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Taxonomic Revision of the Rock-Dwelling Door Snail Genus

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Taxonomic Revision of the Rock-Dwelling Door Snail Genus A peer-reviewed open-access journal ZooKeys 599: 1–137 (2016) Taxonomic revision of Montenegrina 1 doi: 10.3897/zookeys.599.8168 MONOGRAPH http://zookeys.pensoft.net Launched to accelerate biodiversity research Taxonomic revision of the rock-dwelling door snail genus Montenegrina Boettger, 1877 (Mollusca, Gastropoda, Clausiliidae) Zoltán Fehér1,2, Miklós Szekeres3 1 3rd Zoological Department, Natural History Museum Vienna, A-1010 Burgring 7, Vienna, Austria 2 De- partment of Zoology, Hungarian Natural History Museum, H-1088 Baross u 13, Budapest, Hungary 3 Institute of Plant Biology, Biological Research Centre of the Hungarian Academy of Sciences, Temesvári krt. 62, H-6726, Szeged, Hungary Corresponding author: Zoltán Fehér ([email protected]) Academic editor: T. Backeljau | Received 18 February 2016 | Accepted 31 May 2016 | Published 16 June 2016 http://zoobank.org/8BEE967F-7C69-4692-8210-A440AD8E2018 Citation: Fehér Z, Szekeres M (2016) Taxonomic revision of the rock-dwelling door snail genus Montenegrina Boettger, 1877 (Mollusca, Gastropoda, Clausiliidae). ZooKeys 599: 1–137. doi: 10.3897/zookeys.599.8168 Abstract The genusMontenegrina is revised on the basis of material available at the Hungarian Natural History Museum (Budapest), Naturhistorisches Museum Wien (Vienna), and the Naturmuseum Senckenberg (Frankfurt am Main), as well as newly discovered populations. The following new taxa are described: M. haringae sp. n., M. lillae sp. n., M. prokletiana sp. n., M. sturanyana sp. n., M. grammica erosszoltani ssp. n., M. grammica improvisa ssp. n., M. hiltrudae desaretica ssp. n., M. hiltrudae selcensis ssp. n., M. laxa delii ssp. n., M. nana barinai ssp. n., M. prokletiana kovacsorum ssp. n., M. rugilabris golikutensis ssp. n., M. rugilabris gregoi ssp. n., M. skipetarica danyii ssp. n., M. skipetarica gurelurensis ssp. n., M. skipetarica pifkoi ssp. n., M. skipetarica puskasi ssp. n., M. sporadica tropojana ssp. n., M. sturanyana gro- pana ssp. n., M. sturanyana ostrovicensis ssp. n., and M. tomorosi hunyadii ssp. n. A neotype is designated for M. helvola (Küster, 1860), and M. cattaroensis antivaricostata nom. n. was introduced to replace the junior homonym Clausilia umbilicata costata Boettger, 1907 (non Pfeiffer, 1928). Of each taxon types or specimens from the type localities are figured, and distribution maps are provided. Keywords Balkan, Albania, Greece, Montenegro, Macedonia, new taxa, distribution data, types, polytypic species, limestone habitat Copyright Zoltán Fehér, Miklós Szekeres. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. 2 Zoltán Fehér & Miklós Szekeres / ZooKeys 599: 1–137 (2016) Table of contents Introduction ............................................................................................................. 5 Methods ................................................................................................................... 7 Systematic part ....................................................................................................... 12 Genus Montenegrina Boettger, 1877 ................................................................. 12 Montenegrina apfelbecki (Sturany, 1907) ..................................................... 15 Montenegrina attemsi (Wagner, 1914) ......................................................... 17 Montenegrina attemsi attemsi (Wagner, 1914) ............................................. 18 Montenegrina attemsi jakupicensis Fauer, 1993 ............................................ 18 Montenegrina cattaroensis (Rossmässler, 1835) ............................................ 19 Montenegrina cattaroensis cattaroensis (Rossmässler, 1835) .......................... 20 Montenegrina cattaroensis antivaricostata nom. n. ........................................ 21 Montenegrina cattaroensis umbilicata (Boettger, 1879) ................................ 22 Montenegrina chiasma Nordsieck, 1972 ...................................................... 23 Montenegrina dofleini(Wagner, 1928) ........................................................ 24 Montenegrina dofleini dofleini (Wagner, 1928) ............................................ 24 Montenegrina dofleini fagorum Nordsieck, 1974 ......................................... 26 Montenegrina dofleini kastoriae Nordsieck, 1972 ......................................... 27 Montenegrina dofleini pinteri Nordsieck, 1974 ............................................ 28 Montenegrina dofleini prespaensis Nordsieck, 1988 ...................................... 29 Montenegrina dofleini sinosi Páll-Gergely, 2010 ........................................... 