE the lest inlusive lde inluding protosuhids nd rownEgroup roE minrily nd tenttively referred to the ziphodont roE odiles @groodyliAD the wesoeuroodyli is the most inlusive roE odyliform hortodon in some overviews of the olzE odyliform lde exluding protosuhidsD nd the xotosuhi is the zo fossil ssemlge @eFgFD hll ehi et lF PHHIY most inlusive lde inluding xotosuhusD ut exluding rownEgroup hll ehi 8 entor PHHRAD ut remined undeE roodiles @pigF PfAF I snstitut gtlÁde leontologi @sgAD ere de reer del wesozoiD isol sndustril PQD iEHVPHI dellD pinF P wuseo qeologio qiovnni gpelliniD vi moni TQD sERHIPT folognD stlyF QIH hll ehi pFwF 8 gu eF seÂum xtionldu xigerD ximeyD eÂpulique de xigerY wgwD wuE seo dell o di wonflone ± ezione qeopleontologi @former wuseo leontologio gittdino di wonfloneAD qoriziD stlyY D leontologil snstituteD niversity of iennD eustriF qeologil nd pleontologil notes he olzzo fossil site ours in the western porE tion of the stlin setor of the urst lteu nmed grso ssontino9 in stlinF st tully inludes two sitesD nmed site e nd site f nd proly representing two distintD ut loseD strtigrphi setions @see hll eE hi et lF PHHI nd hll ehi 8 entor PHHR for more informtionAF pieldwork t site f where wgw IIUPH ws found hs een undertken systemtilly sine IWWT y the wuseo leontologio gittdino of wonflone @now leontologil etion of the wuseo dell o di wonfloneAF foth site e nd f hve yielded peulir fossil ssemlge omposed minly of perfetly rtiulted pigF I E votion of the olzzo site @poglinoEedipugliD stlyA nd wellEpreserved osteihthyn fishesF he ihthyofuE where wgw IIUPH ws foundF n inludes the pynodontiform olzzodus orontus oytoEerizD PHIHD the lopiform hynhoderetisD smll lupeomorphsD rre ghnideD inhodus spFDn nguilliformD nd some smllEsized eryiform nthopterygins @quidotti IWVQY hll ehi 8 entor PHHRY oytoEeriz PHIHAF errestrilplntreE mins @prenelopsis spFD frhiphyllum spFD giophyllum spFD fF phenolepisD gunninghmites fF elegnsD some prole ginkoles nd indeterminte ngiospermsA re reltively ommon t site eD nd re muh rrer t site f @hll ehi 8 entor PHHRAF snverterte evidene is represented only y few depod rustensD some rudist ivlve shells nd level with tre fossils @hll ehi 8 entor PHHRAF etrpod remins were found nerly exlusively t site e nd onsist of disrtiulted ones of nonEmrine helonins nd worn tooth of roodyliform @hll ehi 8 entor PHHRAF he fossiliferous setion of site f is only IPPFS m thik nd is omposed of thinly lmintedD light gryE pigF P E hylogeneti reltionships mong the min theropod @eA greenish limestone @hll ehi et lF PHHIY hll nd roodyliform @fA ldes mentioned in the textF ine ehi 8 entor PHHRAF wirofies nlysis suggests the position of the hlttosuhi mong the groodyliE tht deposition ws in restrited lgoonl environment formes is ontroversilD n unresolved polytomy is preE with intertidlepisodes @ndro enturiniD persF ommF sented hereF to pF wF hF FAF wgw IIUPH ws found t the oundE ry etween lyers f nd gD PHH nd IWH mm thik erevitions for vrilesF evD pillengthYgeD rown respetively @see hll ehi 8 entor PHHRD figF PUAF ngleY gfvD rown se lengthY gfD rown se widthY gfD rown foth lyers re omposed of sets of stromtolite lmiE se rtio @gfGgfvAY grD rown heightY grD rown height rtio @grGgfvAY heD distlpildentilesdensity @per mmAY hfD distl ne nd thinD orser storm levelsF he tooth ws deE sl dentiles density @per mmAY hgD distl midErown dentiles denE posited t the se of the lmin fT tht is the lowerE sity @per mmAY wfD mesilsldentilesdensity @per mmAY wgD mesil most storm level of lyer fY other fossils found in fT midErown dentiles density @per mmAF inlude the fish hynhoderetisF his setion is prt of thiker strtigrphi setion mde of pltyD wellE snstitutionl revitionsF pqq fD pultte de qeologie sÎi qeofiziD niversitte fuuresÎtiD fuhrestD omniY wxxD wuE edded limestone ropping out est of the villge of he first reord of notosuhin roodyliform from stly QII olzzoD elonging to the thik pper greteous rE onte pltform sequene of the urstF his sequene is dominted y limestone with rudist ivlves nd is known in literture under the nme of glri di wFte n wihele9 @wrtinis IWTPAF he lithostrtiE grphi unit ontining the olzzo setion is lled lolly glri di eurisin9 @known informlly lso s lri udiste9A nd its ge is onsidered uroninE erly enonin @entor et lF IWWRAF intori et lF @IWWQA referred site e to the lower enonin euse of the presene of the forminifer eordiell oni nd the lge grossoell prtenopei ove the fish levels nd the forminifer wonthrmonti pennini elowF wonthrmonti spF ws found lso t site fY he forE minifers wF penniniD seudoylmmin spheroiE deD eordiell spFD nd ndone smnitiD nd the lge grossoell spF were found in setion immediE tely elow the fossiliferous setion of site fF his sE semlge suggests n erly enonin ge @iFeFD goniE inEntoninA lso for site fD in greement with the strtigrphi frmework of the urst lteu @F entuE riniD persF ommF to pF wF hF FAF he urst lteu is formed minly y rontes generlly supposed to hve een deposited in shllow pigF Q E he western ethys during ntonin @vte greteousA mrine environment t tropilltitudesnd dominted timesD sed on ierli et lF @IWWQAD with lotion of the y rudist reefs9 during vte greteous timesF pollowE urst lteu @steriskA on the edritiEhinri groE ing reent pleogeogrphi reonstrution of western nte ltformF hrk gry olor represents deep se on ethys @seeD for exmpleD ierli et lF IWWQY gmoin et oeni rustD light gry epiontinentl se nd mrine lF IWWQY hilip et lF PHHHAD the urst lteu ws situE sins on ontinentlrustD white emergent lndDnd rikEpttern identifies the ronte pltformsF ted t the northern point of wideD isolted introeE ni ronte pltform of fhmin type @edritiEhiE nri ronte pltformA during enonin times @pigF hesriptionF wgw IIUPH is shed rownY the QAF imersion events ffeted lolly or extensively this root ws nerly ompletely reEdsored nd the sl pltform severl times during greteous timesY the side of the rown ppers onve in liolingul view most importnt ourred during the lte gmpninE @pigF ReEfAF foth mesilnd distlutting mrgins er wstrihtin when reltively lrge islnd formed @edriti sslndX ytoniÏr PHHUY hll ehi PHHVD serrted rineF sts ev is IPFPS mm nd gr is IIFQTY PHHWAF he emergent prts of the ronte pltform the gfv is SFVP mmF he gf is not signifitive eE elonged to n rhipelgo of islnds pled etween use the lil side is dmged nd the lingul side is the xorthern
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  • New Transitional Fossil from Late Jurassic of Chile Sheds Light on the Origin of Modern Crocodiles Fernando E

