Late Bajocian–Bathonian Ammonites from Northeast Iran

Home , Cadomites, Garantiana, Normannites, Oppeliinae, Orthogarantiana, Oxycerites

ACTA PALAEONTOLOGICA ROMANIAE (2015) V. 11 (2), P. 25-41

LATE BAJOCIAN–BATHONIAN AMMONITES FROM NORTHEAST IRAN

Mahmoud Reza Majidifard

Received: 19 August 2015 / Accepted: 18 December 2015 / Published online: 22 December 2015

Abstract The present study is the first part of a larger work concerning the systematic descriptions of the fairly rich Middle to Upper Jurassic (Late Bajocian–Early Tithonian) ammonite fauna from the northeast of Iran. The described ammonites come from the lower part of the Dalichai Formation, consisting of an alternation of green marl, grey limestone and marly limestone, and comprise the families Phylloceratidae, Spiroceratidae, Oppeliidae, Sphaeroceratidae, Stephanoceratidae, Tulitidae, Parkinsoniidae, Morphoceratidae and Perisphinctidae. Some of the taxa are recorded for the first time from Iran: Adabofoloceras subobtusum, Spiroceras cf. orbignyi, Spiroceras annulatum, Cadomites (Polyplectites) cf. dorni, Sphaeroceras tutthum, Garantiana (Orthogarantiana) cf. Densicostata, Garantiana (Pseudogarantiana) dichotoma, Morphoceras egrediens, Microbajocisphinctes cf. pseudointerruptus, Homoeoplanulites (Homoeoplanulites) cf. bugesiacus, Homoeoplanulites (Parachoffatia) arkelli. The following Upper Bajocian to Lower Bathonian ammonite zones are present: Niortense, Garantiana, Parkinsoni, Zigzag, Retrocostatum and Discus. Palaeobiogeographically, the described ammonite fauna is closely related to those from the Sub-Mediterranean Province of the northwestern Tethys.

Keywords: Jurassic; Iran; Late Bajocian–Bathonian; ammonites; taxonomy; Dalichai Formation.

INTRODUCTION GEOLOGICAL SETTING

The Dalichai Formation (Upper Bajocian–Lower The Dalichai Formation, named after the village and river Tithonian) is a rather soft-weathering lithostratigraphic Dalichai in the eastern Central Alborz, consists of an unit widely distributed in the Alborz and Binalud alternation of green marl, grey limestone and marly mountains of northern and northeastern Iran (Fig. 1). The limestone. The lower boundary of the Dalichai Formation Dalichai Formation is rich in ammonites which have been is an unconformity due to the Mid-Cimmerian tectonic in part been described by Assereto (1966), Seyed-Emami event (Seyed-Emami & Alavi-Naini, 1990; Fürsich et al., et al. (1985, 1989, 1991, 1994, 1995, 1996, 1997, 2001, 2009a) and the marine transgression of the Dalichai 2013), Schairer et al. (1992) and Majidifard (2003). Formation across the underlying Shemshak Group (Upper

Fig. 1 Location of the Golbini (1, eastern Alborz) and Tooy (2, Binalud mountains) sections of the Middle and Upper Jurassic Dalichai Formation in northeastern Iran.

______25 Research Institute for Earth Sciences, Geological Survey of Iran, Box131851 1494, Teheran, Iran; [email protected] Mahmoud Reza Majidifard

Triassic-Lower Bajocian; Fürsich et al., 2009b) is 1:250.000), with the following co-ordinates N 37°09ʹ14ʺ, diachronous. The upper boundary to the light-coloured E 57°09ʹ13ʺ. The Dalichai Formation has a thickness of and cliff-forming Lar Formation is gradational to sharp. 449 m at the Golbini section and 567 m at the Tooy The depositional environment of the Dalichai Formation section and consists predominantly of alternation of marl, is lower shelf to continental slope (Majidifard, 2003; grey limestone and marly limestone (Majidifard, 2003; Majidifard, 2008). Fig. 2). According to the ammonite fauna, the Dalichai The described ammonites come from two sections Formation is more or less time equivalent to the Chaman (Golbini and Tooy) in the lower part of the Dalichai Bid Formation of the Koppeh Dagh. Formation in the eastern Alborz (Figs. 1 and 2). The At the Golbini and Tooy sections, the Dalichai Formation Golbini section is located 16 km north of Jajarm near has been subdivided, from base to top, into three Golbini farm (quadrangle map of Kuh-e-Khurkhud, members A-C (Majidifard, 2008): 1:250.000), with the following co-ordinates N 37°05ʹ13ʺ, Member A consists of green-grey silty marls. E 56°44ʹ41ʺ (Fig. 1) and the Tooy section is located 4 km Member B consists alternations of well-bedded greyish- north of Tooy village (49 km west of Esferayen city and green marls, greyish to yellow marly limestones and 93 km SW of Bojnourd city; quadrangle map of Bojnourd medium-bedded, grey limestones with chert nodules at some levels.

