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Ревизия На Многоножките От Трибус Brachyiulini

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Revision of the millipede tribe Brachyiulini ɋɈɎɂɃɋɄɂɍɇɂȼȿɊɋɂɌȿɌÄɋȼɄɅɂɆȿɇɌɈɏɊɂȾɋɄɂ³ (Diplopoda: Julida: Julidae) ȻɂɈɅɈȽɂɑȿɋɄɂɎȺɄɍɅɌȿɌ ɄȺɌȿȾɊȺÄɁɈɈɅɈȽɂəɂȺɇɌɊɈɉɈɅɈȽɂə³ by Boyan Vagalinski 3K'7KHVLV 6R¿D8QLYHUVLW\)DFXOW\RI%LRORJ\ 'HSDUWPHQWRI=RRORJ\DQG$QWKURSRORJ\ 'UDJDQ7VDQNRY%OYG6R¿D ȻɨɹɧɅɸɞɦɢɥɨɜȼɚɝɚɥɢɧɫɤɢ 7KHWKHVLVSUHVHQWVWKH¿UVWFRPSUHKHQVLYHVWXG\RIWKHPRVWO\0HGL WHUUDQHDQ MXOLG WULEH %UDFK\LXOLQL 7KLV LQFOXGHV D FRPSOHWH WD[RQRPLF FDWDORJXHRULJLQDOGHVFULSWLRQVRIVHYHUDOQHZWD[DDQGHVWDEOLVKPHQWRI VHYHUDOQHZV\QRQ\PVDSK\ORJHQHWLFDQDO\VLVDQGDELRJHRJUDSKLFDOGLV ɊȿȼɂɁɂəɇȺɆɇɈȽɈɇɈɀɄɂɌȿɈɌɌɊɂȻɍɋ FXVVLRQ7KHZRUNLVDUHVXOWRI\HDUVSHULRGRIUHVHDUFKEDVHGRQPRUH %5$&+<,8/,1, ',3/232'$-8/,'$-8/,'$( WKDQOLWHUDWXUHVRXUFHVDQGRQPDWHULDORIWKHJURXSRULJLQDWLQJIURP WKHFROOHFWLRQVRIRYHUDGR]HQPXVHXPVDQGRWKHULQVWLWXWLRQVDVZHOODV IURPIUHVKVDPSOHVFROOHFWHGIURPGLIIHUHQWUHJLRQVRI%XOJDULD3DUWRIWKH RXWFRPHVDUHDOUHDG\SXEOLVKHGLQ(QJOLVK /D]iQ\LHWDO/D]iQ\L 9DJDOLQVNLDE9DJDOLQVNLHWDO