    New Transitional Fossil from Late Jurassic of Chile Sheds Light on the Origin of Modern Crocodiles Fernando E

    www.nature.com/scientificreports OPEN New transitional fossil from late Jurassic of Chile sheds light on the origin of modern crocodiles Fernando E. Novas1,2, Federico L. Agnolin1,2,3*, Gabriel L. Lio1, Sebastián Rozadilla1,2, Manuel Suárez4, Rita de la Cruz5, Ismar de Souza Carvalho6,8, David Rubilar‑Rogers7 & Marcelo P. Isasi1,2 We describe the basal mesoeucrocodylian Burkesuchus mallingrandensis nov. gen. et sp., from the Upper Jurassic (Tithonian) Toqui Formation of southern Chile. The new taxon constitutes one of the few records of non‑pelagic Jurassic crocodyliforms for the entire South American continent. Burkesuchus was found on the same levels that yielded titanosauriform and diplodocoid sauropods and the herbivore theropod Chilesaurus diegosuarezi, thus expanding the taxonomic composition of currently poorly known Jurassic reptilian faunas from Patagonia. Burkesuchus was a small‑sized crocodyliform (estimated length 70 cm), with a cranium that is dorsoventrally depressed and transversely wide posteriorly and distinguished by a posteroventrally fexed wing‑like squamosal. A well‑defned longitudinal groove runs along the lateral edge of the postorbital and squamosal, indicative of a anteroposteriorly extensive upper earlid. Phylogenetic analysis supports Burkesuchus as a basal member of Mesoeucrocodylia. This new discovery expands the meagre record of non‑pelagic representatives of this clade for the Jurassic Period, and together with Batrachomimus, from Upper Jurassic beds of Brazil, supports the idea that South America represented a cradle for the evolution of derived crocodyliforms during the Late Jurassic. In contrast to the Cretaceous Period and Cenozoic Era, crocodyliforms from the Jurassic Period are predomi- nantly known from marine forms (e.g., thalattosuchians)1.
  • A New Sebecid Mesoeucrocodylian from the Rio Loro Formation (Palaeocene) of North-Western Argentina

    A New Sebecid Mesoeucrocodylian from the Rio Loro Formation (Palaeocene) of North-Western Argentina

    Zoological Journal of the Linnean Society, 2011, 163, S7–S36. With 17 figures A new sebecid mesoeucrocodylian from the Rio Loro Formation (Palaeocene) of north-western Argentina DIEGO POL1* and JAIME E. POWELL2 1CONICET, Museo Paleontológico Egidio Feruglio, Ave. Fontana 140, Trelew CP 9100, Chubut, Argentina 2CONICET, Instituto Miguel Lillo, Miguel Lillio 205, San Miguel de Tucumán CP 4000, Tucumán, Argentina Received 2 March 2010; revised 10 October 2010; accepted for publication 19 October 2010 A new basal mesoeucrocodylian, Lorosuchus nodosus gen. et sp. nov., from the Palaeocene of north-western Argentina is presented here. The new taxon is diagnosed by the presence of external nares facing dorsally, completely septated, and retracted posteriorly, elevated narial rim, sagittal crest on the anteromedial margins of both premaxillae, dorsal crests and protuberances on the anterior half of the rostrum, and anterior-most three maxillary teeth with emarginated alveolar margins. This taxon is most parsimoniously interpreted as a bizarre and highly autapomorphic basal member of Sebecidae, a position supported (amongst other characters) by the elongated bar-like pterygoid flanges, a laterally opened notch and fossa in the pterygoids located posterolaterally to the choanal opening (parachoanal fossa), base of postorbital process of jugal directed dorsally, and palatal parts of the premaxillae meeting posteriorly to the incisive foramen. Lorosuchus nodosus also shares with basal neosuchians a suite of derived characters that are interpreted as convergently acquired and possibly related to their semiaquatic lifestyle. The phylogenetic analysis used for testing the phylogenetic affinities of L. nodosus depicts Sebecidae as the sister group of Baurusuchidae, forming a monophyletic Sebecosuchia that is deeply nested within Notosuchia.
  • Abstracts of the 6Th Meeting of the EAVP 2008