Fig. 2 Stratigraphic logs of the studied sections of the Dalichai Formation. 26

Late Bajocian–Bathonian ammonites from northeast Iran

Member C comprises alternations of medium-bedded, Dimensions (in mm): grey to pinkish limestones with cherts, green-grey. specimen D U% H% W% The ammonites described herein have been collected GZN2011I -S-2-43 31 19 54 42 from the middle and upper part (member B) of the Dalichai Formation. Description: Involute phylloceratid with smooth shell, steep umbilicus and with 5 sigmoidal constrictions on the SYSTEMATIC PALAEONTOLOGY last visible whorl. The constrictions are narrow, start from the umbilical margin, and terminate at the rounded As far as permitted by the preservation of the specimens, venter. measurements of the following parameters are given: Stratigraphic distribution: Holcophylloceras has a diameter (D) in mm, umbilical width (U), whorl height worldwide distribution, from the Lower Jurassic (H), whorl width (W), the latter all in % of diameter; (Hettangian) to the Lower Cretaceous (Valanginian). The numbers of primary ribs per whorl (PR) and secondary present specimen comes from the Bathonian. ribs (SR). [M] = macroconch; [m] = microconch. All specimens are figured in natural size, except Genus Ptychophylloceras Spath, 1927 otherwise indicated. The figured specimens are located at the Bayerische Staatssammlung für Paläontologie und Ptychophylloceras sp. historische Geologie, Munich, Germany (prefix Fig. 3.3a-b GZN2011I); the other letter-number combination indicates the locality (J=Golbini section; S=Tooy section) Material: 1 specimen from Tooy section (GZN2011I-S- and the position in the section. 42). Dimensions (in mm): Superfamily Phylloceratinoidea Zittel, 1884 specimen D U% H% W% Family Phylloceratidae Zittel, 1884 GZN2011I-S-2-42 65 16 54 39 Subfamily Phylloceratinae Zittel, 1884 Genus Adabofoloceras Joly, 1977 Description: Shell involute, whorl section rounded- quadrate with narrow and shallow constrictions which Adabofoloceras subobtusum (Kudernatsch, 1852) start out as concave near the umbilical margin and turn Fig. 3.1a-b convex in a ventral direction. 1852 Ammonites subobtusus n. sp. Kudernatsch, p. 76, pl. Stratigraphic distribution: These specimens come from v. the Bathonian. 1977 Adabofoloceras subobtusum (Kudernatsch, 1852) – Joly, p. 34, pl. 2, fig. 12; pl. 5, figs. 3–5; pl. 41, fig. 1–2, Superfamily Spiroceratoidea Hyatt, 1900 pl. 43, figs. 3, 10 Family Spiroceratidae Hyatt, 1900 1980 Adabofoloceras subobtusum (Kudernatsch, 1852) – Genus Spiroceras Quenstedt, 1858 Galacz, 88, pl. iv, figs. 4–5 2000 Adabofoloceras subobtusum (Kudernatsch, 1852) – Spiroceras cf. orbignyi (Baugier & Sauze, 1843) Joly, p. 27, pl. 9, fig. 16; pl. 10, figs. 1–2. Fig. 3.5 1843 Toxoceras orbignyi sp. nov. – Baugier& Sauze: 6, Material: 1 specimen from Golbini section (GZN2011I-J- pl. 1, figs. 1–4. 32) and 3 specimens from Tooy section (GZN2011I-S- 1978 Spiroceras orbignyi (Baugier & Sauze) – Dietl: 76, 23, 28, 44). pl. 1, figs. 1–5, pl. 2, figs. 1–6, pl. 3, figs. 1–2. Dimensions (in mm): 1980 Spiroceras orbignyi (Baugier & Sauze) – Galacz: specimen D U% H% W% 199, pl. XXII, figs. 1–2 S-2-44 (body chamber) 40 16 55 45 1985 Spiroceras orbignyi (Baugier & Sauze) – GZN2011I -S-2-28 32 19 53 37 Schlegelmilch: 220, pl. 35, fig. 6. GZN2011I -J-2-32 25 12 52 28 1986 Spiroceras orbignyi (Baugier & Sauze) – Dietl: 42, pl. 1, figs. 2–6. Description: Involute with high-ovate whorl section, 2001 Spiroceras orbignyi (Baugier & Sauze) – steep umbilicus, and rounded venter. With typical Hillebrandt: 85, pl. 9, fig. 4. triphyllid phylloceratid suture line. Stratigraphic distribution: The present specimens come Material: 1 fragmented specimen from Tooy section from the Bathonian and is recorded for the first time from (GZN2011I-S-2a). Iran. Description: The fragmentary shell forms an open spiral with rounded whorl section. The ribs are simple with two Subfamily Calliphylloceratinae Spath, 1927 rows of nodes (lateral and ventrolateral) ending at a Genus Holcophylloceras Spath, 1927 ventral smooth band, where they bend slightly forward. Remarks: The single specimen differs from the holotype Holcophylloceras sp. in being more finely ribbed and from the specimen Fig. 3.2 figured by Dietl (1978) in the fainter swing of the ribs. Material: 1 specimen from Tooy section, (GZN2011I-S- Stratigraphic distribution: The present specimen occurs in 43). the Upper Bajocian Garantiana Zone. In France and Spain

Mahmoud Reza Majidifard

Late Bajocian–Bathonian ammonites from northeast Iran

Fig. 3 1a-b Adabofoloceras subobtusum (Kudernatsch), 1911 Oppelia yeovilensis n. sp. – Rollier: 305. specimen (GZN2011I-S-2-44) with body chamber from the 1968 Oxycerites yeovilensis Rollier – Hahn: 29, pl. 2, Tooy section, Bathonian. 2 Holcophylloceras sp., specimen figs. 1–4. (GZN2011I-S-2-43) from the Tooy section, Bathonian. 3a-b 1985 Oxycerites yeovilensis (Rollier) – Seyed-Emami et Ptychophylloceras sp., specimen (GZN2011I-S-2-42) from the al.: 62, pl. 1, fig. 4a–b. Tooy section, Bathonian. 4 Spiroceras annulatum (Deshayes), specimen (GZN2011I-S-1-2b) from the Tooy section, 1989 Oxycerites yeovilensis (Rollier) – Seyed-Emami et Garantiana Zone. 5 Spiroceras cf. orbignyi (Baugier & Sauze), al.: 82, pl. 1, figs. 1–3. specimen (GZN2011I-S-1-2a), from the Tooy section, 1991 Oxycerites yeovilensis (Rollier) – Seyed-Emami et Garantiana Zone, x3. 6 Oxycerites cf. oxus (Buckman), al.: 69, pl. 1, figs. 11–13. specimen (GZN2011I-2b-27) from the Golbini section, 2013 Oxycerites yeovilensis (Rollier) – Seyed-Emami et Tenuiplicatus to Subcontractus Zone. 7 Oecotraustes al.: 49, figs. 5a–b. (Paroecotraustes) aff. serrigerus (Waagen), specimen (GZN2011I-J-2c-23) from the Golbini section, Retrocostatum Material: 4 specimens from Golbini section (GZN2011I- Zone. 8a-b Oxycerites yeovilensis (Rollier), specimen J-24-26, 29). (GZN2011I-J-2b-29) from the Golbini section, Zigzag Zone. 9a-c Cadomites (Polyplectites) sp., specimen (GZN2011I-DH- Dimensions (in mm): 1-6) from the Golbini section, Upper Bajocian-Lower specimen D U% H% W% Bathonian, x1.5. 10a-b Compressed Cadomites sp. B, specimen GZN2011I-J-2b-26 58 9 59 22 (GZN2011I-J-2b-88a) from the Golbini section, Bathonian, x3. GZN2011I-J-2b-29 54 7 59 18 11 Cadomites (Polyplectites) cf. dorni (Roche), specimen GZN2011I-J-2b-24 54 9 56 20 (GZN2011I-S-2a-21) from the Tooy section, Upper Bajocian- Lower Bathonian. 12a-b Parkinsonia radiata Renz, specimen GZN2011I-J-2b-25 40 10 55 22 (GZN2011I-DH-1-1) from the Golbini section, Parkinsoni Zone. 13a-b Strenoceras sp., specimen (GZN2011I-J-1-2a) Description: Shell involute and oxycone, whorl cross- from the Golbini section, Niortense Zone, x1.5. 14a-b section trigonal to high-oval. The ribbing is mostly Bullatimorphites sp., specimen (GZN2011I-J-2a-17) from the restricted to the outer part of the whorl and consists of Golbini section, Zigzag Zone, x1.5. 5a-b, 16a-b. Sphaeroceras distant, rursiradiate ribs that end at a very sharp keel. The tutthum (Buckman), Garantiana Zone, x3. 15 specimen greatest whorl thickness is about mid-flank, from where GZN2011I-S-1-1b from the Tooy section. 16 specimen the flank slopes towards the umbilicus and the sharp keel. GZN2011I-J-1-1c from the Golbini section. Remarks: This species can be distinguished from

Oxycerites limosus Hahn (1968) by its strong and distinct the species occurs in the Parkinsoni Zone, Acris ribs. SubzoneDietl (1986) and are recorded for the first time Stratigraphic distribution: Lower Bathonian Zigzag Zone from Iran. (Parvum and Macrescens subzones) and Aurigerus Zone

(Recinctus Subzone), Fernández-López et al., 2009. Spiroceras annulatum (Deshayes, 1831)

