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California Terrestrial and Vernal Pool Invertebrates of Conservation Priority1

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California Terrestrial and Vernal Pool Invertebrates of Conservation Priority1 Species Federal ESA CA ESA G-Rank S-Rank Other Statuses GASTROPODA (Snails and slugs) Ammonitella yatesii None None G1 S1 IUCN - Vulnerable tight coin (=Yates' snail) Ancotrema voyanum None None G1G2 S1S2 BLM-Sensitive hooded lancetooth Assiminea infima None None G1 S1 IUCN - Vulnerable Badwater snail Binneya notabilis None None G1 S1 IUCN - Data Deficient Santa Barbara shelled slug Eremarionta immaculata None None G1 S1 IUCN - Vulnerable white desertsnail Eremarionta millepalmarum None None G1 S1 IUCN - Vulnerable Thousand Palms desertsnail Eremarionta morongoana IUCN - Near None None G1G3 S1 Morongo (=Colorado) desertsnail Threatened Eremarionta rowelli bakerensis None None G3G4T1 S1 IUCN - Data Deficient Baker's desertsnail Eremarionta rowelli mccoiana None None G3G4T1 S1 IUCN - Data Deficient California Mccoy snail Haplotrema catalinense None None G1 S1 Santa Catalina lancetooth Haplotrema duranti None None G1G2 S1S2 ribbed lancetooth Helminthoglypta allynsmithi None None G1 S1 IUCN - Vulnerable Merced Canyon shoulderband Helminthoglypta arrosa monticola None None G2G3T1 S1 mountain shoulderband Helminthoglypta arrosa pomoensis None None G2G3T1 S1 IUCN - Data Deficient Pomo bronze shoulderband Helminthoglypta ayresiana sanctaecrucis None None G1G2T1T2 S1S2 Ayer's snail Helminthoglypta callistoderma None None G1 S1 IUCN - Endangered Kern shoulderband 1 This list pertains to Scientific Collecting Permit regulations (Title 14, Section 650, California Code of Regulations). The species on this list include vernal pool, terrestrial, and aquatic invertebrates with a terrestrial phase that are listed under the Federal and California Endangered Species Acts, or have a NatureServe rank of G1, G1G2, S1, S1S2, SH or SX. For a full list of invertebrates of conservation concern, see the California Natural Diversity Database Special Animals List: https://www.wildlife.ca.gov/Data/CNDDB/Plants-and-Animals. List generated on June 12, 2017 from a query of the California Natural Diversity Database. California Department of Fish and Wildlife Page 1 of 17 California Terrestrial and Vernal Pool Invertebrates of Conservation Priority Species Federal ESA CA ESA G-Rank S-Rank Other Statuses Helminthoglypta coelata None None G1 S1 IUCN - Vulnerable mesa shoulderband Helminthoglypta concolor None None G1G2 S1S2 whitefir shoulderband Helminthoglypta fontiphila None None G1 S1 Soledad shoulderband Helminthoglypta greggi None None G1 S1 Mohave shoulderband Helminthoglypta hertleini None None G1 S1S2 BLM-Sensitive Oregon shoulderband Helminthoglypta milleri None None G1 S1 peak shoulderband Helminthoglypta mohaveana IUCN - Near None None G1 S1 Victorville shoulderband Threatened Helminthoglypta nickliniana awania None None G3T1 S1 IUCN - Data Deficient Peninsula coast range shoulderband Helminthoglypta nickliniana bridgesi None None G3T1 S1S2 IUCN - Data Deficient Bridges' coast range shoulderband Helminthoglypta sequoicola consors None None G2T1 S1 IUCN - Data Deficient redwood shoulderband Helminthoglypta stiversiana williamsi None None G2G3T1 S1 IUCN - Data Deficient Williams' bronze shoulderband Helminthoglypta taylori None None G1 S1 westfork shoulderband Helminthoglypta traskii pacoimensis None None G1G2T1 S1 Pacoima shoulderband Helminthoglypta traskii traskii None None G1G2T1 S1 Trask shoulderband Helminthoglypta uvasana None None G1 S1 Grapevine shoulderband Helminthoglypta vasquezi None None G1 S1 Vasquez shoulderband Helminthoglypta walkeriana IUCN - Critically Morro shoulderband (=banded dune) Endangered None G1 S1S2 Endangered snail Herpeteros angelus None None G1 S1 Soledad desertsnail Hesperarion plumbeus None None G1 S1 leaden slug Megomphix californicus None None G1G2 S1S2 