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Home , Acronychia pedunculata, Afzelia xylocarpa, Albizia lucidior, Ancistrocladus tectorius, Anisoptera costata, Aporosa octandra

Labelled and other important and in Cát Tiên National Park (Checklist and higher of trees, lianas and larger ) .2 update: 18 August 2021

The purpose of this checklist is to provide an inventory, with higher taxonomy, of a majority of labelled specimen trees and lianas found on the main roads, Botanic, Crocodile-lake trails and the Headquarters area. The Park's 2021 labelling initiative has enormously enhanced the scientific and educational value of the commonly-used trails. The number of names – more than 150 of trees and lianas (woody vines) - can be bewildering to non-specialists: the word "diversity" is commonly used, but with it comes complexity. How best to dissect the problem? Botanists tend to place species in families (Họ), but there are still some 45 names to learn here; to make this more manageable, families can be grouped into (say a dozen) orders (Bộ), easier to cope-with. Up-to-date species names will help visitors obtain further information online – but please be careful to spell them correctly! The original, official Park species list was prepared in 2002 (by Prof. Phạm Hoàng Hộ, author of the definitive work Cây Cỏ Việt Nam). However since then, taxonomy has been revised substantially by the international Angiosperm Phylogeny Group (APG), with whole plant families promoted, demoted or moved to different orders. Furthermore, genera and species have been – and continue to be - revised in light of scientific evidence; this also can be confusing for non-botanists. Two important families, the and have undergone substantial revision, sometimes with reorganisation into new subfamily names (indicated in blue). The is a very large and diverse Order, heavily revised under the APG. Some trees, especially with older labels, may contain original names that have been changed recently (these are highlighted in yellow). Identifying is of ecological interest: "gibbon food" refers to a Park study of wild populations in 2014-151.

Order, species Family (subfamily etc.) VN: loài / Chi Notes bộ Thạch nam (bộ đỗ quyên) collinsiae Craib thị đen lá to - Maa: ddôin Diospyros lanceifolia Roxb. săng đen significant gibbon food Diospyros montana Roxb. thị da đen (>6%): esp. "Diospyros Diospyros variegata Kurz nhọ nồi cf. sumatrana" in paper Careya arborea Roxb. vừng (vừng xoan) Maa: (plai) tự (fruit) acutangula (.) Gaertn. lộc vừng (chiếc) Barringtonia macrostachya (Jack.) Kurz. chiếc tam lang (chùm to) Madhuca cochinchinensis (Pierre ex Dubard) HJ Lam sến nam bộ obovatum (Griff.) Engl. ‡ xây dao, cốt; Maa: kiöt Camellia (7 spp. incl. . cattienensis) ‡ Theaceae Chi trà tea genus

Fabales bộ Đậu Lasiobema scandens (L.) de Wit var. Fabaceae subfamily dây móng bò, móng bò "monkey ladder" : horsfieldii (Miq.) de Wit Cercidoideae leo spp. are usu. syn. Bauhinia scandens L. (=Leguminosae) ornamental trees. Entada cf. rheedii Spreng. (syn. . pursaetha) bàm bàm; Maa: klaac box bean: a large liana Mimosa diplotricra C. Wight. Ex Sauvalle ‡ clade/tribe trinh nữ móc hooked liana . pudica L.; M. pigra L ‡ (basal genera) trinh nữ all invasive alien spp. dasyrrhachis (Miq.) Kurz. (NB: Mimosoideae was lim vàng Xylia xylocarpa (Roxb.) Taubert. previously a subfamily) cẩm xe Adenanthera pavonina L. Accacia clade trạch quạch Albizia spp. (approx.. 8) hợp hoan v.v. Albizia chinensis (Osbeck) Merr. sống rắn Albizia lucidior (Stend.) L. Niels. bản xe (đái bò) Cassia javanica L. - syn. Cassia agnes (de Wit) tribe: Cassieae muồng đào, muồng "peach blossom Brenan: now subsp. C.. agnes (de Wit) .Larsen ‡ , muồng hương shower" Senna siamea (Lam.) Irwin et Barneby muồng đen (~ xiêm) Siamese cassia Senna alata (L.) Roxb. (invasive shrub sp.) ‡ muồng trâu candle bush

1 Thanh Hai Bach, Jin Chen, Minh Hoang, Beng KC, Van Nguyen (2017) Feeding behaviour and activity budget of the southern yellow-cheeked crested gibbons (Nomascus gabriellae) in a lowland tropical forest. Am. J Primatol. 9999:e22667. ‡ labelled specimens not seen © www.namcattien.org 1 xylocarpa (Kusz.) Craib. gõ đỏ also gibbon food siamensis Teysm. Ex Miq. gõ (gụ) mật lanceolaria var. lakhonensis vảy ốc, trắc múi giáo good for replanting on (Gagnep.) Niyomdham & P..Hô (=Papilionoideae) cẩm lai; Maa: lôo degraded land. Gamble ex Prain tribe: (cẩm lai vú , cẩm lai Highly valuable syn. D. bariaensis, D. mammosa bông, trắc Ðồng nai) - Dalbergia nigrescens Kurz. trắc đen, cẩm lai đen, endangered syn. D. . var. anomala (Pierre) Niyomdham cẩm lai giấy Ormosia sumatrana (Miq.) Prain. ràng ràng xanh macrocarpus Kurz giáng, dáng hương Xanthophyllum flavescens Roxb. Polygalaceae (prev. săng ớt cao (vàng syn. Xanthophyllum excelsum Bl. Xanthophyllaceae) chanh)

Gentianales bộ Long đởm Alstonia scholaris (L.) .Br. (hoa) sữa, mò cua Maa: blaang sêr Kibatalia macrophylla (Pierre ex Hua) Woodson thừng mực tiêu croc. lake trail Wrightia pubescens R.Br. lồng mức lông Cyrtophyllum fragrans (Roxb.) DC. trai large trees on Heaven syn. fragrans Roxb. (previously in Loganiaceae) rapids road Fagraea auriculata Jack ‡ trai tai food plant (<1%) for Fagraea ceilanica Thunb ‡ trai tích lan (lậu bình) gibbons Strychnos (4 spp.) ‡ Loganiaceae (họ Mã tiền) mã tiền primate food Adina trichotoma (Zoll. & Moritzi) Benth. & .. vàng vé (gáo lông) Lag. & bot. trails syn. Metadina trichotoma (Zoll. & Moritzi) Bakh.f. Haldina cordifolia (Roxb) Ridsd. gáo trò diversifolia (.Don) Havil. mao thư (cà dăm) Nauclea orientalis (L.) L. gáo vàng Leichhardt tree … Neolamarckia cadamba (Roxb.) Bosser. gáo nước, gáo trắng … often confused Ridsdalea (syn. Rothmannia) eucodon (K Schum.) JT Pereira ‡ găng cơm (găng cao) gibbon food (4.5 %)

Lamiales (also Acanthaceae) bộ Hoa môi notable for stipulata (Wall.) Seem.ex Schum. thiết đinh lá bẹ Oroxylum indicum (L.) Vent. núc nác (sò đo) Stereospermum cylindricum Piere ex Dop. ‡ quao trụ Stereospermum tetragonum D.C. quao (núi) syn. Stereospermum colais (Dillw.) Mabb. Gmelina (5 spp.) ‡ Lamiaceae tu hú (5 loài) Vitex ajugaeflora Dop. Note: genera moved bình linh ngệ VU_lnerable sp. Vitex pinnata L. [V. p. f. ptilota (Dop) Moldenke] from the Verbenaceae bình linh cánh Vitex tripinnata (Lour.) Merr. mắt cáo Tectona grandis L.f. ‡ tếch, giá tỵ teak (usu. planted)