Fig. 3.4 Oxycerites cf. oxus (Buckman, 1926) 1831 Hamites annulatus sp. nov. – Deshayes: 228, pl. 6, Fig. 3.6 fig. 5. cf. 1926 Micromphalites oxus sp. nov. –Buckman: pl. 1978 Spiroceras annulatum (Deshayes) – Dietl: 87, pl. 6, 644. figs. 1–5. cf. 1983 Oxycerites oxus (Buckman) – Dietl & Kapitzke: 1985 Spiroceras annulatum (Deshayes) – Schlegelmilch: 9, pl.1, fig. 4a, b. 223, pl. 36, fig. 2. cf. 1987 Oxycerites oxus (Buckman) – Torrens: 96, pl. 1, 1990 Spiroceras annulatum (Deshayes) – Sandoval: 161, fig. 6. pl. 4, fig. 1. cf. 1991 Oxycerites oxus (Buckman) – Seyed-Emami et 2002 Spiroceras annulatum (Deshayes) – Schweigert: 57, al.: 69, pl. 2, figs. 3–4. pl. 1, fig. 4. cf. 2001 Oxycerites oxus (Buckman) – Mitta: 112, pl. 1,

fig. 5. Material: 3 small fragments from Tooy section cf. 2007 Oxycerites (Oxycerites) oxus (Buckman, 1926) – (GZN2011I-S-2b-d). Kim et al.: 26, pl. 9, fig. 7. Description: The fragmentary shell forms an open spiral with a high-ovate whorl section. The ribbing is strong and Material: 1 specimen from Golbini section (GZN2011I-J- simple. It ends at a ventral smooth band. 27). Stratigraphic distribution: The species is widely Dimensions (in mm): distributed in Europe (e.g. Germany, France, Spain, specimen D U% H% W% England, Italy), North America, and North Africa. The present specimens have been found in the Upper Bajocian GZN2011I-J-2b-27 36 8 57 22 Garantiana Zone and are recorded for the first time from Iran. Description: Shell involute and oxycone, without Family Oppeliidae Bonarelli, 1894 ornamentation on the inner flank. The outer flanks have Subfamily Oppeliinae Bonarelli, 1894 weak falcate ribs that terminate at the ventrolateral Genus Oxycerites Rollier, 1909 shoulder. Remarks: The specimen is referred to Oxycerites oxus Oxycerites yeovilensis (Rollier, 1911) with reservation, because it is broken and the whorl Fig. 3.8a-b cross-section cannot be seen. The specimen is similar to 29

Mahmoud Reza Majidifard

Oxycerites seebachi Hahn (1968) but differs in having a specimen D U% H% W% narrower umbilicus and weaker ribs. GZN2011I -S-2a-21 25 40 32 52 Stratigraphic distribution: Lower Bathonian Zigzag Zone. Torrens (1987) reported the species also from the Description: Umbilical wall steep and flat, venter broad. Tenuiplicatus to Subcontractus Zone. Primary ribs prorsiradiate, fairly strong and dense, starting at the umbilical margin and ending at prominent Genus Oecotraustes Waagen, 1869 sharp nodes at around the ventrolateral shoulder. From Subgenus Paroecotraustes Spath, 1928 this point they divide into 2 or 3 fairly strong secondary ribs that cross the venter. Oecotraustes (Paroecotraustes) aff. serrigerus (Waagen, Remarks: The present specimen is similar to the holotype 1869) but differs in having coarser ribs. Moreover the position Fig. 3.7 of the nodes is closer to the venter. Also, it has finer ribs aff. 1869 Oecotraustes serrigerus n. sp. – Waagen: 230, than Sturani’s (1966) specimen. pl. 20, fig. 7. Stratigraphic distribution: Upper Bajocian to Lower aff. 1966 Oecotraustes (Paroecotraustes) serrigerus Bathonian. Waagen – Stephanov: 48, pl. 3, figs. 12–14. aff. 1968 Oecotraustes (Paroecotraustes) aff. serrigerus Cadomites (Polyplectites) sp. Waagen – Hahn: 60, pl. 5, figs. 4–5. Fig. 3.9a-c aff. 1972 Oecotraustes (Paroecotraustes) serrigerus Material: 1 specimen and 3 fragments from Golbini (Waagen) – Krystyn: 234, pl. 4, fig. 5. section (GZN2011I-DH-6). aff. 1985 Oecotraustes (Paroecotraustes) aff. serrigerus Dimensions (in mm): (Waagen) – Seyed-Emami et al.: 63, pl. 1, fig. 7a, b. specimen D U% H% W% GZN2011I -DH-1-6 28 42 47 58 Material: 1 specimen from Golbini section (GZN2011I-J- 23). Description: Whorl cross-section trapezoidal, venter Dimensions (in mm): broad. The slightly prorsiradiate and dense primaries start specimen D U% H% W% at the umbilical margin and end at prominent nodes near GZN2011I-J-2c-23 33 24 55 24 mid-flank. The bundled secondaries, which cross the venter, are trifurcate with some intercalatory ribs. Description: Shell nearly evolute, whorl section ovate Remarks: The specimens can be compared with with distinct keel, umbilical margin rounded, and Cadomites (Cadomites) extinctus Quenstedt (1887) but umbilical wall steep with falcoid ribs. Ornamentation of differ in the narrower inner coiling. Since the umbilical fine and prorsiradiate primary ribs, secondary ribs strong area is not seen in the present specimens, it is better to and rursiradiate. The secondary ribs terminate on the refrain from a specific identification. venter at a sharp keel. Stratigraphic distribution: Upper Bajocian to Lower Remarks: The present specimen with its fine primary ribs Bathonian. closely resembles the specimens as illustrated by Krystyn (1972) and Stephanov (1966). It differs from the Cadomites sp. specimens figured by Hahn (1968) by its blunt secondary Fig. 3.10a-b ribs and lack of primary ribs. Material: 1 specimen from Golbini section (GZN2011I-J- Stratigraphic distribution: Lower part of the Upper 88a). Bathonian, Retrocostatum Zone. Description: Specimen compressed, venter rounded. Ribbing relatively coarse and strong. The slightly Superfamily Stephanoceratoidea Neumayr, 1875 rectiradiate primary ribs end mostly at small nodes, from Family Stephanoceratidae Neumayr, 1875 where they bifurcate and trifurcate with some Genus Cadomites Munier-Chalmas, 1892 intercalatory ribs, extending across the venter. Subgenus Polyplectites Mascke, 1907 Stratigraphic distribution: Upper Bajocian to Lower Bathonian. Cadomites (Polyplectites) cf. dorni (Roche, 1939) Fig. 3.11 Family Sphaeroceratidae Buckman, 1920 cf. 1939 Normannites dorni n. nov. – Roche: 222. Genus Sphaeroceras Bayle, 1878 cf. 1966 Polyplectites dorni (Roche) – Sturani: 29, pl. 6, fig. 4. Sphaeroceras tutthum Buckman, 1921 cf. 1985 Cadomites (Polyplectites) dorni (Roche) – Figs. 3.15a-b, 16a-b Schlegelmilch: 83, pl. 30, fig. 1. 1921 Sphaeroceras tutthum sp. nov. – Buckman: 258. cf. 2000 Cadomites (Polyplectites) dorni (Roche) – 1937 Sphaeroceras tutthum Buckman – Wetzel: 78, pl. Dietze: 40, pl. 12, fig. 3. 10, fig. 1. 1976 Sphaeroceras (Sphaeroceras) auritum subsp. Material: 1 fragmentary specimen from Tooy section tutthum (S. Buckman) – Parsons: 22, pl. 2, figs. 4–6. (GZN2011I-S-21). Dimensions (in mm):

Late Bajocian–Bathonian ammonites from northeast Iran

Material: 32 specimens from Golbini section (GZN2011I- Description: Shell inflated, with very broad venter and J-1a–z, 44a–e) and 16 specimens from Tooy section broad-ovate whorl section. The ribbing is faint. The ribs (GZN2011I-S-1a–p). are rectiradiate, start at the umbilical margin, and cross Dimensions (in mm): the venter. specimen D U% H% W% Remarks: The present specimens differ from GZN2011I -S-1-1b 10 7 35 100 Bullatimorphites aff. ymir published by Seyed-Emami et GZN2011I -S-1-1h 9 6 37 100 al. (1991) in having a smaller umbilical angle and a larger GZN2011I -S-1-1c 8 6 46 88 diameter. GZN2011I -S-1-1f 9 6 33 89 Stratigraphic distribution: The species has been recorded GZN2011I -J-1-1c 11 9 55 100 by Sandoval (1983) from the Upper Bathonian–Lower Callovian. Seyed-Emami et al. (1991) recorded it from GZN2011I -J-1-1m 7 6 50 93 the Middle Bathonian of Iran. GZN2011I -J-1-1g 5 6 40 80

GZN2011I -J-1-1s 10 9 40 98 Bullatimorphites sp. Fig. 3.14a-b Description: A very small, globose ammonite, shell 1989 Bullatimorphites sp. – Seyed-Emami et al.: 84, pl. involute, shape rounded to ovate, whorl cross-section 2, fig. 4. broad-ovate, venter broad. The very fine and rectiradiate primaries start at the umbilical margin and divide into 2 Material: 2 specimens from Golbini section (GZN2011I- or 3 secondaries around one-fourth of flank height, J-17-18). crossing the venter. Dimensions (in mm): Remarks: The specimens are similar to those described specimen D U% H% W% by Wetzel (1937), but differ in having somewhat stronger GZN2011I -J-2a-17 22 27 45 73 ribs and a slightly more inflated shell. It differs from the GZN2011I -J-2a-18 13 23 27 45 specimens figured by Parsons (1976) by in finer ribs at the umbilical margin and a bigger umbilical angle. Stratigraphic distribution: Upper Bajocian, Garantiana Description: Shell nearly involute with steep and deep zone. umbilical wall and broad venter. The primaries begin at the umbilical margin and around the ventrolateral Family Tulitidae Buckman, 1921 shoulder they divide mainly into two secondaries, which Genus Bullatimorphites Buckman, 1921 cross the venter. Some of the primaries do not divide. Subgenus Kheraiceras Arkell, 1952 Remarks: The present specimen is similar to the specimen figured by Seyed-Emami et al. (1989) except Bullatimorphites (Kheraiceras) bullatus (d´ Orbigny, for its finer and denser ribbing. 1846) Stratigraphic distribution: Lower Bathonian, Zigzag Fig. 4.7a-b Zone. 1846 Ammonites bullatus sp. nov. – dÓrbigny: 12, pl. 142, figs. 1–2. Superfamily Perisphinctoidea Steinmann, 1890 1887 Ammonites bullatus dÓrbigny – Quenstedt: 658, pl. Family Parkinsoniidae Buckman, 1920 77, figs. 7– 12, pl. 78, fig. 1. Genus Strenoceras Hyatt, 1900 1887 Ammonites platystomus globulatus – Quenstedt: 661, pl. 78, fig. 2. Strenoceras sp. 1887 Ammonites platystomus – Quenstedt: 665, pl. 78, Fig. 3.13a-b figs. 22– 25, 28– 30. Material: 2 fragments from Golbini section (GZN2011I- 1958 Bullatimorphites bullatus bullatus (dÓrbigny) – J-2a, b). Westermann: 65, pl. 20. Description: The ribs are very strong, sharp, rectiradiate 1958 Bullatimorphites bullatus hannoveranus to slightly prorsiradiate and mainly simple with lateral (dÓrbigny) – Westermann: 65, pl. 21. and ventral tubercles. The primaries begin at the 1971 Bullatimorphites (Kheraiceras) bullatus umbilical margin and end at prominent lateral tubercles, (dÓrbigny) – Hahn: 99, pl. 7, figs. 1–3, pl. 8, figs. 1-4. from where they continue as secondaries, which are 1985 Bullatimorphites (Kheraiceras) bullatus usually simple, seldom bifurcating. There are also (dÓrbigny) – Schlegelmilch: 143, pl. 52, fig. 4. tubercles on the ventrolateral shoulder. The secondaries 1991 Kheraiceras cf. bullatum (dÓrbigny) – Seyed- terminate at a ventral furrow. Emami et al.: 72, pl. 4, fig. 1. Remarks: The present material is compressed and 1994 Bullatimorphites bullatus (dÓrbigny) – Fischer: deformed. The specimens can be compared with 131, pl. 56, fig. 1. Strenoceras bentzi as illustrated by Dietl (1983) but differ in having somewhat finer ribs and a higher position of Material: 2 specimens and 1 fragment from Tooy section lateral tubercles. (GZN2011I-S-12-14). Stratigraphic distribution: Upper Bajocian, Niortense Dimensions (in mm): Zone.

specimen D U% H% W% Genus Garantiana Mascke, 1907 GZN2011I -S-2b-12 52 22 40 98 Subgenus Orthogarantiana Bentz, 1928 GZN2011I -S-2b-13 61 20 41 84 31

