Ann. Bot. Fennici 49: 248–254 ISSN 0003-3847 (print) ISSN 1797-2442 (online) Helsinki 31 August 2012 © Finnish Zoological and Botanical Publishing Board 2012

Camellia cherryana (), a new species from China

George Orel* & Peter G. Wilson

Royal Botanic Gardens, Mrs Macquaries Road, Sydney, NSW 2000, Australia (*corresponding author’s e-mail: [email protected])

Received 28 Apr. 2011, final version received 29 Feb. 2012, accepted 24 May 2012

Orel, G. & Wilson, P. G. 2012: cherryana (Theaceae), a new species from China. — Ann. Bot. Fennici 49: 248–254.

A new species of Camellia (Theaceae), C. cherryana Orel, endemic to the Sichuan– Yunnan border region, the People’s Republic of China, is described and illustrated. It has variably sized, slightly coriaceous, elliptic to lanceolate leaves with a strongly acu- minate apex and cuneate base; the adaxial leaf surface is glabrous and slightly pitted, mid-green and shiny, the abaxial leaf surface is sparsely pubescent, light green and shiny; the flowers are solitary, pedicellate, 5–7 cm in diam.; the corolla is two-whorled, white and the petals are soft, opaque to translucent, orbicular or almost orbicular, widely reflexed with undulate margins; the filaments are glabrous, up to 1.5 cm long, but mostly shorter; the styles are three, brown, densely pubescent, 1–1.2 cm long, basally joined but otherwise free; and the stigma is of the same colour, indistinct. The morphological evidence supports a placement in Camellia sect. Heterogena.

Introduction In this paper, a new species of Camellia sect. Heterogena from China is described and Camellia is a member of the tea family its taxonomic position assessed in the light of (Theaceae). Some 80% of the Camellia species the treatments of Sealy (1958), Chang and Bar- are found in the People’s Republic of China, tholomew (1984) and Ming and Bartholomew where they are distributed throughout the moun- (2007). While collecting under the auspices of tainous areas of the country, mostly in the prov- the Kunming Institute, in the Sichuan–Yunnan inces of Guandong, Guangxi, Yunnan, Guizhou, border region, the collector and nursery- Hunan and Sichuan. The other major centre of man Mr. Robert ‘Bob’ Cherry (Paradise , diversity for the genus is Vietnam, where almost Kulnura, NSW, Australia), discovered a distinc- 20% of the Camellia species can be found (Sealy tive Camellia with soft, opaque to translucent 1958, Chang & Bartholomew 1984, Gao et al. white petals, glabrous filaments and three styles 2005, Ming & Bartholomew 2007). Representa- that were free almost to the base. Only a single tives of Camellia also occur within the territories plant was found. The senior author was unable of Laos, Cambodia, Myanmar, India, Taiwan, to find another specimen that matched this plant Korea and Japan. One, or possibly two, species in any pertinent herbarium collection and the have also been found in the Philippines (Sealy Bob Cherry specimen is here described as a new 1958, Gao et al. 2005). species, C. cherryana. It is apparently confined Ann. BOT. Fennici Vol. 49 • Camellia cherryana, a new species from China 249 to a mountainous area of rainforest situated at (Sealy 1958). In Sealy’s opinion, the species in an altitude of between 1600–1700 m above sea sect. Heterogena, including the type species C. level. Further searching of the relevant area is henryana, may have originated from plants not required to better determine the distribution of dissimilar to C. flava of sect. Archaecamellia the species. (Sealy 1958). We are of the opinion that it is Analysis of the morphological characteristics beyond the scope of this paper, which only aims places C. cherryana within sect. Heterogena to describe a new species, to discuss the relation- (sensu Sealy 1958). That section was originally ships of Camellia species in sect. Heterogena, established by Sealy (1958) and it contains ten with Camellia species in a number of other sec- species, namely C. granthamiana, C. tennii, C. tions in sufficient depth and detail. However, in henryana, C. yunnanensis, C. paucipunctata, C. Tables 1 and 2 we list the differences between C. fleuryi, C. gaudichaudii, C. crapnelliana, C. cherryana and the two most similar species, C. wardii and C. furfuracea. The ten species of the yunnanensis and C. henryana. then newly constituted section are distinguished by perulate flowers, stamens that are free above the base and, in all but one instance (C. grantha- Notes on the classificatory system miana), styles that are free above their basal of Ming union (Sealy 1958). The proposed new species C. cherryana In Ming’s classificatory system (Ming &- Bar broadly fulfils the required phenotypic specifi- tholomew 2007), Camellia sect. Heterogena cations for sect. Heterogena, as it has perulate contains 13 species of Chinese provenance, flowers, with outer stamens free above the base namely C. szechuanensis, C. pubifurfuracea, C. and free styles that are joined only at the base. yunnanensis, C. tenii, C. mileensis, C. pauci- It should be pointed out that sect. Heterogena punctata, C. gaudichaudii, C. granthamiana, C. was created by Sealy (1958) to accommodate crapnelliana, C. furfuracea, C. pachyandra, C. “an assemblage of otherwise diverse specimens” wardii, C. szemaoensis, plus C. fleurii, which is that did not fit into any existing section. Thus of Vietnamese origin (Sealy 1958, Ming 1999). the morphology of the species in sect. Hetero- In comparison, the classificatory systems of gena is diverse to a degree, where the member Sealy (1958) and Ming (Ming & Bartholomew taxa possess many morphological characteristics 2007) appear to be superficially similar, yet a of Camellia species placed into other sections. number of differences exist. Sealy (1958) based The following example is included to illustrate his sect. Heterogena on C. henryana, which was this inherent diversity and its possible origins. designated as the type species for the section. In Some of the species included in sect. Hetero- Ming’s taxonomic treatment, Camellia species gena possess hairy and some glabrous gynoecia from C. sect. Furfuracea, C. subg. Protoca-

