Ревизия На Многоножките От Трибус Brachyiulini

Ревизия На Многоножките От Трибус Brachyiulini

Revision of the millipede tribe Brachyiulini ɋɈɎɂɃɋɄɂɍɇɂȼȿɊɋɂɌȿɌÄɋȼɄɅɂɆȿɇɌɈɏɊɂȾɋɄɂ³ (Diplopoda: Julida: Julidae) ȻɂɈɅɈȽɂɑȿɋɄɂɎȺɄɍɅɌȿɌ ɄȺɌȿȾɊȺÄɁɈɈɅɈȽɂəɂȺɇɌɊɈɉɈɅɈȽɂə³ by Boyan Vagalinski 3K'7KHVLV 6R¿D8QLYHUVLW\)DFXOW\RI%LRORJ\ 'HSDUWPHQWRI=RRORJ\DQG$QWKURSRORJ\ 'UDJDQ7VDQNRY%OYG6R¿D ȻɨɹɧɅɸɞɦɢɥɨɜȼɚɝɚɥɢɧɫɤɢ 7KHWKHVLVSUHVHQWVWKH¿UVWFRPSUHKHQVLYHVWXG\RIWKHPRVWO\0HGL WHUUDQHDQ MXOLG WULEH %UDFK\LXOLQL 7KLV LQFOXGHV D FRPSOHWH WD[RQRPLF FDWDORJXHRULJLQDOGHVFULSWLRQVRIVHYHUDOQHZWD[DDQGHVWDEOLVKPHQWRI VHYHUDOQHZV\QRQ\PVDSK\ORJHQHWLFDQDO\VLVDQGDELRJHRJUDSKLFDOGLV ɊȿȼɂɁɂəɇȺɆɇɈȽɈɇɈɀɄɂɌȿɈɌɌɊɂȻɍɋ FXVVLRQ7KHZRUNLVDUHVXOWRI\HDUVSHULRGRIUHVHDUFKEDVHGRQPRUH %5$&+<,8/,1, ',3/232'$-8/,'$-8/,'$( WKDQOLWHUDWXUHVRXUFHVDQGRQPDWHULDORIWKHJURXSRULJLQDWLQJIURP WKHFROOHFWLRQVRIRYHUDGR]HQPXVHXPVDQGRWKHULQVWLWXWLRQVDVZHOODV IURPIUHVKVDPSOHVFROOHFWHGIURPGLIIHUHQWUHJLRQVRI%XOJDULD3DUWRIWKH RXWFRPHVDUHDOUHDG\SXEOLVKHGLQ(QJOLVK /D]iQ\LHWDO/D]iQ\L 9DJDOLQVNLDE9DJDOLQVNLHWDO DQGWKHPDLQUHVXOWVRIWKH UHYLVLRQDUHFXUUHQWO\LQSUHSDUDWLRQIRUDIRUWKFRPLQJPRQRJUDSK 9DJD OLQVNL /i]DQ\LLQSUHS ȺȼɌɈɊȿɎȿɊȺɌ 7KHWD[RQRPLFSDUWRIWKHWKHVLVEURXJKWWRWKHUHFRJQLWLRQRIDWRWDO RIEUDFK\LXOLQHVSHFLHVEHORQJLQJWRJHQHUD7KLVPDNHVWKHWULEH