Malpighiales bộ Sơ ri Note: This very large, diverse Order has been especially heavily revised by the APG: genera such as have been moved here from the to form the family and the "Flacourtiaceae", long considered a "wastebasket taxon", is now obsolete. Several genera have been moved from the Euphorbiaceae to the . Hydnocarpus castaneus Hook.f. & Thomson Achariaceae đại phong tử (lọ nồi) syn.H. anthelminthicus Pierre ex Laness. (Flacourtiaceae Họ Mùng quân đã lỗi obsolete) thời Chrysobalanaceae họ Cám moved to this order Parinari anamensis Hance. (previously Rosaceae) cám minor gibbon food Calophyllum spp. cồng Maa: kroec may include C. calaba var. bracteatum (Wight) P.F.Stevens cồng tía C. dongnaiense siamensis (Miq.) T.Anderson (no longer ) trau tráu significant (>1%) syn. Ochrocarpos siamensis (Miq.) T.Anderson gibbon food ferrea Pierre Clusiaceae rõi mật genus Garcinia hanburyi Hook.f. vàng nghệ (fruit: G. mangostana) Garcinia harmandii Pierre. bứa mọi Garcinia oliveri Pierre ‡ bứa núi gibbon food (>2%) ‡ labelled specimens not seen © www.namcattien.org 2 Garcinia vilersiana Pierre. vàng nhựa (bứa lá to) Garcinia xanthochymus Hook.f. bứa mủ vàng Croton delpyi Gagnep. Euphorbiaceae bã đậu lá xanh croc. lake trail Macaranga spp. (5 recorded) ‡ Note: several genera mã rạng, long màng secondary coloniser Mallotus philippensis (Lam.) Müll.Arg. have been moved Cánh kiến (Ba chia) herbal properties Suregada multiflora (Juss) H. Baill. from here to the cổ ngỗng, mân mây syn. Gelonium multiflorum A.Juss. Phyllanthaceae (kén) Cratoxylum cochinchinense (Lour) Bl. Hypericaceae thành ngạnh nam NB "Cratoxilon" is an Cratoxylum formosum (Jack.) Dyer. thành ngạnh (dẹp) orthographic variant Irvingia malayana Oliv.ex Benn. Irvingiaceae cầy (kơ nia); Maa: niör prev. Ixonanthaceae Gaertn. Phyllanthaceae chòi mòi (chua mòi) croc. lake shrubs ficifolia Baill. (Aporusa Blume is .V.) Note: many genera in tai nghé, ngăm lông dày var. octandra this family were thầu tấu syn Aporusa dioica (Roxb.) Müll.Arg. previously placed in Baccaurea ramiflora Lour. the Euphorbiaceae dâu da rừng, dâu ta "Burmese grape": imp. Cleistanthus oblongifolius (Roxb.) Müll.Arg. cọc rào gibbon food (>6%) syn. Cleistanthus myrianthus (Hassk.) Kurz Leptopus australis (Zoll. & Moritzi) Pojark. bụi trườn Phyllanthus reticulatus Poir. phèn đen Carallia brachiata (Lour.) Merr. Rhizophoraceae xăng mã nguyên Significant gibbon food Carallia suffruticosa Ridl. xăng mã răng (>2%)

Magnoliids: bộ Mộc lan 'basal' clade Cananga brandisiana (Pierre) Saff. công chúa lá rộng ylang ylang genus syn. C. latifolia (Hook.f.& Thoms) Fin Huberantha cerasoides (Roxb.) Chaowasku nhọc vàng, quần đầu See trees: syn. Polyalthia cerasoides (Roxb.) Benth. & Hook trái tròn (CCVN) has been reclassified Monoon harmandii (Pierre) .Xue & RMK Saunders nhọc lá lớn (quần đầu See trees: Polyalthia syn. Polyalthia harmandii (Pierre) Fin. & Gagn. Harmand) has been reclassified Monoon thorelii (Pierre) B.Xue & R.M.K.Saunders nhọc lá nhỏ, lèo heo, syn. Polyalthia thorelii (Pierre) Fin. & Gagn. quần đầu Thorel Polyalthiopsis floribunda (Jovet-Ast) Chaowasku nhọc nhiều hoa native range is only syn. Polyalthia floribunda Jovet-Ast Central & . Sageraea elliptica (A.D.C) Hook. & Thoms. săng mây Spondias pinnata (L.f.) Kurz cóc rừng Xylopia vielana Pierre ex Fin giền đỏ Knema lenta Warb. Myristicaceae máu chó thấu kính gibbon food: <1% Knema globularia (Lamk.) Warb. máu chó cầu (~ lá nhỏ)

Magnoliids: bộ Nguyệt quế iners Reinw. ex Blume quế rừng (quế giả) Dehaasia kurzii King. cà đuối Kurz cambodiana Lec. bời lời planted understory tree syn. Litsea pierrei Lec.; L. pierrei var. grandifolia bời lời vàng lá to Litsea monopetala (Roxb.) Pers. bời lời lá tròn Machilus cochinchinensis Lecomte kháo vàng Phoebe pallida Nees. kháo nhớt (cha)

Malvales bộ Cẩm quỳ, bộ Bông guiso (Blco.) Bl. chai Shorea thorelii Pierre. chai Thorel alatus Roxb. dầu nước (dầu rái) Maa: kloong wang Dipterocarpus turbinatus C.F.Gaertn. dầu con rái đỏ (dầu lá bóng) Hopea odorata Roxb. sao đen Maa: nggir Lag. trail good specs. costata Kortn. vên vên tree near Park ferry Vatica odorata (Griff.) Sym. subsp. odorata (làu) táu trắng Malvaceae: note - revised with subfamilies that were previously families ceiba L. subfamily gạo rừng; Maa: blaang (was in family Bombax anceps Pierre. ‡ Bombacoideae gạo hoa đỏ, gạo 2 mặt Bombacaceae)