Table 1. Comparison of selected vegetative characters for Camellia yunnanensis, C. henryana and C. cherryana.

C. yunnanensis* C. henryana* C. cherryana

Bark rusty-brown to purplish-brown smooth or exfoliating medium grey Leaf shape oblong, elliptic to ovate oval, elliptic to oblong elliptic to ovate or oblong lanceolate Leaf apex bluntly acuminate to bluntly acuminate to long strongly acuminate acute, or caudate acuminate Leaf base cuneate to rounded cuneate or rounded or rounded cuneate Leaf margins serrulate or serrate serrulate or crenulate serrate Petiole length sessile or 2.0 to 6.0 mm 3.0 to 7.0 mm 8.0 to 10.0 mm Primary vein puberulate villous villous

* Data from Sealy (1958), Chang and Bartholomew (1984), Gao et al. (2005) and Ming and Bartholomew 2007. 250 Orel & Wilson • Ann. BOT. Fennici Vol. 49 mellia, C. sect. Pseudocamellia, together with ther, when compared with sect. Heterogena sensu Glyptocarpa, were subsumed into sect. Hetero- Sealy, Ming’s modified and enlarged section is gena sensu Ming and Bartholomew (2007). Fur- even more diverse than the original section estab- ther, Ming and Bartholomew (2007) divided the lished by Sealy. Ming’s addition of further three genus Camellia into only two subgenera, namely Camellia species, namely C. szechuanensis, C. subg. Thea and subg. Camellia. pubifurfuracea and C. pachyandra, which pos- If their criteria are recognised, then in general sess three “distinct” types of styles (3–4, 5 and terms the proposed new species, C. cherryana, 3–4 styles respectively), introduces an additional meets the requirements outlined for subg. Thea element of incompatibility into an already too (flowers clearly pedicellate, bracteoles differenti- diverse section and widens its terms of reference. ated from sepals, sepals persistent) but not those In view of these facts, we placed the pro- for subg. Camellia (flowers apparently sessile, posed new species, C. cherryana, within sect. bracteoles undifferentiated from sepals). Ming Heterogena sensu Sealy (1958), as its morphol- (Ming & Bartholomew 2007), however, while ogy tends to follow the requirements for the retaining the sectional name Heterogena, did not section as published by Sealy (1958). It should accept a specific status for C. henryana, merging be noted that in the taxonomic systems proposed it with various other taxa under the varietal name by Chang and Bartholomew (1984), and in the C. yunnanensis var. yunnanensis (Ming & Bar- treatment published by Gao et al. (2005), sect. tholomew 2007). Ming and Bartholomew (2007), Heterogena is not recognized as an independent perhaps to enable them to include a further five section. Chang and Bartholomew (1984) distrib- Camellia species in this section, slightly modified uted the constituent species of sect. Heterogena Sealy’s sectional description. For example: flow- mostly amongst sects. Archaecamellia, Stereo- ers sessile or sub-sessile (Sealy 1958), flowers carpus, Paracamellia, Pseudocamellia, Corra- sub-sessile (Ming & Bartholomew 2007); flowers lina, Furfuracea and Brachyandra. Under this terminal or axillary (Sealy 1958), flowers sub-ter- scheme, the type species of sect. Heterogena, C. minal or axillary (Ming & Bartholomew 2007); henryana, was included in sect. Pseudocamellia. stamens united with petals at the base, then Analysis of pertinent taxonomic data gained free (Sealy 1958), outer stamens basally connate from the evaluation of the newly proposed (Ming & Bartholomew 2007); styles 3–5 free or Camellia species indicates a number of mor- united (Sealy 1958), styles 3–5 distinct (Ming & phological characteristics that suggest affinities Bartholomew 2007). However, the term ‘distinct’ with sects. Furfuracea (Chang 1981, Chang & does not adequately describe the morphology Bartholomew 1984) and Pseudocamellia (Sealy of styles. The essential features of the styles, 1958), both of which Ming and Bartholomew e.g. their structure, shape, length, colour and the (2007) incorporated into their newly rede- quality of their surface are not supplied. Fur- fined sect. Heterogena. The authors considered

