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The Gastropod Family Cancellariidae

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The Gastropod Family Cancellariidae Cainozoic Research, 4(1-2), pp. 97-108, February 2006 Revision of the gastropod family Cancellariidae from the Danian (Early Paleocene) of Fakse, Denmark Kai+Ingemann Schnetler¹ & Richard+E. Petit² 1 Fuglebakken 14, Stevnstrup, DK-8870 Langd, Denmark; e-mail: [email protected] 2 P.O. Box 30, North Myrtle Beach, SC 29597-0030, U.S.A.; e-mail: [email protected] Received 11 June 2004; revised version accepted 18 March 2005 The rich mollusc fauna fromthe MiddleDanian ofFakse, Denmark contains ofthe Cancel- very deposits eight species gastropod family lariidae. Ofthese six Unitas anderseni Unitas Plesiotriton Ad- species are previously undescribed, viz: n. sp., aliceae n. sp., stem n. sp., metula Admetula and Tatara danica A for Semitriton is des- rosenkrantzin. sp., faksensisn. sp. n. sp. lectotype biplicatus (Ravn, 1902) ignated. Key words:: Mollusca, Gastropoda,Cancellariidae, Danian, Early Paleocene, Fakse, Denmark, new species. Introduction Schilder 1928,Rosenkrantz 1960; SchnetlerefaZ. 2001). A rare part of this fauna is the gastropod family Cancellarii- dae, to which Ravn (1933)referred three species only in his monograph on the Fakse fauna. Since additionalmaterial has been collected (Spren Bo Andersen and Sten Lennart Jakobsen in 1972, Alice Rasmussen in the 1990s), a revi- sion ofthe family is possible. Inthe Rosenkrantzcollection additional specimens were located. These specimens were collected by him in 1933 and thus not treated in the revi- sion by Ravn (1933). Eight species of Cancellariidae are recognized, of which six are newly described herein, viz: Unitas anderseni n. sp., Unitas aliceae n. sp., Plesiotriton Admetula rosenkrantzi Admetula jakobseni n. sp., n. sp., faksensis n. sp. and Tatara danica n. sp. Geological setting and stratigraphy The limestone in the quarry east ofthe small town ofFakse in Eastern Zealand has been excavated for several hundred years. Fakse is situatedin the Danish-Polish Trough, which is boundedto the north the FennoscandianShield and to structures by Figure 1. Main and facies ofthe Danish area in the south the A sec- Middle Danian. Compiled by Erik Thomsen, Geologisk In- by Ringkpbing-Fyn High (Figure 1). tion mound is stitut, Aarhus Universitet. (Reproducedfrom Schnetler et through a bryozoan-coral complex exposed al, 2001). The boundary to the north to the Fennoscandian in the quarry. In the complex a vast numberofinterbedded Shield is indicated. microfacies results of may be recognized, as early diagene- sis: Bryzoan limestone, chalky limestone and coral lime- of Denmark The Danian deposits Fakse, containa very rich stone (Bemecker & Weidlich, 1990;Willumsen, 1995).For mollusc fauna (Ravn 1902a; 1902b; 1933; Nielsen 1919; a review see Surlyk & Hakansson (1999). Desor (1847) -98 - introduced the Danian Stage with the typical localities Fakse was discussed by Ravn (1933) and more recently Fakse Quarry and Stevns Klintand consideredthe stage as Schnetler (2001) noted the fauna in comparison with the the of the Cretaceous youngestpart System. For manyyears mollusc fauna ofthe younger (Selandian) Lellinge Green- the ofthe Danian has been discussed, sand. The stratigraphic position very rich faunaofthe “nose-chalk” is character- e.g. by Nielsen (1919), Ravn (1925) and Rosenkrantz ised by e. g. numerous Pleurotomariidae, trochoids, (1938), but is now generally considered as the oldest part Cerithiopsidae, Triphoridae, Ranellidae and. cypraeids. of the Paleocene. Various subdivisions ofthe Danian have Many small species are undescribed. Among the gastro- been and the Fakse has suggested, sequence at been re- pods, supposed sponge eaters were common (Pleurotom- ferred the local bruennichi echinoid to Tylocidaris Zone ariidae, Cerithiopsidae, Triphoridae), while sometrochoids (0dum 1926, Rosenkrantz 1938) ofMiddle Danian age and were herbivorous and others carnivorous (Fretter & Gra- NP3 Re- also to nannoplankton zone (Perch-Nielsen 1979). ham, 1978). Emarginula was very common (sponge cently Thomsen (1995) divided the Danish Danian into 9 and detritus eater; Fretter & Graham, 1978).The Ranelli- calcareous this nannoplankton zones. According to subdivi- dae were carnivorous (Riedel, 1995). Representatives of sion the in Fakse falls into sequence quarry his calcareous the fauna from soft bottom(e.g. Turridae) were very rare. zones 4 and 5 (Middle Danian A Eulima nannoplankton age). was rather common; this genus lives parasitic on echinoids (Muller & Strauch, 1991). Among the bivalves sessile genera were common, cementing forms like Pycno- Previous work donte and Gryphaeostrea_ or byssate, like representatives of _,t the Arcidae and Pectinidae (Muller & Strauch, 1991). Li- Ravn (1902b) recognised no Cancellariidaein