RQHRIWKHPRVWSUROL¿FDQGGLYHUVHDPRQJWKH-XOLGDH0RUHRYHUSOHQW\ RIQHZVSHFLHVDUH\HWWREHGHVFULEHGPRVWO\FRPLQJIURPWKHWHUULWRULHV RI*UHHFH7XUNH\DQGWKH&DXFDVXV 7KHUHVXOWVRIWKHSK\ORJHQHWLFDQDO\VLVDOWKRXJKJHQHUDOO\LQFRQJUX ɇȺȾɂɋȿɊɌȺɐɂəɁȺɉɊɂɋɔɀȾȺɇȿɇȺɈȻɊȺɁɈȼȺɌȿɅɇȺ HQFHZLWKWKHUHYLVHGV\VWHPDWLFVRIWKH%UDFK\LXOLQLDUHRQO\SUHOLPLQDU\ ɂɇȺɍɑɇȺɋɌȿɉȿɇÄȾɈɄɌɈɊ³ DQGQHHGIXUWKHUVXSSRUWZLWKPROHFXODUPHWKRGV 7KH ELRJHRJUDSKLFDO SDWWHUQV WUDFHG LQ WKH EUDFK\LXOLQHV VXJJHVW D ɇɚɭɱɧɚɫɩɟɰɢɚɥɧɨɫɬɁɨɨɥɨɝɢɹ YHU\FOHDQPRGHORIDORSDWKULFVSHFLDWLRQUHVXOWLQJRIFRQVHFXWLYHHYHQWV RIPLJUDWLRQRQGYLFDULDQFH(DVW$QDWROLDHPHUJHGDVWKHPRVWOLNHO\RULJ LQDOWHUULWRU\RIWKHWULEH¶VDQFHVWRUV ɇɚɭɱɟɧɪɴɤɨɜɨɞɢɬɟɥ ɞɨɰɞɪɉɥɚɦɟɧȽɟɧɤɨɜɆɢɬɨɜ ɋɨɮɢɹ ɍɜɨɞ ɋɧɚɞɯɢɥɨɩɢɫɚɧɢɜɢɞɚɩɪɢɧɚɞɥɟɠɚɳɢɤɴɦɫɟɦɟɣɫɬɜɚɢɪɚɡɪɟɞɚ 6LHUZDOG %RQG*RORYDWFK ɦɧɨɝɨɧɨɠɤɢɬɟ ɤɥɚɫ'LSORSRGD ɫɚɬɪɟɬɢ- ɹɬɩɨɜɢɞɨɜɨɛɨɝɚɬɫɬɜɨɤɥɚɫɧɚ$UWKURSRGDɧɚɫɭɲɚɬɚɫɥɟɞ,QVHFWDɢ$UDFKQLGDɤɚɬɨ ɫɟɩɪɟɞɩɨɥɚɝɚɱɟɪɟɚɥɧɢɹɬɛɪɨɣɧɚɜɢɞɨɜɟɬɟɧɚɞɜɢɲɚɜɚɯɢɥȾɧɟɫɞɢɩɥɨɩɨɞɢɬɟ ɧɚɫɟɥɹɜɚɬɜɫɢɱɤɢɤɨɧɬɢɧɟɧɬɢɫɢɡɤɥɸɱɟɧɢɟɧɚȺɧɬɚɪɤɬɢɞɚɤɚɬɨɜɢɞɨɜɨɬɨɪɚɡɧɨɨ- ɛɪɚɡɢɟɟɧɚɣɝɨɥɹɦɨɜɴɜɜɥɚɠɧɢɬɟɬɪɨɩɢɱɧɢɪɚɣɨɧɢɡɚɤɨɢɬɨɫɟɩɪɟɞɩɨɥɚɝɚɱɟɟ ɢɡɜɟɫɬɧɚɟɞɜɚɦɚɥɤɚɱɚɫɬɨɬɞɢɩɥɨɩɨɞɧɚɬɚɮɚɭɧɚ ɋɟɦɟɣɫɬɜɨ-XOLGDHɧɚɪɚɡɪɟɞ-XOLGDɤɴɦɤɨɟɬɨɩɪɢɧɚɞɥɟɠɢɬɪɢɛɭɫ%UDFK\LXOLQL ɟɟɞɧɨɨɬɧɚɣɩɪɨɛɥɟɦɚɬɢɱɧɢɬɟɜɫɢɫɬɟɦɚɬɢɱɧɨɢɮɢɥɨɝɟɧɟɬɢɱɧɨɨɬɧɨɲɟɧɢɟɊɚɡ- ɩɨɡɧɚɬɢɬɟɬɪɢɛɭɫɢɜɪɚɦɤɢɬɟɧɚɫɟɦɟɣɫɬɜɨɬɨɫɚ (QJKRII ɢɥɢ 5HDG ɚɭɩɨɬɪɟɛɚɬɚɧɚɩɨɞɫɟɦɟɣɧɚɬɚɤɚɬɟɝɨɪɢɹɫɟɢɡɛɹɝɜɚɩɨɪɚɞɢɬɜɴɪɞɟɧɟɹɫɧɢɬɟ ɮɢɥɨɝɟɧɟɬɢɱɧɢ ɜɪɴɡɤɢ ɦɟɠɞɭ ɨɬɞɟɥɧɢɬɟ ɬɪɢɛɭɫɢ Ɍɜɴɪɞɟ ɦɚɥɤɨ ɫɚ ɢ ɢɡɫɥɟɞɜɚ- ɧɢɹɬɚɜɬɚɡɢɧɚɫɨɤɚɨɝɪɚɧɢɱɚɜɚɳɢɫɟɫɤɥɚɫɢɱɟɫɤɚɬɚɪɟɜɢɡɢɹɧɚ6WUDVVHU ɜɴɪɯɭ7\SKORLXOLQLɢɩɨɫɴɜɪɟɦɟɧɧɢɬɟɮɢɥɨɝɟɧɟɬɢɱɧɢɪɚɛɨɬɢɧɚ5HDG ɜɴɪɯɭ &\OLQGURLXOLQLɢɧɚ(QJKRII ɜɴɪɯɭ3DHFWRSK\OOLQLɢ&DO\SWRSK\OOLQL Ɍɪɢɛɭɫ%UDFK\LXOLQLHɟɞɢɧɨɬɧɚɣɛɨɝɚɬɢɬɟɜɪɚɦɤɢɬɟɧɚ-XOLGDHɨɛɟɞɢɧɹɜɚɳ ɨɤɨɥɨɜɢɞɚɪɚɡɩɪɨɫɬɪɚɧɟɧɢɩɪɟɞɢɦɧɨɜɘɝɨɢɡɬɨɱɧɚȿɜɪɨɩɚɆɚɥɚȺɡɢɹɢɄɚɜ- ɤɚɡȿɧɞɟɦɢɡɦɴɬɜɝɪɭɩɚɬɚɟɢɡɤɥɸɱɢɬɟɥɧɨɜɢɫɨɤɤɚɬɨɚɪɟɚɥɴɬɧɚɦɧɨɝɨɜɢɞɨɜɟɟ ɫɜɟɞɟɧɞɨɞɚɞɟɧɬɹɫɧɨɨɛɨɫɨɛɟɧɝɟɨɝɪɚɮɫɤɢɪɚɣɨɧɋɬɨɜɚɫɚɫɜɴɪɡɚɧɢɢɦɚɥɤɨɬɨ ɞɚɧɧɢɡɚɝɨɥɹɦɚɱɚɫɬɨɬɬɹɯɤɚɤɬɨɜɬɚɤɫɨɧɨɦɢɱɧɨɮɚɭɧɢɫɬɢɱɟɧɬɚɤɚɢɜɟɤɨɥɨɝɢ- ɱɟɧɚɫɩɟɤɬ %UDFK\LXOLQLɟɜɴɜɟɞɟɧɨɬ9HUKRHII ɤɚɬɨɬɚɤɫɨɧɨɦɢɱɧɢɹɬɫɴɫɬɚɜɢɦɹɫ- ɬɨɬɨ ɧɚ ɬɪɢɛɭɫɚ ɜ ɪɚɦɤɢɬɟ ɧɚ ɫɟɦɟɣɫɬɜɨɬɨ ɫɚ ɧɟɢɡɹɫɧɟɧɢ ɢ ɞɨɫɟɝɚ ɋɩɨɪɟɞ ɪɚɡ- ɥɢɱɧɢɬɟɫɯɜɚɳɚɧɢɹɬɨɣɨɛɯɜɚɳɚɦɟɠɞɭɢɪɨɞɚɤɚɬɨɪɚɡɥɢɱɢɹɬɚɫɟɞɴɥɠɚɬ ɧɚɧɟɞɨɛɪɨɬɨɞɟɮɢɧɢɪɚɧɟɢɧɟɹɫɧɢɹɬɚɤɫɨɧɨɦɢɱɟɧɪɚɧɝɧɚɝɨɥɹɦɚɱɚɫɬɨɬɬɹɯɨɬ ɟɞɧɚɫɬɪɚɧɚɢɧɚɥɢɩɫɚɬɚɧɚɫɢɧɚɩɨɦɨɪɮɧɢɛɟɥɟɡɢɯɚɪɚɤɬɟɪɢɡɢɪɚɳɢɬɪɢɛɭɫɚɤɚɬɨ ɰɹɥɨɨɬɞɪɭɝɚɇɟɩɨɦɚɥɤɢɫɚɢɩɪɨɛɥɟɦɢɬɟɨɬɧɨɦɟɧɤɥɚɬɭɪɧɨɟɫɬɟɫɬɜɨɫɜɴɪɡɚɧɢ ɫɞɴɥɝɨɝɨɞɢɲɧɚɧɟɩɪɚɜɢɥɧɚɢɩɪɟɩɥɢɬɚɳɚɫɟɭɩɨɬɪɟɛɚɧɚɧɹɤɨɥɤɨɪɨɞɚɤɚɤɬɨɢɫ ɧɚɥɢɱɢɟɧɨɧɚɪɟɞɢɰɚɫɢɧɨɧɢɦɧɢɢɥɢɧɟɜɚɥɢɞɧɨɩɪɟɞɥɨɠɟɧɢɩɨɞɪɨɞɨɜɟɧɚɧɚɣɛɨ- ɝɚɬɢɹɧɚɜɢɞɨɜɟɪɨɞMegaphyllum9HUKRHII Ⱦɨɫɟɝɚɫɢɫɬɟɦɚɬɢɤɚɬɚɢɮɢɥɨɝɟɧɢɹɬɚɧɚɬɪɢɛɭɫ%UDFK\LXOLQLɧɟɫɚɛɢɥɢɨɛɟɤɬ ɧɚɨɬɞɟɥɧɨɢɡɫɥɟɞɜɚɧɟɤɨɟɬɨɫɟɨɬɧɚɫɹɢɡɚɩɪɟɞɩɨɥɚɝɚɟɦɢɬɟɪɨɞɨɜɟɜɫɴɫɬɚɜɚɦɭɫ ɢɡɤɥɸɱɟɧɢɟɧɚAnaulaciulus3RFRFNɤɨɣɬɨɟɩɨɞɪɨɛɧɨɪɟɜɢɡɢɪɚɧɨɬ.RUVRV 9HUKRHII $WWHPV ɢ/RKPDQGHU ɢɡɜɴɪɲɜɚɬɱɚɫɬɢɱɧɢɪɟɜɢ- ɡɢɢɢɩɨɞɪɨɞɨɜɢɝɪɭɩɢɪɚɧɢɹɝɥɚɜɧɨɧɚɪɨɞMegaphyllum ɂɫɬɨɪɢɹɧɚɢɡɫɥɟɞɜɚɧɢɹɬɚ Ɂɚɧɚɱɚɥɨɧɚɬɚɤɫɨɧɨɦɢɱɧɢɬɟɢɡɫɥɟɞɜɚɧɢɹɜɴɪɯɭɬɪɢɛɭɫ%UDFK\LXOLQLɭɫɥɨɜɧɨ ɦɨɠɟɞɚɫɟɩɪɢɟɦɟɝɨɞɢɧɚɤɨɝɚɬɨ%HUOHVH ɞɟɮɢɧɢɪɚɬɢɩɨɜɢɹɪɨɞɧɚ ɬɪɢɛɭɫɚ±BrachyiulusɬɨɝɚɜɚɫɴɫɫɬɚɬɭɬɧɚɩɨɞɪɨɞɧɚɪɨɞJulus/LQQDHXVɋɚɦɚɬɚ ɞɟɮɢɧɢɰɢɹɟɫɴɜɫɟɦɤɪɚɬɤɚɛɚɡɢɪɚɧɚɢɡɰɹɥɨɧɚɨɫɨɛɟɧɨɫɬɢɧɚɝɨɧɨɩɨɞɢɬɟ Ɉɫɧɨɜɨɩɨɥɚɝɚɳɚɪɨɥɹɡɚɫɴɜɪɟɦɟɧɧɚɬɚɫɢɫɬɟɦɚɬɢɤɚɧɚɦɧɨɝɨɧɨɠɤɢɬɟɜɱɚɫɬ- ɧɨɫɬɬɚɡɢɧɚɸɥɢɞɢɬɟɢɬɪɢɛɭɫ%UDFK\LXOLQLɢɦɚɬ.:9HUKRHIIɢ&*$WWHPVɢ ɞɜɚɦɚɬɚɪɚɛɨɬɢɥɢɨɬɤɪɚɹɧɚɬɢɞɨɨɤɨɥɨɫɪɟɞɚɬɚɧɚɬɢɜɟɤȿɞɧɚɨɬɪɚɧɧɢɬɟ ɩɭɛɥɢɤɚɰɢɢɧɚɩɴɪɜɢɹɚɜɬɨɪ 9HUKRHIID Hɩɨɫɜɟɬɟɧɚɢɦɟɧɧɨɧɚ-XOLGDHɢɩɪɟɞ- ɫɬɚɜɥɹɜɚɩɴɪɜɢɹɬɫɟɪɢɨɡɟɧɨɩɢɬɡɚɢɡɹɫɧɹɜɚɧɟɧɚɪɨɞɫɬɜɟɧɢɬɟɜɪɴɡɤɢɜɫɟɦɟɣɫɬɜɨ- ɬɨɂɡɜɟɫɬɧɢɬɟɞɨɨɧɡɢɦɨɦɟɧɬ%UDFK\LXOLQLɬɨɣɪɚɡɩɪɟɞɟɥɹɜɞɜɚɩɨɞɪɨɞɚɧɚɪɟɞɟ- 1 ɮɢɧɢɪɚɧɢɹɨɬɧɟɝɨɪɨɞIulus±AnoploiulusɢChromatoiulusȼɫɴɳɚɬɚɝɨɞɢɧɚ9HUKRHII E ɨɩɢɫɜɚɪɨɞMegaphyllumɡɚɟɞɧɨɫɬɢɩɨɜɢɹɦɭɜɢɞ±Megaphyllum projectum ɆɚɥɤɨɫɥɟɞɬɨɜɚɜɫɢɱɤɢɜɢɞɨɜɟɞɨɬɨɝɚɜɚɫɩɚɞɚɳɢɤɴɦɩɨɞɪɨɞChromatoiulusɬɨɣ ɩɪɟɯɜɴɪɥɹ ɤɴɦ Megaphyllum 9HUKRHII 9HUKRHII ɞɟɮɢɧɢɪɚ ɧɚɧɨɜɨ ɩɨɞɪɨɞBrachyiulus%HUOHVHɤɚɬɨɦɭɩɪɢɞɚɜɚɪɨɞɨɜɪɚɧɝȼɟɪɨɹɬɧɨɬɨɜɚɟɦɨɦɟɧɬɴɬ ɨɬɤɨɣɬɨɡɚɩɨɱɜɚɞɜɨɣɧɚɬɚɭɩɨɬɪɟɛɚɧɚBrachyiulus±ɟɞɧɚɬɚsensu%HUOHVHɚɞɪɭ- ɝɚɬɚsensu9HUKRHIIȼɬɚɡɢɜɪɴɡɤɚɧɚɫɥɟɞɜɚɳɚɬɚɝɨɞɢɧɚ9HUKRHII ɜɴɜɟɠɞɚ ɧɨɜɨ ɩɨɞɪɨɞɨɜɨ ɢɦɟ ɜ ɪɚɦɤɢɬɟ ɧɚ Brachyiulus sensu9HUKRHII ± Microbrachyiulus 9HUKRHII D ɩɭɛɥɢɤɭɜɚɫɬɚɬɢɹɜɴɪɯɭɦɧɨɝɨɧɨɠɤɢɬɟɧɚȻɨɫɧɚɏɟɪɰɟɝɨɜɢɧɚɢ Ⱦɚɥɦɚɰɢɹȼɱɚɫɬɬɚɡɚɫɹɝɚɳɚɸɥɢɞɢɬɟɨɬɪɟɝɢɨɧɚɚɜɬɨɪɴɬɫɴɫɬɚɜɹɧɹɤɨɥɤɨɤɥɸɱɚ ɟɞɢɧɨɬɤɨɢɬɨɫɟɨɬɧɚɫɹɞɨɩɨɞɪɨɞɨɜɨɬɨɪɚɡɞɟɥɹɧɟɧɚBrachyiulus sensu9HUKRHII Ʉɴɦ ɜɟɱɟ ɞɟɮɢɧɢɪɚɧɢɬɟ ɨɬ ɧɟɝɨ ɩɨɞɪɨɞɚ Chromatoiulus Microbrachyiulus HeteroiulusɢLeptomastigoiulus ɬɨɣɞɨɛɚɜɹɨɳɟɟɞɢɧ±Pachybrachyiulusɫɬɢɩɨɜ ɜɢɞChromatoiulus podabrus /DW]HO ȼɴɩɪɟɤɢɛɟɡɫɩɨɪɧɢɹɩɪɢɧɨɫɧɚ9HUKRHIIɜɦɨɪɮɨɥɨɝɢɹɬɚɢɫɢɫɬɟɦɚɬɢɤɚɬɚɧɚ ɸɥɢɞɢɬɟɧɟɝɨɜɚɬɚɧɟɩɨɫɥɟɞɨɜɚɬɟɥɧɚɢɫɢɥɧɨɢɡɦɟɧɱɢɜɚɤɨɧɰɟɩɰɢɹɩɨɨɬɧɨɲɟɧɢɟ ɤɥɚɫɢɮɢɤɚɰɢɹɬɚɧɚɝɨɪɟɢɡɛɪɨɟɧɢɬɟɬɚɤɫɨɧɢɩɪɟɡɬɟɡɢɧɹɤɨɥɤɨɝɨɞɢɧɢɡɚɟɞɧɨɫɴɫ ɡɚɟɞɧɨɫɴɫɫɢɫɬɟɦɧɨɬɨɧɚɪɭɲɚɜɚɧɟɧɚɨɫɧɨɜɧɢɩɪɢɧɰɢɩɢɜɡɨɨɥɨɝɢɱɧɚɬɚɧɨɦɟɧɤɥɚ- ɬɭɪɚɢɡɥɢɲɧɨɭɫɥɨɠɧɹɜɚɬɢɨɛɴɪɤɜɚɬɭɩɨɬɪɟɛɚɬɚɧɚ ɩɨɞ ɪɨɞɨɜɢɬɟɧɚɢɦɟɧɨɜɚɧɢɹ ɡɚɪɚɞɢɤɨɟɬɨɟɛɢɥɤɪɢɬɢɤɭɜɚɧɨɬɫɜɨɢɬɟɤɨɥɟɝɢ 3RFRFN$WWHPVɜɠ ɫɴɳɨ/D]iQ\L .RUVyV ȼɫɴɳɢɹɩɟɪɢɨɞɫɚɨɩɢɫɚɧɢɞɜɚɧɨɜɢɪɨɞɚ%UDFK\LXOLQL±Anaulaciulus3RFRFN ɢ Rhamphidoiulus $WWHPV ɧɚ ɛɚɡɚɬɚ ɧɚ ɞɜɚ ɧɨɜɢ ɜɢɞɚ ɫɴɨɬɜɟɬɧɨ A. paludicolaɢR. bujukderensis 3RFRFN$WWHPV Ɉɬɞɨ$WWHPVɨɩɢɫɜɚɞɪɭɝɢɩɪɟɞɫɬɚɜɢɬɟɥɢɧɚɫɴɳɢɹɪɨɞɤɚɤɬɨɢ Brachyiulus varibolinusɩɪɟɡ Ɍɪɢɛɭɫ%UDFK\LXOLQLɟɜɴɜɟɞɟɧɤɚɬɨɬɚɤɫɨɧɨɦɢɱɧɚɤɚɬɟɝɨɪɢɹɨɬ9HUKRHII Ⱦɟɮɢɧɢɰɢɹɬɚ ɟ ɧɚɩɪɚɜɟɧɚ ɩɨɫɪɟɞɫɬɜɨɦ ɤɥɸɱ ɧɚ ɢɡɜɟɫɬɧɢɬɟ ɞɨɬɨɝɚɜɚ ɸɥɢɞɢ ɇɚ ɫɥɟɞɜɚɳɚɬɚ ɝɨɞɢɧɚ ɚɜɬɨɪɴɬ ɩɭɛɥɢɤɭɜɚ ɦɨɧɨɝɪɚɮɢɹ ɜɴɪɯɭ ɧɹɤɨɥɤɨ ɝɪɭɩɢ ɦɧɨ- ɝɨɧɨɠɤɢɫɪɟɞɤɨɢɬɨɢɫɟɦɟɣɫɬɜɨ-XOLGDH 