30 Montenegrina dofleini wagneri Szekeres, 2006 ............................................. 30 Montenegrina drimmeri Fehér & Szekeres, 2006 ......................................... 31 Montenegrina fuchsi Brandt, 1961 .............................................................. 32 Montenegrina fuchsi fuchsi Brandt, 1961 ..................................................... 32 Montenegrina fuchsi klemmi Brandt, 1962 .................................................. 34 Montenegrina fuchsi muranyii Fehér & Szekeres, 2006 ................................ 35 Montenegrina fuchsi pallida Fauer, 1993 ..................................................... 36 Montenegrina grammica Nordsieck, 1988 ................................................... 36 Montenegrina grammica grammica Nordsieck, 1988 ................................... 37 Montenegrina grammica erosszoltani ssp. n. ................................................. 37 Montenegrina grammica improvisa ssp. n. .................................................... 38 Montenegrina haringae sp. n. ...................................................................... 39 Montenegrina helvola (Küster, 1860) ........................................................... 40 Montenegrina helvola helvola (Küster, 1860) ............................................... 41 Montenegrina helvola carinata Erőss & Szekeres, 1999 ................................ 42 Montenegrina helvola magna Fehér & Szekeres, 2006 .................................. 42 Montenegrina helvola ornata Erőss & Szekeres, 1999 .................................. 44 Montenegrina helvola pageti Brandt, 1962 ................................................... 44 Montenegrina hiltrudae Nordsieck, 1972 .................................................... 45 Montenegrina hiltrudae hiltrudae Nordsieck, 1972 ...................................... 45 Montenegrina hiltrudae desaretica ssp. n. ..................................................... 47 Taxonomic revision of Montenegrina 3 Montenegrina hiltrudae costulata Erőss & Szekeres, 2006 ............................ 48 Montenegrina hiltrudae dennisi Gittenberger, 2002 ..................................... 48 Montenegrina hiltrudae densicostulata Nordsieck, 1974 ............................... 49 Montenegrina hiltrudae fusca Fehér & Szekeres, 2006 ................................. 51 Montenegrina hiltrudae maasseni Gittenberger, 2002 .................................. 51 Montenegrina hiltrudae protruda Gittenberger, 2002 .................................. 52 Montenegrina hiltrudae robusta Nordsieck, 2009 ........................................ 53 Montenegrina hiltrudae sattmanni Nordsieck, 1988 .................................... 53 Montenegrina hiltrudae selcensis ssp. n. ........................................................ 54 Montenegrina janinensis (Mousson, 1859) .................................................. 55 Montenegrina laxa (Küster, 1861) ............................................................... 56 Montenegrina laxa laxa (Küster, 1861) ........................................................ 57 Montenegrina laxa dedovi Nordsieck, 2009 ................................................. 61 Montenegrina laxa delii ssp. n. .................................................................... 61 Montenegrina laxa disjuncta Fehér & Szekeres, 2006 .................................. 62 Montenegrina laxa errans Erőss & Szekeres, 2006 ....................................... 62 Montenegrina laxa iba Nordsieck, 1972 ...................................................... 63 Montenegrina laxa kontschani Erőss & Szekeres, 2006 ................................ 64 Montenegrina laxa lakmosensis Nordsieck, 2009 .......................................... 65 Montenegrina laxa miraka Nordsieck, 1996 ................................................ 65 Montenegrina lillae sp. n. ............................................................................ 66 Montenegrina minuscula Erőss & Szekeres, 2006 ........................................ 67 Montenegrina nana Fehér & Szekeres, 2006 ............................................... 68 Montenegrina nana nana Fehér & Szekeres, 2006 ....................................... 68 Montenegrina nana barinai ssp. n. .............................................................. 69 Montenegrina nana gracilis Erőss & Szekeres, 2006 ..................................... 70 Montenegrina okolensis Szekeres, 2006 ........................................................ 71 Montenegrina okolensis okolensis Szekeres, 2006 .......................................... 71 Montenegrina okolensis caesia Fehér & Szekeres, 2006 ................................. 72 Montenegrina perstriata (Wagner,
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