Table 2. Comparison of selected reproductive characters for Camellia yunnanensis, C. henryana and C. cherryana.

C. yunnanensis* C. henryana* C. cherryana

Perule number 8 to 12 9 to 10 14 to 17 Petal number 7 to 12 7 to 10 4 to 6 Androecium 1.4 cm to 2.5 cm 1.8 cm to 2.5 cm 1.5 cm Stamens free or united united partially united Gynoecium length 1.4 cm to 3.5 cm 1.0 cm to 2.1 cm 2.5 cm to 3.5 cm Styles glabrous glabrous or hairy densely pubescent Style number 3 to 7 3 3 Seed capsules diam. 3.5 cm to 8.0 cm 2.2 cm to 4.0 cm 1.5 cm to 2.5 cm Seed capsules shape globose, subglobose globose to oblate globose to ovoid to oblate

* Data from Sealy (1958), Chang and Bartholomew (1984), Gao et al. (2005) and Ming and Bartholomew 2007. Ann. BOT. Fennici Vol. 49 • Camellia cherryana, a new species from China 251 placement of the new Camellia species in sect. variable in size, irregularly but distinctly serrate, Furfuracea, according to the criteria of Chang less so at proximal end, margins slightly undu- (1981) and Chang and Bartholomew (1984), or late, up to 11 cm long and 4.2 cm wide; lamina in sect. Pseudocamellia, according to the criteria slightly coriaceous, elliptic to lanceolate, leaf of Sealy (1958) and Chang and Bartholomew apex strongly acuminate, base cuneate; adax- (1984), but neither was judged as appropriate, ial leaf surface glabrous, slightly pitted, more since the morphological traits in common were so towards margins, pits randomly distributed, few and rather superficial. mid-green and shiny, abaxial leaf surface light green, sparsely pubescent and shiny; primary vein on adaxial leaf surface up to 4 mm wide Camellia cherryana Orel, sp. nova (Fig. 1) proximally (up to 2 mm on abaxial side), less than 1 mm wide distally, light green (yellowish Camelliae henryanae filamentis exterioribus in pressed specimens), dull, villous and sunken partim connatis, stylis 3, liberis praeter bases affi- adaxially, abaxially prominent and pubescent, nis sed petiolis longioribus (8–10 vice 3–7 mm), although less so at distal end; secondary vena- corolla tantum 4–6 mera (vice 7+) et staminibus tion pinnate, brochidodromous, with 6–8(10) brevioribus (usque ad 15 vice 18–20 mm) differt. distally indistinct pairs of veins; veins adaxi- ally slightly sunken, abaxially prominent; terti- Type: People’s Republic of China. Sichuan Province, ary venation poorly defined, sparse and unevenly steep, forested ridge of unnamed mountain on Sichuan- distributed, partially craspedodromous. Mature Yunnan border, ca. 125 km NW of Kunming, 19 Nov. 1992 R. Cherry 912 (holotype NSW; isotype SYS). flowers slightly scented, white, shortly pedi- Etymology. The specific epithet honours Mr. Robert cellate, mostly axillary, sometimes terminal, ‘Bob’ Cherry, the eminent Australian plant collector, breeder always solitary; flower buds numerous, orbicu- and nurseryman (Kulnura, NSW, Australia). lar or nearly so, finely pubescent, up to 12 mm diam., bud scales prominent initially green, then Perennial, small, evergreen shrub to 3 m mostly white, pubescent, brownish; flowers 5–7 high; single stemmed, upright habit; with semi- cm diam., soft, opaque to translucent; petals pendulous outer branches, juvenile branches white, glabrous, orbicular or almost orbicular finely pubescent, yellowish, becoming orange to obovate, widely reflexed, with undulate mar- towards tips, turning entirely light brown and gins, overlapping at proximal end; pedicel ca. slightly pubescent on semi-mature branches; 1 mm long, 1–2 mm wide, light green; petals mature branches glabrous, light gray and smooth arranged in two whorls, unevenly distributed to slightly furrowed; trunk light grey with around common axis, proximally joined for random medium-dark gray blotches and slightly 5–7 mm; outer whorl of 2–3 petals, up to 3–5 furrowed; adult leaf petiole linear or slightly cm long, 3.5–4.0 cm wide, but mostly smaller, falcate, lacking lateral compression, slender, slightly concave, slightly asymmetric, obovate sometimes slightly wider at distal end, shallowly to orbicular in shape, lacking keel, or striations; channelled on adaxial side, uniformly medium- inner whorl of 2–3 petals, up to 3.0–3.5 cm long, green, pubescent with short white hairs, leaf 3.0–3.5 cm wide but mostly smaller, slightly petiole 8–10 mm long and 2–4 mm thick; petiole concave, slightly asymmetric, obovate to orbicu- ascending, at about 45° angle to midrib; axillary lar in shape, lacking keel, or striations; sepals leaf buds dull green to grey, pubescent, elliptic, 5(6) proximally joined forming a circular struc- with a soft, sharp apex, slightly compressed; ture 3.5–4.0 cm diam., variable in size, 10–14 terminal leaf buds also pubescent, laterally flat- mm long, 10–20 mm wide, greenish-white and tened, proximally scaled, with soft, sharp apex; thickened in centre and proximal end, light juvenile leaves shiny, red-coppery in colour, brown on margins, otherwise white in colour, later yellowish-green, soft, slightly waxy, lamina roughly obovate, outer surface finely pubes- slightly lustrous on both sides, variable in size cent with white hairs, inner surface glabrous; and shape, adaxially glabrous, abaxially sparsely perules 9–11, partially and proximally finely pubescent, margins finely serrate; mature leaves pubescent when juvenile, mostly glabrous when 252 Orel & Wilson • Ann. BOT. Fennici Vol. 49