Mahmoud Reza Majidifard

Late Bajocian–Bathonian ammonites from northeast Iran

Fig. 4 1a-b Garantiana (Orthogarantiana) cf. densicostata Subgenus Pseudogarantiana Bentz, 1928 (Quenstedt), specimen (GZN2011I-J-1-3a) from the Golbini section, Niortense Zone, specimen compressed, x3. 2a-b Garantiana (Pseudogarantiana) dichotoma Bentz, 1928 Garantiana (Pseudogarantiana) dichotoma Bentz, specimen Fig. 4.2a-b (GZN2011I-S-1-3a) from the Tooy section, Garantiana Zone, 1928 Garantiana (Pseudogarantiana) dichotoma sp. nov. x1.5. 3a-b Morphoceras multiforme Arkell, specimen (GZN2011I-J-2b-1) from the Golbini section, Zigzag Zone. 4a- – Bentz: 200, pl. 19, figs. 2–6. b Parkinsonia aff. depressa (Quenstedt), specimen (GZN2011I- 1973 Garantiana (Pseudogarantiana) dichotoma Bentz – DH-1-2) from the Golbini section, Upper Bajocian (Parkinsoni Pavia: 109, pl. 19, fig. 5. Zone) to ?Lower Bathonian. 5a-b Parkinsonia parkinsoni 1974 Garantiana (Pseudogarantiana) dichotoma Bentz – (Sowerby), specimen (GZN2011I-S-1-20) from the Tooy Dietl: 12, pl. 2, fig. 3. section, Parkinsoni Zone. 6a-b Morphoceras macrescens 1989 Garantiana (Pseudogarantiana) dichotoma Bentz – (Buckman), specimen (GZN2011I-J-2b-2) from the Golbini Morton & Dietl: 157, figs. 1E, H. section, Zigzag Zone. 7a-b Bullatimorphites (Kheraiceras) 2002 Garantiana (Pseudogarantiana) dichotoma Bentz – bullatus (D’Orbigny), specimen (GZN2011I-S-2b-12) from the Schweigert et.al.: 81, pl. 13, fig. 5. Tooy section, Middle Bathonian. 8 Ebrayiceras cf. sulcatum (Zieten), specimen (GZN2011I-J-2b-20a) with lappet from the Golbini section, Zigzag Zone. 9a-b Microbajocisphinctes cf. Material: 4 specimens and 32 fragments from Tooy pseudointerruptus Fernandez Lopez, specimen (GZN2011I-S-1- section (GZN2011I-S-3a–z, 4a-l). 2) from the Tooy section, Garantiana Zone. 10a-b Morphoceras Dimensions (in mm): egrediens Wetzel, specimen (GZN2011I-J-2b-20) from the specimen D U H% W% PR/2 SR/2 Golbini section, Zigzag Zone. 11a-b Microbajocisphinctes sp. B, specimen (GZN2011I-S-1-3) from the Tooy section, % Garantiana Zone. GZN2011I -S-1-3a 22 41 36 23 15 27 GZN2011I -S-1-3d 12 42 33 25 14 26 Garantiana (Orthogarantiana) cf. densicostata (Quenstedt, 1886) Description: Shell evolute with subquadrangular whorl Fig. 4.1a-b cross-section. Ribs sharp, mostly bifurcating and only a cf. 1886 Ammonites Garantianus densicostatus – few simple. The ribs are prorsiradiate, only slightly Quenstedt: 593, pl. 71, fig. 9. curved, and terminate in small tubercles. Fine and sharp cf. 1974 Garantiana (Orthogarantiana) densicostata nodes developed at the point of bifurcation (around mid- (Quenstedt) – Dietl: 13, pl. 2, fig. 4. flank). On the venter there is a small furrow, which is cf. 1985 Orthogarantiana gr. densicostata (Quenstedt) – typical of Pseudogarantiana. Fernandez-Lopez: pl. 45, fig. 3. Remarks: The present material is very similar to that as illustrated by Dietl (1974) and Morton & Dietl (1989). Material: 39 specimens and 10 fragments from Golbini The specimens differ from Pavia´s figure (1973) by the section, (GZN2011I-J-3a–z, 4a–w). more ventral position of the lateral tubercles and from Dimensions (in mm): Schlegelmilch´s (1985) figure by the dense and more commonly bifurcating ribs. specimen D U% H% W% PR/2 SR/2 Stratigraphic distribution: GZN2011I -J-1-3a 13 38 38 46 9 19 Garantiana (Pseudogarantiana) dichotoma is very GZN2011I-J-1-3m 12 42 37 42 10 22 widely distributed in the Submediterranean and GZN2011I -J-1-4l 10 30 45 50 9 20 Subboreal faunal provinces, from southern Germany and GZN2011I -J-1-4d 11 35 39 40 10 23 Spain to northwest Scotland. The species occurs mainly GZN2011I -J-1-4e 9 33 41 55 9 20 in the lower and middle parts of the Garantiana Zone and GZN2011I -J-1-3j 9 39 36 51 10 23 are recorded for the first time from Iran. GZN2011I -J-1-4g 9 40 39 48 12 26 GZN2011I -J-1-3h 11 34 45 46 9 21 Genus Parkinsonia Bayle, 1878 GZN2011I -J-1-3t 10 32 36 40 10 22 Parkinsonia radiata Renz, 1904 Description: Shell evolute with rounded umbilical margin Fig. 3.12a-b and steep umbilical wall. Ribs sharp, slightly 1886 Ammonites Parkinsonia planulatus – Quenstedt: prorsiradiate and mostly bifurcating, with only a few of 599, pl. 71, fig. 19. them simple or trifurcating. Prominent, sharp nodes 1904 Parkinsonia parkinsonia Sow. var. radiata Renz – developed at the point of bifurcation (at one-third of flank Renz: 77. height). Secondaries prorsiradiate, terminating on the 1913 Parkinsonia radiata Renz – Renz: 690, pl. 28, fig. venter at a small furrow. 3. Remarks: Some of the specimens are compressed and 1985 Parkinsonia radiata Renz – Seyed-Emami et al.: deformed. The present material is similar to the 66, pl. 3, fig. 2. specimens as illustrated by Dietl (1974) and Fernandez- 1985 Parkinsonia (Parkinsonia) radiata Renz – Lopez (1985) except for their lower position of lateral Schlegelmilch: 94, pl. 33, figs. 4–5. tubercles and somewhat stronger ribs. Stratigraphic distribution: Upper Bajocian, Niortense Material: 1 specimen from Golbini section (GZN2011I- Zone and are recorded for the first time from Iran. DH-1).

Mahmoud Reza Majidifard

Dimensions (in mm): 2006 Parkinsonia parkinsoni (Sowerby) – Dietze & specimen D U% H% W% PR/2 SR/2 Dietl: 35, pl. 2, fig. 1. 2010 Parkinsonia (Parkinsonia) parkinsoni (Sowerby) – GZN2011I -DH-1-1 50 46 34 32 16 30 Zaton: 115–213, pl. 3, figs. H–K.

Description: Umbilical wall vertical, whorl cross-section Material: 1 fragment from Tooy section (GZN2011I-S- rectangular-ovate. The inner whorls are wider than high 20). but near the body chamber the width equals height. The Description: Shell relatively evolute, with rectangular- sharp, strong and prorsiradiate primary ribs begin at the oval whorl cross-section. The primary ribs are strong, umbilical margin and terminate at about mid-flank. They sharp and fairly distant, starting at the umbilical margin split into two prorsiradiate secondaries, which end at a and dividing at around two-thirds of flank height into furrow on the venter. prorsiradiate secondary ribs that terminate at a narrow, Remarks: The inner whorls of this specimen are very sharp furrow on the venter. similar to that of the holotype. According to Galacz Stratigraphic distribution: Upper Bajocian, Parkinsoni (1980), Parkinsonia depressa (Quenstedt) and Zone. Parkinsonia subarietis Wetzel (1937) belong to

Parkinsonia rarecostata (Quenstedt) and differ from Family Morphoceratidae Hyatt, 1900 Parkinsonia radiata in being more evolute, having a Genus Morphoceras Douville, 1880 wider whorl cross-section and less terminal growth.

Parkinsonia bigoti of Nicolesco (1928) can be considered Morphoceras multiforme Arkell, 1951 as a synonym of Parkinsonia radiata. Fig. 4.3a-b Stratigraphic distribution: Upper Bajocian, Parkinsoni 1846 Ammonites polymorphus sp. nov. – d' Orbigny: 379, Zone. pl. 124, fig. 1.