‡ labelled specimens not seen © www.namcattien.org 3 auriculata (H, Baill.) Craib. bồ an (lá tai); Maa: yyar common large bush Colona evecta (Pierre.) Burret (was listed as ) chàm ron; Maa: vlal Vietnam endemic tree Microcos tomentosa Sm. cò ke (đầu lõm) syn. Grewia paniculata Roxb. ex DC. Microcos laurifolia (Hook.f. ex Mast.) Burret Sếu lá tre Croc. lake trail syn. Grewia laurifolia Hook.f. ex Mast. Grewia hirsuta Vahl. cò ke lông gibbon food (>2%) syn. G. tomentosa Roxb. ex Wight & Arn. Pterospermum diversifolium Blume Dombeyoideae lòng mang lá lớn (lòng Pterospermum grewiifolium Pierre mang Đa dạng) (P. grewiaefolium in PV Hộ is an orth.var.) lòng mang lá nhỏ Pterospermum lanceifolium Roxb. ex DC. lòng mang tía syn. Pterospermadendron syn: P. jackianum (Wall. ex Mast.) Kuntze; (lòng mang Sài gòn) saigonense (Pierre) Kuntze Pterospermum megalocarpum Tardieu lòng mang quả lớn croc. lake trail macrophyllus Roxb. ex Hornem. bụp lá to 2 common spp. fallen Hibiscus mesnyi Pierre. ex Laness. ‡ (Malvaceae sensu stricto) bụp Mesny flowers Dec. - Feb. Pterocymbium dongnaiense Pierre ‡ dực nang Ðồng nai Previously family macropodum (Miq.) Beumée ex K.Heyne lười ươi high value ; good Maa: ryôom specimens on Croc. Trail Sterculia cochinchinensis Pierre. trôm nam bộ Sterculia foetida L. trôm hôi (type species) Sterculia hypochroa Pierre trôm quạt Sterculia pexa Pierre trôm hoe

Myrtales bộ Đào kim nương Combretum Loefl. (4 spp. recorded) Combretaceae trậm bầu, chưn bầu African sp. labelled Terminalia calamansanai (Bl.) Rolfe. chiêu liêu nước Terminalia phillyreifolia (Van Heurck & Müll.Arg.) Gere & Boatwr. râm (chò nhai) syn. Anogeissus acuminata (DC.) Guill. Ferr. Terminalia nigrovenulosa Pierre chiêu liêu nghệ syn. Terminalia triptera Stapf. calyculata Kurz. bằng lăng ổi 6 calyx lobes () Lagerstroemia floribunda Jack Bằng lăng dị dực syn. Lagerstroemia anisoptera Koehne Maa: rpaa Lagerstroemia ovalifolia Teijsm. & Binn. cau Maa: si crộộ bằng lăng xoan 7–9 sepals; sig. gibbon Lagerstroemia speciosa (L.) Pers. bằng lăng nước food (>7%) Syzygium cumini (L.) Skeels Myrtaceae trâm mốc Java plum, black plum Syzygium jambos (L.) Alston. mận rừng domestic - wax apple Syzygium ripicola (Craib) Merr. & L.M.Perry trâm suối, trâm nước & wild relatives of wax syn. S. cochinchinense (Gagnep.) Merr. & L.M.Perry (trâm nam bộ) apples (quả roi) Syzygium tramnion (Gagn.) Merr. & Perry trâm rim Syzygium zeylanicum (L.) DC. trâm vỏ đỏ Melastoma 5 spp. recorded ‡ họ & chi mua / muôi bushes, damp areas edule Roxb. sầm, (sầm bùi)

Rosales bộ Hoa hồng Streblus ilicifolius (Vidal) Corner duối ô rô understorey bushes Streblus taxoides (Roth) Kurz duối gai - prickly! lacucha Buch.-Ham. chay Jackfruit genus: gibbon syn. Artocarpus lakoocha Roxb. food Artocarpus rigidus Blume mít nài (da xốp) monkey jack subgenus Ficus: F. pandurata Hance sung tì bà croc. lake, Ben Cu subgenus Sycomorus: Ficus racemosa L. sung (cluster fig) cluster fig tree: gibbon Ficus fistulosa Reinw. ex Blume sung giòn food (>1%) Ficus hispida L.f. var. hispida ngái (hairy fig) Ficus subgenus Urostigma (many spp. ‡) incl.: đa (~ tía v.v.) - banyans Hemiepiphytes – see F. altissima, F. costata, F. kurzii, F. sumatrana Maa: jrii tree section: major Ficus benjamina L. Si (Xanh) gibbon food (>20%)