Fig. 1. Camellia cherryana (from the holotype). — A: Juvenile leaves, adaxial and abaxial view. — B: Adult leaf and leaf petiole, adaxial view. — C: Adult leaf with axillary leaf bud, abaxial view. — D: Adult leaf, primary, second- ary, and tertiary venation, adaxial view. — E: Fully developed axillary flower bud. — F: Petal, the first or outer whorl. — G and H: Petals, the second or inner whorl. — I: Sepals, outer surface. — J: Sepals, inner surface. — K: Perules, the first or outer whorl. — L: Perules, the second whorl. — M: Perules, the third pubes- cent whorl. — N: Perules, the fourth and innermost whorl. — O: Corolla and a part of androecium. — P: Adult gynoecium. — Q: Senescing perules, calyx and gynoecium, post anthesis view. — R: Adult fruit. — S: Branch with flowers and leaves. mature or senescent, partially persistent, woody, surface, woody and slightly thickened, tightly irregularly arranged in four whorls, eventually appressed to third whorl, 7–8 mm wide, 4–5 mm forming an irregularly shaped involucre (which long; third whorl: two perules of variable shape is post-anthesis remnant of fourth innermost and size, roughly opposite to each other, slightly whorl) 7–9 mm high; first (outer) whorl: two concave, hairy, light green, woody and slightly perules, unevenly semi-orbicular, roughly oppo- thickened, especially at proximal end, on aver- site to each other, light green, initially tightly age 8–10 mm wide, 4–5 mm long; fourth whorl: adpressed to pedicel and second whorl, slightly three perules, roughly semi-orbicular, slightly reflexed when mature, variable in size, 4.5–5 concave, light green, yellowish and glabrous mm wide, 2–3 mm long; second whorl: two to on inner surface, 6–9 mm wide, 9–11 mm long; three perules, semi-orbicular to amorphous in stamens relatively numerous (200–250), in a cir- shape, slightly concave, light green and finely cular formation, 3.0–3.5 cm diam.; filaments gla- pubescent outside, yellow and glabrous on inner brous, variable in length, up to 1.5 cm long, but Ann. BOT. Fennici Vol. 49 • Camellia cherryana, a new species from China 253 mostly shorter, outer filaments proximally thick- cent and relatively short styles (C. gaudichau- ened and joined for 8–10 mm forming a shallow dii); perulate and solitary white flowers with a cup, inner stamens free, bright yellow, whitish small number of petals and glabrous filaments at base, up to 1 mm thick proximally; anthers (C. crapnelliana); perulate flowers with a small versatile or nearly so, dark yellow turning dark number of petals and three styles (C. wardii) and brown on margins when mature, 1–2 mm long, fragrant, perulate and few-petalled flowers with up to 2 mm wide; styles three, 1–1.2 cm long three styles and glabrous filaments (C. furfura- (which in adult flowers soon decay to variable cea). length, as short as 3–4 mm long), basally joined Camellia cherryana differs from the type for less than 1 mm, otherwise free, brown, species of sect. Heterogena, namely C. henry- densely pubescent with white hairs; stigma of ana, in a number of morphological character- same colour, indistinct; ovary superior, three car- istics. Camellia cherryana is a shrub up to 3 m pellate, irregular in shape, yellowish in colour, high (not a