1951 Morphoceras multiforme nom. nov. – Arkell: 17. Parkinsonia aff. depressa (Quenstedt, 1858) 1954 Morphoceras multiforme Arkell – Arkell: 132, pl. Fig. 4.4a-b 16, figs. 1–2. aff. 1858 Ammonites Parkinsoni depressus sp. nov. – 1970 Morphoceras multiforme Arkell – Hahn: 33, pl. 5, Quenstedt: 472, pl. 63, fig. 9. figs. 1–5. aff. 1973 Parkinsonia (P) cf. depressa (Quenstedt) – 1985 Morphoceras multiforme Arkell – Seyed-Emami et Pavia: 121, pl. 23, fig. 3. al.: 67, pl. 4, figs. 6–7. aff. 1985 Parkinsonia cf. depressa (Quenstedt) – Seyed- 2010 Morphoceras multiforme Arkell – Zaton: 115–213, Emami et al.: 67, pl. 3, fig. 7. pl. 13, figs. A–B, pl. 12, fig. I.

Material: 2 specimens from Golbini section, (GZN2011I- Material: 2 specimens from Golbini section (GZN2011I- DH-2, 20). J-1, 4). Dimensions (in mm): Dimensions (in mm): specimen D U% H% W% PR/2 specimen D U% H% W% GZN2011I -DH-1-2 40 49 34 24 17 GZN2011I -J-2b-1 32 19 50 53 GZN2011I -DH-1-20 47 47 34 28 17 GZN2011I -J-2b-4 45 16 54 57

Description: Whorl cross-section broad-ovate. Primaries Description: Whorl cross-section nearly rectangular. prorsiradiate and sharp, starting at the umbilical margin, Umbilical wall close to vertical. The ribbing consists of dividing into two sharp and prorsiradiate secondaries at 3–5 primary ribs each that originate from small nodes about two-thirds of flank height, and ending at a narrow near the umbilical margin. Most primary ribs divide into and shallow furrow on the venter. two prorsiradiate secondary ribs at about one-fourth of Remarks: The specimens resemble the holotype but differ flank height and terminate at a faint furrow on the venter. in being more involute and having slightly coarser ribs. There are three deep and prorsiradiate constrictions on They also can be compared with Parkinsonia rarecostata the last visible whorl. Schlegelmilch (1985) except for being somewhat more Remarks: The inner whorls are similar to Morphoceras involute and having a wider whorl cross-section. macrescens Buckman (1923) but their width exceeds the Stratigraphic distribution: Upper Bajocian (Parkinsoni height. The beginning body chamber in Morphoceras Zone) to ?Lower Bathonian. macrescens is distinctly evolute.

Stratigraphic distribution: Lower Bathonian, Zigzag Parkinsonia parkinsoni (J. de C. Sowerby, 1823) Zone. Fig. 4.5a-b

1823 Ammonites Parkinsoni sp. nov. – J. De C. Sowerby: Morphoceras macrescens (Buckman, 1923) 1, pl. 307, fig. 1. Fig. 4.6a-b 1980 Parkinsonia (Parkinsonia) parkinsoni (Sowerby) – 1923 Patemorphoceras macrescens sp. nov. – Buckman: Galacz: 93, pl. 22, fig. 5. 376. 1985 Parkinsonia parkinsoni (Sowerby) – Seyed-Emami 1971 Morphoceras macrescens (Buckman) – Hahn: 35, et al.: 67, pl. 3, fig. 1. pl. 5, figs. 10–15. 1985 Parkinsonia (Parkinsonia) parkinsoni (Sowerby) – Schlegelmilch: 94, pl. 33, fig. 7. 34

Late Bajocian–Bathonian ammonites from northeast Iran

1985 Morphoceras macrescens (Buckman) – Seyed- cf. 1830 Ammonites sulcatus sp. nov. – Zieten: 6, pl. 5, Emami: 68, pl. 4, figs. 3–4. fig. 3. 2010 Morphoceras multiforme Arkell – Zaton: 115-213, cf. 1970 Ebrayiceras sulcatum (Zieten) – Hahn: 42, pl. 6, pl. 13, fig. c, pl. 12, fig. h. figs. 1–6. cf. 1970a Ebrayiceras sulcatum (Zieten) – Mangold: 89, Material: 2 fragments from Golbini section (GZN2011I- pl. 7, fig. 6. J-2-3). cf. 1985 Ebrayiceras sulcatum (Zieten) – Seyed-Emami Dimensions (in mm): et al.: 69, pl. 3, figs. 8–10. specimen D U% H% W% cf. 1985 Morphoceras (Ebrayiceras) sulcatum (Zieten) – GZN2011I -J-2b-2 35 21 50 44 Schlegelmilch: 103, pl. 36, fig. 15. GZN2011I -J-2b-3 33 21 45 cf. 1991 Ebrayiceras sulcatum (Zieten) – Seyed-Emami et al.: 73, pl. 4, figs. 3, 8. Description: Whorl cross-section rounded to ovate. The ribbing is fine and dense. The primaries start at the Material: 1 specimen from Golbini section (GZN2011I-J- umbilical margin, divide mostly into two prorsiradiate 20a). secondaries at about one-fourth of flank height, and end Dimensions (in mm): at a shallow furrow on the venter. There are five shallow specimen D U% H% W% PR SR distant constrictions on the last visible whorl. The first rib /2 /2 after a constriction is parallel to it, the subsequent ones GZN2011I -J-2b-20a 28 25 46 31 27 form an angle. Remarks: The inner whorls are similar to Morphoceras Description: Shell relatively evolute, but inner whorls multiforme Arkell (1951) but the height exceeds the nearly involute; with faint constrictions and with lappet. width, and the constrictions are shallow. According to The primary ribs start at faint and regular nodes near the Wetzel (1937), Morphoceras egrediens somewhat differs umbilical margin and divide mainly into two secondary from Morphoceras macrescens in having less terminal ribs at about one-fourth of flank height. They terminate at growth. According to Hahn (1971), Morphoceras a distinct furrow on the venter. egrediens is a variant of M. macrescens. Remarks: The specimen can be compared with the Stratigraphic distribution: Lower Bathonian, Zigzag holotype but differs somewhat in having finer ribs and Zone. nodes. Ebrayiceras cf. sulcatum is similar to E. filicostatum Seyed-Emami et al. (1985) but differs in Morphoceras egrediens Wetzel, 1937 having coarser and more distant ribs. Fig. 4.10 a-b Stratigraphic distribution: Lower Bathonian, Zigzag 1937 Morphoceras inflatum var. egrediens var. nov. – Zone. Wetzel: 132. 1970a Morphoceras egrediens Wetzel – Mangold: 66, pl. Family Perisphinctidae Steinmann, 1890 5, figs. 4–5, 10. Genus Microbajocisphinctes Fernandez-Lopez, 1985