‡ labelled specimens not seen © www.namcattien.org 4 bọ Bồ hòn Mangifera reba Pierre. quéo (~ xoài rừng) Vietnam endemic spp. Mangifera dongnaiensis Pierre. xoài Ðồng nai in the mango genus Swintonia sp. ‡ xuân tôn; Maa: riol Heaven rapids specimen dao (Blanco) Merr. & Rolfe ‡ long cóc xoài Sig. (>1%) gibbon food Canarium album (Lour.) DC. Burseraceae trám trắng poss. called an "olive" Garuga pierrei Guill cốc đá (chua luy) Labelled near HQ Aglaia pleuropteris Pierre Meliaceae gội gà croc. lake trail Aphanamixis polystachya JN Parker gội Croc. trail specimens Chukrasia tabularis A. Juss. lát hoa nam near Ben Cu Dimocarpus subsp. malesianus Leenh. nhãn Mã lai longan (domestic) Glenniea philippinensis (Radlk.) Leenh. chua khẹt Mischocarpus sundaicus Bl. trường kẹn hypoleucum Kurz chôm chôm trắng duối gibbon food: 'korlan' spirei Lecomte trường vải Xerospermum noronhianum (Bl.) Bl. trường gibbon food (>2%) pedunculata (L.) Miq. bưởi bung, bí bái (citrus family) Murraya koenigii (L.) Spreng. nguyệt quế Koenig The ten major groupings (mostly orders) above constitute >90% of the major tree and liana species, with others in the tables below. The Monocots remain as a monophyletic clade distinct from the Magnoliids ('basal' flowering plants) and (new or higher dicots). Not always considered trees per se., large Monocots such as palms form an important part of the forest under-storey layers (2-9 m), with and other grasses covering significant areas of the Park.

Other Orders (Eudicots) bọ khác other trees & lianas Gonocaryum lobbianum (Miers.) Kurz. Cardiopteridaceae cuống vàng (quỳnh Aquifoliales (was in Icacinaceae) lâm) Capparis spp. (6 recorded) ‡ Capparaceae cáp Brassicales Crateva cf. adansonii DC bún trái đỏ Crateva magna (Lour.) DC bún near waterways tectorius (Lour.) Merr. Ancistrocladaceae trung quân lợp nhà, dây trung quân Tetrameles nudiflora R.Br. Tetramelaceae tung; Maa: tung Cucurbitales Dillenia scabrella (D.Don.) Roxb. Dilleniaceae sổ nhám Dilleniales Engelhardia roxburghiana Wall. Juglandaceae chẹo Fagales Lithocarpus dinhensis (Hickel & Camus) A.Camus Fagaceae dẻ núi Dinh Leea aequata L. Vitaceae gối hạc, củ rối bằng Vitales (was Leeaceae)

Gymnospermae Thực vật hạt trần "conifers" Gnetum gnemon L. Margf. ‡ Gnetophyta: gắm, bét (lá nhíp); lianas: 'nhíp' are Gnetum latifolium Blume ‡ Gnetaceae Maa: kloot (chee kloot) popular vegetable Cycas inermis Loureiro Cycadales Cycadaceae thiên tuế (không gai); Maa: kul Endemic, VU

Monocots thực vật một lá mầm – about 10 spp. including: Order : mây; Maa: gool rattans C. dongnaiensis Pierre ex Conrad ‡ : họ Cau mây Ðồng Nai >2% gibbon C. flagellum Griff. ex Mart. ‡ mây song food; another rattan is Calamus poilanei Conr. song bột laciniosa ‡ Caryota mitis Lour. đùng đình (~ ngứa) fish-tail palm bracteata Gagnep mật cật, tơi Botanic trail Licuala cattienensis Henderson, Ban, Dung ‡ mật Cát Tiên Maa: sraa (genus) Licuala spinosa Wurmb. ‡ mật cật gai (ra gai) fan palms saribus (Lour.) Merr.ex Chev. ‡ cọ bầu, lá gồi, kè nam "taraw palm" Maa: triö Pinanga 5 spp. ‡ cau chuột; Maa: mpir Bambusa blumeana Schultes. Poales: Poaceae tre gai, tre la ngà spiny Bambusa procera A. Chev. & A Cam. (grass family) lồ ô Schizostachyum aciculare Gamble mung, nưa nhỏ Musa acuminata Colla ‡ Zingiberales: Musaceae chuối rừng; Maa: juu wild banana

‡ labelled specimens not seen © www.namcattien.org 5