the gastro- mopsis. Nuculana and other representatives ofthe infauna fauna of pod Fakse. He described Tritonium biplicatum, were rare. it referring to the Cymatiidae ( = Ranellidae) in accordance with at that time. customary placement In his 1933 paper Ravn revised the faunaand established numerous new spe- Cancellariidaein other Paleocene and Eocene faunas cies from the so-called “nose-chalk”. This type of lime- stone is an unconsolidatedcoral limestone in which origi- Cancellariidae are diverse and abundantin the Selandianof mollusc shells nally aragonitic are preserved due to a trans- Copenhagen (von Koenen, 1885; Ravn, 1939; Schnetler, formationinto calcite (Ravn, 1933; Bernecker & Weidlich, 2001). In the rich mollusc fauna eight cancellariid species 1990). He consideredthree species to belong to the Cancel- have been recorded. From the Danian and Montian of Bel- lariidae:Admete ? biplicata (Ravn, 1902),Admete (Bonel- gium Glibert (1973) mentionedfive species, all established litia) and Uxia These will be discussed in Brian & Cornet (1877) and all referred the sp. sp. species by to genus the systematic part. Unitas Palmer, 1947. Traub (1979) mentioned two rare Much of the material from the nose-chalk, published by species ofCancellariidaefrom the PaleoceneofHaunsberg, Ravn (1933) was collected by the late Professor Alfred Austria. The Paleocene of Nuussuaq, West Greenland has Rosenkrantz, Copenhagen. However, material collectedin yielded a highly diverse cancellariidfauna, consisting of 13 1933 him by was not published by Ravn, because it was species (Kollmann & Peel, 1983). Furthermore they re- collected later the same year. Rosenkrantz obviously in- ferred one specimen to a new genus, cf. Plesiotriton (p. 67, tended in his later he a publication years as arranged gas- figs 139A-B), indicating placement in Ranellidae. Accord- from the Paleocene of Fakse, to Beu & tropod species Copenhagen ing Maxwell (1987, p. 23, pi. 29, figs g, h) this and Nuussuaq, West Greenland, and hadartists make draw- specimen should most likely be referred to Turehua Mar- under his & These 1943. No of ings supervision (Kollman Peel, 1983). wick, representatives any Nuussuaq generaare hundreds of illustrations housed many prepared are in the present in the Fakse fauna with the possible exception of Rosenkrantz files in the Museum of the Geological Copenha- juvenile listed herein as Unitas sp. The Eocene of the gen. Four of the cancellariids treated in this study were Paris Basin contains a rich cancellariidfauna (Cossmann & drawn those by artists, and Rosenkrantz made a few draw- Pissarro, 1910-13; Pacaud & Le Renard, 1995).The genus with notes offive established ings accompanying species as Unitas is represented by 25 species and the genus Plesiotri- new in this Unitas anderseni, Plesiotriton steni, ton three list 11 in the paper; by species. They species genus Admetula rosenkrantzi n. Admetula and Bonellitia Jousseaume, 1887 which consider sp., faksensis they a syno- Tatara danica. nym of Admetula, a synonymy not accepted herein. The Wenz referred Plesiotriton and Semitritonto the from (1938-44) remaining genera the Fakse faunaare not represented. Beu family Ranelliidae, but & Maxwell (1987) referred The Eocene ofCotentin(North France) contains seven can- both Cancellariidae. genera to Tatara, originally (Fleming, cellariids (Cossmann & Pissarro, 1901), ofwhich six were included in the also referred referred the 1950) family Ranellidae, was to genus Uxia Jousseaume, 1887and one to the to the Cancellariidaeby Beu & Maxwell (1987). genus Sveltella Cossmann, 1889. Unitas (as Palmer, 1947 is a replacement name for Uxia Jousseaume, 1887) [non Walker, 1866]. The British Eocene contains seven species The molluscan fauna referred to the Unitas and 14 Admetula genus (as Uxia) to (as Bonellitia) fide Wrigley 1935. The mollusc fauna of the bryozoan and coral limestone at -99- Palaeogeographic conclusions Subfamily PlesiotritoninaeBeu & Maxwell, 1987 Genus PlesiotritonFischer, 1884 Type species Cancellaria volutella Lamarck, 1803 by Of the rare Cancellariidaefrom the Danian of Fakse some original designation. species demonstrate faunal affinities to the New Zealand region. According to Beu & Maxwell (1987) Tatara is known only from the Eocene to Lower Miocene of New Plesiotriton steni n. sp. Zealand and Semitriton from the Eocene ofAustralia and Figures 4, 11 the Eocene of North France (Fresville). Beu & Maxwell (1987, p. 52) questioned the generic referenceof Semitri- - Fakse Type locality quarry. ton inopinatus Cossmann & Pissarro, 1905 fromFresville but the characters ofthe Fakse species treatedherein agree Type stratum - Coral Limestone, Middle Danian, Paleo- well with the of Semitriton. Plesiotriton is type a more cene. widespread genus: Cretaceous, Paleocene and Eocene of USA, Eocene
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