Natural Bridge megomphix Micrarionta facta None None G1G2 S1S2 IUCN - Vulnerable Santa Barbara islandsnail Micrarionta feralis IUCN - Critically None None G1 S1 San Nicolas islandsnail Endangered Micrarionta gabbi None None G1 S1 IUCN - Vulnerable San Clemente islandsnail California Department of Fish and Wildlife Page 2 of 17 California Terrestrial and Vernal Pool Invertebrates of Conservation Priority Species Federal ESA CA ESA G-Rank S-Rank Other Statuses Micrarionta opuntia None None G1 S1 IUCN - Vulnerable pricklypear islandsnail Monadenia callipeplus None None G1G2 S1S2 downy sideband Monadenia circumcarinata BLM-Sensitive | IUCN None None G1 S1 keeled sideband - Vulnerable Monadenia cristulata None None G1G2 S1S2 crested sideband Monadenia fidelis leonina None None G4G5T1T2 S1S2 A terrestrial snail Monadenia fidelis pronotis None None G4G5T1 S1 rocky coast Pacific sideband Monadenia infumata ochromphalus None None G2T1 S1 yellow-based sideband Monadenia infumata setosa None Threatened G2T2 S2 IUCN - Vulnerable Trinity bristle snail Monadenia marmarotis None None G1 S1 marble sideband Monadenia mormonum buttoni None None G2T1 S1S2 Button's Sierra sideband Monadenia mormonum hirsuta None None G2T1 S1 BLM-Sensitive hirsute Sierra sideband Monadenia troglodytes troglodytes IUCN - Data Deficient | None None G1G2T1T2 S1S2 Shasta sideband USFS - Sensitive Monadenia troglodytes wintu IUCN - Data Deficient | None None G1G2T1T2 S1S2 Wintu sideband USFS - Sensitive Monadenia tuolumneana None None G1 S1 BLM-Sensitive Tuolumne sideband Monadenia yosemitensis None None G1 S1S2 Yosemite Mariposa sideband Pristiloma shepardae None None G1 S1 Shepard's snail Prophysaon coeruleum None None G3G4 S1S2 Blue-gray taildropper slug Punctum hannai None None G1G2 S1S2 Trinity Spot Radiocentrum avalonense IUCN - Critically None None G1 S1 Catalina mountainsnail Endangered Rothelix warnerfontis None None G1 S1 USFS - Sensitive Warner Springs shoulderband Sterkia clementina IUCN - Near None None G1 S1S2 San Clemente Island blunt-top snail Threatened Trilobopsis roperi None None G1 S1 USFS - Sensitive Shasta chaparral Trilobopsis tehamana BLM-Sensitive | USFS None None G1 S1 Tehama chaparral - Sensitive California Department of Fish and Wildlife Page 3 of 17 California Terrestrial and Vernal Pool Invertebrates of Conservation Priority Species Federal ESA CA ESA G-Rank S-Rank Other Statuses Vespericola pressleyi BLM-Sensitive | USFS None None G1 S1 Big Bar hesperian - Sensitive Vespericola scotti None None G1 S1 Benson Gulch hesperian Vespericola shasta None None G1 S1 USFS - Sensitive Shasta hesperian Vespericola sierranus None None G2 S1S2 Siskiyou hesperian Xerarionta intercisa None None G1 S1 IUCN - Vulnerable horseshoe snail Xerarionta redimita None None G1G2 S1 IUCN - Vulnerable wreathed cactussnail Xerarionta tryoni None None G1 S1 IUCN - Vulnerable Bicolor cactussnail ARACHNIDA (Spiders and relatives) Aphrastochthonius grubbsi None None G1G2 S1S2 Grubbs' Cave pseudoscorpion Aphrastochthonius similis None None G1G2 S1S2 Carlow's Cave pseudoscorpion Archeolarca aalbui None None G1G2 S1S2 Aalbu's Cave pseudoscorpion Banksula californica None None GH SH Alabaster Cave harvestman Banksula galilei None None G1 S1 Galile's cave harvestman Banksula grubbsi None None G1 S1 Grubbs' cave harvestman Banksula incredula None None G1 S1 incredible harvestman Banksula martinorum None None G1 S1 Martins' cave harvestman Banksula melones None None G1 S1 IUCN - Vulnerable Melones Cave harvestman Banksula rudolphi None None G1 S1 Rudolph's cave harvestman Banksula tuolumne None None G1 S1 Tuolumne cave harvestman Banksula tutankhamen None None G1 S1 King Tut Cave harvestman Calicina arida None None G1 S1 San Benito harvestman Calicina breva None None G1 S1 Stanislaus harvestman Calicina cloughensis None None G1 S1 Clough Cave harvestman California Department of Fish and Wildlife Page 4 of 17 California Terrestrial and Vernal Pool Invertebrates of Conservation