    Abstracts of the 6Th Meeting of the EAVP 2008

    6TH MEETING OF THE EUROPEAN ASSOCIATION OF VERTEBRATE PALAEONTOLOGISTS 30TH JUNE – 5TH JULY 2008 SPIŠSKÁ NOVÁ VES, SLOVAK REPUBLIC VOLUME OF ABSTRACTS 6TH MEETING OF THE EUROPEAN ASSOCIATION OF VERTEBRATE PALAEONTOLOGISTS SPIŠSKÁ NOVÁ VES, SLOVAK REPUBLIC 30TH JUNE – 5TH JULY 2008 VOLUME OF ABSTRACTS ORGANIZED BY THE MUSEUM OF SPIŠ IN SPIŠSKÁ NOVÁ VES SPIŠSKÁ NOVÁ VES, 2008 SUPPORTED BY DEPARTMENT OF GEOLOGY NATIONAL PARK SLOVAK TOWN UNION OF MUSEUMS AND PALAEONTOLOGY SLOVAK PARADISE GEOLOGICAL SOCIETY SPIŠSKÁ NOVÁ VES IN SLOVAKIA 6TH MEETING OF THE EUROPEAN ASSOCIATION OF VERTEBRATE PALAEONTOLOGISTS, SPIŠSKÁ NOVÁ VES 2008 EDITOR: ZUZANA KREMPASKÁ SCIENTIFIC SUPERVISOR: MARTIN SABOL THE ORGANIZING COMMITTEE: ZUZANA KREMPASKÁ RUŽENA GREGOROVÁ MARTIN SABOL The authors are responsible for the linguistic rendition of the papers. ACKNOWLEDGEMENT: The publication of this abstract volume was partly supported by the Ministry of Culture Slovak Republic Cover Photo: Medvedia jaskyňa (Bear Cave) by L. Cibula © MUSEUM OF SPIŠ, SPIŠSKÁ NOVÁ VES, SLOVAK REPUBLIC, 2008 ISBN: 978-80-85173-03-1 4 6TH MEETING OF THE EUROPEAN ASSOCIATION OF VERTEBRATE PALAEONTOLOGISTS, SPIŠSKÁ NOVÁ VES 2008 CONTENTS INTRODUCTION ...................................................................................................................................................................... 7 ABSTRACTS BARYSHNIKOV G.: Taxonomical Diversity of Pleistocene Bears in Northern Eurasia ..........................................................9 BAYGUSHEVA V. S. & TITOV V. V.:
  • Chubut Province, Argentina) and Its Phylogenetic Position Within Basal Mesoeucrocodylia

    Chubut Province, Argentina) and Its Phylogenetic Position Within Basal Mesoeucrocodylia

    Cretaceous Research 30 (2009) 1376–1386 Contents lists available at ScienceDirect Cretaceous Research journal homepage: www.elsevier.com/locate/CretRes The first crocodyliform from the Chubut Group (Chubut Province, Argentina) and its phylogenetic position within basal Mesoeucrocodylia Juan Martı´n Leardi a,c,*, Diego Pol b,c a Laboratorio de Paleontologı´a de Vertebrados, Departamento de Ciencias Geolo´gicas, Facultad de Ciencias Exactas y Naturales, Pabello´n II, Universidad de Buenos Aires, Intendente Gu¨iraldes 2160, Ciudad Universitaria (C1428EGA), Buenos Aires, Argentina b Museo Paleontolo´gico Egidio Feruglio, Avenue Fontana 140 (9100), Trelew, Chubut, Argentina c CONICET- Consejo Nacional de Investigaciones Cientı´ficas y Te´cnicas, Avenue Rivadavia 1917, Buenos Aires, Argentina article info abstract Article history: A new crocodyliform specimen is presented here found in the Cerro Castan˜o Member of the Cerro Received 27 February 2009 Barcino Formation (Chubut Group). The material consists of cranial and postcranial remains that Accepted in revised form 4 August 2009 represent a new taxon that has strong affinities with Peirosauridae, but also shares derived features Available online 11 August 2009 present in Araripesuchus. The phylogenetic relationships of this new taxon were tested through a cladistic analysis depicting it as a member of the Peirosauridae. The inclusion of Barcinosuchus within this clade of Keywords: basal mesoeucrocodylians is supported by the presence of hypapophyses up to the third or fourth dorsal Crocodyliformes vertebrae, anterolateral facing edge on postorbital, quadrate dorsal surface divided in two planes by Peirosauridae Lower Cretaceous a ridge; mandibular symphysis tapering anterirorly in ventral view, lateral surface of dentary convex Central Patagonia anterior to mandibular fenestra, distal body of quadrate well developed, anteroposteriorly thin and Argentina lateromedially broad.