shrub or a small tree 2–7 m high); each carpel not always bi-locular; ovary densely leaf petiole 8–10 mm long (not 3–7 mm long); pubescent, 2.5–3.5 mm high and 3–4 mm wide, flowers with 4–6 petals (not 7–8, or sometimes basally connected to a narrow platform 4–6 mm 10, petals); two outermost petals not transitional diam. Mature fruit globose to unevenly ovoid, to the perules (two outermost petals transitional 15–20 mm long, 15–25 mm wide, capsule light to the perules) and androecium up to 15 mm long to medium brown, surface finely textured, finely (not 18–20 mm long) (fide Sealy 1958). pubescent, more so distally, and usually three Camellia cherryana also differs from the chambered, with one or two seeds per chamber, remaining species in sect. Heterogena sensu but capsule sometimes consists of only one large Sealy (1958) as follows: chamber that contains a single globose seed; columella pronounced and persistent after dehis- 1. From C. henryana: flowers always solitary cence, dehisced seed capsules may persist on rather than geminate; styles senescing to a plant until next season’s flowering; seed roughly variable length and as short as 3–4 mm long, ovoid, globose to sub-globose, sometimes later- not permanently 1.4–2 cm long. ally compressed, medium to dark brown, finely 2. From C. granthamiana: 4–6 rather than 8 pubescent, 10–15 mm long, 10–15 mm wide. petals; 3 styles 1–1.2 cm long and senescing Phenology. The new species was collected to as short as 3–4 mm long, not one stout and at early flowering in October. The species was tapering style 2 cm long. in full flower in November, the fruit matured in 3. From C. tenii: shrub to 3 m high, with semi- January–February, and the capsules opened in pendulous outer branches and densely pubes- late June. cent styles, rather than an erect shrub 1 m Distribution. Camellia cherryana is known high, with glabrous styles. only from the type collection. The species’ pre- 4. From C. yunnanensis: 4–6 rather than 8–12 cise provenance details are withheld for conser- petals; three mostly free, pubescent styles vation reasons. rather than 5 (or 4) joined and glabrous Camellia cherryana shares a number of mor- styles. phological characteristics with members of sect. 5. From C. paucipunctata: shrub to 3 m high Heterogena. These include partially united outer rather than a tree 6–12 m high; adult leaves filaments and three styles (C. henryana); soli- devoid of scattered cork warts on their lower tary and perulate flowers with a small number surface rather than adult leaves with scattered of petals, (C. granthamiana); relatively short cork warts on their lower surface; gynoecium styles (C. tenii); perulate and solitary flowers densely pubescent, not glabrous; fruit sub- and sub-globose seed capsules (C. yunnanensis); globose to unevenly ovoid, never pyriform. flowers with a small number of petals and three 6. From C. fleuryi: white and opaque to trans- styles that are basally united (C. paucipunctata); lucent rather than yellowish corolla; fila- perulate flowers with a small number of petals ments relatively numerous, with outer fila- with three short styles (C. fleuryi); three pubes- ments proximally thickened and joined for 254 Orel & Wilson • Ann. BOT. Fennici Vol. 49