Material: 1 specimen from Golbini section (GZN2011I-J- Microbajocisphinctes cf. pseudointerruptus Fernandez- 20). Lopez, 1985 Dimensions (in mm): Fig. 4.9a-b specimen D U% H% W% PR SR cf. 1985 Microbajocisphinctes pseudointerruptus sp. nov. /2 /2 – Fernandez-Lopez: 20, pl. 49, fig. 9. GZN2011I -J-2b-20 65 18 51 23 47 Material: 1 specimen and 9 fragments from Tooy section Description: Whorl cross-section oval, venter arched. The (GZN2011I-S-2a-I). narrow inner flank is covered by coarse, blunt ribs. The Dimensions (in mm): ribs divide at around one-fourth of flank height into two specimen D U% H% W% PR/2 or three prorsiradiate and dense secondaries that end at a GZN2011I -S-1-2 39 46 34 32 15 faint furrow on the venter. There are three relatively shallow constrictions on the last visible whorl and also Description and remarks: Whorl section sub-rounded. some intercalatory ribs. Primaries strong, coarse, fairly distant and prorsiradiate, Remarks: The specimen can be compared with starting at the umbilical margin and dividing into two and Morphoceras macrescens Buckman (1923), but differ in occasionally three prorsiradiate secondaries at around a greater height/width ratio and blunt primaries at the two-thirds of flank height. Ribbing denser in the posterior umbilical margin. half of the anterior part of the last visible whorl. Stratigraphic distribution: Lower Bathonian, Zigzag Zone Remarks: The present specimens are very similar to the and is recorded for the first time from Iran. specimen described and figured by Fernandez-Lopez (1985) but differ somewhat in being smaller and having Genus Ebrayiceras Buckman, 1920 slightly prorsiradiate ribs. Stratigraphic distribution: According to Fernandez-Lopez Ebrayiceras cf. sulcatum (Zieten, 1830) (1985), Microbajocisphinctes pseudointerruptus occurs Fig. 4.8 35

Mahmoud Reza Majidifard in the Upper Bajocian, Garantiana Zone and are recorded Genus Procerites Siemiradzki, 1898 for the first time from Iran. Procerites sp. Microbajocisphinctes sp. A Fig. 5.9a-b Fig. 5.8a-b Material: 1 specimen from Golbini section (GZN2011I-J- 1985 Microbajocisphinctes sp. – Fernandez-Lopez: 20, 21). pl. 49, fig. 6. Description: Whorl cross-section high-oval, umbilical wall steep, shell medium-sized. The primary ribs are Material: 2 specimens from Tooy section (GZN2011I-S- relatively coarse, distant and prorsiradiate. Most of them 4, 7). split into two prorsiradiate secondary ribs at around three- Dimensions (in mm): fourth of the flank height and cross the venter. specimen D U% H% W PR SR/2 Stratigraphic distribution: Bathonian. % /2 GZN2011I -S-1-4 50 54 29 26 19 Genus Planisphinctes Buckman, 1922 GZN2011I -S-1-7 33 41 36 35 18 36 Planisphinctes sp. Description: Whorl cross-section rounded, venter fairly Fig. 5.4a-b broad. The primaries are strong and prorsiradiate, starting Material: 1 specimen from Golbini section (GZN2011I-J- at the umbilical margin and dividing into two 12). prorsiradiate secondaries at about three-fourth of flank Description: Whorl cross-section nearly oval, shell height, crossing prorsiradiate over the venter. Towards medium-sized, umbilical wall steep, venter rounded. The the aperture ribs become more prorsiradiate. There are ribbing is dense and fine on the inner whorls. The dense two prorsiradiate constrictions on the last visible whorl. primaries divide at around three-fourth of flank height Remarks: Microbajocisphinctes sp. A resembles into two prorsiradiate secondaries which, together with specimens as illustrated by Fernandez-Lopez (1985), but some intercalated ribs, cross the venter. The secondaries differs in having constrictions. are denser and fainter than the primaries. Stratigraphic distribution: According to Fernandez-Lopez Stratigraphic distribution: Bathonian. (1985), the taxon occurs in the Upper Bajocian, Garantiana Zone. Genus Homoeoplanulites Buckman, 1922 Subgenus Homoeoplanulites Buckman, 1922 Microbajocisphinctes sp. B Fig. 4.11a-b Homoeoplanulites (Homoeoplanulites) cf. bugesiacus Material: 1 specimen and 7 fragments from Tooy section (Dominjon, 1969) (GZN2011I-S-3a-h). Figs. 5.2, 5 Dimensions (in mm): cf. 1969 Choffatia (Homoeoplanulites) bugesiaca sp. specimen D U% H% W% PR SR/ nov. – Dominjon: 16, pl. 2, figs. 4–6. /2 2 cf. 1970b Homoeoplanulites (Homoeoplanulites) GZN2011I -S-1-3a 35 47 31 31 19 40 bugesiacus (Dominjon) – Mangold: 59, pl. 2, figs. 2, 7–9. cf. 1994 Homoeoplanulites (Homoeoplanulites) cf.

bugesiacus Dominjon – Dietl: pl. 5, fig. 2. Description: Whorl section rounded. The prorsiradiate primaries divide at around two-thirds of flank height into Material: 1 specimen and 1 fragment from Tooy section two prorsiradiate secondaries; rarely intercalated ribs (GZN2011I-S-47, 49). occur. Dimensions (in mm): Remarks: Microbajocisphinctes sp. B is similar to Microbajocisphinctes sp. A (Fig. 5.8a-b), but lacks specimen D U% H% W% PR/2 constrictions and has fainter prorsiradiate ribs. GZN2011I -S-2b-47 70 37 32 26 21 Stratigraphic distribution: Upper Bajocian, Garantiana GZN2011I -S-2b-49 63 37 39 23 25 Zone. Description: Ribs irregular. The ribbing is fine and dense Microbajocisphinctes sp. C in the posterior part of the last visible whorl. The coarse Fig. 5.3a-b primaries divide around two-thirds of flank height into Material: 4 fragmentary specimens from Tooy section two or three secondaries which cross the venter. Some of (GZN2011I-S-1-9). the ribs do not divide. Rarely, ribs divide about mid- Description: Whorl section high-ovate, primary ribs flank. The ribbing is slightly prorsiradiate on the anterior strong, coarse, distant and prorsiradiate. Most of them part of the last visible whorl. split into two prorsiradiate secondary ribs at around three- Remarks: The specimens from Iran differ from those fourth of the flank height and cross the venter. studied by Dietl (1994) with respect to possessing a flat Remarks: Microbajocisphinctes sp. C differs from flank, more irregular ribs and a more compressed shell. Microbajocisphinctes sp. A and B in having a higher The specimen GZN2011I-S-2-49 (Fig. 5.5) is smaller whorl section and distant ribs. than the other specimen and also the ribbing is denser on Stratigraphic distribution: Upper Bajocian, Garantiana the posterior part of the last visible whorl. Zone. 36

Late Bajocian–Bathonian ammonites from northeast Iran

Fig. 5 1a-b Homoeoplanulites (Parachoffatia) arkelli Mangold, specimen (GZN2011I-S-2b-46) from the Tooy section, Discus Zone. 2, 5 Homoeoplanulites (Homoeoplanulites) cf. bugesiacus (Dominjon) from the Tooy section, Discus Zone; 2 specimen GZN2011I-S-2b-47 with body chamber; 5 specimen GZN2011I-S-2b-49, Discus Zone. 3a-b Microbajocisphinctes sp. C, specimen (GZN2011I-S-1-9) from the Tooy section, Garantiana Zone. 4a-b Planisphinctes sp., specimen (GZN2011I-J-2b-12) from the Golbini section, Bathonian. 6a-b Siemiradzkia sp., specimen (GZN2011I-J-2b-10) from the Golbini section, Bathonian. 7a-b Homoeoplanulites (Homoeoplanulites) sp., microconch [m], specimen (GZN2011I-J-2c-201) from the Golbini section, Upper Bathonian. 8a-b Microbajocisphinctes sp. A, specimen (GZN2011I-S-1-7) from the Tooy section, Garantiana Zone. 9a-b Procerites sp., microconch [m], specimen (GZN2011I-J-2b-21) from the Golbini section, Bathonian. 37