9HUKRHII Ɉɬɞɟɥɟɧɨɟɫɩɟɰɢɚɥɧɨ ɜɧɢɦɚɧɢɟɧɚ³%UDFK\LXOXV%HUO9HUK´ɤɚɬɨɞɜɚɨɬɞɨɬɨɝɚɜɚɲɧɢɬɟɦɭɩɨɞɪɨɞɨɜɟ± MicrobrachyiulusɢHeteroiulus±ɫɚɢɡɞɢɝɧɚɬɢɜɨɬɞɟɥɧɢɪɨɞɨɜɟ $WWHPV ɩɭɛɥɢɤɭɜɚ ɨɛɨɛɳɚɜɚɳɚ ɦɨɧɨɝɪɚɮɢɹ ɜɴɪɯɭ ɦɧɨɝɨɧɨɠɤɢɬɟ ɧɚ ɉɚɥɟɚɪɤɬɢɤɚȼɧɟɹɟɧɚɩɪɚɜɟɧɚɩɴɪɜɚɬɚɪɟɜɢɡɢɹɧɚɪɨɞMegaphyllum Brachyiulus sensu9HUKRHII ɱɢɢɬɨɜɢɞɨɜɟɩɨɨɧɨɜɚɜɪɟɦɟɜɟɱɟɫɚɧɚɞɯɜɴɪɥɢɥɢɋɚɦɚɬɚɪɟ- ɜɢɡɢɹɨɛɯɜɚɳɚɨɤɨɥɨôɨɬɬɹɯɤɨɢɬɨɫɚɩɪɟɞɫɬɚɜɟɧɢɫɧɨɜɢɪɢɫɭɧɤɢɨɩɢɫɚɬɟɥɧɢ ɞɨɩɴɥɧɟɧɢɹɢɨɩɪɟɞɟɥɢɬɟɥɟɧɤɥɸɱ Ɍɪɢ ɝɨɞɢɧɢ ɩɨɤɴɫɧɨ ɢɡɥɢɡɚ ɩɨɪɟɞɧɚɬɚ ɱɚɫɬ ɨɬ ɦɨɧɨɝɪɚɮɢɹɬɚ ɧɚ 9HUKRHII ɡɚ ɦɧɨɝɨɧɨɠɤɢɬɟ ɤɥɚɫ 'LSORSRGD Ɍɪɢɛɭɫɢɬɟ %UDFK\LXOLQL ɢ 8QFLJHULQL 2QFRLXOLQL DXFW 9HUKRHII ɨɛɟɞɢɧɹɜɚ ɜ ɩɨɞɫɟɦɟɣɫɬɜɨ %UDFK\LXOLQDH ɧɚ ɫɟɦɟɣɫɬɜɨ -XOLGDHȼɞɟɮɢɧɢɰɢɢɬɟɧɚɪɨɞɨɜɟɬɟɬɨɣɫɟɩɪɢɞɴɪɠɚɤɴɦɩɪɟɞɢɲɧɢɬɟɫɢɬɪɭɞɨɜɟ ɫɦɚɥɤɢɞɨɩɴɥɧɟɧɢɹɈɩɢɫɚɧɢɫɚɞɜɚɧɨɜɢɩɨɞɪɨɞɚɧɚMegaphyllum Brachyiulus sensu9HUKRHII ±ByzantorhopalumɢSyriobrachyiulus /RKPDQGHU ɩɭɛɥɢɤɭɜɚɫɬɚɬɢɹɜɴɪɯɭɞɢɩɥɨɩɨɞɧɚɬɚɮɚɭɧɚɧɚɄɚɜɤɚɡȼ ɧɟɹɬɨɣɨɩɢɫɜɚɧɨɜɜɢɞɫɩɚɞɚɳɤɴɦɧɨɜɪɨɞ%UDFK\LXOLQL±Grusiniulus redikorzevi ɤɚɤɬɨɢɧɹɤɨɥɤɨɧɨɜɢ ɩɨɞ ɜɢɞɚMegaphyllum ChromatoiulusDXFW ɩɪɢɧɚɞɥɟɠɚ- ɳɢɤɴɦɧɨɜɨɞɟɮɢɧɢɪɚɧɢɹɜɫɴɳɚɬɚɩɭɛɥɢɤɚɰɢɹɩɨɞɪɨɞOmobrachyiulusɢɟɞɢɧɜɢɞ ɤɴɦ ɆHJDSK\OOXP V VWU Chromatoiulus V VWU DXFW Ⱦɟɮɢɧɢɪɚɧɢ ɫɚ ɢ ɬɪɢ ɞɪɭ- ɝɢɩɨɞɪɨɞɚɧɚɫɴɳɢɹɪɨɞɩɨɜɟɱɟɨɩɢɫɚɧɢɜɢɞɨɜɟColchiobrachyiulusɫɟɞɢɧɫɬɜɟɧ ɜɢɞ M. dioscoriadis /LJQDX ArmeniobrachyiulusɢDonbrachyiulus ɫɴɳɨ ɦɨɧɨɬɢɩɧɢɫɴɫɫɴɨɬɜɟɬɧɢɜɢɞɨɜɟM. sevangense /RKPDQGHU ɢɆURVVLFXP 7LPRWKHHZ Ⱥɜɬɨɪɴɬɪɚɡɝɥɟɠɞɚɬɪɢɛɭɫ%UDFK\LXOLQLɤɚɬɨɟɞɢɧɫɬɜɟɧɜɪɚɦɤɢ- ɬɟɧɚɩɨɞɫɟɦɟɣɫɬɜɨ%UDFK\LXOLQDHɌɪɢɛɭɫ8QFLJHULQDHɟɨɛɟɞɢɧɟɧɫ/HXFRJHRUJLLQLɜ ɩɨɞɫɟɦɟɣɫɬɜɨ8QFLJHULQDHɉɨɞɪɨɛɧɢɬɟɨɩɢɫɚɧɢɹɩɨɞɤɪɟɩɟɧɢɫɩɪɟɰɢɡɧɢɞɟɬɚɣɥ- ɧɢɪɢɫɭɧɤɢɩɪɚɜɹɬɬɚɡɢɪɚɛɨɬɚɧɚ/RKPDQGHUɨɫɨɛɟɧɨɰɟɧɧɚ ȼ ɟɞɧɚ ɨɬ ɫɜɨɢɬɟ ɩɭɛɥɢɤɚɰɢɢ ɜɴɪɯɭ ɦɧɨɝɨɧɨɠɤɢɬɟ ɧɚ Ȼɴɥɝɚɪɢɹ 9HUKRHII ɨɩɢɫɜɚ ɦɨɧɨɬɢɩɧɢɹ ɪɨɞ Balkanophoenix ɫ ɬɢɩɨɜ ɜɢɞ B. borisi ɩɪɢɱɢɫɥɹ- ɜɚɣɤɢ ɝɨ ɤɴɦ ɬɪɢɛɭɫ %UDFK\LXOLQL ȼɢɞɴɬ ɟ ɨɩɢɫɚɧ ɨɬ ɞɜɟ ɧɚɯɨɞɢɳɚ ± ɪɚɣɨɧɚ ɧɚ ɝɚɪɚɅɚɤɚɬɧɢɤɢɩɨɪɟɱɢɟɬɨɧɚɪɑɭɤɭɪɫɤɚɜɪɚɣɨɧɚɧɚȼɟɥɢɧɝɪɚɞɎɚɤɬɴɬɱɟɧɢ- ɤɨɝɚ ɧɟ ɟ ɫɴɨɛɳɚɜɚɧ ɨɬɧɨɜɨ ɫɥɟɞ ɞɚɬɚɬɚ ɧɚ ɨɩɢɫɜɚɧɟɬɨ ɦɭ ɩɨɤɚɡɜɚ ɱɟ ɩɨ ɜɫɹɤɚ ɜɟɪɨɹɬɧɨɫɬ ɫɬɚɜɚ ɞɭɦɚ ɡɚ ɢɡɜɴɧɪɟɞɧɨ ɪɹɞɴɤ ɟɧɞɟɦɢɬ ȼ ɩɭɛɥɢɤɚɰɢɹɬɚ ɟ ɨɩɢɫɚɧ ɢ Brachyiulus (Byzantorhopalum) strandschanus ɞɧɟɫ ɜɚɥɢɞɟɧ ɤɚɬɨ Megaphyllum rossicum strandschanum /RKPDQGHU ɨɩɢɫɜɚɧɨɜɩɨɞɜɢɞɧɚMegaphyllum annulatum $WWHPV ±M ChromatoiulusDXFW annulatum samsunensisɩɨɤɨɣɬɨɞɟɮɢɧɢɪɚɧɨɜɩɨɞɪɨɞ PontobrachyiulusȾɨɞɧɟɫɬɨɣɨɫɬɚɜɚɦɨɧɨɬɢɩɟɧɧɨɫɜɨɟɨɛɪɚɡɧɚɬɚɦɨɪɮɨɥɨɝɢɹɧɚ ɜɢɞɚɞɚɜɚɞɨɛɪɢɨɫɧɨɜɚɧɢɹɩɨɞɪɨɞɴɬɞɚÄɨɰɟɥɟɟ´ɢɫɥɟɞɟɜɟɧɬɭɚɥɧɚɬɚɪɟɜɢɡɢɹɧɚ ɪɨɞMegaphyllum $WWHPV ɩɪɟɞɫɬɚɜɹɨɛɨɛɳɚɜɚɳɬɪɭɞɜɴɪɯɭɫɢɫɬɟɦɚɬɢɤɚɬɚɧɚɫɟɦɟɣɫɬɜɨ -XOLGDH ɜ ɤɨɣɬɨ ɨɬɞɟɥɹ ɫɩɟɰɢɚɥɧɨ ɜɧɢɦɚɧɢɟ ɧɚ ɪɨɞ Megaphyllum Chromatoiulus DXFW ɤɨɦɟɧɬɢɪɚɣɤɢ ɜɚɥɢɞɧɨɫɬɬɚ ɧɚ ɜɫɟɤɢ ɟɞɢɧ ɨɬ ɩɨɞɪɨɞɨɜɟɬɟ ɦɭ ȼ ɨɩɢɬ ɡɚ ɢɡɝɪɚɠɞɚɧɟ ɧɚ ɟɞɧɚ ɩɨɫɬɪɨɣɧɚ ɢ ɟɫɬɟɫɬɜɟɧɚ ɤɥɚɫɢɮɢɤɚɰɢɹ ɚɜɬɨɪɴɬ ɪɟɞɟɮɢɧɢ- ɪɚ ɢɡɜɟɫɬɧɢɬɟ ɞɨɬɨɝɚɜɚ ɩɨɞɪɨɞɨɜɟ ɢ ɫɚɦ ɜɴɜɟɠɞɚ ɨɳɟ ɬɪɢ Diaxylus Dittozusɢ Phauloiulus 9HUKRHII ɩɭɛɥɢɤɭɜɚɫɬɚɬɢɹɜɴɪɯɭɦɢɪɢɚɩɨɞɧɚɬɚɮɚɭɧɚɧɚɌɭɪɰɢɹȼɧɟɹ ɬɨɣɞɟɮɢɧɢɪɚɧɨɜɩɨɞɪɨɞɧɚMegaphyllum±Anatolicusɤɴɦɤɨɣɬɨɩɪɢɱɢɫɥɹɜɚɧɨ- ɜɨɨɩɢɫɚɧɢɹɜɫɴɳɚɬɚɩɭɛɥɢɤɚɰɢɹMegaphyllum BrachyiulusDXFW ulunumɡɚɟɞɧɨ ɫɨɳɟɞɜɚɩɨɪɚɧɨɨɩɢɫɚɧɢɨɬɧɟɝɨɜɢɞɚ±M. kosswigiɢM. anatolicumȾɜɟɝɨɞɢɧɢ ɩɨɤɴɫɧɨɚɜɬɨɪɴɬɩɭɛɥɢɤɭɜɚɧɨɜɚɫɬɚɬɢɹɡɚɫɹɝɚɳɚɬɭɪɫɤɢɬɟɦɧɨɝɨɧɨɠɤɢ 9HUKRHII

View Full Text

Details

  • File Type
    pdf
  • Upload Time
    -
  • Content Languages
    English
  • Upload User
    Anonymous/Not logged-in
  • File Pages
    37 Page
  • File Size
    -

Download

Channel Download Status
Express Download Enable

Copyright

We respect the copyrights and intellectual property rights of all users. All uploaded documents are either original works of the uploader or authorized works of the rightful owners.

  • Not to be reproduced or distributed without explicit permission.
  • Not used for commercial purposes outside of approved use cases.
  • Not used to infringe on the rights of the original creators.
  • If you believe any content infringes your copyright, please contact us immediately.

Support

For help with questions, suggestions, or problems, please contact us