8–10 mm forming a shallow cup, rather than back and more or less tawny and velutinous filaments very numerous and free to the base; on the face; capsules finely textured and gynoecium densely pubescent, not glabrous. pubescent, not furfuraceous (cf. Sealy 1958). 7. From C. gaudichaudii: adult leaves devoid of 4. From C. pubifurfuracea (sect. Heterogena scattered cork warts on their lower surface. sensu Ming & Bartholomew 2007): petals 8. From C. crapnelliana: shrub to 3 m high 4–6, slightly concave and slightly asymmet- rather than a small tree 5–7 m high; stamens ric, obovate to orbicular rather than up to 10, relatively numerous, not very numerous; obovate-oblong; and 3 styles, not 5 (Ming & styles soon senescing to 3–4 mm long, not Bartholomew 2007). permanently 1.5–1.6 cm long. 5. From C. mileensis (sect. Heterogena sensu 9. From C. wardii: shrub to 3 m high rather than Ming & Bartholomew 2007): flowers 5–7 cm a tree 6–12 m high; 9–11 rather than ca. 8 rather than 2–3 cm diam.; petals 4–6 petals, perules; styles densely pubescent rather than not 7–8; styles 1–1.2 cm, not 1.7 cm long glabrous or hairy. (Ming & Bartholomew 2007). 10. From C. furfuracea: flowers always solitary, 6. From C. pachyandra (sect. Heterogena sensu never geminate; styles 1–1.2 cm long rather Ming & Bartholomew 2007): petals white, than 1.85 cm long. never pink; styles 3, densely pubescent with white hairs, rather than 3 (or 4), glabrous and Camellia cherryana is also similar to other medium to dark brown; seeds finely pubes- perulate Camellia species with free styles. These cent, not smooth and grayish-brown (Ming & include species within sects. Paracamellia, Bartholomew 2007). Pseudocamellia and at least one species in sect. 7. From C. szemaoensis (sect. Heterogena sensu Camellia (Sealy 1958). Camellia cherryana dif- Ming & Bartholomew 2007): styles densely fers from them as follows: white-pubescent rather than distinctly gla- 1. From C. kissii (type of sect. Paracamellia): brous; capsules light to medium brown, not shrub to 3 m high rather than a tree up to reddish (Ming & Bartholomew 2007). 13 m high; flowers always solitary flowers rather than one or two flowers at the end of the shoots; petals 4–6 and proximally joined References rather than 7–8 and free to the base; styles Chang, H. T. 1981: A taxonomy of the genus Camellia. — basally joined for less than 1 mm, otherwise Acta Scientiarum Naturalium Universitatis Sunyatseni, free, rather than sometimes trifid or divided Monogr. Ser. 1: 1–180. half its length or more into three arms (Sealy Chang, H. T. & Bartholomew, B. 1984: . — Timber 1958). Press, Portland, Oregon. 2. From C. szechuanensis (type of sect. Pseu- Gao, J., Parks, C. R. & Du, Y. 2005: Collected species of the genus Camellia: an illustrated outline. — Zhejiang Sci- docamellia): flowers always solitary, never ence and Technology Press, Hangzhou. geminate; 4–6 rather than 8 petals; 3 styles, Ming, T. L. 1999: A systematic synopsis of the genus Camel- 1–1.2 cm long, not 3 or 4 and 8 cm long (cf. lia. — Acta Botanica Yunnanica 21: 149–159. Sealy 1958). Ming, T. L. & Bartholomew, B. 2007: Theaceae. — In: Wu, 3. From C. hongkongensis (sect. Camellia): Z.-Y. &. Raven, P. H. (eds.), , vol. 12: 366-478. Science Press, Beijing and Missouri Botanical shrub to 3 m high rather than a tree up to Garden Press, St. Louis. 10 m high; perules mostly glabrous when Sealy, J. R. 1958: A revision of the genus Camellia. — The mature, not grayish and velutinous on the Royal Horticultural Society, London.

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