Priority Species Federal ESA CA ESA G-Rank S-Rank Other Statuses Calicina conifera None None G1 S1 Crane Flat harvestman Calicina diminua None None G1 S1 Marin blind harvestman Calicina dimorphica None None G1 S1 Watts Valley harvestman Calicina macula None None G1 S1 marbled harvestman Calicina mesaensis None None G1 S1 Table Mountain harvestman Calicina minor None None G1 S1 Edgewood blind harvestman Calicina piedra None None G1 S1 Piedra harvestman Calileptoneta briggsi None None G1 S1 Briggs' leptonetid spider Calileptoneta oasa None None G1 S1 Andreas Canyon leptonetid spider Calileptoneta ubicki None None G1 S1 Ubick's leptonetid spider Calileptoneta wapiti None None G1 S1 Mendocino leptonetid spider Fissilicreagris imperialis None None G1 S1 IUCN - Vulnerable Empire Cave pseudoscorpion Hubbardia idria None None G1 S1 Idria short-tailed whipscorpion Hubbardia secoensis None None G1 S1 Arroyo Seco short-tailed whipscorpion Hubbardia shoshonensis None None G1 S1 BLM-Sensitive Shoshone Cave whip-scorpion Larca laceyi None None G1G2 S1 Lacey's Cave pseudoscorpion Meta dolloff None None G1 S1 IUCN - Vulnerable Dolloff Cave spider Microcina edgewoodensis None None G1 S1 Edgewood Park micro-blind harvestman Microcina homi None None G1 S1 Hom's micro-blind harvestman Microcina jungi None None G1 S1 Jung's micro-blind harvestman Microcina leei None None G1 S1 Lee's micro-blind harvestman Microcina lumi None None G1 S1 Lum's micro-blind harvestman Microcina tiburona None None G1 S1 Tiburon micro-blind harvestman California Department of Fish and Wildlife Page 5 of 17 California Terrestrial and Vernal Pool Invertebrates of Conservation Priority Species Federal ESA CA ESA G-Rank S-Rank Other Statuses Neochthonius imperialis None None G1 S1 Empire Cave pseudoscorpion Pseudogarypus
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  • Helminthoglypta Walkeriana COMMON NAME: Morro Shoulderband Snail CLASS, FAMILY: Gastropoda, Helminthoglyptidae

    Helminthoglypta Walkeriana COMMON NAME: Morro Shoulderband Snail CLASS, FAMILY: Gastropoda, Helminthoglyptidae

    SCIENTIFIC NAME: Helminthoglypta walkeriana COMMON NAME: Morro shoulderband snail CLASS, FAMILY: Gastropoda, Helminthoglyptidae ORIGINAL DESCRIPTION: Hemphill, H. 1911. Descriptions of some varieties of shells, with short notes on the geographical range and means of distribution of land snails. Transactions of the San Diego Society of Natural History 1(3):102, pl. 2 (two views of shells). (Described as Helix walkeriana, with morroensis described as a variety.) TYPE MATERIAL: Roth and Sadeghian (2003) list the syntypes as follows: Academy of Natural Sciences, Philadelphia #112424 (4 specimens), California Academy of Sciences #058838 (6), #065523 (2), #065524 (3), Santa Barbara Museum of Natural History #33958 (22), University of Colorado, Boulder #20178 (4), and United States National Museum of Natural History (Smithsonian Institution) #174679-174682 (8). RANKING/STATUS: Federally Endangered (1994), G1S1 (NatureServe – CNDDB), CR/A1ce, B1+2bc (IUCN). GENERAL DESCRIPTION: Moderately large helminthoglyptid snails with globose, helicoid shells and brown bodies. DIAGNOSTIC CHARACTERS: Only three helminthoglyptid species occur in coastal San Luis Obispo County; until recently H. morroensis was considered a subspecies of H. walkeriana (Walgren 2003). The third species, H. umbilicata, has distinctive malleated shell sculpture (Roth and Tupen 2004). Detailed morphometric analysis of shells (Roth and Tupen 2004) revealed that H. walkeriana and H. morroensis are separate species. Shells of H. walkeriana are more globose and tightly coiled, with more whorls and less papillation than those of H. morroensis. The skin color of morroensis is blackish in life, whereas in walkeriana it is medium brown, and the mantle pigmentation is more extensive in morroensis. Penial morphology also differs, with the penis of walkeriana being slender and hourglass-shaped, with simple, smooth pilasters.