Mahmoud Reza Majidifard

Stratigraphic distribution: Upper Bathonian, Discus Zone finer than the primaries. There are four weak nodes on the and Subzone and are recorded for the first time from Iran last visible whorl. Stratigraphic distribution: Bathonian. Homoeoplanulites (Homoeoplanulites) sp. [m] A Fig. 5.7a-b CONCLUDING REMARKS Material: 3 specimens from Golbini section (GZN2011I- J-201-203). The follwing Upper Bajocian families have been Dimensions (in mm): identified from the lower part of the Dalichai Formation specimen D U% H% W% PR/2 at the Golbini and Tooy sections in the eastern part of the GZN2011I -J-2c-201 33 36 29 26 19 Alborz Mountains of Iran: Parkinsoniidae (53%), GZN2011I -J-2c-202 26 42 35 23 17 Sphaeroceratidae (28%), Perisphinctidae (14%) and other GZN2011I -J-2c-203 26 42 35 31 less than (5%) (Fig. 6). During the Bathonian following family are present: families Phylloceratidae (33.4%), Description and remarks: Whorl cross-section nearly Stephanoceratidae (14.3%), Oppeliidae (14.3%), rectangular, umbilical wall vertical. On inner whorls the Morphoceratidae (14%), Perisphinctidae (12%) and ribbing is denser than on outer whorls. The strong and Tulitidae (12%) (Fig. 6). slightly prorsiradiate primaries usually divide at around three-fourth of flank height into two secondaries that LATE BAJOCIAN cross the venter. Compared with specimens studied by Mangold (1970b) the present specimens are microconchs. Stratigraphic distribution: Upper Bathonian.

Subgenus Parachoffatia Mangold, 1970b

Homoeoplanulites (Parachoffatia) arkelli Mangold, 1970b Fig. 5.1a-b 1970b Homoeoplanulites (M. Parachoffatia) arkelli sp. nov. – Mangold: 77, pl. 3, fig. 2. 1994 Homoeoplanulites (Parachoffatia) arkelli Mangold – Dietl: pl. 6, fig. 2, pl. 9, figs. 1–2.

BATHONIAN Material: 1 specimen from Tooy section (GZN2011I-S-

46). Dimensions (in mm): specimen D U% H% W% PR/2 GZN2011I -S-2b-46 60 45 31 34 19

Description: Umbilical wall vertical, venter broad- rounded. Primary ribs dense, coarse and bullate, dividing near mid-flank into two or three finer secondaries. Remarks: The Iranian specimen resembles the holotype of Mangold (1970b) but is smaller. It also differs from specimens studied by Dietl (1994) in having denser ribs on the inner whorls.

Stratigraphic distribution: Upper Bathonian, Discus Zone and Subzone Mangold (1970b) and is recorded for the Fig. 6 Relative abundances of ammonite families at the studied first time from Iran. sections from the Late Bajocian to the Bathonian.

Genus Siemiradzkia Hyatt, 1898 Palaeobiogeographically the studied ammonites belong to Sub-Mediterranean Province, as elsewhere in north and Siemiradzkia sp. central Iran (Seyed-Emami et al. 1985, 1989, 1991, 1994, Fig. 5.6a-b 1995, 1996, 2001, 2013). This is supported by the Material: 1 specimen from Golbini section (GZN2011I-J- occurrence of Submediterranean ammonites such as 10). Garantiana and Morphoceras and some cosmopolitan Description: Shell medium-sized, umbilical wall low and taxa such as Cadomites and Oxycerites. steep, venter broad. The fine but sharp primaries divide at In order to unravel the origin of the faunal elements and around two-thirds of flank height into secondaries which, their migration routes, the relationship of the ammonite together with some intercalated ribs, cross straight over fauna of Iran to that of other regions was evaluated. On the venter. Some of the ribs do not divide. Primary and the whole, at the species level, the Toarcian to Early secondary ribs are equally coarse, but the secondaries are Bajocian ammonite faunas of northern and central Iran show a close relationship to that of northwestern Europe 38

Late Bajocian–Bathonian ammonites from northeast Iran

(Seyed-Emami, 1988; Seyed-Emami et al., 2001). A Dietl, G., 1974. Zur Stratigraphie und Ammonitenfauna characteristic feature of this fauna is the scarcity of des Dogger, insbesondere des Oberbajocium der Phylloceratidae (accounting for less than 1% up to 3%) westlichen Keltiberischen Ketten (Spanien): and the absence of Lytoceratidae. Remarkably, from Late Stuttgarter Beiträge zur Naturkunde, 14: 1–21. Bathonian onward to Kimmeridgian, Phylloceratidae Dietl, G., 1978. Die heteromorphen Ammoniten des account for more than 50% of the ammonites fauna. Dogger: Stuttgarter Beiträge zur Naturkunde, 33: 1– Palaeogeographic reconstructions show the position of 97. the Iranian plate (North and Central Iran) during the Dietl, G., 1983. Die Ammoniten-Gattung Strenoceras aus Middle Jurassic time at the southern margin of Eurasia at dem südwestdeutschen Subfurcaten-Oolith a palaeo-latitude of around 30° N which rather (Bajocium, Mittl. Jura): Stuttgarter Beiträge zur corresponds to European areas (Enay & Cariou, 1997; Naturkunde 90: 1–37. Wilmsen et al., 2010). The open migration routes across Dietl, G., 1986. Die entrollten Ammoniten des pericontinental shelf seas along the northern Tethyan Schwäbischen Juras: Stuttgarter Beiträge zur margin that were approximately parallel to palaeo- Naturkunde, 4: 36–44. latitudes may explain the strong affinities of the Late Dietl, G., 1994. Der hochstetteri-Horizont – ein Bajocian – Bathonian ammonites of northern and Central Ammonitenfaunen-Horizont (Discus-Zone, Ober- Iran to those of the the Sub-Mediterranean Province. Bathonium, Dogger) aus dem Schwäbischen Jura: Stuttgarter Beiträge zur Naturkunde, 202: 1–39. ACKNOWLEDGEMENTS Dietl, G., 1983. Die Ammoniten-Gattung Strenoceras aus dem südwestdeutschen Subfurcaten-Oolith I thank Prof. F.T. Fürsich (Erlangen), who provided the (Bajocium, Mittl. Jura): Stuttgarter Beiträge zur opportunity for me to study in Würzburg and Erlangen. I Naturkunde 90: 1–37. am very much indebted to Prof. K. Seyed-Emami Dietze, V., 2000. Feinstratigraphie und Ammonitenfauna (Tehran) for helpful comments and valuable discussions. der Acris-Subzone Parkinsoni-Zone, Ober-Bajocium, I thank to the reviewers Prof. Dr. Sixto Rafael Fernandez- Mittlerer Jura (östliche Schwäbische Alb, Lopez and PD Dr. Markus Wilmsen, for useful comments Süddeutschland): Stuttgarter Beiträge zur Naturkunde, and corrections which improved my manuscript. I would 295: 1–43. like to extend my thanks to Mrs. H. Schoenig, B. Dietze, V., Dietl, G., 2006. Feinstratigraphie und Schroeder, and M. Strifler (all Würzburg) for Ammoniten-Faunenhorizonte im Ober-Bajocium und photographic work. 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