DQGWKHPDLQUHVXOWVRIWKH UHYLVLRQDUHFXUUHQWO\LQSUHSDUDWLRQIRUDIRUWKFRPLQJPRQRJUDSK 9DJD OLQVNL /i]DQ\LLQSUHS ȺȼɌɈɊȿɎȿɊȺɌ 7KHWD[RQRPLFSDUWRIWKHWKHVLVEURXJKWWRWKHUHFRJQLWLRQRIDWRWDO RIEUDFK\LXOLQHVSHFLHVEHORQJLQJWRJHQHUD7KLVPDNHVWKHWULEH RQHRIWKHPRVWSUROL¿FDQGGLYHUVHDPRQJWKH-XOLGDH0RUHRYHUSOHQW\ RIQHZVSHFLHVDUH\HWWREHGHVFULEHGPRVWO\FRPLQJIURPWKHWHUULWRULHV RI*UHHFH7XUNH\DQGWKH&DXFDVXV 7KHUHVXOWVRIWKHSK\ORJHQHWLFDQDO\VLVDOWKRXJKJHQHUDOO\LQFRQJUX ɇȺȾɂɋȿɊɌȺɐɂəɁȺɉɊɂɋɔɀȾȺɇȿɇȺɈȻɊȺɁɈȼȺɌȿɅɇȺ HQFHZLWKWKHUHYLVHGV\VWHPDWLFVRIWKH%UDFK\LXOLQLDUHRQO\SUHOLPLQDU\ ɂɇȺɍɑɇȺɋɌȿɉȿɇÄȾɈɄɌɈɊ³ DQGQHHGIXUWKHUVXSSRUWZLWKPROHFXODUPHWKRGV 7KH ELRJHRJUDSKLFDO SDWWHUQV WUDFHG LQ WKH EUDFK\LXOLQHV VXJJHVW D ɇɚɭɱɧɚɫɩɟɰɢɚɥɧɨɫɬɁɨɨɥɨɝɢɹ YHU\FOHDQPRGHORIDORSDWKULFVSHFLDWLRQUHVXOWLQJRIFRQVHFXWLYHHYHQWV RIPLJUDWLRQRQGYLFDULDQFH(DVW$QDWROLDHPHUJHGDVWKHPRVWOLNHO\RULJ LQDOWHUULWRU\RIWKHWULEH¶VDQFHVWRUV ɇɚɭɱɟɧɪɴɤɨɜɨɞɢɬɟɥ ɞɨɰɞɪɉɥɚɦɟɧȽɟɧɤɨɜɆɢɬɨɜ ɋɨɮɢɹ ɍɜɨɞ ɋɧɚɞɯɢɥɨɩɢɫɚɧɢɜɢɞɚɩɪɢɧɚɞɥɟɠɚɳɢɤɴɦɫɟɦɟɣɫɬɜɚɢɪɚɡɪɟɞɚ 6LHUZDOG %RQG*RORYDWFK ɦɧɨɝɨɧɨɠɤɢɬɟ ɤɥɚɫ'LSORSRGD ɫɚɬɪɟɬɢ- ɹɬɩɨɜɢɞɨɜɨɛɨɝɚɬɫɬɜɨɤɥɚɫɧɚ$UWKURSRGDɧɚɫɭɲɚɬɚɫɥɟɞ,QVHFWDɢ$UDFKQLGDɤɚɬɨ ɫɟɩɪɟɞɩɨɥɚɝɚɱɟɪɟɚɥɧɢɹɬɛɪɨɣɧɚɜɢɞɨɜɟɬɟɧɚɞɜɢɲɚɜɚɯɢɥȾɧɟɫɞɢɩɥɨɩɨɞɢɬɟ ɧɚɫɟɥɹɜɚɬɜɫɢɱɤɢɤɨɧɬɢɧɟɧɬɢɫɢɡɤɥɸɱɟɧɢɟɧɚȺɧɬɚɪɤɬɢɞɚɤɚɬɨɜɢɞɨɜɨɬɨɪɚɡɧɨɨ- ɛɪɚɡɢɟɟɧɚɣɝɨɥɹɦɨɜɴɜɜɥɚɠɧɢɬɟɬɪɨɩɢɱɧɢɪɚɣɨɧɢɡɚɤɨɢɬɨɫɟɩɪɟɞɩɨɥɚɝɚɱɟɟ ɢɡɜɟɫɬɧɚɟɞɜɚɦɚɥɤɚɱɚɫɬɨɬɞɢɩɥɨɩɨɞɧɚɬɚɮɚɭɧɚ ɋɟɦɟɣɫɬɜɨ-XOLGDHɧɚɪɚɡɪɟɞ-XOLGDɤɴɦɤɨɟɬɨɩɪɢɧɚɞɥɟɠɢɬɪɢɛɭɫ%UDFK\LXOLQL ɟɟɞɧɨɨɬɧɚɣɩɪɨɛɥɟɦɚɬɢɱɧɢɬɟɜɫɢɫɬɟɦɚɬɢɱɧɨɢɮɢɥɨɝɟɧɟɬɢɱɧɨɨɬɧɨɲɟɧɢɟɊɚɡ- ɩɨɡɧɚɬɢɬɟɬɪɢɛɭɫɢɜɪɚɦɤɢɬɟɧɚɫɟɦɟɣɫɬɜɨɬɨɫɚ (QJKRII ɢɥɢ 5HDG ɚɭɩɨɬɪɟɛɚɬɚɧɚɩɨɞɫɟɦɟɣɧɚɬɚɤɚɬɟɝɨɪɢɹɫɟɢɡɛɹɝɜɚɩɨɪɚɞɢɬɜɴɪɞɟɧɟɹɫɧɢɬɟ ɮɢɥɨɝɟɧɟɬɢɱɧɢ ɜɪɴɡɤɢ ɦɟɠɞɭ ɨɬɞɟɥɧɢɬɟ ɬɪɢɛɭɫɢ Ɍɜɴɪɞɟ ɦɚɥɤɨ ɫɚ ɢ ɢɡɫɥɟɞɜɚ- ɧɢɹɬɚɜɬɚɡɢɧɚɫɨɤɚɨɝɪɚɧɢɱɚɜɚɳɢɫɟɫɤɥɚɫɢɱɟɫɤɚɬɚɪɟɜɢɡɢɹɧɚ6WUDVVHU ɜɴɪɯɭ7\SKORLXOLQLɢɩɨɫɴɜɪɟɦɟɧɧɢɬɟɮɢɥɨɝɟɧɟɬɢɱɧɢɪɚɛɨɬɢɧɚ5HDG ɜɴɪɯɭ &\OLQGURLXOLQLɢɧɚ(QJKRII ɜɴɪɯɭ3DHFWRSK\OOLQLɢ&DO\SWRSK\OOLQL Ɍɪɢɛɭɫ%UDFK\LXOLQLHɟɞɢɧɨɬɧɚɣɛɨɝɚɬɢɬɟɜɪɚɦɤɢɬɟɧɚ-XOLGDHɨɛɟɞɢɧɹɜɚɳ ɨɤɨɥɨɜɢɞɚɪɚɡɩɪɨɫɬɪɚɧɟɧɢɩɪɟɞɢɦɧɨɜɘɝɨɢɡɬɨɱɧɚȿɜɪɨɩɚɆɚɥɚȺɡɢɹɢɄɚɜ- ɤɚɡȿɧɞɟɦɢɡɦɴɬɜɝɪɭɩɚɬɚɟɢɡɤɥɸɱɢɬɟɥɧɨɜɢɫɨɤɤɚɬɨɚɪɟɚɥɴɬɧɚɦɧɨɝɨɜɢɞɨɜɟɟ ɫɜɟɞɟɧɞɨɞɚɞɟɧɬɹɫɧɨɨɛɨɫɨɛɟɧɝɟɨɝɪɚɮɫɤɢɪɚɣɨɧɋɬɨɜɚɫɚɫɜɴɪɡɚɧɢɢɦɚɥɤɨɬɨ ɞɚɧɧɢɡɚɝɨɥɹɦɚɱɚɫɬɨɬɬɹɯɤɚɤɬɨɜɬɚɤɫɨɧɨɦɢɱɧɨɮɚɭɧɢɫɬɢɱɟɧɬɚɤɚɢɜɟɤɨɥɨɝɢ- ɱɟɧɚɫɩɟɤɬ %UDFK\LXOLQLɟɜɴɜɟɞɟɧɨɬ9HUKRHII ɤɚɬɨɬɚɤɫɨɧɨɦɢɱɧɢɹɬɫɴɫɬɚɜɢɦɹɫ- ɬɨɬɨ ɧɚ ɬɪɢɛɭɫɚ ɜ ɪɚɦɤɢɬɟ ɧɚ ɫɟɦɟɣɫɬɜɨɬɨ ɫɚ ɧɟɢɡɹɫɧɟɧɢ ɢ ɞɨɫɟɝɚ ɋɩɨɪɟɞ ɪɚɡ- ɥɢɱɧɢɬɟɫɯɜɚɳɚɧɢɹɬɨɣɨɛɯɜɚɳɚɦɟɠɞɭɢɪɨɞɚɤɚɬɨɪɚɡɥɢɱɢɹɬɚɫɟɞɴɥɠɚɬ ɧɚɧɟɞɨɛɪɨɬɨɞɟɮɢɧɢɪɚɧɟɢɧɟɹɫɧɢɹɬɚɤɫɨɧɨɦɢɱɟɧɪɚɧɝɧɚɝɨɥɹɦɚɱɚɫɬɨɬɬɹɯɨɬ ɟɞɧɚɫɬɪɚɧɚɢɧɚɥɢɩɫɚɬɚɧɚɫɢɧɚɩɨɦɨɪɮɧɢɛɟɥɟɡɢɯɚɪɚɤɬɟɪɢɡɢɪɚɳɢɬɪɢɛɭɫɚɤɚɬɨ ɰɹɥɨɨɬɞɪɭɝɚɇɟɩɨɦɚɥɤɢɫɚɢɩɪɨɛɥɟɦɢɬɟɨɬɧɨɦɟɧɤɥɚɬɭɪɧɨɟɫɬɟɫɬɜɨɫɜɴɪɡɚɧɢ ɫɞɴɥɝɨɝɨɞɢɲɧɚɧɟɩɪɚɜɢɥɧɚɢɩɪɟɩɥɢɬɚɳɚɫɟɭɩɨɬɪɟɛɚɧɚɧɹɤɨɥɤɨɪɨɞɚɤɚɤɬɨɢɫ ɧɚɥɢɱɢɟɧɨɧɚɪɟɞɢɰɚɫɢɧɨɧɢɦɧɢɢɥɢɧɟɜɚɥɢɞɧɨɩɪɟɞɥɨɠɟɧɢɩɨɞɪɨɞɨɜɟɧɚɧɚɣɛɨ- ɝɚɬɢɹɧɚɜɢɞɨɜɟɪɨɞMegaphyllum9HUKRHII Ⱦɨɫɟɝɚɫɢɫɬɟɦɚɬɢɤɚɬɚɢɮɢɥɨɝɟɧɢɹɬɚɧɚɬɪɢɛɭɫ%UDFK\LXOLQLɧɟɫɚɛɢɥɢɨɛɟɤɬ ɧɚɨɬɞɟɥɧɨɢɡɫɥɟɞɜɚɧɟɤɨɟɬɨɫɟɨɬɧɚɫɹɢɡɚɩɪɟɞɩɨɥɚɝɚɟɦɢɬɟɪɨɞɨɜɟɜɫɴɫɬɚɜɚɦɭɫ ɢɡɤɥɸɱɟɧɢɟɧɚAnaulaciulus3RFRFNɤɨɣɬɨɟɩɨɞɪɨɛɧɨɪɟɜɢɡɢɪɚɧɨɬ.RUVRV 9HUKRHII $WWHPV ɢ/RKPDQGHU ɢɡɜɴɪɲɜɚɬɱɚɫɬɢɱɧɢɪɟɜɢ- ɡɢɢɢɩɨɞɪɨɞɨɜɢɝɪɭɩɢɪɚɧɢɹɝɥɚɜɧɨɧɚɪɨɞMegaphyllum ɂɫɬɨɪɢɹɧɚɢɡɫɥɟɞɜɚɧɢɹɬɚ Ɂɚɧɚɱɚɥɨɧɚɬɚɤɫɨɧɨɦɢɱɧɢɬɟɢɡɫɥɟɞɜɚɧɢɹɜɴɪɯɭɬɪɢɛɭɫ%UDFK\LXOLQLɭɫɥɨɜɧɨ ɦɨɠɟɞɚɫɟɩɪɢɟɦɟɝɨɞɢɧɚɤɨɝɚɬɨ%HUOHVH ɞɟɮɢɧɢɪɚɬɢɩɨɜɢɹɪɨɞɧɚ ɬɪɢɛɭɫɚ±BrachyiulusɬɨɝɚɜɚɫɴɫɫɬɚɬɭɬɧɚɩɨɞɪɨɞɧɚɪɨɞJulus/LQQDHXVɋɚɦɚɬɚ ɞɟɮɢɧɢɰɢɹɟɫɴɜɫɟɦɤɪɚɬɤɚɛɚɡɢɪɚɧɚɢɡɰɹɥɨɧɚɨɫɨɛɟɧɨɫɬɢɧɚɝɨɧɨɩɨɞɢɬɟ Ɉɫɧɨɜɨɩɨɥɚɝɚɳɚɪɨɥɹɡɚɫɴɜɪɟɦɟɧɧɚɬɚɫɢɫɬɟɦɚɬɢɤɚɧɚɦɧɨɝɨɧɨɠɤɢɬɟɜɱɚɫɬ- ɧɨɫɬɬɚɡɢɧɚɸɥɢɞɢɬɟɢɬɪɢɛɭɫ%UDFK\LXOLQLɢɦɚɬ.:9HUKRHIIɢ&*$WWHPVɢ ɞɜɚɦɚɬɚɪɚɛɨɬɢɥɢɨɬɤɪɚɹɧɚɬɢɞɨɨɤɨɥɨɫɪɟɞɚɬɚɧɚɬɢɜɟɤȿɞɧɚɨɬɪɚɧɧɢɬɟ ɩɭɛɥɢɤɚɰɢɢɧɚɩɴɪɜɢɹɚɜɬɨɪ 9HUKRHIID Hɩɨɫɜɟɬɟɧɚɢɦɟɧɧɨɧɚ-XOLGDHɢɩɪɟɞ- ɫɬɚɜɥɹɜɚɩɴɪɜɢɹɬɫɟɪɢɨɡɟɧɨɩɢɬɡɚɢɡɹɫɧɹɜɚɧɟɧɚɪɨɞɫɬɜɟɧɢɬɟɜɪɴɡɤɢɜɫɟɦɟɣɫɬɜɨ- ɬɨɂɡɜɟɫɬɧɢɬɟɞɨɨɧɡɢɦɨɦɟɧɬ%UDFK\LXOLQLɬɨɣɪɚɡɩɪɟɞɟɥɹɜɞɜɚɩɨɞɪɨɞɚɧɚɪɟɞɟ- 1 ɮɢɧɢɪɚɧɢɹɨɬɧɟɝɨɪɨɞIulus±AnoploiulusɢChromatoiulusȼɫɴɳɚɬɚɝɨɞɢɧɚ9HUKRHII E ɨɩɢɫɜɚɪɨɞMegaphyllumɡɚɟɞɧɨɫɬɢɩɨɜɢɹɦɭɜɢɞ±Megaphyllum projectum ɆɚɥɤɨɫɥɟɞɬɨɜɚɜɫɢɱɤɢɜɢɞɨɜɟɞɨɬɨɝɚɜɚɫɩɚɞɚɳɢɤɴɦɩɨɞɪɨɞChromatoiulusɬɨɣ ɩɪɟɯɜɴɪɥɹ ɤɴɦ Megaphyllum 9HUKRHII 9HUKRHII ɞɟɮɢɧɢɪɚ ɧɚɧɨɜɨ 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ɡɚɪɚɞɢɤɨɟɬɨɟɛɢɥɤɪɢɬɢɤɭɜɚɧɨɬɫɜɨɢɬɟɤɨɥɟɝɢ 3RFRFN$WWHPVɜɠ ɫɴɳɨ/D]iQ\L .RUVyV ȼɫɴɳɢɹɩɟɪɢɨɞɫɚɨɩɢɫɚɧɢɞɜɚɧɨɜɢɪɨɞɚ%UDFK\LXOLQL±Anaulaciulus3RFRFN ɢ Rhamphidoiulus $WWHPV ɧɚ ɛɚɡɚɬɚ ɧɚ ɞɜɚ ɧɨɜɢ ɜɢɞɚ ɫɴɨɬɜɟɬɧɨ A. paludicolaɢR. bujukderensis 3RFRFN$WWHPV Ɉɬɞɨ$WWHPVɨɩɢɫɜɚɞɪɭɝɢɩɪɟɞɫɬɚɜɢɬɟɥɢɧɚɫɴɳɢɹɪɨɞɤɚɤɬɨɢ Brachyiulus varibolinusɩɪɟɡ Ɍɪɢɛɭɫ%UDFK\LXOLQLɟɜɴɜɟɞɟɧɤɚɬɨɬɚɤɫɨɧɨɦɢɱɧɚɤɚɬɟɝɨɪɢɹɨɬ9HUKRHII Ⱦɟɮɢɧɢɰɢɹɬɚ ɟ ɧɚɩɪɚɜɟɧɚ ɩɨɫɪɟɞɫɬɜɨɦ ɤɥɸɱ ɧɚ ɢɡɜɟɫɬɧɢɬɟ ɞɨɬɨɝɚɜɚ ɸɥɢɞɢ ɇɚ ɫɥɟɞɜɚɳɚɬɚ ɝɨɞɢɧɚ ɚɜɬɨɪɴɬ ɩɭɛɥɢɤɭɜɚ ɦɨɧɨɝɪɚɮɢɹ ɜɴɪɯɭ ɧɹɤɨɥɤɨ ɝɪɭɩɢ ɦɧɨ- ɝɨɧɨɠɤɢɫɪɟɞɤɨɢɬɨɢɫɟɦɟɣɫɬɜɨ-XOLGDH 9HUKRHII Ɉɬɞɟɥɟɧɨɟɫɩɟɰɢɚɥɧɨ ɜɧɢɦɚɧɢɟɧɚ³%UDFK\LXOXV%HUO9HUK´ɤɚɬɨɞɜɚɨɬɞɨɬɨɝɚɜɚɲɧɢɬɟɦɭɩɨɞɪɨɞɨɜɟ± MicrobrachyiulusɢHeteroiulus±ɫɚɢɡɞɢɝɧɚɬɢɜɨɬɞɟɥɧɢɪɨɞɨɜɟ $WWHPV ɩɭɛɥɢɤɭɜɚ ɨɛɨɛɳɚɜɚɳɚ ɦɨɧɨɝɪɚɮɢɹ ɜɴɪɯɭ ɦɧɨɝɨɧɨɠɤɢɬɟ ɧɚ ɉɚɥɟɚɪɤɬɢɤɚȼɧɟɹɟɧɚɩɪɚɜɟɧɚɩɴɪɜɚɬɚɪɟɜɢɡɢɹɧɚɪɨɞMegaphyllum Brachyiulus sensu9HUKRHII ɱɢɢɬɨɜɢɞɨɜɟɩɨɨɧɨɜɚɜɪɟɦɟɜɟɱɟɫɚɧɚɞɯɜɴɪɥɢɥɢɋɚɦɚɬɚɪɟ- ɜɢɡɢɹɨɛɯɜɚɳɚɨɤɨɥɨôɨɬɬɹɯɤɨɢɬɨɫɚɩɪɟɞɫɬɚɜɟɧɢɫɧɨɜɢɪɢɫɭɧɤɢɨɩɢɫɚɬɟɥɧɢ ɞɨɩɴɥɧɟɧɢɹɢɨɩɪɟɞɟɥɢɬɟɥɟɧɤɥɸɱ Ɍɪɢ ɝɨɞɢɧɢ ɩɨɤɴɫɧɨ ɢɡɥɢɡɚ ɩɨɪɟɞɧɚɬɚ ɱɚɫɬ ɨɬ ɦɨɧɨɝɪɚɮɢɹɬɚ ɧɚ 9HUKRHII ɡɚ ɦɧɨɝɨɧɨɠɤɢɬɟ ɤɥɚɫ 'LSORSRGD Ɍɪɢɛɭɫɢɬɟ %UDFK\LXOLQL ɢ 8QFLJHULQL 2QFRLXOLQL DXFW 9HUKRHII ɨɛɟɞɢɧɹɜɚ ɜ ɩɨɞɫɟɦɟɣɫɬɜɨ %UDFK\LXOLQDH ɧɚ ɫɟɦɟɣɫɬɜɨ -XOLGDHȼɞɟɮɢɧɢɰɢɢɬɟɧɚɪɨɞɨɜɟɬɟɬɨɣɫɟɩɪɢɞɴɪɠɚɤɴɦɩɪɟɞɢɲɧɢɬɟɫɢɬɪɭɞɨɜɟ ɫɦɚɥɤɢɞɨɩɴɥɧɟɧɢɹɈɩɢɫɚɧɢɫɚɞɜɚɧɨɜɢɩɨɞɪɨɞɚɧɚMegaphyllum Brachyiulus sensu9HUKRHII ±ByzantorhopalumɢSyriobrachyiulus /RKPDQGHU ɩɭɛɥɢɤɭɜɚɫɬɚɬɢɹɜɴɪɯɭɞɢɩɥɨɩɨɞɧɚɬɚɮɚɭɧɚɧɚɄɚɜɤɚɡȼ ɧɟɹɬɨɣɨɩɢɫɜɚɧɨɜɜɢɞɫɩɚɞɚɳɤɴɦɧɨɜɪɨɞ%UDFK\LXOLQL±Grusiniulus redikorzevi ɤɚɤɬɨɢɧɹɤɨɥɤɨɧɨɜɢ ɩɨɞ ɜɢɞɚMegaphyllum ChromatoiulusDXFW ɩɪɢɧɚɞɥɟɠɚ- ɳɢɤɴɦɧɨɜɨɞɟɮɢɧɢɪɚɧɢɹɜɫɴɳɚɬɚɩɭɛɥɢɤɚɰɢɹɩɨɞɪɨɞOmobrachyiulusɢɟɞɢɧɜɢɞ ɤɴɦ ɆHJDSK\OOXP V VWU Chromatoiulus V VWU DXFW Ⱦɟɮɢɧɢɪɚɧɢ ɫɚ ɢ ɬɪɢ ɞɪɭ- ɝɢɩɨɞɪɨɞɚɧɚɫɴɳɢɹɪɨɞɩɨɜɟɱɟɨɩɢɫɚɧɢɜɢɞɨɜɟColchiobrachyiulusɫɟɞɢɧɫɬɜɟɧ ɜɢɞ M. dioscoriadis /LJQDX ArmeniobrachyiulusɢDonbrachyiulus ɫɴɳɨ ɦɨɧɨɬɢɩɧɢɫɴɫɫɴɨɬɜɟɬɧɢɜɢɞɨɜɟM. sevangense /RKPDQGHU ɢɆURVVLFXP 7LPRWKHHZ Ⱥɜɬɨɪɴɬɪɚɡɝɥɟɠɞɚɬɪɢɛɭɫ%UDFK\LXOLQLɤɚɬɨɟɞɢɧɫɬɜɟɧɜɪɚɦɤɢ- ɬɟɧɚɩɨɞɫɟɦɟɣɫɬɜɨ%UDFK\LXOLQDHɌɪɢɛɭɫ8QFLJHULQDHɟɨɛɟɞɢɧɟɧɫ/HXFRJHRUJLLQLɜ ɩɨɞɫɟɦɟɣɫɬɜɨ8QFLJHULQDHɉɨɞɪɨɛɧɢɬɟɨɩɢɫɚɧɢɹɩɨɞɤɪɟɩɟɧɢɫɩɪɟɰɢɡɧɢɞɟɬɚɣɥ- ɧɢɪɢɫɭɧɤɢɩɪɚɜɹɬɬɚɡɢɪɚɛɨɬɚɧɚ/RKPDQGHUɨɫɨɛɟɧɨɰɟɧɧɚ ȼ ɟɞɧɚ ɨɬ ɫɜɨɢɬɟ ɩɭɛɥɢɤɚɰɢɢ ɜɴɪɯɭ ɦɧɨɝɨɧɨɠɤɢɬɟ ɧɚ Ȼɴɥɝɚɪɢɹ 9HUKRHII ɨɩɢɫɜɚ ɦɨɧɨɬɢɩɧɢɹ ɪɨɞ Balkanophoenix ɫ ɬɢɩɨɜ ɜɢɞ B. borisi ɩɪɢɱɢɫɥɹ- ɜɚɣɤɢ ɝɨ ɤɴɦ ɬɪɢɛɭɫ %UDFK\LXOLQL ȼɢɞɴɬ ɟ ɨɩɢɫɚɧ ɨɬ ɞɜɟ ɧɚɯɨɞɢɳɚ ± ɪɚɣɨɧɚ ɧɚ ɝɚɪɚɅɚɤɚɬɧɢɤɢɩɨɪɟɱɢɟɬɨɧɚɪɑɭɤɭɪɫɤɚɜɪɚɣɨɧɚɧɚȼɟɥɢɧɝɪɚɞɎɚɤɬɴɬɱɟɧɢ- ɤɨɝɚ ɧɟ ɟ ɫɴɨɛɳɚɜɚɧ ɨɬɧɨɜɨ ɫɥɟɞ ɞɚɬɚɬɚ ɧɚ ɨɩɢɫɜɚɧɟɬɨ ɦɭ ɩɨɤɚɡɜɚ ɱɟ ɩɨ ɜɫɹɤɚ ɜɟɪɨɹɬɧɨɫɬ ɫɬɚɜɚ ɞɭɦɚ ɡɚ ɢɡɜɴɧɪɟɞɧɨ ɪɹɞɴɤ ɟɧɞɟɦɢɬ ȼ ɩɭɛɥɢɤɚɰɢɹɬɚ ɟ ɨɩɢɫɚɧ ɢ Brachyiulus (Byzantorhopalum) strandschanus ɞɧɟɫ ɜɚɥɢɞɟɧ ɤɚɬɨ Megaphyllum rossicum strandschanum /RKPDQGHU ɨɩɢɫɜɚɧɨɜɩɨɞɜɢɞɧɚMegaphyllum annulatum $WWHPV ±M ChromatoiulusDXFW annulatum samsunensisɩɨɤɨɣɬɨɞɟɮɢɧɢɪɚɧɨɜɩɨɞɪɨɞ PontobrachyiulusȾɨɞɧɟɫɬɨɣɨɫɬɚɜɚɦɨɧɨɬɢɩɟɧɧɨɫɜɨɟɨɛɪɚɡɧɚɬɚɦɨɪɮɨɥɨɝɢɹɧɚ ɜɢɞɚɞɚɜɚɞɨɛɪɢɨɫɧɨɜɚɧɢɹɩɨɞɪɨɞɴɬɞɚÄɨɰɟɥɟɟ´ɢɫɥɟɞɟɜɟɧɬɭɚɥɧɚɬɚɪɟɜɢɡɢɹɧɚ ɪɨɞMegaphyllum $WWHPV ɩɪɟɞɫɬɚɜɹɨɛɨɛɳɚɜɚɳɬɪɭɞɜɴɪɯɭɫɢɫɬɟɦɚɬɢɤɚɬɚɧɚɫɟɦɟɣɫɬɜɨ -XOLGDH ɜ ɤɨɣɬɨ ɨɬɞɟɥɹ ɫɩɟɰɢɚɥɧɨ ɜɧɢɦɚɧɢɟ ɧɚ ɪɨɞ Megaphyllum Chromatoiulus DXFW ɤɨɦɟɧɬɢɪɚɣɤɢ ɜɚɥɢɞɧɨɫɬɬɚ ɧɚ ɜɫɟɤɢ ɟɞɢɧ ɨɬ ɩɨɞɪɨɞɨɜɟɬɟ ɦɭ ȼ ɨɩɢɬ ɡɚ ɢɡɝɪɚɠɞɚɧɟ ɧɚ ɟɞɧɚ ɩɨɫɬɪɨɣɧɚ ɢ ɟɫɬɟɫɬɜɟɧɚ ɤɥɚɫɢɮɢɤɚɰɢɹ ɚɜɬɨɪɴɬ ɪɟɞɟɮɢɧɢ- ɪɚ ɢɡɜɟɫɬɧɢɬɟ ɞɨɬɨɝɚɜɚ ɩɨɞɪɨɞɨɜɟ ɢ ɫɚɦ ɜɴɜɟɠɞɚ ɨɳɟ ɬɪɢ Diaxylus Dittozusɢ Phauloiulus 9HUKRHII ɩɭɛɥɢɤɭɜɚɫɬɚɬɢɹɜɴɪɯɭɦɢɪɢɚɩɨɞɧɚɬɚɮɚɭɧɚɧɚɌɭɪɰɢɹȼɧɟɹ ɬɨɣɞɟɮɢɧɢɪɚɧɨɜɩɨɞɪɨɞɧɚMegaphyllum±Anatolicusɤɴɦɤɨɣɬɨɩɪɢɱɢɫɥɹɜɚɧɨ- ɜɨɨɩɢɫɚɧɢɹɜɫɴɳɚɬɚɩɭɛɥɢɤɚɰɢɹMegaphyllum BrachyiulusDXFW ulunumɡɚɟɞɧɨ ɫɨɳɟɞɜɚɩɨɪɚɧɨɨɩɢɫɚɧɢɨɬɧɟɝɨɜɢɞɚ±M. kosswigiɢM. anatolicumȾɜɟɝɨɞɢɧɢ ɩɨɤɴɫɧɨɚɜɬɨɪɴɬɩɭɛɥɢɤɭɜɚɧɨɜɚɫɬɚɬɢɹɡɚɫɹɝɚɳɚɬɭɪɫɤɢɬɟɦɧɨɝɨɧɨɠɤɢ 9HUKRHII
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